Sabah, Borneo

26th March – 9th April 2016

Leader: Rob Hutchinson

Participants:Heather Bagg, Jan-Joost Bouwman, Mike & Denice Coverdale, Gordon Ellis, Erkki Holopainen

Whitehead’s Broadbill © Rob Hutchinson / Birdtour Asia

Borneo has one of the finest collection of spectacular endemics in Asia and the vast majority can be encountered in Sabah thanks to good transport links, and particularly well placed lodges. The endemics rarely give themselves up easily and this tour with Borneo in the grip of a substantial El Niño induced drought was particularly difficult and it was amazing to see Mount Kinabalu with rivers and waterfalls running dry. Fortunately, bird activity was surprisingly good and we had nothing more than a few drops of rain during the whole two weeks. We managed to find a total of 311 species which included an impressive 48 island endemic, while the Sarawak pre-tour extension added a further 38 species and 4 endemics not seen on the main tour. We started in the Crocker range which got us off to a great start with Golden-naped, Mountain and Bornean Barbets, Bornean Leafbird, Bornean Bulbul, Whitehead’s Spiderhunter, Mountain Serpent Eagle and Bare-headed Laughingthrush. Moving on to the impressive Mount Kinabalu we easily completed the ‘Whitehead’s Trio’ with excellent encounters with Whitehead’s Broadbill and Trogon. Fruit-hunter performed nicely as did Bornean Stubtail, Mountain Wren Babbler, Eye-browed Jungle Flycatcher, Mountain Black-eye and with some work both Red-breasted and Crimson-headed Partridge, a brief Everett’s Thrush and daytime views of Mountain Scops Owl and Sunda Collared Owlet. A morning at Poring was quite slow although Hose’s Broadbill showed briefly, then it

was time to head into the lowlands. Our first stop was Sepilok where Black-crowned Pitta stood out among the many lowland additions, the Gomantong Caves where amazing Orangutan encounters stole the show. The Kinabatangan River didn’t disappoint with an impressive set of hornbills including the important Wrinkled and White-crowned Hornbills, Storm’s Stork, Jerdon’s Baza and the ever-entertaining Proboscis Monkeys. Bornean Ground Cuckoo made us work hard as always but the reward was stupendous views, while night-time excursions gave amazing looks at both Large Frogmouth and Oriental Bay Owl. We finished the tour at the fabulous Borneo Rainforest Lodge in Danum Valley where Bristlehead was definitely the highlight but we also reveled in Black-throated Wren Babbler, Bornean Ground Babbler, Chestnut-necklaced Partridge, ‘Bornean’ Banded Kingfisher, White-necked Babbler, Bornean Banded and Blue-headed Pitta.

Mountain Black-eye and Whitehead’s Broadbill

Bornean Brown Barbet and Mountain Serpent Eagle Departing Kota Kinabalu early on the first morning found us in the Crocker Mountain Range just before dawn and we had already polished off our breakfast by the time that the first birds appeared. ‘Sunda’ Ashy Drongo and Mountain Barbet were the first, both heavily silhouetted but we were to see plenty of both in excellent light later. Once good light appeared we started on a high when a flowering tree we had spotted attracted a Whitehead’s Spiderhunter. A Bornean Forktail out on the roadside was nice but things were generally rather quiet. We plugged away and found wintering Mugimaki Flycatchers in good numbers, Bornean Treepie, Sunda Cuckooshrike, Gold-whiskered Barbet, Black-and-crimson Oriole, Cinereous Bulbul, Blyth’s Hawk Eagle and an excellent close flyover Mountain Serpent Eagle. Some species, notably Bornean Barbet and Bornean Leafbird were notable by the

absence so we dropped down in elevation and were soon enjoying the leafbirds as well as the wacky Long-tailed Broadbill, Black-and-yellow Broadbill, Bar-winged Flycatcher Shrike, Asian Drongo Cuckoo, Banded Woodpecker, affine Blyth’s Hawk Eagle, and at long last a Bornean Barbet which still wasn’t singing but gave fine views nevertheless. We started the afternoon at higher elevations where Mountain Black-eyes appeared at very close range to check us out, a single Golden-naped Barbet posed in the Swaro ‘scope, several Sunda Cuckoos teased us before finally a vocal individual was found calling from a distant tree. The distinctive boop boop of a Bare-headed Laughingthrush group was heard and we teased them closer with one perched up beautifully with his bare neck patches bulging with each note. We finished with a noisy group of Bornean Brown Barbets and another Sunda Cuckooshrike before making the drive to our base for the coming days on the lower slopes of Mount Kinabalu.

Indigo Flycatcher and Fruit-hunter

Mountain Scops Owl and Whitehead’s Trogon Our first full day on the mountain began before light with a Mountain Scops Owl calling but not seen. A pair of Orange-headed Thrushes ventured onto the roadside in the early morning then as things brightened up we got off to a flying start with excellent views of an adult and immature Whitehead’s Broadbill, Fruit-hunter for some, and a frustrating heard only Everett’s Thrush. Another area gave us fine views of Eye-browed Jungle Flycatcher (though it’s actually a robin!), Sunda Cuckooshrike, Bornean Forktail and with some patience we tracked down a calling Whitehead’s Trogon spotted by Mike through the tiniest of gaps, which went on to perform splendidly and end our morning on a high. Indigo and Blue-and-white Flycatchers, Black-sided Flowerpecker and Temminck’s Sunbirds all joined us for lunch at the hotel restaurant balcony. Another trail in the afternoon gave us an energetic group of

Mountain Wren Babblers bouncing around in the undergrowth, then a series of 3 Fruit-hunters which performed very well. We finished the afternoon trying again for the Mountain Scops Owl which we found at day roost but was mostly obscured, and perhaps the same 2 Whitehead’s Broadbills from the morning.

Red-bearded Bee-eater and Temminck’s Sunbird The next day would always be an anti-climax with just a few specific targets to search for but after breakfast things picked up with our largest feeding flock so far containing both Checker-throated and Orange-backed Woodpeckers, our first Bornean Green Magpies and further singles of Whitehead’s Broadbill and Fruit-hunter. A calling Sunda Collared Owlet led us a merry dance and we never manage to see him in his chosen calling tree. The diminutive Bornean Stubtail gave us nice clear views although the colourful leg rings were a distraction! In the afternoon we quickly located two vocal Pale-faced Bulbuls, a single Giant Swiftlet overhead (having heard them earlier when we were under the forest canopy), Mountain Tailorbird and Mountain Black-eye. Some colourful Orange-headed Thrushes livened up our last walk and we again enjoyed the large feeding flock that now included White-throated Fantail and Yellow-breasted Warbler.

Red Giant Flying Squirrel and Brown Boobook No less than three Orange-headed Thrushes started things the following day but again the Everett’s Thrushes were heard only so we tried another trail where another fantastic male Whitehead’s Trogon was sallying actively and a single Red-breasted Partridge was seen by some of the group while checking a quiet side trail. The last walk of the morning found a smart female White-browed Shortwing which we were able to track through the forest for some time.

The highlight of the early afternoon was a mammal rather than a bird, with an orgy of 3 Kinabalu Squirrels so engulfed in their romantic encounter that they barely noticed us watching. We were also able to find the Pale-faced Bulbuls again and at least 2 Golden-naped Barbets feasting in a fruiting tree. A Sunda Collared Owlet was calling excitedly and when we eventually tracked it down in the canopy it was clutching a freshly killed lizard. The last stop of the day finally gave a sighting of Everett’s Thrush, although only for those at the front of the line and it never reappeared.

Bornean Orang-utans

Scarlet-rumped Trogon and Ruddy Kingfisher We headed to lower elevations the next day for a morning at Poring Hot Springs. It was nice to have a change of scene but Poring is quiet at the best of times but in this year’s drought conditions the cicada’s were near deafening and it was particularly tough going. We stuck at it though and found Grey-headed Babbler, Yellow-bellied Warblers, Grey-chested Jungle Flycatcher, Purple-naped Spiderhunter, White-crowned Forktail and a posing Hill Blue Flycatcher, no an easy bird in Sabah and a likely future split with it’s entirely orange underparts, small size and distinct song. A calling Blue-banded Pitta didn’t want to come any closer and a singing Hose’s Broadbill disappeared not long after Mike spotted it and wasn’t seen again. The Cicada’s had actually quietened down as we headed back down the trail and we were entertained by a group of Crested Jays, Chestnut-breasted Malkoha, Chestnut-backed Scimitar Babbler, Orange-breasted Trogon and a stunning male Rufous-collared Kingfisher. Back at the resort we met up with the rest of the group who had a more relaxing time at the fruiting trees around the resort where highlights included Scaly-breasted Bulbul and after lunch visited an impressive flowering Rafflesia

keithii. Next stop was some scrubby habitat which is the favoured habitat of Pygmy White-eye and sure enough we had seen several within just a few minutes of trying, some at close range with the delicate spotted crown clearly visible. We finished off with another couple of hours on the mountain with a group of Giant Swiftlets overhead getting us looking skywards at the right time to see a vagrant Asian House Martin but other than the now familiar Eye-browed Jungle Flycatcher and a single Bornean Stubtail there was nothing new. We had time for a final few hours in the morning before we needed to leave so naturally we headed back to the forest for another session. A pair of Mountain Scops Owls called well but were never captured in the torch light so we continued on but the only thrush making use of the trail in the early morning was the orange-headed variety. We did though score with a pair of Crimson-headed Partridges crossing the trail ahead of us and then we located one of the Mountain Scops Owl at it’s day roost for mega views. It was then time to make our way eastwards and into the lowlands through the vast oil palm plantations that sadly now dominate the landscape. Our next stop at Sepilok is something of a forest haven among the bird-less oil palm and we had an enjoyable afternoon session on the trails adding many lowland species like Chestnut-breasted and Raffle’s Malkoha, Green Iora, Red-throated Sunbird and Thick-billed Spiderhunter. The highlights species were undoubtedly a stunning male Red-naped Trogon and a Black-crowned Pitta which enjoyed a bathe in the nearby stream. We finished the day on the impressive canopy walkway where a pair of Wallace’s Hawk Eagles were performing nicely and as dusk approached so did half a dozen Red Giant Flying Squirrels, a couple of which glided right over our heads. Brown Boobooks were very vocal and some long views finished the day nicely.

Bornean Ground Cuckoo and Storm’s Stork

Oriental Bay Owl and Large Frogmouth

A few more hours in the forest next morning add the amazing vocalisation of Bornean Black Magpie, Crimson and Van Hasselt’s Sunbird, Scarlet-backed Flowerpecker, Fiery Minivet, Indian Cuckoo, Grey-and-buff, Buff-rumped and and Sunda Woodpeckers. Black Hornbills were appreciated but they were outshone by a flyover from an impressive pair of Rhinoceros Hornbills. Next stop for the day was Gomantong Caves where the birding began even before we parked with White-crowned Hornbill twice in flight over the road and Banded Bay Cuckoo seen well. A Bat Hawk made a brief flyover, presumably one of the pair that we found at a nest a short time later. The forest here gave us Rufous-backed Kingfisher, a stunning male Scarlet-rumped Trogon, White-crowned Shama and a small feeding group that included Spotted Fantail. Even better though was a male Orangutan that appeared out of a Jack Fruit tree with two large fruits stuffed in his mouth to keep all hands and feet free while he found a comfortable place to sit to scoff them down, as we watched from just a few meters away. He was later joined in the same tree by a younger Orangutan and there was also a female with a baby very nearby. The swiftlets were never going to compete but we did find all four species at their nests including the pure white nests of the Edible-nest Swiftlets which demand such a high price on the Chinese market. We were also lucky to see the collection in full swing with collectors suspended 50 meters above at the roof of the cave.

Malaysian Hawk Cuckoo and Rhinoceros Hornbill It was then time to make our way to our lodge on the banks of the mighty Kinabatangan River and even the short boat ride into our lodge produced our first Lesser Adjutants circling menacingly overhead. Back out on the river again in the afternoon we easily added one of the river’s main specialities with several Storm’s Storks perched up, and not long later a group of handsome Wrinkled Hornbills. Grey-headed Fish Eagle, Rufous-bellied Eagle, many Oriental Pied and Black Hornbills, Brown-backed Needletails and the prehistoric-looking Oriental Darter were all seen and we found our first groups of the big-nose, pot-bellied Proboscis Monkeys. We stayed out after dark and were quickly rewarded with an extremely close encounter with the immense Large Frogmouth and not long later an Oriental Bay Owl gave walkaway views. Buffy Fish Owl finished things off nicely although we couldn’t track down a calling Sunda Frogmouth. We had one big aim for the next morning, the enigmatic Bornean Ground Cuckoo, but although we heard at least three they weren’t very responsive and although one did come very close on one occasion we failed to catch even a glimpse before they became quiet. There were still plenty of nice birds to see though with a male Olive-backed Woodpecker a surprise find, a single Blue-rumped Parrot, yet more Storm’s Storks, a group of Bushy-crested Hornbill, Bold-striped Tit Babbler, Changeable Hawk Eagle and a stunning pair of Ruddy Kingfisher. In the afternoon we headed further downriver and this proved fruitful with two pairs of White-fronted Falconet, the first very far, but the second pair were very close by. A Large Frogmouth sat on a nest was a surprise treat, we found our first Malaysian Blue Flycatcher, and a confiding Lesser Fish Eagle beside yet more entertainment from the Proboscis Monkeys. A magnificent pair of Rhinoceros Hornbills were the first to greet us on our final morning, crossing the river before perching nicely, and not longer later a pair of Jerdon’s Baza. In a quiet tributary a Malaysian Hawk Cuckoo posed nicely, followed by a large gathering of hornbills raining the nearby oil palm nuts which included some Wrinkled Hornbills and after a long wait a pair of calling White-crowned Hornbills popped up for prolonged views. Back out on the main river it was already heating up when the Bornean Ground Cuckoos started calling in the same area as the previous day. This time though we were much more successful as one was teased closer and closer to the bank and finally hopped into a tree where it’s wagging tail betrayed its presence and other body parts could gradually be located through the foliage. Already happy at having seen the bird albeit typically skulking we were stunned when it hopped up further to give completely unobscured views before retreating slightly but still giving us fabulous views of the bright blue eye-patch, intricate barring on the underparts and blue-violet iridescence on

the wings and tail. In all we were able to watch it for more than 30 minutes and it’s no surprise it scored bird-of the-trip! If this great success under the belt it was time to depart the river and head for our final destination but this was no hardship as we were headed for the Danum Valley Conservation Area, an extensive area of lowland rainforest which can be explored from the fabulous Borneo Rainforest Lodge, our home for the next four nights. On the drive in we made some stops which included eye-level Whiskered Treeswifts, but it was our last stop where we struck lucky with one of the biggest targets of the trip; a group of Bristleheads cavorting in the nearby treetops, uttering their weird vocalisations as they foraged and posed in the ‘scope for as long as we wanted. We arrived at the lodge in time to get first views of a group of Bornean Crested Firebacks stalking through the gardens and later roosting above our cabins. At night we took our first of three night-drives finding some nice mammals some of which were regular features like Lesser Mouse Deer, Red Giant, Thomas’s and Black Flying Squirrels, a brief Bornean Loris and Small-toothed Palm Civet. On two nights we enjoyed nice views of a Blyth’s Frogmouth and on another a magnificent Barred Eagle Owl.

Bornean Banded Pitta and Black-crowned Pitta

Bristlehead and Hooded Pitta Our first full day in the forest was as quiet as predicted in the very dry El Niño induced conditions but we were rewarded for our efforts by a singing Black-throated Wren Babbler which proved quite viewable once we realised it was up in the canopy and not near the ground as they usually are! Bornean Banded Pitta eventually cooperated for the whole group, Bornean Blue Flycatchers were very vocal with several nice views, a Grey-chested Jungle

Flycatcher gave very close looks and we found two Large-billed Blue Flycatchers, very much a Bornean speciality being rarely seen in Sumatra. A female ‘Bornean’ Banded Kingfisher was ‘scoped high in the canopy as a nice insurance for this recently proposed split and down in the understorey an inquisitive Chestnut-necklaced Partridge was twice encourage to parade across the trail in front of us. There were plenty of more widespread species of course and these included Scarlet-rumped and Diard’s Trogon, Spotted Fantail, Buff-necked, Maroon and Rufous Woodpeckers, Chestnut-backed Scimitar Babbler, Chestnut-rumped Babbler and a manic group of Dusky Broadbills. The gardens of the lodge were typically productive even during the midday with Bornean Spiderhunter on the flowers, our first of several Yellow-rumped Flowerpeckers and fruiting trees with a variety of bulbuls including the attractive Grey-bellied Bulbul which kept us company each lunch. In the afternoon a tame female Blue-headed Pitta gave extended views, a Little Bronze Cuckoo was located high in the canopy, White-crowned Hornbills sat exposed on a far hill as they belted out their territorial calls and Red-throated Barbet was found from the canopy walkway. We finished the day in a colourful fashion with a roosting male Blue-headed Pitta after the night-drive.

Barred Eagle Owl and Blyth’s Frogmouth

Yellow-rumped Flowerpecker and Chestnut-necklaced Partridge Another morning of deafening cicadas was bit frustrating but we had two noisy groups of Bushy-crested Hornbills, several Rhinoceros Hornbills and two groups of Helmeted Hornbills passing over. Both Striped Wren Babbler and Short-tailed Babbler were new and there was another Bornean Banded Pitta to enjoy. Over lunch a Great-billed Heron stalked into view on the river below the restaurant and an excellent afternoon included Chestnut-naped

Forktail on the trickle that remained of its favourite river, Red-naped Trogon, Banded Broadbill, a singing male White-tailed Blue Flycatcher, Violet Cuckoo and those who rushed up to the viewpoint to enjoy the late afternoon view were treated to eye-level views of both Rhinoceros and Helmeted Hornbills in a fruiting tree. Horsfield’s Wren Babbler was our first new species on our last full day although a flock of 16 Pygmy White-eyes in a flowering tree was impressive. Next followed Ferruginous Babblers which had been unusually quite until now, then an obliging pair of Black-capped Babblers sneaking around on the ground. Another Bornean Banded Pitta was seen before finally, after two days of trying, we finally heard the sweet double whistle of a Bornean Ground Babbler and within a few minutes we were enjoying enthralling views of this delightfully striped endemic calling away below us. Not a bird but equally thrilling was a cryptically patterned Sunda Colugo clinging to a tree truck near the path, then Helmeted, Rhinoceros and Bushy-crested Hornbills from a viewpoint where a Grey-headed Fish Eagle was nesting far below. A Grey-headed Babbler was particularly appreciated by those who hadn’t done the long trek at Poring and a male Rufous-collared Kingfisher came in and showed nicely. Some roadside birding made a pleasant change in the afternoon and we found Yellow-crowned Barbet besides some other commoner special like Bronzed Drongo and Large Woodshrike which made their first appearance on the trip list. There were few new birds possible by our last morning but we did have more views of Chestnut-naped Forktail, a pair of White-tailed Blue Flycatchers with an interestingly plumaged immature male, a Cinnamon-rumped Trogon sadly didn’t stick around with only flight views as it continued on its way but a White-necked Babbler performed superbly, and before departing we had fine views of Great-billed Heron feeding to the river in front of the lodge to bring the tour to a nice end. We finished the tour with a total of 311 species recorded which included 10 species heard only and 47 amazing Borneo endemics. For information regarding our tours to Borneo please click here. Alternatively please contact us via e-mail or phone +44 1332 516254 Bird of the trip 1 = Bornean Ground Cuckoo = Oriental Bay Owl 2 = Bristlehead 3 = Whitehead’s Broadbill 4 = Barred Eagle Owl 5 = Red-bearded Bee-eater

Other tour photos

Blyth’s Hawk Eagle and Black-and-yellow Broadbill

Sunda Bush Warbler and Eye-browed Jungle Flycatcher

Bornean Forktail and White-browed Shortwing

Bornean Whistling Thrush and Bornean Stubtail

Bornean Whistler and Mountain Scops Owl

Red-bellied Sculptor Squirrel and Mountain Wren Babbler

Pygmy White-eye and Sunda Laughingthrush

Black-and-red Broadbill and Blue-eared Kingfisher

Bold-striped Tit Babbler and Buff-vented Bulbul

Oriental Bay Owl and Buffy Fish Owl

Lesser Fish Eagle and Oriental Darter

Proboscis Monkey and White-fronted Falconet

Wrinkled Hornbill and White-crowned Hornbill

Blue-headed Pitta (female & roosting male)

Bornean Blue Flycatcher and Large-billed Blue Flycatcher

Dusky Broadbill and Grey-bellied Bulbul

Buff-necked Woodpecker and Velvet-fronted Nuthatch

Grey-chested Jungle Flycatcher and Grey-headed Canary Flycatcher

Orange-bellied Flowerpecker and Red-throated Barbet

Whiskered Treeswift and Rufous-collared Kingfisher

White-tailed Blue Flycatcher (Immature & adult)

Great-billed Heron and Bornean Crested Fireback

Sunda Colugo and Rhinoceros Hornbill

Bornean Swiftlet and Kinabalu Squirrel

Systematic List GALLIFORMES: Phasianidae

Great Argus Argusianus argus

Heard only Red-breasted Partridge Arborophila hyperythra e

Chestnut-necklaced Partridge Tropicoperdix charltonii graydoni Crimson-headed Partridge Haematortyx sanguiniceps e

Bornean [Crested] Fireback Lophura ignita nobilis e

CICONIIFORMES: Ciconiidae

Storm's Stork Ciconia stormi Lesser Adjutant Leptoptilos javanicus

CICONIIFORMES: Ardeidae

Great-billed Heron Ardea sumatrana Purple Heron Ardea purpurea Eastern Great Egret Ardea modestus Intermediate Egret Ardea intermedia intermedia Eastern Cattle Egret Ardea coromandus Little Egret Egretta garzetta Striated Heron Butorides striata Black-crowned Night Heron Nycticorax nycticorax Yellow Bittern Ixobrychus sinensis

PELECANIFORMES: Anhingidae

Oriental Darter Anhinga melanogaster

FALCONIFORMES: Accipitridae

Jerdon's Baza Aviceda jerdoni Oriental Honeybuzzard Pernis ptilorhynchus torquatus Bat Hawk Macheiramphus alcinus alcinus Brahminy Kite Haliastur indus White-bellied Fish Eagle Ichthyophaga leucogaster Lesser Fish Eagle Ichthyophaga humilis Grey-headed Fish Eagle Ichthyophaga ichthyaetus Mountain Serpent Eagle Spilornis kinabaluensis e

Crested Serpent Eagle Spilornis cheela Crested Goshawk Tachyspiza trivirgatus Black Eagle Ictinaetus malayensis Rufous-bellied Eagle Lophotriorchis kienerii Changeable Hawk Eagle Nisaetus limnaetus Blyth's Hawk Eagle Nisaetus alboniger Wallace's Hawk Eagle Nisaetus nanus

GRUIFORMES: Rallidae

White-breasted Waterhen Amaurornis phoenicurus

CHARADRIIFORMES: Scolopacidae

Wood Sandpiper Tringa glareola Common Sandpiper Actitis hypoleucos

CHARADRIIFORMES: Laridae

Whiskered Tern Chlidonias hybridus

COLUMBIFORMES: Columbidae

Spotted Dove Streptopelia chinensis Little Cuckoo Dove Macropygia ruficeps nana Asian Emerald Dove Chalcophaps indica indica

Zebra Dove Geopelia striata I Little Green Pigeon Treron olax

Thick-billed Green Pigeon Treron curvirostra Green Imperial Pigeon Ducula aenea Mountain Imperial Pigeon Ducula badia

CUCULIFORMES: Cuculidae

Bock's Hawk Cuckoo Hierococcyx [sparverioides] bocki

Heard only Malaysian Hawk Cuckoo Hierococcyx fugax

Himalayan / Oriental Cuckoo Cuculuc saturatus / optatus Sunda Cuckoo Cuculus lepidus Indian Cuckoo Cuculus micropterus Banded Bay Cuckoo Cacomantis sonneratii Plaintive Cuckoo Cacomantis merulinus Little Bronze Cuckoo Chrysococcyx minutillus aheneus Violet Cuckoo Chrysococcyx xanthorhynchus Asian Drongo Cuckoo Surniculus lugubris brachyurus Raffles's Malkoha Rhinortha chlorophaeus Chestnut-breasted Malkoha Phaenicophaeus curvirostris Bornean Ground Cuckoo Carpococcyx radiatus e

Short-toed Coucal Centropus rectunguis

Heard only Greater Coucal Centropus sinensis bubutus

STRIGIFORMES: Tytonidae

Oriental Bay Owl Phodilus badius badius

STRIGIFORMES: Strigidae

Mountain Scops Owl Otus spilocephalus luciae Barred Eagle Owl Bubo sumatranus Buffy Fish Owl Bubo ketupu Brown Wood Owl Strix leptogrammica vaga Brown Boobook Ninox scutulata borneensis 'Sunda' Collared Owlet Glaucidium [brodiei] borneense

CAPRIMULGIFORMES: Podargidae

Large Frogmouth Batrachostomus auritus Blyth's Frogmouth Batrachostomus affinis affinis Sunda Frogmouth Batrachostomus cornutus

Heard only

CAPRIMULGIFORMES: Caprimulgidae

Malaysian Eared Nightjar Lyncornis temminckii

APODIFORMES: Hemiprocnidae

Grey-rumped Treeswift Hemiprocne longipennis harterti Whiskered Treeswift Hemiprocne comata comata

APODIFORMES: Apodidae

Giant Swiftlet Hydrochous gigas Glossy Swiftlet Collocalia esculenta cyanoptila Bornean Swiftlet Collocalia dodgei e

Mossy-nest Swiftlet Aerodramus salangana natunae Black-nest Swiftlet Aerodramus maximus lowi Edible-nest Swiftlet Aerodramus fuciphagus vestita Silver-rumped Spinetail Rhaphidura leucopygialis Brown-backed Needletail Hirundapus giganteus House Swift Apus [affinis] nipalensis

TROGONIFORMES: Trogonidae

Red-naped Trogon Harpactes kasumba impavidus Diard's Trogon Harpactes diardii diardii Whitehead's Trogon Harpactes whiteheadi e

Scarlet-rumped Trogon Harpactes duvaucelii Orange-breasted Trogon Harpactes oreskios dulitensis

CORACIIFORMES: Coraciidae

Dollarbird Eurystomus orientalis

CORACIIFORMES: Alcedinidae

Common Kingfisher Alcedo atthis bengalensis Blue-eared Kingfisher Alcedo meninting verreauxii Rufous-backed Kingfisher Ceyx rufidorsa motleyi 'Bornean Banded Kingfisher' Lacedo pulchella melanops Stork-billed Kingfisher Pelargopsis capensis inominata Ruddy Kingfisher Halcyon coromanda minor Collared Kingfisher Todirhamphus chloris laubmannianus Rufous-collared Kingfisher Actenoides concretus borneanus

CORACIIFORMES: Meropidae

Red-bearded Bee-eater Nyctyornis amictus Blue-throated Bee-eater Merops viridis viridis

CORACIIFORMES: Bucerotidae

Oriental Pied Hornbill Anthracoceros albirostris convexus Black Hornbill Anthracoceros malayanus Rhinoceros Hornbill Buceros rhinoceros borneoensis Helmeted Hornbill Rhinoplax vigil Bushy-crested Hornbill Anorrhinus galeritus White-crowned Hornbill Berenicornis comatus Wrinkled Hornbill Aceros corrugatus Wreathed Hornbill Rhyticeros undulatus

PICIFORMES: Capitonidae

Gold-whiskered Barbet Psilopogon chrysopogon chrysopsis Red-throated Barbet Psilopogon mystacophanos Mountain Barbet Psilopogon monticola e

Yellow-crowned Barbet Psilopogon henricii Golden-naped Barbet Psilopogon pulcherrima e

Blue-eared Barbet Psilopogon australis duvaucelii Bornean Barbet Psilopogon eximia e

Bornean Brown Barbet Calorhamphus tertius e

PICIFORMES: Picidae

Sunda Woodpecker Dendrocopos moluccensis Rufous Woodpecker Celeus brachyurus badiosus Banded Woodpecker Chrysophlegma mineaceus malaccensis Crimson-winged Woodpecker Chrysophlegma puniceus observandus Checker-throated Woodpecker Chrysophlegma mentalis humii Olive-backed Woodpecker Dinopium rafflesii dulitense Maroon Woodpecker Blythipicus rubiginosus Orange-backed Woodpecker Reinwardtipicus validus xanthopygius Buff-rumped Woodpecker Meiglyptes tristis grammithorax Buff-necked Woodpecker Meiglyptes tukki tukki Grey-and-buff Woodpecker Hemicircus concretus sordidus Great Slaty Woodpecker Mulleripicus pulverulentus

Heard only

FALCONIFORMES: Falconidae

White-fronted Falconet Microhierax latifrons e

Peregrine Falcon Falco peregrinus ernesti

PSITTACIFORMES: Psittacidae

Blue-rumped Parrot Psittinus cyanurus Long-tailed Parakeet Psittacula longicauda Blue-crowned Hanging Parrot Loriculus galgulus

PASSERIFORMES:Calyptomenidae

Green Broadbill Calyptomena viridis Hose's Broadbill Calyptomena hosii e

Whitehead's Broadbill Calyptomena whiteheadi e

PASSERIFORMES: Eurylaimidae

Dusky Broadbill Corydon sumatranus Black-and-red Broadbill Cymbirhynchus macrorhynchos Banded Broadbill Eurylaimus javanicus Black-and-yellow Broadbill Eurylaimus ochromalus Long-tailed Broadbill Psarisomus dalhousiae

PASSERIFORMES: Pittidae

Giant Pitta Hydrornis caerulea hoesi

Heard only Bornean Banded Pitta Hydrornis schwaneri e

Blue-headed Pitta Hydrornis baudii e Hooded Pitta Erythropitta sordida mulleri

Blue-banded Pitta Erythropitta arquata e Heard only Black-crowned Pitta Erythropitta ussheri e

PASSERIFORMES: Acanthizidae

Golden-bellied Gerygone Gerygone sulphurea sulphurea

PASSERIFORMES: Tephrodornithidae

Bar-winged Flycatcher-shrike Hemipus picatus intermedius Black-winged Flycatcher-shrike Hemipus hirundinaceus Large Woodshrike Tephrodornis gularis frenatus Rufous-winged Philentoma Philentoma pyrhopterum Maroon-breasted Philentoma Philentoma velatum

Heard only

PASSERIFORMES: Pityriaseidae

Bristlehead Pityriasis gymnocephala e

PASSERIFORMES: Artamidae

White-breasted Woodswallow Artamus leucorynchus

PASSERIFORMES: Aegithinidae

Common Iora Aegithina tiphia aequanimis Green Iora Aegithina viridissima

PASSERIFORMES: Campephagidae

Sunda Cuckooshrike Coracina larvata normani Pied Triller Lalage nigra nigra Fiery Minivet Pericrocotus igneus igneus Scarlet Minivet Pericrocotus speciosus insulanus Grey-chinned Minivet Pericrocotus solaris cinereigula

PASSERIFORMES: Pachycephalidae

Bornean Whistler Pachycephala hypoxantha e

PASSERIFORMES: Laniidae Long-tailed Shrike Lanius schach

Crested Jay Platylophus galericulatus coronatus

PASSERIFORMES: Vireonidae

Blyth's Shrike Babbler Pteruthius aeralatus robinsoni Erpornis Erpornis zantholeuca

PASSERIFORMES: Oriolidae

Dark-throated Oriole Oriolus xanthonotus consobrinus Black-and-crimson Oriole Oriolus cruentus vulneratus

PASSERIFORMES: Dicruridae

Sunda [Ashy] Drongo Dicrurus leucophaeus stigmatops Bronzed Drongo Dicrurus aeneus malayensis Hair-crested Drongo Dicrurus [hottentottus] borneensis Greater Racket-tailed Drongo Dicrurus paradiseus brachyphorus

PASSERIFORMES: Rhipiduridae

White-throated Fantail Rhipidura albicollis kinabalu Pied Fantail Rhipidura javanica Spotted Fantail Rhipidura perlata

PASSERIFORMES: Monarchidae

Black-naped Monarch Hypothymis azurea prophata Asian Paradise Flycatcher Terpsiphone paradisi borneensis

PASSERIFORMES: Corvidae

Bornean Black Magpie Platysmurus aterrimus e Bornean Green Magpie Cissa jefferyi e Bornean Treepie Dendrocitta cinerascens e Slender-billed Crow Corvus enca compilator

PASSERIFORMES: Stenostiridae

Grey-headed Canary Flycatcher Culicicapa ceylonensis

PASSERIFORMES: Pycnonotidae

Black-headed Bulbul Pycnonotus atriceps atriceps Bornean Bulbul Pycnonotus montis e

Scaly-breasted Bulbul Pycnonotus squamatus borneensis Grey-bellied Bulbul Pycnonotus cyaniventris Puff-backed Bulbul Pycnonotus eutilotus Pale-faced Bulbul Pycnonotus [flavescens] leucops e

Yellow-vented Bulbul Pycnonotus goiavier gourdini Olive-winged Bulbul Pycnonotus plumosus plumosus Cream-vented Bulbul Pycnonotus simplex perplexus Red-eyed Bulbul Pycnonotus brunneus brunneus Spectacled Bulbul Pycnonotus erythropthalmos Ochraceous Bulbul Alophoixus ochraceus ruficrissus Grey-cheeked Bulbul Alophoixus bres gutturalis Yellow-bellied Bulbul Alophoixus phaeocephalus Hairy-backed Bulbul Tricholestes criniger Buff-vented Bulbul Iole olivacea charlottae Cinereous Bulbul Hemixos cinereus connectens

PASSERIFORMES: Hirundinidae

Barn Swallow Hirundo rustica gutturalis Pacific Swallow Hirundo tahitica

Asian House Martin Delichon dasypus

PASSERIFORMES: Cettidae

Yellow-bellied Warbler Abroscopus superciliaris schwaneri Bornean Stubtail Urosphena whiteheadi e

Aberrant (Sunda) Bush Warbler Horornis flavolivaceus oreophila Mountain Tailorbird Phyllergates cuculatus cinereicollis

PASSERIFORMES: Phylloscopidae

Mountain Leaf Warbler Phylloscopus trivirgatus kinabaluensis Yellow-breasted Warbler Seicercus montis floris

PASSERIFORMES: Locustellidae

Striated Grassbird Megalurus palustris

PASSERIFORMES: Cisticolidae

Dark-necked Tailorbird Orthotomus atrogularis atrogularis Rufous-tailed Tailorbird Orthotomus sericeus sericeus Ashy Tailorbird Orthotomus ruficeps borneoensis Yellow-bellied Prinia Prinia flaviventris latrunculus

PASSERIFORMES: Timaliidae

Bold-striped Tit Babbler Mixornis bornensis Fluffy-backed Tit Babbler Macronous ptilosus reclusus Chestnut-winged Babbler Cyanoderma erythropterum bicolor Rufous-fronted Babbler Cyanoderma rufifrons sarawacensis Chestnut-backed Scimitar Babbler Pomatorhinus montanus bornensis Grey-throated Babbler Stachyris nigriceps borneensis Grey-headed Babbler Stachyris poliocephala White-necked Babbler Stachyris leucotis obscurata Black-throated Babbler Stachyris nigricollis

Heard only Chestnut-rumped Babbler Stachyris maculata maculata

PASSERIFORMES: Pellorneidae

Moustached Babbler Malacopteron magnirostre cinereocapilla Sooty-capped Babbler Malacopteron affine phoeniceum Scaly-crowned Babbler Malacopteron cinereum cinereum Rufous-crowned Babbler Malacopteron magnum saba White-chested Babbler Pellorneum rostratum macropterum Ferruginous Babbler Pellorneum bicolor Short-tailed Babbler Pellorneum malaccense poliogenys Temminck's Babbler Pellorneum pyrrogenys canicapillus Black-capped Babbler Pellorneum capistratum morrelli Striped Wren Babbler Kenopia striata Bornean Ground Babbler Ptilocichla leucogrammica e

Horsfield's Wren Babbler Turdinus sepiarius harterti Black-throated Wren Babbler Turdinus atrigularis e

Mountain Wren Babbler Turdinus crassus e

PASSERIFORMES: Leiothrichidae Brown Fulvetta Alcippe brunneicauda

Sunda Laughingthrush Garrulax palliatus Bare-headed Laughingthrush Garrulax calvus e

Rufous-hooded Laughingthrush Ianthocincla treacheri treacheri e

PASSERIFORMES: Zosteropidae

Black-capped White-eye Zosterops atricapillus Everett's White-eye Zosterops everetti Mountain Black-eye Zosterops emiliae e

Pygmy White-eye Heleia squamifrons e

Chestnut-crested Yuhina Yuhina everetti e

PASSERIFORMES: Sittidae

Velvet-fronted Nuthatch Sitta frontalis corallipes

PASSERIFORMES: Sturnidae

Asian Glossy Starling Aplonis panayensis Common Hill Myna Gracula religiosa Crested Myna Acridotheres cristatellus I

Javan Myna Acridotheres javanicus I

PASSERIFORMES: Turdidae

Orange-headed Thrush Geokichla citrina aurata Everett's Thrush Geokichla everetti e

Eye-browed Thrush Turdus obscurus Fruit-hunter Chlamydochaera jefferyi e

PASSERIFORMES: Muscicapidae

Oriental Magpie Robin Copsychus saularis adamsi White-crowned Shama Copsychus stricklandii e

Rufous-tailed Shama Copsychus pyrropyga Dark-sided Flycatcher Muscicapa sibirica Asian Brown Flycatcher Muscicapa dauurica dauurica Narcissus Flycatcher Ficedula narcissina narcissina Mugimaki Flycatcher Ficedula mugimaki Taiga Flycatcher Ficedula albicilla Snowy-browed Flycatcher Ficedula hyperythra sumatrana Little Pied Flycatcher Ficedula westermanni Blue-and-white Flycatcher Cyanoptila cyanomelana Verditer Flycatcher Eumyias thalassina Indigo Flycatcher Eumyias indigo Grey-chested Jungle Flycatcher Cyornis umbratilis White-tailed Blue Flycatcher Cyornis concretus everetti Pale Blue Flycatcher Cyornis unicolor herterti Hill Blue Flycatcher Cyornis [banyumas] coeruleatus Large-billed Blue Flycatcher Cyornis caerulatus caerulatus Malaysian Blue Flycatcher Cyornis turcosus Bornean Blue Flycatcher Cyornis superbus e

White-browed Shortwing Brachypteryx montana erythrogyna Eye-browed Jungle Flycatcher Vauriella gularis e

Chestnut-naped Forktail Enicurus ruficapillus White-crowned Forktail Enicurus leschenaulti frontalis Bornean Forktail Enicurus borneensis e

Bornean Whistling Thrush Myophonus borneensis e

PASSERIFORMES: Chloropseidae

Greater Green Leafbird Chloropsis sonnerati zosterops Lesser Green Leafbird Chloropsis cyanopogon cyanopogon Bornean Leafbird Chloropsis kinabaluensis e

PASSERIFORMES: Dicaeidae

Yellow-breasted Flowerpecker Prionochilus maculatus maculatus Yellow-rumped Flowerpecker Prionochilus xanthopygius e

Yellow-vented Flowerpecker Dicaeum chrysorrheum Orange-bellied Flowerpecker Dicaeum trigonostigma dayakanum Black-sided Flowerpecker Dicaeum monticolum e

Scarlet-backed Flowerpecker Dicaeum cruentatum nigrimentum

PASSERIFORMES: Nectariniidae

Ruby-cheeked Sunbird Chalcoparia singalensis Plain Sunbird Anthreptes simplex Brown-throated Sunbird Anthreptes malacensis Red-throated Sunbird Anthreptes rhodolaema Olive-backed Sunbird Cinnyris jugularis Copper-throated Sunbird Leptocoma calcostetha Van Hasselt's Sunbird Leptocoma brasiliana Crimson Sunbird Aethopyga siparaja Temminck's Sunbird Aethopyga temminckii Purple-naped Spiderhunter Hypogramma hypogrammicum Thick-billed Spiderhunter Arachnothera crassirostris Spectacled Spiderhunter Arachnothera flavigaster Long-billed Spiderhunter Arachnothera robusta

Heard only Little Spiderhunter Arachnothera longirostra

Yellow-eared Spiderhunter Arachnothera chrysogenys Bornean Spiderhunter Arachnothera everetti e

Whitehead's Spiderhunter Arachnothera juliae e

PASSERIFORMES: Passeridae

Eurasian Tree Sparrow Passer montanus malaccensis

PASSERIFORMES: Estrildidae

Dusky Munia Lonchura fuscans e White-bellied Munia Lonchura leucogastra smythiesi

Black-headed Munia Lonchura atricapilla jagori

PASSERIFORMES: Motacillidae

Grey Wagtail Motacilla cinerea cinerea Paddyfield Pipit Anthus rufulus

MAMMALS Mountain Treeshrew Tupaia montana baluensis e

Lesser Treeshrew Tupaia minor minor Large Treeshrew Tupaia tana paitana Sunda Colugo Cynocephalus variegatus Bornean Loris Nycticebus borneanus e

Maroon Leaf Monkey Presbytis rubicunda e Sunda Silvered Leaf Monkey Presbytis cristata

Proboscis Monkey Nasalis larvatus e Long-tailed Macaque Macaca fascicularis

Sunda Pig-tailed Macaque Macca nemestrina Bornean Gibbon Hylobates muelleri e Heard only

Bornean Orang-utan Pongo pygmaeus e 'Bornean' Giant Squirrel Ratufa affinis

Prevost's Squirrel Callosciurus prevostii pluto Kinabalu Squirrel Callosciurus baluensis e

Plantain Squirrel Callosciurus notatus Bornean Black-banded Squirrel Callosciurus orestes e

Low's Squirrel Sundasciurus lowii Jentink's Squirrel Sundasciurus jentinki e

Red-bellied Sculptor Squirrel Glyphotes simus e Four-striped Ground Squirrel Lariscus hosei e Bornean Mountain Ground Squirrel Dremomys everetti e Plain Pygmy Squirrel Exilisciurus exilis e Black Flying Squirrel Aeromys tephromelas

Thomas's Flying Squirrel Aeromys thomasi e Red Giant Flying Squirrel Petaurista petaurista

Small-toothed Palm Civet Arctogalidia trivirgata Short-tailed Mongoose Herpestes brachyurus

Collared Mongoose Herpestes semitorquatus Bearded Pig Sus barbatus Lesser Mouse-Deer Tragulus kanchil Sambar Deer Cervus unicolor

e = Species endemic to Borneo I = Introduced species