Sabah, Borneo

31st August – 13th September 2016

Leader: Dave Bakewell

Participants: Dennis Nick and Holly Faithfull

Everett’s Thrush, Mount Kinabalu © Dave Bakewell / Birdtour Asia Sabah, the north-eastern corner of Borneo, has a surfeit of attractions for anyone with an interest in the region’s natural history, whether it is the many stunning bird species (with over 50 endemic on the island), its mammals, or the amazing vistas, including the sunsets off Tanjung Aru, the towering Mount Kinabalu, the tranquil beauty of the Kinabatangan River and the verdant, vibrant hues of the lowland rainforest. This relaxed tour, aimed at encompassing a memorable selection of all three, also connected with such sought-after birds as Whitehead’s Trogon and Broadbill, Giant and Blue-banded Pitta, Everett’s Thrush and the unique Bristlehead among the over 260 species recorded (including 41 endemics); nine primates, five civets and 14 squirrels and flying squirrels were among the 45 mammal species (13 endemics) seen. The tour got of to a cracking start as we pulled off the road in the Crocker Range to get great views of a perched, calling Mountain Serpent Eagle, being mobbed by Temminck’s Sunbirds and Sunda Ashy Drongos. This was followed by point blank views of Sunda Bush Warbler, one of many belting out their car alarm songs, and a pair of Chestnut-crested Yuhinas busy collecting nest material. A few other montane birds put in appearances as the day wore one, including Bornean Bulbul, Mountain Barbet, and a Bornean Swiftlet, seen perched on the eaves of a restaurant among the more numerous nesting Glossy Swiftlets. After checking into our accommodation in the imposing shadow of Mount Kinabalu in late afternoon, a short walk produced excellent eye-level views of Pygmy Heleia to round off the day. The following day saw us in the foothills of majestic Mount Kinabalu, where we were greeted by fine clear weather. A couple of Mountain Black-eye were followed by views of Golden-naped Barbet, Pale-faced Bulbuls, a pair of ever-confiding Snowy-browed Flycatchers, and Bornean Whistling Thrush, while a trek down one of the

trails produced a Crimson-headed Partridge. After rain in the early afternoon, we were fortunate to find a pair of Everett’s Thrushes feeding by the roadside, oblivious to the passing traffic. These were the highlight of the afternoon, but we also had overflying Sunda Cuckoo, Blyth’s Hawk Eagle and Oriental Honey Buzzard, as well as a number of rodents, including Bornean Black-banded and Brooke’s Squirrels, Mountain Treeshrew and Lesser Gymnure. We made a pre-dawn start the next day to try our luck with Mountain Scops Owls. Despite getting tantalisingly close, no views were forthcoming. Once it got light we began connecting with some quality birds, including a pair of stunning emerald-green Whitehead’s Broadbills and a Bornean Stubtail pulsating with effort as it issued its near supersonic song. Hitting the trails again, we added Bornean Shortwing and Temminck’s Babbler before our second Whitehead’s of the morning - a pair of Trogons, including an adult male in his sunburn-inducing brilliance. The afternoon was a wash-out, so we went out after dinner to try for owls again, adding Sabah Giant Rat to our mammal list before finally tracking down a Mountain Scops Owl - a fitting reward for our perseverance.

Mountain Serpent Eagle and Pygmy Heleia

Whitehead’s Broadbill and Whitehead’s Trogon

The following day we opted for a slightly more relaxed pace, starting off in the Botanical Gardens, and enjoying the incredible diversity of mountain’s plant life showcased therein. It proved a good place to get close to some of the commoner montane birds; Yellow-breasted Warbler, Eyebrowed Jungle Flycatcher (though it’s a robin, and named Bornean Shade-dweller in the new field guide to the region) and Bornean Whistler among them. A Red-bellied Sculptor Squirrel was the highlight, and we also observed several Brooke’s Squirrels. Higher up the

mountain we were enshrouded in low cloud, but still managed point blank views of Jentink’s and Bornean Black-banded Squirrels, as well as a party of seven Red-breasted Partridges foraging below us on the slopes. Rain once again curtailed the afternoon’s planned activities.

Mountain Scops Owl and Red-breasted Partridge The next morning was taken up with the long drive east to Sepilok, and the journey was enlivened by our first few lowland species, notably Black-and-yellow Broadbill and Blue-crowned Hanging Parrot. Long-tailed Shrike is an increasingly commonly encountered species in Sabah these days; a very early migrant Brown Shrike was more of a surprise. At the Rainforest Discovery Centre, we enjoyed prolonged views of two Orang-utan mothers with their youngsters, as well as a fine crepuscular performance by several Red Giant Flying Squirrels. Wallace’s Hawk Eagles were in attendance as always, and we also encountered Green Ioras, Black-winged Flycatcher Shrikes and Fiery Minivets, as well as Black and Bushy-crested Hornbills.

Black-and-Yellow Broadbill and Black-and-Red Broadbill

Dawn the following morning saw us once again at the Rainforest Discovery Centre. Bird activity was rather low, with the highlights being several fly-past Cinnamon-headed Green Pigeons, Copper-throated Sunbird, displaying Crested Goshawk and Bornean Brown Barbets. The spectacular Gomantong Caves did not disappoint, despite a recent harvest of nests, and we were able to locate a few Edible-nest Swiftlets at nests among the more numerous Black-nest Swiftlets. A calling Short-toed Coucal declined to show itself, but a Black-and-red Broadbill performed to an appreciative audience. An afternoon trip along the Kinabatangan River produced a magnificent male Orang-utan, as well as several Proboscis Monkey troops, and one or two Silvered Langurs as the sun set. Black and Oriental Pied Hornbills, and a Jerdon’s Baza were the avian highlights.

We were on the river again before first light, but despite trawling the smaller tributaries extensively, there was no sign or sound of our main target - Bornean Ground Cuckoo. Compensations were to be had in the form of pairs of Storm’s Storks and Rhinoceros Hornbills, a reasonably close White-fronted Falconet, Hooded Pitta and Blue-eared Kingfisher. In the evening we watched a large troop of Proboscis Monkeys crossing the river, and after dark we had tremendous looks at an oddly-vocalising Large Frogmouth, Small-toothed Palm Civets and several Buffy Fish Owls.

Large Frogmouth and Hooded Pitta

Wrinkled Hornbill and White-crowned Hornbill Our last morning on the river turned up a few noteworthy additions, including a fine pair of White-crowned Hornbills, a basking Estuarine Crocodile and a Sacred Kingfisher, an uncommon Austral winter visitor to Borneo. Our transfer to Tabin Wildlife Reserve took up much of the middle of the day, but there was still time to go exploring in late afternoon. We were treated to a family party of five Great Slaty Woodpeckers, before chancing upon no fewer than three Giant Pittas calling in a small triangle of undergrowth. Some strategic positioning enabled close views of one of the birds, a female, in the fading light. After dark a night drive brought us views of Brown Wood Owl and Sunda Frogmouth, as well as Black, Thomas’s and Red Giant Flying Squirrels, Leopard Cat, Common Palm and Malay Civets. Our first full morning at Tabin began in the best possible way, with a family of five Bornean Gibbons calling in the trees above us. This was followed shortly afterwards by a pair of Helmeted Hornbills flying overhead, their maniacal laughing calls were a constant accompaniment throughout the morning, at the same time spectacular and poignant given the current severe threat facing the species at the hands of illegal ivory poachers – the

species has recently been uplisted to Critically Endangered. The three Giant Pittas were once again in full voice, and this time a male gave close but tantalisingly brief views. A calling Olive-backed Woodpecker obligingly flew into a nearby tree, and Whiskered Treeswifts were admired as they landed and shut their wings with clipped precision. Already in pitta heaven, we trekked up a steep slope to eventually get decent views of a resplendent Blue-banded Pitta, but our fortune ran out with a briefly calling Bornean Banded Pitta, which chose the thickest undergrowth imaginable to call from and refused to show. A splendid male Bornean Crested Fireback was some compensation. The afternoon was spent in relaxed mode, with just Black-capped Babbler and Rufous-winged Philentoma of note, before we set off on an extended night drive. Having been entertained by a Buffy Fish Owl hunting in mid-stream, we went on to see a pair of Bat Hawks and a good selection of mammals, of which Greater Mouse Deer was new for the trip. After an early breakfast, and a walk back to the chalet that was interrupted by a juvenile Wallace’s Hawk Eagle doing its best to grab a squirrel out of a thick bush, we were again serenaded by Bornean Gibbons, before visiting a mud volcano. Birds en route included White-crowned Forktail, Striped Wren Babbler, Short-tailed Babbler and Checker-throated Woodpecker, as well as Wreathed Hornbill, our eighth species for the trip. Lesser Mouse Deer and Sambar were also seen, but best of all were three Sabah Grey Langurs coming down to feed on the mineral-rich mud. Our departure from Tabin in the midst of a deluge was enlivened by a couple of Storm’s Storks feeding on the waterlogged road out. We arrived at Danum Valley just in time for a night drive, which added Binturong and Masked Palm Civet to our mammal list. Our first full day at Danum was very hard work. The canopy walkway, only part-open due to a lightning strike earlier in the year, produced a few bulbuls and always entertaining Black-and-yellow Broadbills, but the highlight was a close-feeding Plain Pygmy Squirrel. Lunchtime was enlivened by a Great-billed Heron and a Lesser Fish Eagle, and in the afternoon we added Black-throated Wren Babbler. A search for Blyth’s Frogmouth after dark produced three calling birds but none were responsive, and an aggressive ‘Bornean Wood Owl’ (a split from Brown) put paid to our chances of seeing a briefly calling Oriental Bay Owl.

North Bornean Gibbon and Leopard Cat A small troop of Red Langurs began proceedings the next morning, followed by three White-fronted Falconets warming up in the early morning rays. Another hunt for Bornean Banded Pitta produced a male Diard’s Trogon, and on our way back to the lodge we spent some time admiring a pregnant female Orang-utan just waking up from a snooze. A pair of Wallace’s Hawk Eagles flushed a Yellow-throated Marten which ran vertically down a tree-trunk, leaving us wondering how on earth it could do that without falling. A roosting red morph Colugo was appreciated, and further highlights were a Brown Wood Owl flushed from a day roost while we were admiring a Black-headed Pitta, a large male Orang-utan and a couple of male Bornean Crested Firebacks walking to roost. Our penultimate day in the field began by dividing our attentions between the river and the steeper slopes above the lodge. On the latter, Cinnamon-rumped Trogon and White-necked Babbler were a good return for the laborious uphill climb. A Scarlet-rumped Trogon and Malaysian Hawk Cuckoo were seen in the afternoon, before a deluge curtailed the rest of the day’s activities. Our last morning started early, and as we watched a fruiting tree attracting a variety of birds, we were surprised to hear a Blyth’s Frogmouth calling softly in broad daylight. It bamboozled all attempts to locate it until Holly brilliantly spotted the bird, a male, sitting on a nest about 30 feet up. Having enjoyed this late bonus, we then ran into a mixed flock of Dusky Broadbills, Bornean Black Magpies and at least 10 Bristleheads. The latter are developing a bit of a reputation on our tours as ‘last day birds’, but we did not complain as the

foraging flock gave us prolonged views. Our last addition was a pair of Bornean Ground Babblers, which gave excellent views down to a few metres. Bird of the Tour

1. Blue-banded Pitta 2. Giant Pitta 3. Bristlehead 4. Black-and-Yellow Broadbill 5. Helmeted Hornbill

For information regarding our tours to Malaysia please click here. Alternatively please contact us via e-mail or phone +44 1332 516254 regarding organising a custom tour to Malaysia. More photos from the tour

Black-throated Wren Babbler and Great-billed Heron

Bornean Crested Fireback and Brown Wood Owl

Blyth’s Frogmouth and Bornean Ground Babbler

Wallace’s Hawk Eagle and Buffy Fish Owl

Pale-faced Bulbul and Sunda Bush Warbler

Indigo Flycatcher and Whitehead’s Trogon

Bornean Orang-utan and Proboscis Monkey

Black Hornbill and Helmeted Hornbill

Stork-billed Kingfisher and Sacred Kingfisher

Lesser Fish Eagle and Red Langur

Red Giant Flying Squirrel and Plain Pygmy Squirrel

Sambar and Bristlehead

Lesser Green Leafbird and Draco cornatus

Systematic List

GALLIFORMES: Phasianidae Great Argus Argusianus argus heard only Red-breasted Partridge Arborophila hyperythra e Chestnut-necklaced Partridge Tropicoperdix charltonii graydoni Crimson-headed Partridge Haematortyx sanguiniceps e Bornean Crested Fireback Lophura ignita nobilis e COLUMBIFORMES: Columbidae Spotted Dove Streptopelia chinensis Little Cuckoo Dove Macropygia ruficeps nana Asian Emerald Dove Chalcophaps indica indica Zebra Dove Geopelia striata I Little Green Pigeon Treron olax Pink-necked Green Pigeon Treron vernans Cinnamon-headed Green Pigeon Treron fulvicollis Large Green Pigeon Treron capellei

Green Imperial Pigeon Ducula aenea Mountain Imperial Pigeon Ducula badia CUCULIFORMES: Cuculidae Bock's Hawk Cuckoo Hierococcyx [sparverioides] bocki heard only Moustached Hawk Cuckoo Hierococcyx vagans Malaysian Hawk Cuckoo Hierococcyx fugax Sunda Cuckoo Cuculus lepidus Indian Cuckoo Cuculus micropterus Banded Bay Cuckoo Cacomantis sonneratii Plaintive Cuckoo Cacomantis merulinus Violet Cuckoo Chrysococcyx xanthorhynchus Asian Drongo Cuckoo Surniculus lugubris brachyurus Raffles's Malkoha Rhinortha chlorophaeus Chestnut-breasted Malkoha Phaenicophaeus curvirostris Short-toed Coucal Centropus rectunguis heard only Greater Coucal Centropus sinensis bubutus heard only Lesser Coucal Centropus bengalensis javanensis APODIFORMES: Hemiprocnidae Grey-rumped Treeswift Hemiprocne longipennis harterti Whiskered Treeswift Hemiprocne comata comata APODIFORMES: Apodidae Glossy Swiftlet Collocalia esculenta cyanoptila Bornean Swiftlet Collocalia dodgei e Mossy-nest Swiftlet Aerodramus salangana natunae Black-nest Swiftlet Aerodramus maximus lowi Edible-nest Swiftlet Aerodramus fuciphagus vestita Silver-rumped Spinetail Rhaphidura leucopygialis Brown-backed Needletail Hirundapus giganteus House Swift Apus [affinis] nipalensis CAPRIMULGIFORMES: Podargidae Large Frogmouth Batrachostomus auritus Blyth's Frogmouth Batrachostomus affinis affinis Sunda Frogmouth Batrachostomus cornutus GRUIFORMES: Rallidae White-breasted Waterhen Amaurornis phoenicurus CHARADRIIFORMES: Scolopacidae Common Sandpiper Actitis hypoleucos PELECANIFORMES: Anhingidae Oriental Darter Anhinga melanogaster CICONIIFORMES: Ciconiidae Storm's Stork Ciconia stormi CICONIIFORMES: Ardeidae Great-billed Heron Ardea sumatrana Purple Heron Ardea purpurea Eastern Great Egret Ardea modesta Intermediate Egret Ardea intermedia intermedia Eastern Cattle Egret Ardea coromandus Little Egret Egretta garzetta Black-crowned Night Heron Nycticorax nycticorax

ACCIPITRIFORMES: Accipitridae Jerdon's Baza Aviceda jerdoni Oriental Honeybuzzard Pernis ptilorhynchus Bat Hawk Macheiramphus alcinus alcinus Brahminy Kite Haliastur indus Lesser Fish Eagle Ichthyophaga humilis Grey-headed Fish Eagle Ichthyophaga ichthyaetus Mountain Serpent Eagle Spilornis kinabaluensis e Crested Serpent Eagle Spilornis cheela Crested Goshawk Lophospiza trivirgatus Black Eagle Ictinaetus malayensis Changeable Hawk Eagle Nisaetus limnaetus Wallace's Hawk Eagle Nisaetus nanus STRIGIFORMES: Tytonidae Oriental Bay Owl Phodilus badius badius heard only STRIGIFORMES: Strigidae Mountain Scops Owl Otus spilocephalus luciae Buffy Fish Owl Bubo ketupu Bornean Wood Owl Strix leptogrammica vaga Brown Boobook Ninox scutulata borneensis heard only Sunda (Collared) Owlet Glaucidium [brodiei] borneense heard only TROGONIFORMES: Trogonidae Red-naped Trogon Harpactes kasumba impavidus heard only Diard's Trogon Harpactes diardii diardii Whitehead's Trogon Harpactes whiteheadi e Cinnamon-rumped Trogon Harpactes orrhophaeus vidua Scarlet-rumped Trogon Harpactes duvaucelii CORACIIFORMES: Bucerotidae Oriental Pied Hornbill Anthracoceros albirostris convexus Black Hornbill Anthracoceros malayanus Rhinoceros Hornbill Buceros rhinoceros borneoensis Helmeted Hornbill Rhinoplax vigil Bushy-crested Hornbill Anorrhinus galeritus White-crowned Hornbill Berenicornis comatus Wrinkled Hornbill Aceros corrugatus Wreathed Hornbill Rhyticeros undulatus PICIFORMES: Capitonidae Gold-whiskered Barbet Psilopogon chrysopogon chrysopsis Red-throated Barbet Psilopogon mystacophanos heard only Mountain Barbet Psilopogon monticola e Yellow-crowned Barbet Psilopogon henricii heard only Golden-naped Barbet Psilopogon pulcherrima e Blue-eared Barbet Psilopogon australis duvaucelii Bornean Barbet Psilopogon eximia e heard only Bornean Brown Barbet Calorhamphus [fuliginosus] tertius e PICIFORMES: Picidae Rufous Piculet Sasia abnormis heard only Rufous Woodpecker Celeus brachyurus badiosus White-bellied Woodpecker Dryocopus javensis javensis Banded Yellownape Chrysophlegma mineaceus malaccensis heard only Checker-throated Yellownape Chrysophlegma mentalis humii heard only

Olive-backed Woodpecker Dinopium rafflesii dulitense Maroon Woodpecker Blythipicus rubiginosus Orange-backed Woodpecker Reinwardtipicus validus xanthopygius heard only Buff-rumped Woodpecker Meiglyptes tristis grammithorax Buff-necked Woodpecker Meiglyptes tukki tukki heard only Grey-and-buff Woodpecker Hemicircus concretus sordidus Great Slaty Woodpecker Mulleripicus pulverulentus CORACIIFORMES: Alcedinidae Blue-eared Kingfisher Alcedo meninting verreauxii Rufous-backed Kingfisher Ceyx rufidorsa motleyi Stork-billed Kingfisher Pelargopsis capensis inominata Ruddy Kingfisher Halcyon coromanda minor Collared Kingfisher Todirhamphus chloris laubmannianus Sacred KIngfisher Todirhamphus sanctus Rufous-collared Kingfisher Actenoides concretus borneanus heard only CORACIIFORMES: Meropidae Red-bearded Bee-eater Nyctyornis amictus Blue-throated Bee-eater Merops viridis viridis CORACIIFORMES: Coraciidae Common Dollarbird Eurystomus orientalis FALCONIFORMES: Falconidae White-fronted Falconet Microhierax latifrons e PSITTACIFORMES: Psittacidae Blue-naped Parrot Tanygnathus lucionensis I heard only Long-tailed Parakeet Psittacula longicauda Blue-crowned Hanging Parrot Loriculus galgulus PASSERIFORMES: Calyptomenidae Whitehead's Broadbill Calyptomena whiteheadi e PASSERIFORMES: Eurylaimidae Dusky Broadbill Corydon sumatranus Black-and-red Broadbill Cymbirhynchus macrorhynchos Banded Broadbill Eurylaimus javanicus heard only Black-and-yellow Broadbill Eurylaimus ochromalus PASSERIFORMES: Pittidae Giant Pitta Hydrornis caerulea hoesi Bornean Banded Pitta Hydrornis schwaneri e heard only Blue-headed Pitta Hydrornis baudii e heard only Hooded Pitta Erythropitta sordida mulleri Blue-banded Pitta Erythropitta arquata e Black-crowned Pitta Erythropitta ussheri e PASSERIFORMES: Pardalotidae Golden-bellied Gerygone Gerygone sulphurea sulphurea heard only PASSERIFORMES: Oriolidae Dark-throated Oriole Oriolus xanthonotus consobrinus Black-and-crimson Oriole Oriolus cruentus vulneratus heard only PASSERIFORMES: Pachycephalidae Bornean Whistler Pachycephala hypoxantha e

PASSERIFORMES: Vangidae Black-winged Flycatcher-shrike Hemipus hirundinaceus Large Woodshrike Tephrodornis gularis frenatus Rufous-winged Philentoma Philentoma pyrhopterum Maroon-breasted Philentoma Philentoma velatum heard only PASSERIFORMES: Pityriaseidae Bristlehead Pityriasis gymnocephala e PASSERIFORMES: Aegithinidae Common Iora Aegithina tiphia aequanimis heard only Green Iora Aegithina viridissima PASSERIFORMES: Artamidae White-breasted Woodswallow Artamus leucorynchus PASSERIFORMES: Campephagidae Fiery Minivet Pericrocotus igneus igneus Grey-chinned Minivet Pericrocotus solaris cinereigula PASSERIFORMES: Rhipiduridae White-throated Fantail Rhipidura albicollis kinabalu Pied Fantail Rhipidura javanica Spotted Fantail Rhipidura perlata PASSERIFORMES: Dicruridae Sunda [Ashy] Drongo Dicrurus leucophaeus stigmatops Bronzed Drongo Dicrurus aeneus malayensis Hair-crested Drongo Dicrurus [hottentottus] borneensis Greater Racket-tailed Drongo Dicrurus paradiseus brachyphorus PASSERIFORMES: Monarchidae Black-naped Monarch Hypothymis azurea prophata PASSERIFORMES: Laniidae Crested Jay Platylophus galericulatus coronatus heard only Brown Shrike Lanius cristatus Long-tailed Shrike Lanius schach PASSERIFORMES: Corvidae Bornean Black Magpie Platysmurus aterrimus e Bornean Green Magpie Cissa jefferyi e Bornean Treepie Dendrocitta cinerascens e Slender-billed Crow Corvus enca compilator PASSERIFORMES: Stenostiridae Grey-headed Canary-flycatcher Culicicapa ceylonensis PASSERIFORMES: Hirundinidae Barn Swallow Hirundo rustica gutturalis Pacific Swallow Hirundo tahitica PASSERIFORMES: Pycnonotidae Straw-headed Bulbul Pycnonotus zeylanicus Black-headed Bulbul Pycnonotus atriceps atriceps Bornean Bulbul Pycnonotus montis e Grey-bellied Bulbul Pycnonotus cyaniventris

Puff-backed Bulbul Pycnonotus eutilotus heard only Pale-faced Bulbul Pycnonotus [flavescens] leucops e Yellow-vented Bulbul Pycnonotus goiavier gourdini Olive-winged Bulbul Pycnonotus plumosus plumosus Cream-vented Bulbul Pycnonotus simplex perplexus Red-eyed Bulbul Pycnonotus brunneus brunneus Spectacled Bulbul Pycnonotus erythropthalmos Ochraceous Bulbul Alophoixus ochraceus ruficrissus Grey-cheeked Bulbul Alophoixus bres gutturalis Yellow-bellied Bulbul Alophoixus phaeocephalus Hairy-backed Bulbul Tricholestes criniger Buff-vented Bulbul Iole olivacea charlottae Streaked Bulbul Ixos malaccensis PASSERIFORMES: Timaliidae Bold-striped Tit Babbler Mixornis bornensis Fluffy-backed Tit Babbler Macronous ptilosus reclusus Chestnut-winged Babbler Cyanoderma erythropterum bicolor Chestnut-backed Scimitar Babbler Pomatorhinus montanus bornensis heard only Grey-throated Babbler Stachyris nigriceps borneensis White-necked Babbler Stachyris leucotis obscurata Black-throated Babbler Stachyris nigricollis heard only Chestnut-rumped Babbler Stachyris maculata maculata heard only PASSERIFORMES: Pellorneidae Moustached Babbler Malacopteron magnirostre cinereocapilla Sooty-capped Babbler Malacopteron affine phoeniceum Scaly-crowned Babbler Malacopteron cinereum cinereum Rufous-crowned Babbler Malacopteron magnum saba White-chested Babbler Pellorneum rostratum macropterum Ferruginous Babbler Pellorneum bicolor Short-tailed Babbler Pellorneum malaccense sordidum Temminck's Babbler Pellorneum pyrrogenys canicapillus Black-capped Babbler Pellorneum capistratum morrelli Striped Wren Babbler Kenopia striata Black-throated Wren Babbler Turdinus atrigularis e Mountain Wren Babbler Turdinus crassus e heard only Bornean Ground Babbler Ptilocichla leucogrammica e PASSERIFORMES: Leiothrichidae Brown Fulvetta Alcippe brunneicauda Sunda Laughingthrush Garrulax palliatus Chestnut-hooded Laughingthrush Ianthocincla treacheri treacheri e PASSERIFORMES: Zosteropidae Chestnut-crested Yuhina Yuhina everetti e Pygmy Heleia Heleia squamifrons e Black-capped White-eye Zosterops atricapillus Mountain Black-eye Zosterops emiliae e PASSERIFORMES: Phylloscopidae Arctic Warbler Seicercus borealis Mountain Leaf Warbler Seicercus trivirgatus kinabaluensis Yellow-breasted Warbler Seicercus montis floris PASSERIFORMES: Cettidae Bornean Stubtail Urosphena whiteheadi e Sunda Bush Warbler Horornis flavolivaceus oreophila

Mountain Tailorbird Phyllergates cuculatus cinereicollis PASSERIFORMES: Locustellidae Striated Grassbird Megalurus palustris PASSERIFORMES: Cisticolidae Rufous-tailed Tailorbird Orthotomus sericeus sericeus Ashy Tailorbird Orthotomus ruficeps borneoensis Yellow-bellied Prinia Prinia flaviventris latrunculus PASSERIFORMES: Sturnidae Asian Glossy Starling Aplonis panayensis Common Hill Myna Gracula religiosa Javan Myna Acridotheres javanicus I PASSERIFORMES: Turdidae Everett's Thrush Zoothera everetti e PASSERIFORMES: Muscicapidae Oriental Magpie Robin Copsychus saularis adamsi White-crowned Shama Copsychus stricklandii e Dark-sided Flycatcher Muscicapa sibirica Asian Brown Flycatcher Muscicapa dauurica dauurica Malaysian Blue Flycatcher Cyornis turcosus Bornean Blue Flycatcher Cyornis superbus e Verditer Flycatcher Eumyias thalassina Indigo Flycatcher Eumyias indigo Bornean Shade-dweller Vauriella gularis e White-browed Shortwing Brachypteryx montana erythrogyna White-crowned Forktail Enicurus leschenaulti frontalis Bornean Forktail Enicurus borneensis e Bornean Whistling Thrush Myophonus borneensis e Snowy-browed Flycatcher Ficedula hyperythra sumatrana Rufous-chested Flycatcher Ficedula dumetoria heard only Little Pied Flycatcher Ficedula westermanni PASSERIFORMES: Chloropseidae Greater Green Leafbird Chloropsis sonnerati zosterops Lesser Green Leafbird Chloropsis cyanopogon cyanopogon PASSERIFORMES: Irenidae Asian Fairy Bluebird Irena puella crinigera PASSERIFORMES: Dicaeidae Yellow-breasted Flowerpecker Prionochilus maculatus maculatus Yellow-rumped Flowerpecker Prionochilus xanthopygius e Yellow-vented Flowerpecker Pachyglossa chrysorrhea Orange-bellied Flowerpecker Dicaeum trigonostigma dayakanum Black-sided Flowerpecker Dicaeum monticolum e Scarlet-backed Flowerpecker Dicaeum cruentatum nigrimentum PASSERIFORMES: Nectariniidae Ruby-cheeked Sunbird Chalcoparia singalensis Plain Sunbird Anthreptes simplex Brown-throated Sunbird Anthreptes malacensis Olive-backed Sunbird Cinnyris jugularis Copper-throated Sunbird Leptocoma calcostetha Crimson Sunbird Aethopyga siparaja

Temminck's Sunbird Aethopyga temminckii Purple-naped Spiderhunter Arachnothera hypogrammicum Spectacled Spiderhunter Arachnothera flavigaster Little Spiderhunter Arachnothera longirostra Bornean Spiderhunter Arachnothera everetti e PASSERIFORMES: Passeridae Eurasian Tree Sparrow Passer montanus malaccensis PASSERIFORMES: Estrildidae Dusky Munia Lonchura fuscans e White-bellied Munia Lonchura leucogastra smythiesi Black-headed Munia Lonchura atricapilla jagori PASSERIFORMES: Motacillidae Grey Wagtail Motacilla cinerea cinerea Paddyfield Pipit Anthus rufulus MAMMALS Asian Whiskered Myotis Myotis muricola Wrinkle-lipped Bat Tadarica plicata Lesser Gymnure Hylomys suillus Mountain Treeshrew Tupaia montana baluensis e Lesser Treeshrew Tupaia minor minor Large Treeshrew Tupaia tana paitana Smooth-tailed Treeshrew Dendrogale melanura baluensis Sunda Colugo Cynocephalus variegatus Bornean Loris Nycticebus borneanus e Red Langur Presbytis rubicunda e Sabah Grey Langur Presbytis sabana e Silvered Langur Presbytis cristata Proboscis Monkey Nasalis larvatus e Long-tailed Macaque Macaca fascicularis Sunda Pig-tailed Macaque Macca nemestrina North Bornean Gibbon Hylobates funereus e Bornean Orang-utan Pongo pygmaeus e 'Bornean' Giant Squirrel Ratufa affinis Prevost's Squirrel Callosciurus prevostii pluto Plantain Squirrel Callosciurus notatus Ear-spot Squirrel Callosciurus adamsi Bornean Black-banded Squirrel Callosciurus orestes e Low's Squirrel Sundasciurus lowii Jentink's Squirrel Sundasciurus jentinki e Brooke's Squirrel Sundasciurus brookei Red-bellied Sculptor Squirrel Glyphotes simus e Bornean Mountain Ground Squirrel Dremomys everetti e Plain Pygmy Squirrel Exilisciurus exilis e Black Flying Squirrel Aeromys tephromelas Thomas's Flying Squirrel Aeromys thomasi e Red Giant Flying Squirrel Petaurista petaurista Tajuddin’s Maxomys Maxomys tajuddini Sabah Giant Rat Leopoldamys sabanus Yellow-throated Martin Martes flavigula saba Oriental Small-clawed Otter Aonyx cinerea Malay Civet Viverra tangalunga Common Palm Civet Paradoxurus hermaphroditus Masked Palm Civet Paguma larvata Binturong Arctictis binturong

Small-toothed Palm Civet Arctogalidia trivirgata Leopard Cat Felis bengalensis Bearded Pig Sus barbatus Lesser Mouse-Deer Tragulus kanchil Greater Mouse-Deer Tragulus napu Sambar Deer Cervus unicolor