Enumeratio nova Coleopterorum Fennoscandiae, Daniae et ... · Enumeratio nova Coleopterorum...

1© Sahlbergia Vol. 9: 1-111, 2004

Enumeratio nova Coleopterorum Fennoscandiae, Daniae et BaltiaeHans Silfverberg

Silfverberg, H. 2004: Enumeratio nova Coleopterorum Fennoscandiae, Daniaeet Baltiae. — Sahlbergia 9:1-111. Helsinki, Finland, ISSN 1237-3273.A revised check-list is presented of the Coleoptera of northern Europe (Fin-land, Sweden, Norway, Denmark, Estonia, Latvia, Lithuania and theFennoscandian parts of Russia).Hans Silfverberg, Zoological Museum, P.O.Box 17, FI-00014 Universityof Helsinki; E-mail: [email protected]

EsipuheEdellisen pohjoismaisen kovakuoriaisluettelonilmestymisestä on kulunut toistakymmentä vuotta,ja niin systematiikan kuin faunistiikankin alalla ontapahtunut paljon sen jälkeen. Näin ollen aika onkypsä uutta luetteloa varten. Uuden luettelonjärjestys perustuu maailmalla nyt jo laajaltiomaksuttuun systeemiin, jonka Lawrence & New-ton (1995) esittivät. Uutuudet luettelossaperustuvat pääasiassa jo julkaistuihin tietoihin,Suomen kohdalla on kuitenkin mukana joukkolajeja jotka on ilmoitettu kokouksissa (Liite 1),mutta joiden löydöistä ei vielä ole painettua tietoa.

Maat ilmoitetaan samoin kuin ennen, näin ollenF = Suomi, K = Venäjän fennoskandiset osat, S= Ruotsi, N = Norja, D = Tanska, E = Viro, A =Latvia ja I = Liettua (Kuva 1). Erona aikaisempaanvakiintumattomat importtilajit ilmaistaan kyseisenmaan pienellä kirjaimella, siten f = importtilajiSuomessa; näissä on pääasiassa huomioitu useitakertoja kulkeutuneet, sekä lähialueilta tulleet lajit.

FörordSedan den föregående nordiskaskalbaggskatalogen utkom har över ett decenniumgått, och mycket har hänt inom såväl systematiksom faunistik. Detta gör att tiden är mogen för enny katalog. Denna är ordnad enligt det systemLawrence & Newton (1995) uppställt, och somredan accepterats i stora delar av världen. De nyauppgifterna i katalogen baseras i huvudsak påpublicerad information, för Finlands del ingår

emellertid ett antal arter som anmälts på möten(Appendix 1), men för vilka uppgifter ännu intekommit i tryck.

Länderna anges på samma sätt som förr, d.v.s.F = Finland, K = ryska delarna av Fennoskandien,S = Sverige, N = Norge, D = Danmark, E =Estland, A = Lettland och I = Litauen (Bild 1). Enskillnad är att icke etablerade importarter nu angesmed liten bokstav för respektive land (t.ex. f =import till Finland); av dessa har huvudsakligenmedtagits ofta inkomna arter, samt sådana somkommit från närliggande områden.

IntroductionA check-list of northern European Coleoptera waspublished in 1992 (Silfverberg 1992), and afterthat much has happened both in systematics andin faunistics. A new list is thus appropriate. In gen-eral structure this list agrees with the previous one,but some comments on different aspects may bein order.

The countries or regions are indicated in thesame way as before, that is F = Finland, K = Rus-sian part of Fennoscandia, S = Sweden, N =Norway, D = Denmark, E = Estonia, A = Latviaand I = Lithuania (Figure 1). As a change from theprevious arrangement, imported, unestablishedspecies are now indicated with a lower case letterfor the area in question (f = imported to Finland,etc.). There has been no attempt to include allimported species; mainly the frequent ones andthose imported from nearby areas are mentioned.

2 Silfverberg: Enumeratio Coleopterorum

Kuva 1. Luettelossa käytetyt maakohtaiset lyhenteet.Bild 1. De landsvisa förkortningarna som används ikatalogen.Figure 1. Abbreviations of the countries as used in thecatalogue.

The previous list was arranged systematicallyusing the system gradually created by R. Crowsonin numerous works. A number of changes wereintroduced by Lawrence & Newton (1995), andthis revised system has already been used widely.It is generally accepted here, too. In some casesthe intrafamilial classification is based on still morerecent works, so for instance in Adephaga the newPalaearctic catalogue (Löbl & Smetana 2003), inStaphylinidae the catalogue of Herman (2001), inErotylidae the revision by WÞgrzynowicz (2002)and in Curculionidae to a large extent the cata-logue of Alonso-Zarazaga & Lyal (1999).

In Staphylinidae many generally used nameshave been found to be invalid for various reasons(Herman 2001). In order to avoid confusion ap-plications were made (particularly Herman 2002)to the International Commission on ZoologicalNomenclature for the preservation a large numberof such names, and accordingly customary use hasbeen retained in this catalogue.

There are no objective criteria for decidingthe extent of genera, and therefore we can see

quite different approaches in different works. Re-cent work in some families has led to heavy split-ting, and although such an arrangement can beassumed to give a more detailed picture of sys-tematic relationships, I prefer to have the genericlevel more equal in different families. Thus the treat-ment of, for instance, Cerambycidae andApionidae is relatively conservative .

Authors of family-group names were dis-cussed by Lawrence & Newton (1995), and thislist generally follows them. Yet many tribal nameswere not included in their work, and it is quitepossible that some of such names will be found tohave another author than the one suggested here.

The changes to the previous catalogue areprimarily based on published papers. Most of themhave been mentioned in Silfverberg (1997) orSilfverberg (2003), some further are listed in ref-erences below. The status of some Danish spe-cies has been changed in accordance with Hansen(1996). There have been many recent advancesin Latvian coleopterology, resulting in a check-listof Latvian beetles (Telnov et al. 1997, revised2002), and the Latvian fauna is mostly listed ac-cording to that catalogue. In the previous list therewere several species with a B for country, mean-ing either Estonia or Latvia. Those records thatcould not be assigned to one country or the otherhave now been omitted. Concerning the Latvianfauna, the assistance given by A. Barševskis shouldbe specially acknowledged.

Some records that have not yet been pub-lished are also included. A few species are reportedas new to Estonia in a paper by I. Rutanen, soonto be published. A number of species have beenreported from Finland in meetings of scientific so-cieties, but have not yet been published in print.They are listed in the Appendix 1, with a refer-ence to the reporter of the species.

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3Silfverberg: Enumeratio Coleopterorum

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MYXOPHAGASphaeriusidae Erichson, 1845

Microsporidae Crotch, 1873Sphaerius Waltl, 1838Microsporus Kolenati, 1846

acaroides Waltl, 1838 FKS-D-AIobsidianus (Kolenati, 1846)

ADEPHAGAGyrinidae Latreille, 1810Gyrininae Latreille, 1810Gyrinus Geoffroy, 1762minutus Fabricius, 1798 FKSNDEAIopacus Sahlberg, 1819 FKSN-EAIaeratus Stephens, 1835 FKSNDEAIpullatus Zaitzev, 1908 FKS--EAImarinus Gyllenhal, 1808 FKSNDEAIcolymbus Erichson, 1837 ----D---distinctus Aubé, 1836 FKSNDEAIsuffriani Scriba, 1855 F-SNDE--natator (Linnaeus, 1758) FKSNDEAIsubstriatus Stephens, 1828 FKSNDEAIurinator Illiger, 1807 -------Icaspius Ménétriés, 1832 --SNDEAIpaykulli Ochs, 1927 FKSNDEAI

Orectochilinae Régimbart, 1882Orectochilus Dejean, 1833villosus (Müller, 1776) FKSNDEAI

Haliplidae Brullé, 1835Brychius Thomson, 1859elevatus (Panzer, 1793)ssp. elevatus (Panzer, 1793) ----D---ssp. norvegicus Munster, 1922 --SN----ssp. cristatus J.Sahlberg, 1875 FKSN-EAI

Peltodytes Régimbart, 1879caesus (Duftschmid, 1805) --S-D--I

Haliplus Latreille, 1802lineatocollis (Marsham, 1802) --S-D--Ilaminatus (Schaller, 1783) --S-D--Iflavicollis Sturm, 1834 FKSNDEAIfulvus (Fabricius, 1801) FKSNDEAI

lapponum Thomson, 1856variegatus Sturm, 1834 F-SNDEAIconfinis Stephens, 1828 FKSNDEAIvarius Nicolai, 1822 FK---E-Iobliquus (Fabricius, 1787) F-SNDEAIapicalis Thomson, 1868 --SNDE-I

furcatus Seidlitz, 1887 F-S-DEAIfulvicollis Erichson, 1837 FKSNDEA-fluviatilis Aubé, 1836 FKS-DEAIlineolatus Mannerheim, 1844 FKSNDEA-

nomax Balfour-Browne, 1911interjectus Lindberg, 1937 FK----A-sibiricus Motschulsky, 1860 FKSNDEAI

wehnckei Gerhardt, 1877sahlbergi Falkenström, 1940lineolatus auct. nec Mannerheim, 1844

ruficollis (DeGeer, 1774) FKSNDEAIheydeni Wehncke, 1875 FKSNDEAIimmaculatus Gerhardt, 1877 FKSNDEAI

Trachypachidae Thomson, 1857Trachypachus Motschulsky, 1844zetterstedtii (Gyllenhal, 1827) FKSN-EA-

Noteridae Thomson, 1860Noterus Clairville, 1806clavicornis (DeGeer, 1774) F-SNDEAIcrassicornis (Müller, 1776) FKSNDEAI

Dytiscidae Leach, 1815Copelatinae Van den Branden, 1885Copelatus Erichson, 1832haemorrhoidalis (Fabricius, 1787) FKS-DEAI

ruficollis (Schaller, 1783) nec (DeGeer, 1774)

Laccophilinae Gistel, 1856Laccophilus Leach, 1815hyalinus (DeGeer, 1774) FKSNDEAIminutus (Linnaeus, 1758) FKSNDEAIbiguttatus Kirby, 1837 FKSN----

strohmi Thomson, 1874poecilus Klug, 1834 --SN-EAI

ponticus Sharp, 1882variegatus (Germar, 1812) nec (Geoffroy, 1785)obsoletus auct. nec Westhoff, 1881

Hydroporinae Aubé, 1836Laccornini Wolfe & Roughley, 1990Laccornis Des Gozis, 1914oblongus (Stephens, 1835) FKSNDEA-

Hydrovatini Sharp, 1882Hydrovatus Motschulsky, 1853cuspidatus (Kunze, 1818) ----D---

Bidessini Sharp, 1882Bidessus Sharp, 1882unistriatus (Goeze, 1777) FKSNDEAIgrossepunctatus Vorbringer, 1907 FKSND-AI


Hydroglyphus Motschulsky, 1853Guignotus Houlbert, 1934

geminus (Fabricius, 1792) FKSNDEAIpusillus (Fabricius, 1781) nec (Müller, 1776)

hamulatus (Gyllenhal, 1813) FKS-DEAI

Hyphydrini Sharp, 1882Hyphydrus Illiger, 1802ovatus (Linnaeus, 1761) FKSNDEAI

Hygrotini Portevin, 1929Hygrotus Stephens, 1828decoratus (Gyllenhal, 1810) FKSNDEAIinaequalis (Fabricius, 1777) FKSNDEAIversicolor (Schaller, 1783) FKSNDEAIquinquelineatus (Zetterstedt, 1828) FKSND-A-

Coelambus Thomson, 1860polonicus (Aubé, 1842) FK---EAIimpressopunctatus (Schaller, 1783) FKSNDEAIparallelogrammus (Ahrens, 1812) F-SNDE-Inovemlineatus (Stephens, 1829) FKSNDE--marklini (Gyllenhal, 1813) FKSNDEA-nigrolineatus (Steven, 1808) F-S-D---

lautus (Schaum, 1843)confluens (Fabricius, 1787) --SNDE--

Hydroporini Aubé, 1836Hydroporus Clairville, 1806Hydrocolus Roughley & Nilsson, 2000

nigrita (Fabricius, 1792) FKSNDEAIbrevis F.Sahlberg, 1834 FKSN-EA-acutangulus Thomson, 1856 FKSN----neuter Fairmaire & Laboulbène, 1854 F-SNDEA-

discretus Fairmaire, 1859pubescens (Gyllenhal, 1808) FKSNDEA-fuscipennis Schaum, 1868 FKSNDEAIplanus (Fabricius, 1781) FKSNDEAItessellatus (Drapiez, 1819) ------A-obscurus Sturm, 1835 FKSNDEAIelongatulus Sturm, 1835 FKSNDEAI

eugeniae Zaitzev, 1909semenowi Jakowlew, 1897 FKSN----

longitarsis J.Sahlberg, 1910afflatus Scholtz, 1917tartaricus auct. nec LeConte, 1850eugeniae auct. nec Zaitzev, 1909

geniculatus Thomson, 1854 FKSN----subseriatus J.Sahlberg, 1910pilipes J.Sahlberg, 1921

nigellus Mannerheim, 1853 FKSND-A-

submuticus Thomson, 1874 FKSN----notabilis LeConte, 1850ssp. arcticus Thomson, 1856 FKSN----

tomentosus Poppius, 1905erythrocephalus (Linnaeus, 1758) FKSNDEAIrufifrons (Müller, 1776) FKSNDEAI

piceus Stephens, 1828scalesianus Stephens, 1828 FKSNDE--angustatus Sturm, 1835 FKSNDEAIlapponum (Gyllenhal, 1808) FKSN----puberulus LeConte, 1850 FKS-----

levanderi J.Sahlberg, 1889glabriusculus Aubé, 1838 FKSND-AIumbrosus (Gyllenhal, 1808) FKSNDEAImorio Aubé, 1838 FKSNDEAI

valliger Hellén, 1929valliger Hellén, 1929melanocephalus auct. nec (Marsham, 1802)

tristis (Paykull, 1798) FKSNDEAIgyllenhalii Schiödte, 1841 FKSND---

piceus auct. nec Stephens, 1828notatus Sturm, 1835 FKS-DEAIincognitus Sharp, 1869 FKSND-AIstriola (Gyllenhal, 1826) FKSNDEAIpalustris (Linnaeus, 1761) FKSNDEAIneglectus Schaum, 1845 FKSNDEAIkraatzii Schaum, 1868 -----E--melanarius Sturm, 1835 FKSNDEAIlongicornis Sharp, 1871 FKSND-AImemnonius Nicolai, 1822 FKSNDEAIobsoletus Aubé, 1838 --SND---sahlbergi (Nilsson, 2001) FKSN----

picicornis J.Sahlberg, 1875, nec (Chevrolat, 1863)

Porhydrus Guignot, 1945lineatus (Fabricius, 1775) FKSNDEAI

Graptodytes Seidlitz, 1887granularis (Linnaeus, 1767) FKSNDEAIbilineatus (Sturm, 1835) F-S-DEAIpictus (Fabricius, 1787) FKSNDEAI

Oreodytes Seidlitz, 1887alpinus (Paykull, 1798) FKSN----septentrionalis (Gyllenhal, 1826) FKSN----sanmarkii (Sahlberg, 1826) FKSND--I

rivalis (Gyllenhal, 1827)

Suphrodytes Des Gozis, 1914dorsalis (Fabricius, 1787) FKSNDEAI

eljasi (Wirén, 1968)


Deronectes Sharp, 1882latus (Stephens, 1829) FKSNDEAI

Scarodytes Des Gozis, 1914halensis (Fabricius, 1787) FKSNDEAI

Nebrioporus Régimbart, 1906Potamonectes Zimmermann, 1921

canaliculatus (Lacordaire, 1835) --S-D---assimilis (Paykull, 1798) FKSN-EAIdepressus (Fabricius, 1775) FKSNDEAI

Stictotarsus Zimmermann, 1919griseostriatus (DeGeer, 1774)ssp. griseostriatus (DeGeer, 1774) FKSN-E--ssp. strandi (Brinck, 1943) -K-N----multilineatus (Falkenström, 1922) FKSN----duodecimpustulatus (Fabricius, 1792) --SND---

Colymbetinae Erichson, 1837Agabini Thomson, 1867Platambus Thomson, 1859maculatus (Linnaeus, 1758) FKSNDEAI

Agabus Leach, 1817unguicularis (Thomson, 1867) FKS-DEA-affinis (Paykull, 1798) FKSNDEAIbiguttulus (Thomson, 1867) FKSN-EAIpaludosus (Fabricius, 1801) FKSNDEAIadpressus Aubé, 1837 FKSN----didymus (Olivier, 1795) --S-D--Iguttatus (Paykull, 1798) FKSNDEAImelanarius Aubé, 1837 FKSNDE-I

tarsatus Zetterstedt, 1838bipustulatus (Linnaeus, 1767) FKSNDEAI

solieri Aubé, 1837conspersus (Marsham, 1802) --S-D---nebulosus (Forster, 1771) --SNDEAIstriolatus (Gyllenhal, 1808) FKS-DEAIsturmii (Gyllenhal, 1808) FKSNDEAIarcticus (Paykull, 1798) FKSN----zetterstedti Thomson, 1856 FKSN----elongatus (Gyllenhal, 1826) FKSN----moestus (Curtis, 1835) F-------

nigripalpis (J.Sahlberg, 1881)congener (Thunberg, 1794) FKSNDEAI

lapponicus (Thomson, 1867) FKSN----

thomsoni (J.Sahlberg, 1871) FKSN----

pseudoclypealis Scholtz, 1933 FK----A-haraldi Håkan Lindberg, 1933

discolor (Harris, 1828) FKSN----levanderi Hellén, 1929approximatus auct. nec Fall, 1923

clypealis (Thomson, 1867) FKS-D-A-setulosus (J.Sahlberg, 1895) FKSN--A-confinis (Gyllenhal, 1808) FKSN--A-infuscatus Aubé, 1838 FK-N----fuscipennis (Paykull, 1798) FKSNDEAIuliginosus (Linnaeus, 1761) FKSNDEAIundulatus (Schrank, 1776) --SNDEAIlabiatus (Brahm, 1791) FKSNDEAIserricornis (Paykull, 1799) FKSN----

Ilybius Erichson, 1832wasastjernai (Sahlberg, 1824) FKSND-A-opacus (Aubé, 1837) FKSN----chalconatus (Panzer, 1796) F-S-DEAImontanus (Stephens, 1828) ----D-AI

melanocornis (Zimmermann, 1915)neclectus (Erichson, 1837) --S-DEAIsubtilis (Erichson, 1837) FKSNDEAIerichsoni (Gemminger & Harold, 1868) FKSNDEAI

nigroaeneus (Erichson, 1837) nec (Marsham, 1802)vittiger (Gyllenhal, 1827) FKSN----picipes (Kirby, 1837) F-SN----angustior (Gyllenhal, 1808) FKSNDEAIcrassus Thomson, 1856 FKSND--Isubaeneus Erichson, 1837 FKSNDEAIater (DeGeer, 1774) FKSNDEAIguttiger (Gyllenhal, 1808) FKSNDEAIquadriguttatus (Lacordaire, 1835) FKSNDEAI

obscurus ((Marsham, 1802)similis Thomson, 1856 FKSNDEAIaenescens Thomson, 1870 FKSNDEAIfuliginosus (Fabricius, 1792) FKSNDEAIfenestratus (Fabricius, 1781) FKSNDEAI

Colymbetini Erichson, 1837Rhantus Dejean, 1833Nartus Zaitzev, 1907

grapii (Gyllenhal, 1808) FKSNDEAIsuturalis (MacLeay, 1825) FKSNDEAI

pulverosus (Stephens, 1828)frontalis (Marsham, 1802) FKSNDEAI

notatus (Fabricius, 1781) nec (Bergsträsser, 1778)notaticollis (Aubé, 1837) FKSNDEAIsuturellus (Harris, 1828) FKSNDEAIbistriatus (Bergsträsser, 1778) FKS-DEAIconsputus (Sturm, 1834) -------I


incognitus Scholz, 1927 ------AIexsoletus (Forster, 1771) FKSNDEAIlatitans Sharp, 1882 -----EAIfennicus Huldén, 1982 FKS-----

Colymbetes Clairville, 1806fuscus (Linnaeus, 1758) FKSNDEAIpaykulli Erichson, 1837 FKSNDEAIstriatus (Linnaeus, 1758) FKSNDEAIdolabratus (Paykull, 1798) FKSN----

Dytiscinae Leach, 1815Hydaticini Sharp, 1882Hydaticus Leach, 1817transversalis (Pontoppidan, 1763) FKSNDEAIcontinentalis Balfour-Browne, 1944 --S-DEAI

stagnalis (Fabricius, 1787) nec (Geoffroy, 1785)modestus auct. nec Sharp, 1882

aruspex Clark, 1864 FKSNDEA-laevipennis Thomson, 1867

seminiger (DeGeer, 1774) FKSNDEAI

Aciliini Thomson, 1867Thermonectini Sharp, 1882

Graphoderus Dejean, 1833austriacus (Sturm, 1834) --S-D-AIbilineatus (DeGeer, 1774) FKSNDEAIcinereus (Linnaeus, 1758) FKSNDEAIzonatus (Hoppe, 1795)ssp. zonatus (Hoppe, 1795) FKSNDEAIssp. verrucifer (Sahlberg, 1824) FKSN----

piciventris Thomson, 1868

Acilius Leach, 1817sulcatus (Linnaeus, 1758) FKSNDEAIcanaliculatus (Nicolai, 1822) FKSNDEAI

Dytiscini Leach, 1815Dytiscus Linnaeus, 1758semisulcatus Müller, 1776 --SNDEAIdimidiatus Bergsträsser, 1778 FKS-DEAImarginalis Linnaeus, 1758 FKSNDEAIlatissimus Linnaeus, 1758 FKSNDEAIcircumcinctus Ahrens, 1811 FKSNDEAIlapponicus Gyllenhal, 1808 FKSNDEAIcircumflexus Fabricius, 1801 --S-DEAI

Cybistrini Sharp, 1882Cybister Curtis, 1827lateralimarginalis (DeGeer, 1774) --S-D-AI

Carabidae Latreille, 1802Omophroninae Bonelli, 1810Omophron Latreille, 1802limbatum (Fabricius, 1777) --S-DEAI

Carabinae Latreille, 1802Pelophilini Kavanaugh, 1996Pelophila Dejean, 1821borealis (Paykull, 1790) FKSN-EA-

Nebriini Laporte de Castelnau, 1834Leistus Frölich, 1799rufomarginatus (Duftschmid, 1812) --S-D-AIterminatus (Panzer, 1793) FKSNDEAI

rufescens (Fabricius, 1775) nec (Ström, 1768)ferrugineus (Linnaeus, 1758) FKSNDEAIpiceus Frölich, 1799 -----EAI

Nebria Latreille, 1802livida (Linnaeus, 1758) FKSNDEAIrufescens (Ström, 1768) FKSN-EAI

gyllenhali (Schönherr, 1806)nivalis (Paykull, 1790) FKSN----brevicollis (Fabricius, 1792) F-SNDEAIsalina Fairmaire & Laboulbène, 1854 --SND---

Notiophilini Motschulsky, 1850Notiophilus Duméril, 1806aestuans Dejean, 1826 FKSNDEAI

pusillus Waterhouse, 1833 nec (Schreber, 1759)aquaticus (Linnaeus, 1758) FKSNDEAIpalustris (Duftschmid, 1812) FKSNDEAIgerminyi Fauvel, 1863 FKSNDEAI

stipraisi Barševskis, 1993rufipes Curtis, 1829 --S-D---reitteri Spaeth, 1900 FKSN----biguttatus (Fabricius, 1779) FKSNDEAI

Carabini Latreille, 1802Calosoma Weber, 1801sycophanta (Linnaeus, 1758) --S-DE-Iinquisitor (Linnaeus, 1758) F-SNDEAIauropunctatum (Herbst, 1784) --SNDEAIdenticolle Gebler, 1833 F-------investigator (Illiger, 1798) --S-----reticulatum (Fabricius, 1787) --S-D---

Carabus Linnaeus, 1758monilis Fabricius, 1792 ---N----arcensis Herbst, 1784 FKSNDEAIgranulatus Linnaeus, 1758 FKSNDEAI


menetriesi Hummel, 1827 -K---EAIclathratus Linnaeus, 1761 FKSNDEAIcancellatus Illiger, 1798ssp. cancellatus Illiger, 1798 --SND---

rauterbergi Kolbe, 1912munsteri Born, 1926

ssp. tuberculatus Dejean, 1826 FK--DEAIsajanensis Reitter, 1896kuntzeni Eidam, 1930

auratus Linnaeus, 1761 --sND---nemoralis Müller, 1764 FKSNDEAIhortensis Linnaeus, 1758 FKSNDEAIglabratus Paykull, 1790 FKSNDEAIproblematicus Herbst, 1786ssp. harcyniae Sturm, 1815 -KSND---

muelverstedti Reitter, 1896wockei Born, 1898relictus Hellén, 1936

ssp. strandi Born, 1926 FKSN----nitens Linnaeus, 1758 FKSNDEAIconvexus Fabricius, 1775 FKSNDEAIintricatus Linnaeus, 1761 --S-DEA-violaceus Linnaeus, 1758 FKSNDEAIcoriaceus Linnaeus, 1758 --SNDEAI

Cychrini Laporte de Castelnau, 1834Cychrus Fabricius, 1794caraboides (Linnaeus, 1758) FKSNDEAI

Cicindelinae Latreille, 1802Cicindela Linnaeus, 1758sylvatica Linnaeus, 1758 FKSNDEAIhybrida Linnaeus, 1758 FKSNDEAImaritima Latreille & Dejean, 1822 FKSNDEAIcampestris Linnaeus, 1758 FKSNDEAI

Cylindera Westwood, 1831germanica (Linnaeus, 1758) -----EAIarenaria (Fuessly, 1775)ssp. viennensis (Schrank, 1781) ------A-

Loricerinae Bonelli, 1810Loricera Latreille, 1802pilicornis (Fabricius, 1775) FKSNDEAI

Elaphrinae Latreille, 1802Blethisa Bonelli, 1810multipunctata (Linnaeus, 1758) FKSNDEAI

Diacheila Motschulsky, 1845arctica (Gyllenhal, 1810) FKSN----polita (Faldermann, 1835) -K-N----

Elaphrus Fabricius, 1775lapponicus Gyllenhal, 1810 FKSN----uliginosus Fabricius, 1792 FKSNDEAIcupreus Duftschmid, 1812 FKSNDEAIriparius (Linnaeus, 1758) FKSNDEAIaureus Müller, 1821 -----EAIangusticollis F. Sahlberg, 1844 -K----A-

Scaritinae Bonelli, 1810Clivinini Rafinesque, 1815Clivina Latreille, 1802fossor (Linnaeus, 1758) FKSNDEAIcollaris (Herbst, 1784) --S-DEAI

Dyschirius Bonelli, 1810digitatus (Dejean, 1825) ------AIthoracicus (Rossi, 1790) FKSNDEAI

arenosus Stephens, 1827obscurus (Gyllenhal, 1827) FKSNDEAIangustatus (Ahrens, 1830) FKSNDEAInitidus (Dejean, 1825) FK---EAIneresheimeri Wagner, 1915 --S--EAIpolitus (Dejean, 1825) FKSNDEAIimpunctipennis Dawson, 1854 FKSNDEAIchalceus Erichson, 1837 --S-DE--salinus Schaum, 1843 F-SNDEA-aeneus (Dejean, 1825) F-SNDEAItristis Stephens, 1827 FKSNDEAI

luedersi Wagner, 1915nigricornis Motschulsky, 1844 FKSN-EA-

helleni Müller, 1922septentrionum Munster, 1923

melancholicus Putzeys, 1866 FKSN----nigricornis auct. nec Motschulsky, 1844

intermedius Putzeys, 1846 -KS-D-AIlaeviusculus Putzeys, 1846 --S-DEAIglobosus (Herbst, 1784) FKSNDEAI

Trechinae Bonelli, 1810Psydrinae LeConte, 1853

Broscini Hope, 1838Broscus Panzer, 1813cephalotes (Linnaeus, 1758) FKSNDEAI

Miscodera Eschscholtz, 1830arctica (Paykull, 1798) FKSNDEAI

Patrobini Kirby, 1837Patrobus Dejean, 1821septentrionis Dejean, 1828 FKSNDEAI

australis J.Sahlberg, 1875


assimilis Chaudoir, 1844 FKSNDEAIatrorufus (Ström, 1768) FKSNDEAI

Rhysodini Laporte de Castelnau, 1840Rhysodes Dalman, 1823sulcatus (Fabricius, 1787) --S-----

Trechini Bonelli, 1810Perileptus Schaum, 1860areolatus (Creutzer, 1799) -KSN--A-

Aepus Leach, 1819marinus (Ström, 1788) --SN----

Trechus Clairville, 1806secalis (Paykull, 1790) FKSNDEAIrivularis (Gyllenhal, 1810) FKSNDEAIrubens (Fabricius, 1792) FKSNDEAIfulvus Dejean, 1831 ---N----quadristriatus (Schranck, 1781) FKSNDEAIobtusus Erichson, 1837 --SND-A-austriacus Dejean, 1831 ------AI

Trechoblemus Ganglbauer, 1891micros (Herbst, 1784) FKSNDEAI

Blemus Dejean, 1821discus (Fabricius, 1792) FKSNDEAI

Pogonini Laporte de Castelnau, 1834Pogonus Dejean, 1821luridipennis (Germar, 1823) --SND---chalceus (Marsham, 1802) ----D--I

Bembidiini Stephens, 1827Bembidiina Stephens, 1827Asaphidion Des Gozis, 1886caraboides (Schrank, 1781) -------Ipallipes (Duftschmid, 1812) FKSNDEAIflavipes (Linnaeus, 1761) FKSNDEAIcurtum (Heyden, 1870) --S-D-A-austriacum Schweiger, 1975 ------A-

Bembidion Latreille, 1802striatum (Fabricius, 1792) FK--DEAIvelox (Linnaeus, 1761) FKSN-EAIlapponicum Zetterstedt, 1828 FKSN----argenteolum Ahrens, 1812 -KSN-EAIlitorale (Olivier, 1791) FKSNDEAInigricorne Gyllenhal, 1827 FKSNDEAIpygmaeum (Fabricius, 1792) FK---EAIlampros (Herbst, 1784) FKSNDEAIproperans (Stephens, 1828) FKSNDEAIobtusum Audinet-Serville, 1821 --SNDEAI

punctulatum Drapiez, 1820 FK---EAIbipunctatum (Linnaeus, 1761) FKSNDEAIruficolle (Panzer, 1797) FKS-DEAIpallidipenne (Illiger, 1802) --SNDEAIilligeri Netolitzky, 1914 F-S-DEAI

genei auct. nec Küster, 1847fellmanni (Mannerheim, 1823) FKSN----difficile (Motschulsky, 1844) FKSN----crenulatum F.Sahlberg, 1844ssp. ponojense J.Sahlberg, 1875 -K------prasinum (Duftschmid, 1812) FKSN----hyperboraeorum Munster, 1923 FKSN----virens Gyllenhal, 1827 FKSN----hastii Sahlberg, 1827 FKSN----hirmocaelum Chaudoir, 1850 -K------tibiale (Duftschmid, 1812) ---N----mckinleyi Fall, 1926ssp. scandicum Lindroth, 1943 F-SN----monticola Sturm, 1825 FK--DEAIdeletum Audinet-Serville, 1821 FKSNDEAI

nitidulum (Marsham, 1802) nec (Schrank, 1781)stephensi Crotch, 1869 FKSNDEA-lunatum (Duftschmid, 1812) --SND-AIgrapii Gyllenhal, 1827 FKSN----yukonum Fall, 1926 FKSN----

grapeioides Munster, 1930dauricum (Motschulsky, 1844) F-SN----tetracolum Say, 1823 FKSNDEAI

ustulatum auct. nec (Linnaeus, 1758)bruxellense Wesmaël, 1835 FKSNDEAI

rupestre auct. nec (Linnaeus, 1758)petrosum Gebler, 1833ssp. siebkei Sparre Schneider, 1910 FKSN----ssp. carlhlindrothi Kangas, 1980 -K------cruciatum Dejean, 1831

andreae auct. nec (Fabricius, 1787)ssp. polonicum J.Müller, 1930 FKS-DEAIfemoratum Sturm, 1825 FKSNDEAImaritimum (Stephens, 1835) ----D---obscurellum (Motschulsky, 1845) -KS-D---saxatile Gyllenhal, 1827 FKSNDEAIdecorum (Panzer, 1799) ----D---dentellum (Thunberg, 1787) FKSNDEAItinctum Zetterstedt, 1828 FKS-----varium (Olivier, 1795) F-SNDEAIsemipunctatum (Donovan, 1806) FKSNDEAIobliquum Sturm, 1825 FKSNDEAIephippium (Marsham, 1802) ----D---


minimum (Fabricius, 1792) FKSNDE-Inormannum Dejean, 1831 ----D---azurescens (Dalla Torre, 1877) FKS--EAItenellum Erichson, 1837 --S-DEAIarticulatum (Panzer, 1796) FKSNDEAIoctomaculatum (Goeze, 1777) F-S-D-AIdoris (Panzer, 1797) FKSNDEAIschuppelii Dejean, 1831 FKSNDEAIchaudoirii Chaudoir, 1850 -K------gilvipes Sturm, 1825 FKSNDEAIfumigatum (Duftschmid, 1812) --SNDEAIassimile Gyllenhal, 1810 F-SNDEAIclarkii (Dawson, 1849) --S-D---transparens (Gebler, 1829) FKSNDEA-quadrimaculatum (Linnaeus, 1761) FKSNDEAIhumerale Sturm, 1825 FKS-DEAIquadripustulatum Audinet-Serville, 1821 --S-D--Ibiguttatum (Fabricius, 1779) FKS-DEAIiricolor Bedel, 1879 ----D---lunulatum (Geoffroy, 1785) --S-D-AIaeneum Germar, 1824 FKSND-AIguttula (Fabricius, 1792) FKSNDEAIneresheimeri Müller, 1929 ------AImannerheimii Sahlberg, 1827 FKSNDEAI

unicolor Chaudoir, 1850

Ocys Stephens, 1828harpaloides (Audinet-Serville, 1821) ---N----quinquestriatus (Gyllenhal, 1810) --SNDEA-

Cillenus Leach, 1819lateralis Samouelle, 1819 ----D---

Tachyina Motschulsky, 1862Tachys Dejean, 1821Paratachys Casey, 1918

bistriatus (Duftschmid, 1812) FK--D-AI

Porotachys Netolitzky, 1914bisulcatus (Nicolai, 1822) FKSND-A-

Tachyura Motschulsky, 1862parvula (Dejean, 1831) --S-----

Tachyta Kirby, 1837nana (Gyllenhal, 1810) FKSNDEAI

Harpalinae Bonelli, 1810Pterostichini Bonelli, 1810Stomis Clairville, 1806pumicatus (Panzer, 1796) FKSNDEAI

Poecilus Bonelli, 1810punctulatus (Schaller, 1783) --S-DEAI

lepidus (Leske, 1785) FKSNDEAIkugelanni (Panzer, 1797) ----D---

dimidiatus (Olivier, 1795) nec (Rossi, 1790)cupreus (Linnaeus, 1758) FKSNDEAIversicolor (Sturm, 1824) FKSNDEAI

caerulescens auct. nec (Linnaeus, 1758)

Pedius Motschulsky, 1850longicollis (Duftschmid, 1812) ----D---

inaequalis (Marsham, 1802) nec (Panzer, 1796)

Pterostichus Bonelli, 1810crenatus (Duftschmid, 1812) FKSNDEAI

vernalis (Panzer, 1796) nec (Müller, 1776)macer (Marsham, 1802) ----DEAIaterrimus (Herbst, 1784) FKSNDEAIaethiops (Panzer, 1796) FK--dEAImadidus (Fabricius, 1775) F---D---oblongopunctatus (Fabricius, 1787) FKSNDEAIadstrictus Eschscholtz, 1823 FKSN----quadrifoveolatus Letzner, 1852 FKSNDEAI

angustatus (Duftschmid, 1812) nec (Fabricius, 1787)niger (Schaller, 1783) FKSNDEAImelanarius (Illiger, 1798) FKSNDEAI

vulgaris auct. nec (Linnaeus, 1758)nigrita (Paykull, 1790) FKSNDEAIrhaeticus Heer, 1837 FKSNDEAIanthracinus (Illiger, 1798) -KS-DEAIgracilis (Dejean, 1828) FKS-DEAIminor (Gyllenhal, 1827) FKSNDEAIstrenuus (Panzer, 1796) FKSNDEAIdiligens (Sturm, 1824) FKSNDEAIbrevicornis Kirby, 1837 -K------middendorffi (J.Sahlberg, 1875) -K------melas (Creutzer, 1799) ----D---burmeisteri Heer, 1841 --S-d---

metallicus (Fabricius, 1792) nec (Scopoli, 1763)

Abax Bonelli, 1810parallelepipedus (Piller & Mitterpacher, 1783) --SND---

Sphodrini Laporte de Castelnau, 1834Calathus Bonelli, 1810fuscipes (Goeze, 1777) FKSNDEAIerratus (Sahlberg, 1827) FKSNDEAIambiguus (Paykull, 1790) FKSNDEAImelanocephalus (Linnaeus, 1758) FKSNDEAIcinctus Motschulsky, 1850 --S-D-A-mollis (Marsham, 1802) --SNDEAI

ochropterus auct. nec (Duftschmid, 1812)micropterus (Duftschmid, 1812) FKSNDEAI


rotundicollis Dejean, 1828 --S-D---piceus (Marsham, 1802) nec (Linnaeus, 1758)

Sphodrus Clairville, 1806leucophthalmus (Linnaeus, 1758) F-S-DEAI

Laemostenus Bonelli, 1810Pristonychus Dejean, 1828

terricola (Herbst, 1784) FKSNDEAI

Dolichus Bonelli, 1810halensis (Schaller, 1783) --S-D--I

Synuchus Gyllenhal, 1810vivalis (Illiger, 1798) FKSNDEAI

nivalis (Panzer, 1797) nec (Paykull, 1790)

Platyderus Stephens, 1828depressus (Audinet-Serville, 1821) ---N----

ruficollis (Marsham, 1802) nec (Fabricius, 1801)

Platynini Bonelli, 1810Olisthopus Dejean, 1828rotundatus (Paykull, 1790) FKSNDEAI

Sericoda Kirby, 1837bogemannii (Gyllenhal, 1813) FKS-----quadripunctata (DeGeer, 1774) FKSNDEAI

Anchomenus Bonelli, 1810dorsalis (Pontoppidan, 1763) F-SNDEAI

Oxypselaphus Chaudoir, 1843obscurus (Herbst, 1784) FKSNDEAI

Paranchus Lindroth, 1974albipes (Fabricius, 1796) FKSNDEAI

ruficornis (Goeze, 1777) nec (DeGeer, 1774)

Platynus Bonelli, 1810livens (Gyllenhal, 1810) FKS-DEAImannerheimii (Dejean, 1828) FKSN-EAIassimilis (Paykull, 1790) FKSNDEAIkrynickii (Sperk, 1835) --S-DEAIlongiventris Mannerheim, 1825 --S--EAI

Agonum Bonelli, 1810scitulum Dejean, 1828 -----EA-micans Nicolai, 1822 FKSNDEAIpiceum (Linnaeus, 1758) FKSNDEAIgracile Sturm, 1824 FKSNDEAI

latvicum Barševskis, 1993fuliginosum (Panzer, 1809) FKSNDEAIthoreyi Dejean, 1828 FKSNDEAI

pelidnum (Paykull, 1790) nec (Herbst, 1784)consimile (Gyllenhal, 1810) FKSN----

munsteri (Hellén, 1935) FKSNDEA-exaratum (Mannerheim, 1853) -K------

aldanicum Poppius, 1906gracilipes (Duftschmid, 1812) FKSNDEAImarginatum (Linnaeus, 1758) FKSNDEAIsexpunctatum (Linnaeus, 1758) FKSNDEAIimpressum (Panzer, 1796) -K---EAIericeti (Panzer, 1809) FKSNDEAIviridicupreum (Goeze, 1777) -----E--muelleri (Herbst, 1784) FKSNDEAIchalconotum Ménétriés, 1832 -K----A-

sahlbergii (Chaudoir, 1850)archangelicum (J.Sahlberg, 1875)

dolens (Sahlberg, 1827) FKSNDEAIversutum Sturm, 1824 FKSNDEAIhypocrita (Apfelbeck, 1904) FKS-D-A-viduum (Panzer, 1796) FKSNDEAIafrum (Duftschmid, 1812) FKSNDEAI

moestum auct. nec (Duftschmid, 1812)duftschmidi Schmidt, 1994 --S-D-AI

moestum (Duftschmid, 1812) nec (Gmelin, 1790)lugens (Duftschmid, 1812) F-S-DEAI

Zabrini Bonelli, 1810Amarina Zimmermann, 1831Amara Bonelli, 1810strenua Zimmermann, 1832 ----D-A-tricuspidata Dejean, 1831 --S-D---plebeja (Gyllenhal, 1810) FKSNDEAIconcinna Zimmermann, 1832 -------Isimilata (Gyllenhal, 1810) FKSNDEAIovata (Fabricius, 1792) FKSNDEAImontivaga Sturm, 1825 FKSN----nitida Sturm, 1825 FKSNDEAIcommunis (Panzer, 1797) FKSNDEAIpulpani Kult, 1949 ------A-

pseudocommunis Burakowski, 1957convexior Stephens, 1828 FKSND-AInigricornis Thomson, 1857 FKSN----lunicollis Schiödte, 1837 FKSNDEAIcurta Dejean, 1828 FKSNDEAIlittorea Thomson, 1857 FKSNDEAIaenea (DeGeer, 1774) FKSNDEAIspreta Dejean, 1831 FKSNDEAIfamelica Zimmermann, 1832 FKSNDEAIeurynota (Panzer, 1796) FKSNDEAIfamiliaris (Duftschmid, 1812) FKSNDEAIanthobia Villa & Villa, 1833 --S-D-A-lucida (Duftschmid, 1812) FKSNDEAI


tibialis (Paykull, 1798) FKSNDEAIerratica (Duftschmid, 1812) FKSN-EAIinterstitialis Dejean, 1828 FKSN----ingenua (Duftschmid, 1812) FKSNDEAIfusca Dejean, 1828 --S-D---cursitans Zimmermann, 1832 FKSNDEAImunicipalis (Duftschmid, 1812) FKSNDEAIbifrons (Gyllenhal, 1810) FKSNDEAIinfima (Duftschmid, 1812) FKSNDEAIpraetermissa (Sahlberg, 1827) FKSNDEAIbrunnea (Gyllenhal, 1810) FKSNDEAIquenseli (Schönherr, 1806) FKSNDEAIcrenata Dejean, 1828 F-S-D-AIapricaria (Paykull, 1790) FKSNDEAIfulva (Müller, 1776) FKSNDEAImajuscula (Chaudoir, 1850) FKSNDEAIconsularis (Duftschmid, 1812) FKSNDEAIaulica (Panzer, 1796) FKSNDEAIgebleri Dejean, 1831 FKSNDEAI

helleri Gredler, 1868convexiuscula (Marsham, 1802) FKSNDEA-torrida (Panzer, 1796) FKSN----alpina (Paykull, 1790) FKSN----hyperborea Dejean, 1831 FKSN----equestris (Duftschmid, 1812) FKSNDEAI

Zabrina Bonelli, 1810Zabrus Clairville, 1806tenebrioides (Goeze, 1777) --S-D---

Panagaeini Bonelli, 1810Panagaeus Latreille, 1802cruxmajor (Linnaeus, 1758) FKSNDEAIbipustulatus (Fabricius, 1775) FKSNDEAI

Chlaeniini Brullé, 1834Chlaenius Bonelli, 1810tristis (Schaller, 1783) FKSNDEAIspoliatus (Rossi, 1790) -------Inigricornis (Fabricius, 1787) FKSNDEAInitidulus (Schrank, 1781) FKS-DEAItibialis Dejean, 1826 ------A-

kindermanni auct. nec Chaudoir, 1856vestitus (Paykull, 1790) --S-D-AIsulcicollis (Paykull, 1798) F-S-DEAIquadrisulcatus (Paykull, 1790) -KS-DEA-costulatus (Motschulsky, 1859) FKS--EAI

Oodini LaFerté-Sénectère, 1851Oodes Bonelli, 1810helopioides (Fabricius, 1792) FKSNDEAI

gracilis Villa & Villa, 1833 --S--EAI

Licinini Bonelli, 1810Badister Clairville, 1806unipustulatus Bonelli, 1813 FKS-DEAIbullatus (Schrank, 1798) FKSNDEAI

bipustulatus (Fabricius, 1792) nec (Fabricius, 1775)meridionalis Puel, 1925 --S-DEAIlacertosus Sturm, 1815 FKSNDEAIsodalis (Duftschmid, 1812) -KSNDEAIdorsiger (Duftschmid, 1812) ----D-A-peltatus (Panzer, 1796) FKSNDEAIdilatatus Chaudoir, 1837 FKSNDEAIcollaris Motschulsky, 1844 --S-DEAI

anomalus (Perris, 1866)striatulus Hansen, 1944

Licinus Latreille, 1802depressus (Paykull, 1790) F-SNDEAI

Harpalini Bonelli, 1810Harpalina Bonelli, 1810Ophonus Stephens, 1828rupicola (Sturm, 1818) --S-D-A-laticollis (Mannerheim, 1825) FKS-DEAI

nitidulus Stephens, 1828punctatulus (Duftschmid, 1812) nec (Fabricius, 1792)

puncticollis (Paykull, 1798) FKSNDEAImelletii (Heer, 1837) --S-D---rufibarbis (Fabricius, 1792) FKSNDEAI

seladon (Schauberger, 1926)schaubergerianus (Puel, 1937) F-----A-puncticeps Stephens, 1828 -KS-D-A-azureus (Fabricius, 1775) --S-DEA-

Harpalus Latreille, 1802Pseudoophonus Motschulsky, 1844Semiophonus Schauberger, 1933

griseus (Panzer, 1796) FKSNDEAIrufipes (DeGeer, 1774) FKSNDEAI

pubescens (Müller, 1776)calceatus (Duftschmid, 1812) FKSNDEAImelancholicus Dejean, 1829 --S-D---signaticornis (Duftschmid, 1812) --S-DE-Iaffinis (Schrank, 1781) FKSNDEAI

aeneus (Fabricius, 1775) nec (DeGeer, 1774)distinguendus (Duftschmid, 1812) FKSNDEAIsmaragdinus (Duftschmid, 1812) FKSNDEAIcaspius (Steven, 1806) -------I

roubali Schauberger, 1928serripes (Quensel, 1806) --S-D--I


autumnalis (Duftschmid, 1812) --S---AIatratus Latreille, 1804 -------Isolitaris Dejean, 1829 FKSNDEAI

fuliginosus (Duftschmid, 1812) nec (Panzer, 1809)latus (Linnaeus, 1758) FKSNDEAInigritarsis Sahlberg, 1827 FKS-----luteicornis (Duftschmid, 1812) FKSN-EAIxanthopus Gemminger & Harold, 1868ssp. winkleri Schauberger, 1923 FKSNDEAIlaevipes Zetterstedt, 1828 FKSNDEAI

quadripunctatus Dejean, 1829rubripes (Duftschmid, 1812) FKSNDEAIprogrediens Schauberger, 1922 -----EA-rufipalpis Sturm, 1818 F-S-DEAI

rufitarsis (Duftschmid, 1812) nec (Illiger, 1802)neglectus Audinet-Serville, 1821 --S-D--Iservus (Duftschmid, 1812) --S-D-AItardus (Panzer, 1796) FKSNDEAImodestus Dejean, 1829 -------Ianxius (Duftschmid, 1812) FKS-DEAIpicipennis (Duftschmid, 1812) --S-DEAIpumilus Sturm, 1818 --S-D-AI

vernalis (Fabricius, 1801) nec (Panzer, 1796)froelichii Sturm, 1818 -KS-DEAIhirtipes (Panzer, 1797) --S-DEAIflavescens (Piller & Mitterpacher, 1783)FKS-D-AI

rufus Brüggemann, 1873

Microderes Faldermann, 1835brachypus (Steven, 1809) F-------

Anisodactylina Lacordaire, 1854Anisodactylus Dejean, 1829poeciloides (Stephens, 1828) --S-D---binotatus (Fabricius, 1787) FKSNDEAInemorivagus (Duftschmid, 1812) FK--D--Isignatus (Panzer, 1797) ----D-AI

Diachromus Erichson, 1837germanus (Linnaeus, 1758) ----D-AI

Stenolophina Kirby, 1837Stenolophus Dejean, 1821teutonus (Schrank, 1781) --S-D-AIskrimshiranus Stephens, 1828 --S-D---mixtus (Herbst, 1784) FKSNDEAI

Bradycellus Erichson, 1837Tetraplatypus Tschitschérine, 1897

ruficollis (Stephens, 1828) FKSNDEAIsimilis (Dejean, 1829)

verbasci (Duftschmid, 1812) F-SND---

harpalinus (Audinet-Serville, 1821) -KSNDEAIcsikii Laczó, 1912 F-S-D---caucasicus (Chaudoir, 1846) FKSNDEAI

collaris (Paykull, 1798) nec (Herbst, 1784)

Dicheirotrichus Jacquelin du Val, 1857Trichocellus Ganglbauer, 1892

gustavii Crotch, 1871 -KSND---pubescens (Paykull, 1790) nec (Müller, 1776)

mannerheimii (F.Sahlberg, 1844) -K------rufithorax (Sahlberg, 1827) FKSN-EAIcognatus (Gyllenhal, 1827) FKSNDEAIplacidus (Gyllenhal, 1827) FKSNDEAI

Acupalpus Dejean, 1829flavicollis (Sturm, 1825) FKSNDEAIbrunnipes (Sturm, 1825) --SN-E-Imeridianus (Linnaeus, 1761) FKSNDEAIsuturalis Dejean, 1829 ------A-parvulus (Sturm, 1825) FKSNDEAI

dorsalis (Fabricius, 1787) nec (Pontoppidan, 1763)dubius Schilsky, 1888 --S-DEA-exiguus Dejean, 1829 FKS-DEAIelegans (Dejean, 1829) ----D---interstitialis Reitter, 1884 -----EA-

Anthracus Motschulsky, 1850consputus (Duftschmid, 1812) F-S-DEAI

Perigonini Horn, 1881Perigona Laporte de Castelnau, 1835Trechicus LeConte, 1853

nigriceps (Dejean, 1831) F-SND-A-

Cyclosomini Laporte de Castelnau, 1834Masoreus Dejean, 1821wetterhallii (Gyllenhal, 1813) --SNDEAI

Odacanthini Laporte de Castelnau, 1834Odacantha Paykull, 1798melanura (Linnaeus, 1767) FKSNDEAI

Lebiini Bonelli, 1810Lebiina Bonelli, 1810Lebia Latreille, 1802cyanocephala (Linnaeus, 1758) FKSNDEAIchlorocephala (Hoffmannsegg, 1803) FKSNDEAIcruxminor (Linnaeus, 1758) FKSNDEAI

Demetriina Bates, 1886Demetrias Bonelli, 1810monostigma Samouelle, 1819 F-S-DEAIatricapillus (Linnaeus, 1758) --S-D-A-imperialis (Germar, 1824) F-SNDEAI


Dromiina Bonelli, 1810Paradromius Fowler, 1887longiceps (Dejean, 1826) F-S-DEA-linearis (Olivier, 1795) FKSNDEAI

Dromius Bonelli, 1810agilis (Fabricius, 1787) FKSNDEAIangustus Brullé, 1834 --SND---quadraticollis Morawitz, 1862 FKS--EAImeridionalis Dejean, 1825 ----D---schneideri Crotch, 1871 FKSNDEAI

marginellus (Fabricius, 1794) nec (Herbst, 1784)fenestratus (Fabricius, 1794) FKSNDEAIquadrimaculatus (Linnaeus, 1758) F-SNDEAI

Calodromius Reitter, 1905spilotus (Illiger, 1798) F-SNDEAI

quadrinotatus (Panzer, 1800) nec (Fabricius, 1798)

Philorhizus Hope, 1838quadrisignatus (Dejean, 1825) --S-D---sigma (Rossi, 1790) FKSNDEAImelanocephalus (Dejean, 1825) --S-D---notatus (Stephens, 1827) F-SNDEA-

nigriventris (Thomson, 1857)

Microlestes Schmidt-Goebel, 1846minutulus (Goeze, 1777) FKS-DEAImaurus (Sturm, 1827) FKS-DEAI

Lionychina Jeannel, 1948Syntomus Hope, 1838truncatellus (Linnaeus, 1761) FKSNDEAI

ai Barševskis, 1993foveatus (Geoffroy, 1785) FKSNDEAI

Lionychus Wissmann, 1846quadrillum (Duftschmid, 1812) F-S-----

tuberculatus Gyllenhal, 1808 FKSNDEAIsibiricus (Motschulsky, 1860) FKSN----

fennicus auct. nec Paykull, 1798aquaticus (Linnaeus, 1758) FK--DEAIaequalis Thomson, 1868 F-SND-A-grandis Illiger, 1798 FKSNDEAIstrandi Angus, 1970 FKSN----brevipalpis Bedel, 1881 FKSNDEAIarvernicus Mulsant, 1846 FK--DEAIglacialis Villa, 1833 FKSN----granularis (Linnaeus, 1761) FKSNDEAIdiscrepans Rey, 1885 FK---E--minutus Fabricius, 1775 FKSNDEAIlapponicus Thomson, 1853 FKSNDE--fulgidicollis Motschulsky, 1860 F-SND---griseus Herbst, 1793 F-SNDEAInanus Sturm, 1836 FKSNDEAIredtenbacheri Kuwert, 1885 --S-DE-Ipumilio Erichson, 1837 ----D---pallidus Gebler, 1830 FKSN-E--laticollis Thomson, 1853 FKSNDE--strigifrons Thomson, 1868 FKSNDEAIasperatus Rey, 1885 --S-D---flavipes Fabricius, 1792 FKSNDEAIobscurus Mulsant, 1844 --SND---

walkeri Sharp, 1916

Georissinae Laporte de Castelnau, 1840Georissus Latreille, 1809crenulatus (Rossi, 1794) FKSNDEAI

Hydrochinae Thomson, 1859Hydrochus Leach, 1817elongatus (Schaller, 1783) F-S-D-A-ignicollis Motschulsky, 1860 FKSNDEAI

elongatus auct. nec (Schaller, 1783)crenatus (Fabricius, 1792) F-S-DEAI

carinatus Germar, 1824brevis (Herbst, 1793) FKSNDEAImegaphallus van Berge Henegouwen, 1988 F-SNDE--angustatus Germar, 1824 -----EA-

Spercheinae Erichson, 1837Spercheus Illiger, 1798emarginatus (Schaller, 1783) -KS-DEAI

Cymindina Laporte de Castelnau, 1834Cymindis Latreille, 1806humeralis (Geoffroy, 1785) --S-D-AIangularis Gyllenhal, 1810 FKSNDEAImacularis Mannerheim, 1824 FKSNDEAIvaporariorum (Linnaeus, 1758) FKSNDEAIBrachininae Bonelli, 1810Brachinus Weber, 1801crepitans (Linnaeus, 1758) F-S-DEA-

POLYPHAGAHYDROPHILOIDEAHydrophilidae Latreille, 1802Helophorinae Leach, 1815Helophorus Fabricius, 1775nubilus Fabricius, 1777 FKSNDEAI

Hydrophilinae Latreille, 1802Berosini Mulsant, 1844Berosus Leach, 1817spinosus (Steven, 1808) F-SNDE--fulvus Kuwert, 1888 --S-D---


signaticollis (Charpentier, 1825) F-S?DE-Iluridus (Linnaeus, 1761) FKSNDEAI

Chaetarthriini Bedel, 1881 (1844)Chaetarthria Stephens, 1832seminulum (Herbst, 1797) FKSNDEAI

Anacaenini Hansen, 1991Paracymus Thomson, 1867aeneus (Germar, 1824) --SND---

Anacaena Thomson, 1859globulus (Paykull, 1798) F-SNDEA-lutescens (Stephens, 1829) FKSNDEAI

limbata auct. nec (Fabricius, 1792)limbata (Fabricius, 1792) F-S-D-A-

Laccobiini Bertrand, 1954Oocyclini Hansen, 1991

Laccobius Erichson, 1837decorus (Gyllenhal, 1827) F-S--E--minutus (Linnaeus, 1758) FKSNDEAIcolon (Stephens, 1829) FKSND-A-

biguttatus Gerhardt, 1877sinuatus Motschulsky, 1849 FKS-D--Istriatulus (Fabricius, 1801) -KSNDEAIbipunctatus (Fabricius, 1775) FKSNDEAI

Hydrophilini Latreille, 1802Helochares Mulsant, 1844punctatus Sharp, 1869 ----D--I

lividus auct. nec (Forster, 1771)obscurus (Müller, 1776) FKSNDEAI

griseus auct. nec (Fabricius, 1787Enochrus Thomson, 1859melanocephalus (Olivier, 1792) FKSNDEAIochropterus (Marsham, 1802) FKSNDEAI

frontalis (Erichson, 1837)fuscipennis (Thomson, 1884) FKSND-AI

sahlbergi (Fauvel, 1887)quadripunctatus (Herbst, 1797) FKSNDEAIhalophilus (Bedel, 1878) --S-D---bicolor (Fabricius, 1792) FKSNDEAItestaceus (Fabricius, 1801) FKSNDEAIaffinis (Thunberg, 1794) FKSNDEAIcoarctatus (Gredler, 1863) FKSNDEAI

Cymbiodyta Bedel, 1881marginella (Fabricius, 1792) FKSNDEAI

Hydrobius Leach, 1815fuscipes (Linnaeus, 1758) FKSNDEAI

subrotundus Stephens, 1829rottenbergi Gerhardt, 1872

arcticus Kuwert, 1890 FKSN----

Limnoxenus Motschulsky, 1853niger (Gmelin, 1790) ----D---

Hydrochara Berthold, 1827caraboides (Linnaeus, 1758) FKSNDEAIflavipes (Steven, 1808) ------A-

Hydrophilus Geoffroy, 1762Hydrous Linnaeus, 1775

piceus (Linnaeus, 1758) F-SND-AIaterrimus Eschscholtz, 1822 FKS-DEAI

Sphaeridiinae Latreille, 1802Coelostomatini Heyden, Reitter & Weise, 1891Coelostoma Brullé, 1835orbiculare (Fabricius, 1775) FKSNDEAI

Megasternini Mulsant, 1844Cercyon Leach, 1817ustulatus (Preyssler, 1790) FKSNDEAIlaminatus Sharp, 1873 F-S-DEAIlittoralis (Gyllenhal, 1808) FKSNDEAIdepressus Stephens, 1829 FKSNDE--obsoletus (Gyllenhal, 1808) FKSNDEAI

lugubris auct. nec (Olivier, 1790)impressus (Sturm, 1807) FKSNDEAIhaemorrhoidalis (Fabricius, 1775) FKSNDEAImelanocephalus (Linnaeus, 1758) FKSNDEAIemarginatus Baranowski, 1985 F-SN----borealis Baranowski, 1985 F-SN----lateralis (Marsham, 1802) FKSNDEAIbifenestratus Küster, 1851 FKSNDEAImarinus Thomson, 1853 FKSNDEAIunipunctatus (Linnaeus, 1758) FKSNDEAIquisquilius (Linnaeus, 1761) FKSNDEAIterminatus (Marsham, 1802) FKSNDEAIpygmaeus (Illiger, 1801) FKSNDEAInigriceps (Marsham, 1802) FKSNDEA-

atricapillus (Marsham, 1802)granarius Erichson, 1837 -K--D-AItristis (Illiger, 1801) FKSNDEAIconvexiusculus Stephens, 1829 FKSNDEAIsternalis Sharp, 1918 F-S-D-A-analis (Paykull, 1798) FKSNDEAI

Megasternum Mulsant, 1844concinnum (Marsham, 1802) FKSNDE-I

obscurum (Marsham, 1802)

Cryptopleurum Mulsant, 1844subtile Sharp, 1884 F-SND---


minutum (Fabricius, 1775) FKSNDEAIcrenatum (Panzer, 1794) FKSNDEAI

Sphaeridiini Latreille, 1802Sphaeridium Fabricius, 1775bipustulatum Fabricius, 1781 FKSNDEAImarginatum Fabricius, 1787 F-SND--Isubstriatum Faldermann, 1838 --S-D--Ilunatum Fabricius, 1792 FKSNDEAIscarabaeoides (Linnaeus, 1758) FKSNDEAI

HISTEROIDEASphaeritidae Shuckard, 1839Sphaerites Duftschmid, 1805glabratus (Fabricius, 1792) FKSNDEAI

Histeridae Gyllenhal, 1808Abraeinae MacLeay, 1819Abraeini MacLeay, 1819Chaetabraeus Portevin, 1929globulus (Creutzer, 1799) --S-DE-I

Abraeus Leach, 1817granulum Erichson, 1839 --S-D---perpusillus (Marsham, 1802) --S-DEAI

globosus (Hoffmann, 1803)parvulus Aubé, 1842 --S-----

Plegaderini Portevin, 1929Plegaderus Erichson, 1834saucius Erichson, 1834 FKSN-EAIvulneratus (Panzer, 1797) FKSNDEAIcaesus (Herbst, 1792) F-SN--AIdissectus Erichson, 1839 --S-D--Isanatus Truqui, 1852ssp. gobanzi Müller, 1902 --S-----

planiusculus Motschulsky, 1849 FKSNDEAIcuspidatus Ihssen, 1949

semistriatus (Scriba, 1790) FKSNDEAIsubnitescens Bickhardt, 1909 ----D-A-tenuistrius Marseul, 1855 ------A-virescens (Paykull, 1798) --S-DEAIimmundus (Gyllenhal, 1827) FKS-DEAIaeneus (Fabricius, 1775) F-SNDEAI

Chalcionellus Reichardt, 1932decemstriatus (Rossi, 1792) --S-DEAI

conjungens (Paykull, 1798)

Hypocacculus Bickhardt, 1916rufipes (Kugelann, 1792) --S-D-AI

Hypocaccus Thomson, 1867Baeckmanniolus Reichardt, 1926

rugiceps (Duftschmid, 1805) FKSND-AImetallicus (Herbst, 1792) --SNDEAIrugifrons (Paykull, 1798) FKSNDEAIdimidiatus (Illiger, 1807)ssp. maritimus (Stephens, 1830) --S-D---

Gnathoncus Jacquelin du Val, 1858rotundatus (Kugelann, 1792) FKSNDEAI

nanus (Scriba, 1790) nec (Piller & Mitterpacher, 1783)punctulatus Thomson, 1862

nannetensis (Marseul, 1862) FKSNDEAIcommunis (Marseul, 1862) FKSND---

schmidti Reitter, 1894buyssoni Auzat, 1917 FKSND--Inidorum Stockmann, 1957 F-S---AI

Myrmetes Marseul, 1862paykulli Kanaar, 1979 FKSNDEAI

piceus (Paykull, 1809) nec (Marsham, 1802)Acritini Wenzel, 1944Acritus LeConte, 1853minutus (Herbst, 1792) FKSNDEAInigricornis (Hoffmann, 1803) FKSNDEA-komai Lewis, 1879 ----D---homoeopathicus Wollaston, 1857 ---ND-A-

Aeletes Horn, 1873atomarius (Aubé, 1842) --S-D---

Teretriini Bickhardt, 1914Teretrius Erichson, 1834fabricii Mazur, 1972 F-S--EAI

picipes (Fabricius, 1792) nec (Olivier, 1789)Saprininae Blanchard, 1845Saprinus Erichson, 1834rugifer (Paykull, 1809) FKSND-AI

quadristriatus auct. nec (Thunberg, 1794)

Dendrophilinae Reitter, 1909Dendrophilini Reitter, 1909Dendrophilus Leach, 1817corticalis (Paykull, 1798) FKSNDEAI

punctatus (Herbst, 1792) nec (Müller, 1776)pygmaeus (Linnaeus, 1758) FKSNDEAI

Paromalini Reitter, 1909Carcinops Marseul, 1855pumilio (Erichson, 1834) F-SNDEAI

quatuordecimstriata (Stephens, 1835)

Platylomalus Cooman, 1948complanatus (Panzer, 1797) -K-N--A-


Paromalus Erichson, 1834Microlomalus Lewis, 1907

flavicornis (Herbst, 1792) F-SNDEA-parallelepipedus (Herbst, 1792) FKSNdEAI

Onthophilinae MacLeay, 1819Onthophilus Leach, 1817striatus (Forster, 1771) --S-DE--punctatus (Müller, 1776) ----D---

sulcatus (Moll, 1784)

Histerinae Gyllenhal, 1808Histerini Gyllenhal, 1808Margarinotus Marseul, 1853Paralister Bickhardt, 1917Stenister Reichardt, 1926Grammostethus Lewis, 1906

brunneus (Fabricius, 1775) FKSNDEAIimpressus (Fabricius, 1798)cadaverinus (Hoffmann, 1803)

striola (Sahlberg, 1819) FKSNDEAIsuccicola (Thomson, 1862)

merdarius (Hoffmann, 1803) FKSNDEAIterricola (Germar, 1824) F-S-DEAIbipustulatus (Schrank, 1781) ----D-AIobscurus (Kugelann, 1792) F-SNDEAI

stercorarius (Hoffmann, 1803)ventralis (Marseul, 1854) FKSNDEAIneglectus (Germar, 1813) FKSNDEAIpurpurascens (Herbst, 1792) FKSNDEAIcarbonarius (Hoffmann, 1803) -KSNDEAImarginatus (Erichson, 1834) --S-DEA-

Hister Linnaeus, 1758quadrimaculatus Linnaeus, 1758 -----EA-unicolor Linnaeus, 1758 FKSNDEAIquadrinotatus Scriba, 1790 ----DEAIilligeri Duftschmid, 1805 -----E--

uncinatus Illiger, 1807helluo Truqui, 1852 FK---EAIbissexstriatus Fabricius, 1801 FKSNDEAIfunestus Erichson, 1834 FKSNDEA-

Atholus Thomson, 1859bimaculatus (Linnaeus, 1758) FKSNDEAIduodecimstriatus (Schrank, 1781) FKSNDEAIpraetermissus (Peyron, 1856) ----DE--corvinus (Germar, 1817) F-SNDEAI

Platysomatini Bickhardt, 1914Platysoma Leach, 1817Eurylister Bickhardt, 1920Cylister Cooman, 1941

minus (Rossi, 1792) FKSNDEAIfrontale (Paykull, 1798)

compressum (Herbst, 1783) FKS-D-AIdeplanatum (Gyllenhal, 1808) FKSN-EAIangustatum (Hoffmann, 1803) FKSNDEAI

ferrugineum (Thunberg, 1794) nec (Olivier, 1789)lineare Erichson, 1834 FKSNDEAIelongatum (Thunberg, 1787) FKS---AI

oblongum (Fabricius, 1792)

Hololeptini Hope, 1840Hololepta Paykull, 1811plana (Sulzer, 1776) FK-NDEAI

Hetaeriinae Marseul, 1857Hetaerius Erichson, 1834ferrugineus (Olivier, 1789) --S-D-AI

STAPHYLINOIDEAHydraenidae Mulsant, 1844Hydraeninae Mulsant, 1844Limnebius Leach, 1815truncatellus (Thunberg, 1794) FKSNDEAIpapposus Mulsant, 1844 --S-D-AIcrinifer Rey, 1885 FKS-DE-Iparvulus (Herbst, 1797) FKSNDEAI

truncatulus Thomson, 1853nitidus (Marsham, 1802) F-S-DE-I

picinus (Marsham, 1802)aluta Bedel, 1881 FKSND---

Hydraena Kugelann, 1794testacea Curtis, 1830 --S-D---palustris Erichson, 1837 FKSNDEA-britteni Joy, 1907 FKSNDE-Iriparia Kugelann, 1794 FKSNDEAIreyi Kuwert, 1888 ------A-

sternalis auct. nec Rey, 1893nigrita Germar, 1824 --SNDE--pulchella Germar, 1824 FKS-DEA-gracilis Germar, 1824 FKSNDEAIexcisa Kiesenwetter, 1849 F-----A-

Ochthebiinae Thomson, 1859Ochthebius Leach, 1815auriculatus Rey, 1886 --S-D---dilatatus Stephens, 1829 --S-D---


bicolon Germar, 1824 F-SND---czwalinai Kuwert, 1887 F-------

stockmanni Balfour-Browne, 1948kaninensis Poppius, 1909 F-------minimus (Fabricius, 1792) FKSNDEAI

impressus (Marsham, 1802)remotus Reitter, 1887 ------A-alpinus (Ieniåtea, 1979) FKSN----

rugulosus auct. nec (Wollaston, 1857)hungaricus Endrödy-Younga, 1967 FK---E--nilssoni Hebauer, 1986 --S-----metallescens Rosenhauer, 1847 -------Ipusillus Stephens, 1835 -------Imarinus (Paykull, 1798) FKSNDEA-lenensis Poppius, 1907 -K-N----viridis Peyron, 1858 --S-D---evanescens J.Sahlberg, 1875 -K------

Ptiliidae Heer, 1843Ptiliinae Heer, 1843Nossidium Erichson, 1845pilosellum (Marsham, 1802) ----D---

Ptenidium Erichson, 1845gressneri Erichson, 1845 --S-D-AIlaevigatum Erichson, 1845 F-SND-A-turgidum Thomson, 1855 --SND-AIintermedium Wankowicz, 1869 --SND-AIfuscicorne Erichson, 1845 FKSND--Ipunctatum (Gyllenhal, 1827) F-SNDE--formicetorum Kraatz, 1851 FKSNDEA-

myrmecophilum (Motschulsky, 1845) nec (Allibert, 1844)pusillum (Gyllenhal, 1808) FKSNDEAIlongicorne (Fuss, 1868) ------A-nitidum (Heer, 1841) FKSNDEA-

Actidium Matthews, 1868boudieri (Allibert, 1844) ----D-A-coarctatum (Haliday, 1855) --S-D---

Oligella Motschulsky, 1868foveolata (Allibert, 1844) FKSND-A-nana (Strand, 1946) --SN----intermedia Besuchet, 1971 --S-----

Micridium Motschulsky, 1868angulicolle (Fairmaire, 1857) --S-----halidaii (Matthews, 1868) F-SND-A-

affine Erichson, 1845 -KS-D---caesum Erichson, 1845 --S-----myrmecophilum (Allibert, 1844) FKSND-A-exaratum (Allibert, 1844) FKSNDEA-horioni Rosskothen, 1934 -KSND---modestum Wankowicz, 1869 F-SND--I

Euryptilium Matthews, 1872gillmeisteri Flach, 1889 ---ND-A-saxonicum (Gillmeister, 1845) FKSND---

marginatum auct. nec (Aubé, 1850)

Ptiliola Haldeman, 1848Nanoptilium Flach, 1889

kunzei (Heer, 1841) FKSNDEA-brevicollis (Matthews, 1860) F-SND-A-

Ptiliolum Flach, 1888stockmanni Besuchet, 1971 F-------caledonicum (Sharp, 1871) FKSN-E--

croaticum (Matthews, 1872)schwarzi (Flach, 1887) F-SNDE--wuesthoffi Rosskothen, 1934 F-SND---spencei (Allibert, 1844) F-SNDE--fuscum (Erichson, 1845) F-SND---

flachi (Reitter, 1909)marginatum (Aubé, 1850) --S-D---

lederi (Flach, 1888)sahlbergi (Flach, 1888) FKSN-EA-

Microptilium Matthews, 1872palustre Kuntzen, 1914 ----D---

Nanosellinae Barber, 1924Baranowskiella Sörensson, 1997ehnstromi Sörensson, 1997 F-SN----

Acrotrichinae Reitter, 1909 (1856)Ptinella Motschulsky, 1844limbata (Heer, 1841) F-SND--I

testacea (Heer, 1841)denticollis (Fairmaire, 1858) FKS-D---johnsoni Rutanen, 1985 F-S-----aptera (Guérin-Ménéville, 1839) F-SND---britannica Matthews, 1858 ----D---tenella (Erichson, 1845) FKSND---microscopica (Gillmeister, 1845) FKS-----

Pteryx Matthews, 1858suturalis (Heer, 1841) FKSND-A-splendens Strand, 1960 F-SN----Ptilium Gyllenhal, 1827

Millidium Motschulsky, 1855minutissimum (Ljungh, 1804) FKSNDEA-

Nephanes Thomson, 1859titan (Newman, 1834) FKSND-AI


Smicrus Matthews, 1872filicornis (Fairmaire & Laboulbène, 1855) FKSND-A-

Baeocrara Thomson, 1859variolosa (Mulsant & Rey, 1873) FKSND--I

littoralis (Thomson, 1855) nec (Motschulsky, 1845)japonica (Matthews, 1884) F-SND-A-

Acrotrichis Motschulsky, 1848grandicollis (Mannerheim, 1844) FKSNDEAImontandonii (Allibert, 1844) FKSNDEAIchevrolatii (Allibert, 1844) FKSND---thoracica (Waltl, 1838) FKSND-A-sericans (Heer, 1841) FKSNDEA-

picicornis auct. nec (Mannerheim, 1843)dispar (Matthews, 1865) FKSNDEAIbrevipennis (Erichson, 1845) FKSNDEA-pumila (Erichson, 1845) FKSNDE--

longicornis auct. nec (Mannerheim, 1844)silvatica Rosskothen, 1935 FKSND---parva Rosskothen, 1935 FKSND-A-volans (Motschulsky, 1845) FKSN----

fennica Renkonen, 1939cognata (Matthews, 1877) FKSNDE--

platonoffi Renkonen, 1945insularis (Mäklin, 1852) F-SND---norvegica Strand, 1941 FKSND---arnoldi Rosskothen, 1935 --S-D---intermedia (Gillmeister, 1845) FKSNDEAIrosskotheni Sundt, 1971 F-SND---

fraterna Johnson, 1975henrici (Matthews, 1872) ---ND---

fratercula (Matthews, 1878)atomaria (DeGeer, 1774) FKSNDEAIlucidula Rosskothen, 1935 F-SND---danica Sundt, 1958 --S-D---sitkaensis (Motschulsky, 1845) FKSNDEA-

fratercula auct. nec (Matthews, 1878)sjobergi Sundt, 1958 F-SN----fascicularis (Herbst, 1793) FKSNDEAIrugulosa Rosskothen, 1935 FKSNDE--strandi Sundt, 1958 FKSND-A-suecica Sundt, 1958 --SND---

Actinopteryx Matthews, 1872fucicola (Allibert, 1844) --S-D---

Agyrtidae Thomson, 1859Pterolominae Thomson, 1862Pteroloma Gyllenhal, 1827forsstromii (Gyllenhal, 1810) FKSN-E--

Agyrtinae Thomson, 1859Ecanus Stephens, 1833glaber (Fabricius, 1787) F-S-----

Agyrtes Frölich, 1799bicolor Laporte de Castelnau, 1840 --S-D---

Leiodidae Fleming, 1821Leiodinae Fleming, 1821Sogdini Lopatin, 1961Triarthron Märkel, 1840maerkelii Märkel, 1840 FKSND---

Sogda Lopatin, 1961Trichohydnobius Vogt, 1961

suturalis (Zetterstedt, 1828) FKSND--Iperrisii (Fairmaire, 1855)

ciliaris (Thomson, 1874) FKSN----hyperborea (Strand, 1943)

Hydnobius Schmidt, 1841septentrionalis Thomson, 1874 FKSN----punctatus (Sturm, 1807) FK--D-A-latifrons (Curtis, 1840) FKSN----

strigosus Schmidt, 1841tibialis J.Sahlberg, 1903 F--N----spinipes (Gyllenhal, 1813) FKSN-E--edentatus (Sahlberg, 1833) F-------

orientalis Hlisnikovsky, 1967claviger Strand, 1943 --SN----multistriatus (Gyllenhal, 1813) --S-D---

Leiodini Fleming, 1821Leiodes Latreille, 1796punctulata (Gyllenhal, 1810) FKSN----inordinata (J.Sahlberg, 1898) FKSN----silesiaca (Kraatz, 1852) FKSND-AIhybrida (Erichson, 1845) F-SN----rugosa Stephens, 1829 F-S-DEA-brandisi Holdhaus, 1902 F-------ciliaris (Schmidt, 1841) FKSND--Ipallens (Sturm, 1807) -K------triepkii (Schmidt, 1841) FKSND-A-rubiginosa (Schmidt, 1841) F-S-D---polita (Marsham, 1802) FKSNDEAI

calcarata (Erichson, 1845)lucens (Fairmaire, 1855) F-SN----cinnamomea (Panzer, 1793) ??--D---oblonga (Erichson, 1845) FKSNDE-Ifracta (Seidlitz, 1874) FKSN--A-

rhaetica auct. nec (Erichson, 1845)


furva (Erichson, 1845) FKS-DE--longipes (Schmidt, 1841) F-SND---

curta (Fairmaire & Laboulbène, 1854)picea (Panzer, 1797) FKSND-AIflavescens (Schmidt, 1841) F-SNDE-Iruficollis J.Sahlberg, 1898 FKS-DE--

nigrita auct. nec (Schmidt, 1841)litura Stephens, 1832 FKSND---obesa (Schmidt, 1841) FKSNDE-Idubia (Kugelann, 1794) FKSN-EAIrufipennis (Paykull, 1798) F-S-D---bicolor (Schmidt, 1841) F------Igallica Reitter, 1884 --S-D---ferruginea (Fabricius, 1787) FKSNDEA-

ovalis (Schmidt, 1841)sparreschneideri Strand, 1943 F-SN----puncticollis (Thomson, 1862) F-SN----

sahlbergi Fleischer, 1907badia (Sturm, 1807) FKSND--Igyllenhalii Stephens, 1829 FKSNDEAI

parvula (Sahlberg, 1833)

Xanthosphaera Fairmaire, 1859Liocyrtusa Daffner, 1982

minuta (Ahrens, 1812) FKSN--AIvittata (Curtis, 1840) FKSND---

pauxilla (Schmidt, 1841)

Cyrtusa Erichson, 1842subtestacea (Gyllenhal, 1813) FKSNDEAI

Pseudoliodini Portevin, 1926Colenis Erichson, 1842immunda (Sturm, 1807) FKSNDEA-

Agaricophagus Schmidt, 1841cephalotes Schmidt, 1841 --S-D-AI

Agathidiini Westwood, 1838Anisotoma Panzer, 1797humeralis (Fabricius, 1792) FKSNDEAIaxillaris Gyllenhal, 1810 FKSNDEAIcastanea (Herbst, 1792) FKSNDEAIglabra (Kugelann, 1794) FKSNDEAIorbicularis (Herbst, 1792) FKSNDEAI

Agathidium Panzer, 1797marginatum Sturm, 1807 FKSNDEAIhaemorrhoum Erichson, 1845 --S-D-A-varians Beck, 1817 F-SND--Iconvexum Sharp, 1866 --S-DEAI

piceum Erichson, 1845, nec Melsheimer, 1844mandibulare Sturm, 1807 FKSNDEA-pulchellum Wankowicz, 1869 FKS----Ipallidum (Gyllenhal, 1827) FKSN----rotundatum (Gyllenhal, 1827) FKSNDEAIconfusum Brisout de Barneville, 1863 FKSND-AIplagiatum (Gyllenhal, 1810) --S---A-nigrinum Sturm, 1807 FKSNDE--arcticum Thomson, 1862 FKSN----discoideum Erichson, 1845 FKSN----nigripenne (Fabricius, 1792) FKSNDEAIatrum (Paykull, 1798) FKSNDEAIseminulum (Linnaeus, 1758) FKSNDEAIlaevigatum Erichson, 1845 FKSNDEAIbadium Erichson, 1845 FKSNDEAIpisanum Brisout de Barneville, 1872 FKSN--AI

bicolor J.Sahlberg, 1881

Coloninae Horn, 1880 (1859)Colon Herbst, 1797latum Kraatz, 1850 FKSNDE--pseudolatum Palm, 1941 F-SN----rufescens Kraatz, 1850 F-SND-A-delarouzei Tournier, 1863 --SN----politum Peck & Stephan, 1996 --S-----curvipes Mäklin, 1880 F-S-----

cordigerum Szymczakowski, 1971arcticum Munster, 1911 F-SN----brundini Palm, 1941 F-SN----angulare Erichson, 1837 FKSND---dentipes (Sahlberg, 1822) F-SND---barnevillei Kraatz, 1858 F-S-D---

dubiosum Ihssen, 1951brunneum (Latreille, 1807) FKSNDEAIappendiculatum (Sahlberg, 1822) FKSND---

denticulatum Kraatz, 1850Liodopria Reitter, 1909serricornis (Gyllenhal, 1813) F-SN-EAI

Amphicyllis Erichson, 1845globus (Fabricius, 1792) FKSNDEAIglobiformis (Sahlberg, 1833) F-SNDEAI

calcaratum Erichson, 1837 -------Iserripes (Sahlberg, 1822) FKSND-A-puncticolle Kraatz, 1850 FKS-DE--viennense Herbst, 1797 FKSNDEAIbidentatum (Sahlberg, 1822) FKSNDEA-


Cholevinae Kirby, 1837Ptomaphagini Jeannel, 1911Ptomaphagus Illiger, 1798variicornis (Rosenhauer, 1847) --SND-AIsubvillosus (Goeze, 1777) FKSNDEAIsericatus (Chaudoir, 1845) F-SNDE-I

medius (Rey, 1889)

Nemadini Jeannel, 1936Nemadus Thomson, 1867colonoides (Kraatz, 1851) F-SNDEAI

Eocatops Peyerimhoff, 1924lapponicus Szymczakowski, 1975 F-S-----

Cholevini Kirby, 1837Nargus Thomson, 1867velox (Spence, 1815) FKS-DE--wilkinii (Spence, 1815) --SND-A-nikitanus (Reitter, 1884) --s-----anisotomoides (Spence, 1815) --S-DE--

Choleva Latreille, 1796spadicea (Sturm, 1839) --S-D---paskoviensis Reitter, 1913 ----D---agilis (Illiger, 1798) F-S-DEAIseptentrionis Jeannel, 1923 ---N----lederiana Reitter, 1902 FKS--E--

aquilonia Krogerus, 1926oblonga Latreille, 1807 FKS-DEAInivalis Kraatz, 1856 ------A-reitteri Petri, 1915 --S-D---cisteloides (Frölich, 1799) -----E-Iglauca Britten, 1918 FKS-DE--angustata (Fabricius, 1781) FKSNDEAIsturmii Brisout de Barneville, 1863 F-SN----fagniezi Jeannel, 1922 F-SND---spinipennis Reitter, 1890 FK---EA-jeanneli Britten, 1922 --S-D---elongata (Paykull, 1798) FKSNDEA-

Catopini Chaudoir, 1845Dreposcia Jeannel, 1922brevipalpis (Reitter, 1901)ssp. colletti (Münster, 1912) FK-N----umbrina (Erichson, 1837) --S-DEAI

Sciodrepoides Hatch, 1933Sciodrepa auct. nec Thomson, 1859

watsoni (Spence, 1815) FKSNDEAIfumatus (Spence, 1815) FKSNDEAIalpestris Jeannel, 1934 F-S-----

Catops Paykull, 1798subfuscus Kellner, 1846 FKSND-AIalpinus Gyllenhal, 1827 FKSN-E--luteipes Thomson, 1884 F-SN----longulus Kellner, 1846 F-SN----coracinus Kellner, 1846 FKSNDE-Igrandicollis Erichson, 1837 ----D-A-kirbii (Spence, 1815) --S-D--Itristis (Panzer, 1793) FKSND--Ichrysomeloides (Panzer, 1798) --S-D---neglectus Kraatz, 1852 --S-D---morio (Fabricius, 1787) FKSND-AInigrita Erichson, 1837 FKSNDEAInigriclavis Gerhardt, 1900 F-SND-A-

dohrni Reitter, 1913westi Krogerus, 1931 FKSND-AIfuscus (Panzer, 1794) FKSNDEAIfuliginosus Erichson, 1837 FKSNDE-Iborealis Krogerus, 1931 FKSN----nigricans (Spence, 1815) FKSND-AIpicipes (Fabricius, 1787) --SNDEAI

Platypsyllinae Ritsema, 1869Leptininae LeConte, 1872

Leptinus Müller, 1817testaceus Müller, 1817 --SND---

Platypsyllus Ritsema, 1869castoris Ritsema, 1869 --SN--A-

Scydmaenidae Leach, 1815Scydmaeninae Leach, 1815Eutheiini Casey, 1897Euthiconus Reitter, 1882conicicollis (Fairmaire & Laboulbène, 1855) --SND---

Eutheia Stephens, 1830plicata (Gyllenhal, 1813) --SNDEAIschaumii Kiesenwetter, 1858 --S-D---linearis Mulsant, 1861 FKSND-A-scydmaenoides Stephens, 1830 FKSNDEAI

Veraphis Casey, 1897engelmarki Franz, 1971 --S-----

Cephenniini Reitter, 1882Cephennium Müller & Kunze, 1822thoracicum Müller & Kunze, 1822 ----D-A-laticolle Aubé, 1842 ------A-majus Reitter, 1882 -----EAIgallicum Ganglbauer, 1899 --S-D---


Cyrtoscydmini Schaufuss, 1889Stenichnini Ganglbauer, 1899

Nevraphes Thomson, 1859elongatulus (Müller & Kunze, 1822) --SNDEAIangulatus (Müller & Kunze, 1822) FKSNDEAIcarinatus (Mulsant, 1861) -----E--perssoni Franz, 1975 --S-----ruthenus Machulka, 1926 --SND---talparum Lokay, 1920 FKS----Iplicicollis Reitter, 1879 F-SND---coronatus J.Sahlberg, 1881 FKSN-E--praeteritus Rye, 1872 -----E--

Scydmoraphes Reitter, 1891sparshalli (Denny, 1825) F-S-D-A-helvolus (Schaum, 1844) F-SND-AIminutus (Chaudoir, 1845) FKSNDE-I

Stenichnus Thomson, 1859scutellaris (Müller & Kunze, 1822) F-SNDEAIpusillus (Müller & Kunze, 1822) --S-----godarti (Latreille, 1806) --SND-AIfoveola Rey, 1888 ------A-collaris (Müller & Kunze, 1822) FKSNDEAIbicolor (Denny, 1825) FKSNDEAI

exilis (Erichson, 1837)poweri (Fowler, 1884) --SND---

harwoodianus Williams, 1927

Microscydmus Saulcy & Croissandeau, 1893nanus (Schaum, 1844) FKSNDE-Iminimus (Chaudoir, 1845) F-SNDEA-

Euconnus Thomson, 1859pubicollis (Müller & Kunze, 1822) -------Iclaviger (Müller & Kunze, 1822) FKSNDEAIpragensis (Machulka, 1923) F-SN----maklinii (Mannerheim, 1844) FKSNDEAIwetterhallii (Gyllenhal, 1813) F-SNDE-Identicornis (Müller & Kunze, 1822) --S-DEAIrutilipennis (Müller & Kunze, 1822) F-S-D--Ihirticollis (Illiger, 1798) FKSNDEA-fimetarius (Chaudoir, 1845) FKSNDEAI

Scydmaenini Leach, 1815Scydmaenus Latreille, 1802tarsatus Müller & Kunze, 1822 FKSNDEAIrufus Müller & Kunze, 1822 F-SNDEAIperrisii (Reitter, 1882) --S-----hellwigii (Herbst, 1792) FKSN-EA-

Silphidae Latreille, 1807Silphinae Latreille, 1807Necrodes Leach, 1815littoralis (Linnaeus, 1758) FKSNDEAI

Thanatophilus Leach, 1815lapponicus (Herbst, 1793) FKSN-E--rugosus (Linnaeus, 1758) FKSNDEAIsinuatus (Fabricius, 1775) FKSNDEAItrituberculatus (Kirby, 1837) -K------

baicalicus Motschulsky, 1860dispar (Herbst, 1793) FKSNDEAI

Oiceoptoma Leach, 1815thoracica (Linnaeus, 1758) FKSNDEAI

Aclypea Reitter, 1884Blitophaga Reitter, 1884

opaca (Linnaeus, 1758) FKSNDEAIundata (Müller, 1776) FKS-DEAI

Dendroxena Motschulsky, 1858Xylodrepa Thomson, 1859

quadrimaculata (Scopoli, 1772) F-SNDEAIquadripunctata (Linnaeus, 1761) nec (Linnaeus, 1758)

Silpha Linnaeus, 1758carinata Herbst, 1783 FKSNDEAIobscura Linnaeus, 1758 FKSNDEAItristis Illiger, 1798 FKSNDEAI

Ablattaria Reitter, 1884laevigata (Fabricius, 1775) -----E--

Phosphuga Leach, 1817atrata (Linnaeus, 1758) FKSNDEAI

Nicrophorinae Kirby, 1837Nicrophorus Fabricius, 1775germanicus (Linnaeus, 1758) --S-D-A-humator (Gleditsch, 1767) F-SNDEAIsepultor Charpentier, 1825 --S-DEAIinvestigator Zetterstedt, 1824 FKSNDEAIfossor Erichson, 1837 --S-DEAI

interruptus Stephens, 1830, nec Brullé, 1822vespilloides Herbst, 1784 FKSNDEAIvespillo (Linnaeus, 1758) FKSNDEAIvestigator Herschel, 1807 --S-DEAI

Staphylinidae Latreille, 1802Omaliinae MacLeay, 1825Eusphalerini Hatch, 1957Eusphalerum Kraatz, 1857Anthobium auct., nec Leach, 1819

longipenne (Erichson, 1839) -K----AI


primulae (Stephens, 1834) ----D-AIminutum (Fabricius, 1792) FKSNDEAIsorbicola (Y.Kangas, 1941) FKSNDE--lapponicum (Mannerheim, 1830) FKSN-EAIluteum (Marsham, 1802) FKSNDEAI

ophthalmicum (Paykull, 1800) nec (Scopoli, 1763)limbatum (Erichson, 1840) -----E--pseudaucupariae (E.Strand, 1916) -K------

aucupariae (Kiesenwetter, 1866) nec (Gistel, 1857)sorbi (Gyllenhal, 1810) FKSNDE--torquatum (Marsham, 1802) -KSND---tenenbaumi (Bernhauer, 1932) ----DE-I

florale auct. nec (Panzer, 1793)

Omaliini MacLeay, 1825Acrulia Thomson, 1858inflata (Gyllenhal, 1813) FKSNDEA-

Pycnoglypta Thomson, 1858lurida (Gyllenhal, 1813) FKSNDEAI

Acrolocha Thomson, 1858minuta (Olivier, 1795) FKS-DE--

striata (Gravenhorst, 1802)pliginskii Bernhauer, 1912 FKS-D---sulcula (Stephens, 1834) FKS-DE-I

Phyllodrepa Thomson, 1859melanocephala (Fabricius, 1787) FKSNDEA-salicis (Gyllenhal, 1810) FKSND---nigra (Gravenhorst, 1806) FKSNDEAIpuberula Bernhauer, 1903 F-SND--Ifloralis (Paykull, 1789) FKSNDEAImelis Hansen, 1940 F-S-D--Isahlbergi Luze, 1906 FKSN----

baicalensis auct. nec Bernhauer, 1903linearis (Zetterstedt, 1828) FKSN-E--clavigera Luze, 1906 FKSN-E--ioptera (Stephens, 1832) FKSNDEAIvilis (Erichson, 1840) --SNDE--gracilicornis (Fairmaire & Laboulbène, 1856) --S-D---

Hapalaraea Thomson, 1858pygmaea (Paykull, 1800) FKSNDEA-

Hypopycna Mulsant & Rey, 1880rufula (Erichson, 1840) ----D---

Omalium Gravenhorst, 1802rugulipenne Rye, 1864 ----D---laeviusculum Gyllenhal, 1827 -KSND---riparium Thomson, 1857 F-SNDE--validum Kraatz, 1858 --S--E--

rivulare (Paykull, 1789) FKSNDEAIseptentrionis Thomson, 1857 FKSNDE-Iallardii Fairmaire & Brisout de Barneville, 1859 --SND---oxyacanthae Gravenhorst, 1806 --S-DE-Istrigicolle Wankowicz, 1869 FKSN----

brevicolle Thomson, 1869, nec Heer, 1839laticolle Kraatz, 1858 FKSND---exiguum Gyllenhal, 1810 F-SNDE--imitator Luze, 1906 F-------littorale Kraatz, 1858 F-SNDE-Icaesum Gravenhorst, 1806 FKSNDEAIrugatum Rey, 1880 F-SNDE--muensteri Bernhauer, 1900 F-SN----excavatum Stephens, 1834 FKSNDEAI

Xylostiba Ganglbauer, 1895monilicornis (Gyllenhal, 1810) FKSN-EAI

Phloeostiba Thomson, 1858plana (Paykull, 1792) FKSNDEAIlapponica (Zetterstedt, 1838) FKSNDEAI

Phloeonomus Heer, 1839pusillus (Gravenhorst, 1806) FKSNDEAIsjobergi Strand, 1937 FKSN-E--punctipennis Thomson, 1867 FKSNDEAI

Xylodromus Heer, 1839depressus (Gravenhorst, 1802) FKSNDEA-affinis (Gerhardt, 1877) --S-D---brunnipennis (Stephens, 1832) FKSNDEAI

concinnus (Marsham, 1802) nec (Gravenhorst, 1802)testaceus (Erichson, 1840) -KS-D---

Micralymma Westwood, 1838marinum (Ström, 1783) -KSN----

Anthophagini Thomson, 1859Philorinum Kraatz, 1857sordidum (Stephens, 1834) ----D---

Cylletron Thomson, 1859nivale Thomson, 1859 FKSN----

Porrhodites Kraatz, 1857fenestralis (Zetterstedt, 1828) FKSN----

Orochares Kraatz, 1857angustatus (Erichson, 1840) --S-D-A-

Mannerheimia Mäklin, 1880arctica (Erichson, 1840) FKSN----brevipennis (Motschulsky, 1858) FKS-----

divergens (Mäklin, 1877)


Phyllodrepoidea Ganglbauer, 1895crenata Ganglbauer, 1895 --SND---

Deliphrum Erichson, 1839tectum (Paykull, 1789) FKSNDEAI

Anthobium Leach, 1819Lathrimaeum Erichson, 1839

melanocephalum (Illiger, 1794) --SN-EA-atrocephalum (Gyllenhal, 1827) FKSNDEAIunicolor (Marsham, 1802) --SND---fusculum (Erichson, 1839) F-SND---

Olophrum Erichson, 1839piceum (Gyllenhal, 1810) F-SNDEAIfuscum (Gravenhorst, 1806) FKSNDEAIassimile (Paykull, 1800) FKSNDEAIboreale (Paykull, 1792) FKSN----

florae Scheerpeltz, 1935consimile (Gyllenhal, 1810) FKSN-E-Irotundicolle (Sahlberg, 1830) FKSN---I

Arpedium Erichson, 1839quadrum (Gravenhorst, 1806) FKSNDEAI

Eucnecosum Reitter, 1909puncticolle (J.Sahlberg, 1880) FK------brachypterum (Gravenhorst, 1802) FKSNDEAItenue (LeConte, 1863) FKSN----

norvegicum (Munster, 1933)brunnescens (J.Sahlberg, 1871) FKSN-E--

Deliphrosoma Reitter, 1909macrocephalum (Eppelsheim, 1873) ------A-

pechlaneri Lohse, 1963prolongatum (Rottenberg, 1873) -----E--

Acidota Stephens, 1829crenata (Fabricius, 1793) FKSNDEAIcruentata Mannerheim, 1830 FKSND-AIquadrata (Zetterstedt, 1838) FKSN----

Lesteva Latreille, 1797punctata Erichson, 1839 F-SNDE--sicula Erichson, 1840ssp. heeri Fauvel, 1872 --SND---longoelytrata (Goeze, 1777) FKSNDEAImonticola Kiesenwetter, 1847 FKSN----hanseni Lohse, 1953 --S-D---

fontinalis auct. nec Kiesenwetter, 1850pubescens Mannerheim, 1830 FKSND-A-

Geodromicus Redtenbacher, 1857Psephidonus Gistel, 1856

plagiatus (Fabricius, 1798) FKSNDE--

nigrita (Müller, 1821) FKSN----longipes (Mannerheim, 1830) FKSN----hoejeri Palm, 1961 --S-----

Anthophagus Gravenhorst, 1802alpinus (Paykull, 1790) FKSN----angusticollis (Mannerheim, 1830) FK---EAI

abbreviatus auct. nec (Fabricius, 1779)omalinus Zetterstedt, 1828 FKSN-EA-caraboides (Linnaeus, 1758) FKSNDEAI

Coryphiini Jacobson, 1908Coryphium Stephens, 1834angusticolle Stephens, 1834 FKSNDEAI

Coryphiomorphus Zerche, 1988hyperboreus (Mäklin, 1880) F-SN----

norvegicus (Munster, 1911)

Eudectus Redtenbacher, 1856giraudi Redtenbacher, 1856 FKSN-E--

Boreaphilus Sahlberg, 1832henningianus Sahlberg, 1832 FKSN----

Proteininae Erichson, 1839Metopsia Wollaston, 1854Phloeobium auct., nec Dejean, 1829

similis Zerche, 1998 --SND---clypeata auct. nec (Müller, 1821)

clypeata (Müller, 1821) --S-----retusa (Stephens, 1835)gallica (Koch, 1938)

Megarthrus Curtis, 1829prosseni Schatzmayr, 1904 FKSNDEAI

depressus auct., nec (Paykull, 1789)strandi Scheerpeltz, 1931 FKSN-E--nigrinus J.Sahlberg, 1876 FKSN----depressus (Paykull, 1789) FKSNDEAI

sinuatocollis auct., nec (Lacordaire,1835)denticollis (Beck, 1817) FKSNDEAInitidulus Kraatz, 1857 FKSN-EA-hemipterus (Illiger, 1794) -----EAIfennicus Lahtinen, 1938 FKSN----

Proteinus Latreille, 1797Pteronius Blackwelder, 1952

brachypterus (Fabricius, 1792) FKSNDEAIcrenulatus Pandelle, 1867 --SN----altaicus Reitter, 1905 FKSN----

apicidens Jansson & Sjöberg, 1929hyperboreus Muona, 1978 FKSN----

altaicus auct. nec Reitter, 1905


laevigatus Hochhuth, 1872 FKSNDE-Imacropterus (Gyllenhal, 1810) nec (Gravenhorst, 1806)serrifer (Muona, 1977)

atomarius Erichson, 1840 FKSNDEAI

Micropeplinae Leach, 1815Arrhenopeplus Koch, 1937tesserula (Curtis, 1828) FKSND--I

Micropeplus Latreille, 1809staphylinoides (Marsham, 1802) -----EA-latus Hampe, 1861 --S-----fulvus Erichson, 1840 F-SNDE--porcatus (Paykull, 1789) FKSNDEA-caelatus Erichson, 1839 --S-D-AI

Pselaphinae Latreille, 1802Euplectini LeConte, 1861Bibloporus Thomson, 1859ultimus Guillebeau, 1892 --S-----bicolor (Denny, 1825) FKSNDEAIminutus Raffray, 1914 FKSND-A-

hoglundi Palm, 1948

Pseudoplectus Reitter, 1881perplexus (Jacquelin du Val, 1854) -KS-D---

Bibloplectus Reitter, 1881tenebrosus (Reitter, 1880) F-SND---spinosus Raffray, 1914 F-SNDEA-ambiguus (Reichenbach, 1816) FKSNDEAIminutissimus (Aubé, 1833) F-SND-A-pusillus (Denny, 1825) ----DEA-

Meliceria Raffray, 1898tragardhi Palm, 1938 --S--A--

Euplectus Leach, 1817nanus (Reichenbach, 1816) FKSNDEAIkirbii Denny, 1825ssp. kirbii Denny, 1825 --SND---ssp. revelierei Reitter, 1884 F----EA-piceus Motschulsky, 1835 F-SNDEAIdecipiens Raffray, 1910 FKSN-E--infirmus Raffray, 1910 --S-D---bescidicus Reitter, 1881 F-SNDE--

bohemicus Machulka, 1930sanguineus Denny, 1825 FKSNDEAIsignatus (Reichenbach, 1816) FKSNDEAIbonvouloiri Reitter, 1881ssp. rosae Raffray, 1910 --S-D---punctatus Mulsant, 1861 FKSND-A-tholini Guillebeau, 1888 --S-D---

karstenii (Reichenbach, 1816) FKSNDEAImutator Fauvel, 1895 FKSNDE--

fauveli Guillebeau, 1888, nec Raffray, 1882falsus Bedel, 1906tomlini Joy, 1906

brunneus (Grimmer, 1841) --SND---duponti Aubé, 1833 --S-D-A-

Leptoplectus Casey, 1908Grammoplectus Jeannel, 1950

spinolai (Aubé, 1844) --SN----

Plectophloeus Reitter, 1891nitidus (Fairmaire, 1857) F-S-DE--nubigena (Reitter, 1876) --S-D---fischeri (Aubé, 1833) -------I

Saulcyella Reitter, 1901schmidtii (Märkel, 1844) --S-D---

Trimium Aubé, 1833brevicorne (Reichenbach, 1816) FKSNDEAI

Batrisini Reitter, 1882Batrisus Aubé, 1833formicarius Aubé, 1833 -------I

Batrisodes Reitter, 1882delaporti (Aubé, 1833) --S-----venustus (Reichenbach, 1816) FKSNDEAIhubenthali Reitter, 1913 FKS--E--adnexus (Hampe, 1863) FKS-DE--

Goniacerini Reitter, 1882 (1872)Bythinus Leach, 1817macropalpus Aubé, 1833 F-S-DE-I

distinctus Chaudoir, 1845burrellii Denny, 1825 --SNDE--securiger (Reichenbach, 1816) -------I

Bryaxis Kugelann, 1794puncticollis (Denny, 1825) FKSNDEAI

validus (Aubé, 1844)nigripennis (Aubé, 1844) ------A-bulbifer (Reichenbach, 1816) FKSNDEAIclavicornis (Panzer, 1806) --S-D---curtisii (Leach, 1817) --SND-A-

Trichonyx Chaudoir, 1845sulcicollis (Reichenbach, 1816) F-SND--I

Amauronyx Reitter, 1881maerkelii (Aubé, 1844) --S-D---

Tychus Leach, 1817niger (Paykull, 1800) FKSNDEAI


monilicornis Reitter, 1880 --S-D---normandi Jeannel, 1950 --S-D---

Rybaxis Saulcy, 1876Bryaxis Leach, 1817, nec Kugelann, 1794

longicornis (Leach, 1817) FKSNDEAIlaminata (Motschulsky, 1836) ---ND---

Brachygluta Thomson, 1859fossulata (Reichenbach, 1816) FKSNDEAIhaematica (Reichenbach, 1816)ssp. sinuata (Aubé, 1833) F-SNDEA-helferi (Schmidt-Goebel, 1836) F-SND---paludosa (Peyron, 1858) ----d---

hanseni Besuchet, 1954

Reichenbachia Leach, 1826juncorum (Leach, 1817) --SNDEAI

Trissemus Jeannel, 1949impressus (Panzer, 1803) F-SNDEAI

Pselaphini Latreille, 1802Pselaphaulax Reitter, 1909dresdensis (Herbst, 1792) FKSNDEAI

Pselaphus Herbst, 1792heisei Herbst, 1792 FKSNDEAI

Chennium Latreille, 1807bituberculatum Latreille, 1807 --S-----

Centrotoma Heyden, 1849lucifuga Heyden, 1849 -------I

Tyrus Aubé, 1833mucronatus (Panzer, 1803) FKSNDEAI

Clavigerini Leach, 1815Claviger Preyssler, 1790testaceus Preyssler, 1790 FKSNDEAIlongicornis Müller, 1818 --S-D---

Phloeocharinae Erichson, 1839Phloeocharis Mannerheim, 1830subtilissima Mannerheim, 1830 FKSNDEAI

Olisthaerinae Thomson, 1859Olisthaerus Dejean, 1833megacephalus (Zetterstedt, 1828) FKSN-E-Isubstriatus (Paykull, 1790) FKSN---I

tenuis Mulsant & Rey, 1853 FKSN-E-Imulsanti Ganglbauer, 1895

altaicus Luze, 1901 FKSN----monticola Fowler, 1888 FKSN----piceolus Rey, 1882 -----E--erichsonanus Fagel, 1965 FKSNDE-I

baudueri auct. nec Mulsant & Rey, 1875baudueri Mulsant & Rey, 1875 --SNDE-I

hellieseni Strand, 1950lepidus (Gravenhorst, 1806) FKSNDEAI

brunneus (Marsham, 1802) nec (Fabricius, 1798)longulus Mannerheim, 1830 FKSNDEAIbimaculatus Lacordaire, 1835 FKSNDEA-

ruficornis Kraatz, 1857despectus Strand, 1969 FKSN----

bimaculatus auct. nec Lacordaire, 1835nigricollis Stephens, 1835 ????????

splendens (Marsham, 1802) nec (Fabricius, 1793)glaber (Sperk, 1835) F-S-----dispersus Schülke & Kocian, 2000 --S-----inaris Luze, 1901 FKSN-E--maeklini Bernhauer, 1906 FKSN-E--forticornis Fauvel, 1875 F-SNDE--clavicornis (Stephens, 1832) FKSND-AIsolidicornis Wollaston, 1864 -----E--

reichei Pandellé, 1869aequalis Thomson, 1868 FKSNDE--niger Fairmaire & Laboulbène, 1856 FKSND---angularis Mulsant & Rey, 1854 FK------boreellus J.Sahlberg, 1876 FKSN----

nigrans auct. nec Mäklin, 1853maerkeli Kraatz, 1857 FKSN----rufescens (Stephens, 1832) FKSNDEAIeppelsheimianus Fagel, 1968 FKSND---

brucki auct. nec Pandellé, 1869bruckii (Pandellé, 1869) FKSN----

laevicollis Eppelsheim, 1878punctus (Gravenhorst, 1806) FKSNDEAI

Ischnosoma Stephens, 1829bergrothi (Hellén, 1925) FKSN-EAIlongicorne (Mäklin, 1847) FKSNDEA-splendidum (Gravenhorst, 1806) FKSNDEAI

Bryoporus Kraatz, 1857cernuus (Gravenhorst, 1806) FKSNDE-I

merdarius (Olivier, 1794) nec (Fabricius, 1775)

Tachyporinae MacLeay, 1825Mycetoporini Thomson, 1859

Bolitobiini Horn, 1877Mycetoporus Mannerheim, 1830debilis Mäklin, 1847 FK------

Bryophacis Reitter, 1909punctipennis (Thomson, 1861) FKSN----


rugipennis (Pandellé, 1869) FKSN-E--rufus (Erichson, 1839) -------Icrassicornis (Mäklin, 1847) FKSNDE--

Carphacis Des Gozis, 1886striatus (Olivier, 1795) FKS-D---

angularis (Paykull, 1800)

Lordithon Thomson, 1859Bolitobius auct., nec Leach, 1819

thoracicus (Fabricius, 1777) FKSNDEAIexoletus (Erichson, 1839) FKSNDEAI

bernhaueri (Wanka, 1929)trinotatus (Erichson, 1839) F-SNDEAItrimaculatus (Fabricius, 1793) FKSN--AIlunulatus (Linnaeus, 1761) FKSNDEAIpulchellus (Mannerheim, 1830) FKSN-EAIspeciosus (Erichson, 1840) FKSN-EA-

Bolitobius Leach, 1819Bryocharis Lacordaire, 1835

cingulatus Mannerheim, 1830 FKSNDEAIcastaneus (Stephens, 1832) FKSNDEAI

analis auct. nec (Fabricius, 1787)inclinans (Gravenhorst, 1806) --SND---formosus (Gravenhorst, 1806) FK---EAI

Tachyporini MacLeay, 1825Sepedophilus Gistel, 1856Conosoma auct., nec Kraatz, 1857

littoreus (Linnaeus, 1758) FKSNDEAItestaceus (Fabricius, 1793) FKSNDEAImarshami (Stephens, 1832) F-SNDE-Iconstans (Fowler, 1888) FKSN--A-

strigosus (J.Sahlberg, 1911)stoeckli (Lokay, 1913)

lusitanicus Hammond, 1973 --S-----immaculatus (Stephens, 1832) FKSNDEAIbipunctatus (Gravenhorst, 1802) FKSNDEAIbipustulatus (Gravenhorst, 1802) --S-DEAIpedicularius (Gravenhorst, 1802) FKSNDE-Inigripennis (Stephens, 1832) ----D---

hypnorum (Fabricius, 1775) FKSNDEAIchrysomelinus (Linnaeus, 1758) FKSNDEAIdispar (Paykull, 1789) FKSND---tersus Erichson, 1840 F-SNDEA-atriceps Stephens, 1832 FKSNDEAIduplex Luze, 1901 ---N----quadriscopulatus Pandellé, 1869 --SN-EAI

signifer Pandellé, 1872ruficollis Gravenhorst, 1802 ----DEA-transversalis Gravenhorst, 1806 FKSNDEAIpulchellus Mannerheim, 1843 FKSNDE-Ipusillus Gravenhorst, 1806 FKSNDEAI

macropterus Stephens, 1832scitulus Erichson, 1839 FKSNDEAIcorpulentus J.Sahlberg, 1876 FKSN-EAI

Lamprinodes Luze, 1901saginatus (Gravenhorst, 1806) FKSNDEAI

Lamprinus Heer, 1839erythropterus (Panzer, 1796) -----EA-

Tachinus Gravenhorst, 1802signatus Gravenhorst, 1802 FKSNDEAI

rufipes auct. nec (Linnaeus, 1758)elegans Eppelsheim, 1893 FKSN----pallipes (Gravenhorst, 1806) FKSNDEAImarginatus (Fabricius, 1793) FKSN-EAIscapularis Stephens, 1832 F-------bipustulatus (Fabricius, 1793) --S-DEAIhumeralis Gravenhorst, 1802 ----DEAIproximus Kraatz, 1855 FKSNDEAIrufipennis Gyllenhal, 1810 FKSND-AIatripes J.Sahlberg, 1876 F-SN----subterraneus (Linnaeus, 1758) FKSND-AIfimetarius Gravenhorst, 1802 FKSNDEAIlignorum (Linnaeus, 1758) FKSNDEAIbasalis Erichson, 1840 FKS----Icorticinus Gravenhorst, 1802 FKSNDEAIlaticollis Gravenhorst, 1802 FKSNDEAIschneideri Luze, 1900 F-----A-marginellus (Fabricius, 1781) FKSNDEAIelongatus Gyllenhal, 1810 FKSND-AIpunctipennis (J.Sahlberg, 1876) FK---E--

Tachyporus Gravenhorst, 1802nitidulus (Fabricius, 1781) FKSNDEAIobscurellus Zetterstedt, 1838 FKSN----obtusus (Linnaeus, 1767) FKSNDEAIabdominalis (Fabricius, 1781) FKSN-EAIformosus Matthews, 1838 --S-D---solutus Erichson, 1839 FKSNDEAIpallidus Sharp, 1871 FKSNDEAI

scutellaris Rye, 1871, nec Lacordaire, 1835

Coproporus Kraatz, 1857Erchomus Motschulsky, 1858

colchicus Kraatz, 1858 --SN--Aheterocerus auct. nec (Fauvel, 1898)


Cilea Jacquelin du Val, 1856Leucoparyphus Kraatz, 1857

silphoides (Linnaeus, 1767) FKSNDEAI

Habrocerinae Mulsant & Rey, 1877Habrocerus Erichson, 1839capillaricornis (Gravenhorst, 1806) F-SNDEAI

Trichophyinae Thomson, 1859Trichophya Mannerheim, 1830pilicornis (Gyllenhal, 1810) FKSNDEAI

Aleocharinae Fleming, 1821Aleocharini Fleming, 1821Aleochara Gravenhorst, 1802curtula (Goeze, 1777) FKSNDEAIhaematoptera Kraatz, 1858

haematodes Schaum, 1862ssp. ripicola Mulsant & Rey, 1874 F----E--

crassicornis Lacordaire, 1835, nec Gyllenhal, 1827spissicornis Erichson, 1839 --S-----brevipennis Gravenhorst, 1806 FKSNDEAIpuberula Klug, 1833 F-------intricata Mannerheim, 1830 FKSNDEAItristis Gravenhorst, 1806 FKSNDEAImoesta Gravenhorst, 1802 FKSNDE-Igaudiuscula Tottenham, 1939 --s-----

pulchra Bernhauer, 1901, nec Gravenhorst, 1806inconspicua Aubé, 1850 FKSNDEA-sparsa Heer, 1839 FKSNDEAIstichai Likovský, 1965 F-SN---Ikamila Likovský, 1984 F-SN-E-I

diversa (J.Sahlberg, 1876) nec Mulsant & Rey, 1853funebris Wollaston, 1864 FKSNDE-I

albovillosa Bernhauer, 1901peeziana Lohse, 1961 F-SN----lanuginosa Gravenhorst, 1802 FKSNDEAIlygaea Kraatz, 1862 FKSNDE--villosa Mannerheim, 1830 FKSNDE--sanguinea (Linnaeus, 1758) FKSND-A-fumata Gravenhorst, 1802 FKSN-EAImoerens Gyllenhal, 1827 FKSNDEAImaculata Brisout de Barneville, 1863 F-------laevigata Gyllenhal, 1810 FKSNDEAIhaemoptera Kraatz, 1856 FK------cuniculorum Kraatz, 1858 --S-DE--spadicea (Erichson, 1837) FKSNDE--ruficornis Gravenhorst, 1802 F---D---erythroptera Gravenhorst, 1806 -------Ibilineata Gyllenhal, 1810 FKSNDEAI

binotata Kraatz, 1856 FKSNDEAIverna auct. nec Say, 1836

verna Say, 1836 --S-D--Ibrundini Bernhauer, 1936 FKSN----

suffusa auct. nec (Casey, 1906)bipustulata (Linnaeus, 1761) FKSNDEAIgrisea Kraatz, 1856 F-SNDE-Iobscurella Gravenhorst, 1806 --SND---

algarum Fauvel, 1862punctatella Motschulsky, 1858 F-SNDE--

obscurella auct. nec Gravenhorst, 1806

Tinotus Sharp, 1883morion (Gravenhorst, 1802) FKSNDEAI

Oxypodini Thomson, 1859Oxypodina Thomson, 1859Oxypoda Mannerheim, 1830lugubris Kraatz, 1856 FKSN----elongatula Aubé, 1850 FKSNDEAIprocerula Mannerheim, 1830 FKSNDEAIopaca (Gravenhorst, 1802) FKSNDEAInigricornis Motschulsky, 1860 FKSN----operta Sjöberg, 1950 FKS-----longipes Mulsant & Rey, 1861 FKSNDE-Ivittata Märkel, 1842 FKSND-AIacuminata (Stephens, 1832) FKSNDEAI

lividipennis auct. nec Mannerheim, 1830spectabilis Märkel, 1844 FKSNDEAIfunebris Kraatz, 1856 FKSNDEA-

laticollis (Thomson, 1871)lentula Erichson, 1837 FKS-D--Iinduta Mulsant & Rey, 1861 ----D---skalitzkyi Bernhauer, 1902 FKSN---Ibrevicornis (Stephens, 1832) FKSNDE-I

umbrata (Gyllenhal, 1810) nec (Gravenhorst, 1802)doderoi Bernhauer, 1902 FKS-----hansseni Strand, 1946 F-SN----

lapponica Lundberg, 1980vicina Kraatz, 1856 FKSNDEA-abdominalis (Mannerheim, 1830) FKSN-EAIstrandi Scheerpeltz, 1957 F-SND---rufa Kraatz, 1856 --S-DE--islandica Kraatz, 1857 FKSN-E--uhligi Zerche, 1987 ---N----togata Erichson, 1837 FKSNDE-Iexoleta Erichson, 1839 F-SNDE-Irecondita Kraatz, 1856 F-SNDE-Ialternans (Gravenhorst, 1802) FKSNDEAI


serpentata Kangas, 1983 F-------praecox Erichson, 1839 FKSNDE-Iarborea Zerche, 1994 F-SND---

lucens auct. nec Mulsant & Rey, 1853testacea Erichson, 1837 F-SNDE-Itarda Sharp, 1871 --S-D---brachyptera (Stephens, 1832) FKSNDE--annularis (Mannerheim, 1830) FKSNDE-Iflavicornis Kraatz, 1856 FKSND-A-

amoena Fairmaire & Laboulbène, 1856advena Mäklin, 1846 FKSN-E-Ifiliformis Redtenbacher, 1849 -------Iferruginea Erichson, 1839 -----E-Isoror Thomson, 1855 FKSNDEA-formiceticola Märkel, 1841 FKSNDEAIpratensicola Lohse, 1970 --S-----

scanica Lundberg, 1980rugicollis Kraatz, 1856 F-SN-E--

sjobergi Bernhauer, 1927bicolor Mulsant & Rey, 1853 F-SN-E--haemorrhoa (Mannerheim, 1830) FKSNDEAIcarbonaria (Heer, 1841) ----D---

Devia Blackwelder, 1952Dasyglossa Kraatz, 1856, nec Illiger, 1807

prospera (Erichson, 1839) FKSN-EAI

Hygropora Kraatz, 1856cunctans (Erichson, 1837) FKSNDE--longicornis Palm, 1949 FKSN----

Pyroglossa Bernhauer, 1901pulcherrima Bernhauer, 1901 F-SN----

Acrostiba Thomson, 1858borealis Thomson, 1858 FKSN----

Ocyusa Kraatz, 1856Deubelia Bernhauer, 1899

maura (Erichson, 1837) FKSNDEA-picina (Aubé, 1850) FKS-DE-I

Chanoma Blackwelder, 1952Pseudaphana Bernhauer, 1907, nec Westwood, 1842

vorbringeri (Bernhauer, 1907) --SN----tullgreni (Palm, 1939)

Cephalocousya Lohse, 1971nivicola (Thomson, 1871) FKSN----

Chilomorpha Krása, 1914longitarsis (Thomson, 1867) F-SN-E--

laticollis auct. nec (Thomson, 1871)hibernica (Rye, 1876)

Calodera Mannerheim, 1830Ityocara Thomson, 1867

nigrita Mannerheim, 1830 F-SNDEA-protensa Mannerheim, 1830 FKS-DE--rufescens Kraatz, 1856 F-S-DE--aethiops (Gravenhorst, 1802) FKSND-AIlapponica J.Sahlberg, 1871 F-SN----uliginosa Erichson, 1837 FKS-DE-Iriparia Erichson, 1837 FKSNDE-Irubens Erichson, 1837 F-S-DEA-

Parocyusa Bernhauer, 1902Chilopora Kraatz, 1867, nec Haime, 1854

rubicunda (Erichson, 1837) FKSNDEA-crebrepunctata (Strand, 1962) F-SN----longitarsis (Erichson, 1839) ----DE--

Stichoglossa Fairmaire & Laboulbène, 1856semirufa (Erichson, 1839) ----D---

Ischnoglossa Kraatz, 1856prolixa (Gravenhorst, 1802) FKSNDEA-elegantula (Mannerheim, 1830) FKSN----obscura Wunderle, 1990 F-S-D---

Dexiogyia Thomson, 1858corticina (Erichson, 1837) FKS-D-AIforticornis (Strand, 1939) ---N----

Thiasophila Kraatz, 1856Thyasophila Fairmaire & Laboulbène, 1856

angulata (Erichson, 1837) FKSNDEAIlohsei Zerche, 1987 --S-----canaliculata Mulsant & Rey, 1874 FKSNDE--wockii (Schneider, 1862) FKSND---

rotundicollis Jansson, 1943bercionis Bernhauer, 1926 FKS-----inquilina (Märkel, 1842) FKSNDE--

Crataraea Thomson, 1858suturalis (Mannerheim, 1830) FKSNDEAI

Haploglossa Kraatz, 1856Microglotta Kraatz, 1862

gentilis (Märkel, 1844) --SNDEAIvillosula (Stephens, 1832) FKSNDEAI

pulla (Gyllenhal, 1827) nec (Gravenhorst, 1802)nidicola (Fairmaire, 1852) FKSNDE-Ipicipennis (Gyllenhal, 1827) FKSND-AImarginalis (Gravenhorst, 1806) FKSNDE-I

Mniusa Mulsant & Rey, 1875incrassata (Mulsant & Rey, 1852) FKSNDE-Igrandiceps (J.Sahlberg, 1876) FKSN----


Poromniusa Ganglbauer, 1895Eurymniusa Ganglbauer, 1895

crassa (Eppelsheim, 1883) ----D-A-procidua (Erichson, 1837) FKSN---I

Ocalea Erichson, 1837badia Erichson, 1837 FKSNDEAIpicata (Stephens, 1832) FKSNDEAIconcolor Kiesenwetter, 1847 ----DE--rivularis Miller, 1851 -KS-DE-Ilatipennis Sharp, 1870 F-SND---

Ilyobates Kraatz, 1856bennetti Donisthorpe, 1914 FKSNDEAI

subopacus Palm, 1953nigricollis (Paykull, 1800) FKSNDEAI

Pentanota Bernhauer, 1905meuseli Bernhauer, 1905 F-SN----

Amarochara Thomson, 1858umbrosa (Erichson, 1837) F-SND---bonnairii (Fauvel, 1865) --S-D---forticornis (Lacordaire, 1835) F-S-DEA-

Phloeopora Erichson, 1837Phloeodroma Kraatz, 1856

testacea (Mannerheim, 1830) FKSNDE-Initidiventris Fauvel, 1900 FKSN-E--corticalis (Gravenhorst, 1802) FKSNDEAI

teres auct. nec (Gravenhorst, 1802)angustiformis auct. nec Baudi, 1869

bernhaueri Lohse, 1984 ----D---teres (Gravenhorst, 1802) ----D---

teres auct. nec (Gravenhorst, 1802)opaca Bernhauer, 1902 F-------concolor (Kraatz, 1856) FKSNDEAI

Dinardina Mulsant & Rey, 1873Dinarda Samouelle, 1819dentata (Gravenhorst, 1806) FKSND-A-hagensii Wasmann, 1889 FKSND---maerkelii Kiesenwetter, 1843 FKSNDEA-

filiformis (Motschulsky, 1860) FKSNDE--capitalis Mulsant & Rey, 1873apicalis Benick, 1953clavata Benick, 1953

exillima Sharp, 1915 ----D---exilis (Knoch, 1806) FKSNDE--

exiliformis Joy, 1915winkleri Benick, 1954 ----D---

Tachyusina Thomson, 1859Dasygnypeta Lohse, 1974velata (Erichson, 1837) FKSNDEAI

Gnypeta Thomson, 1858caerulea (Sahlberg, 1831) FKSN-E--sellmani Brundin, 1929 FKSN----brincki Palm, 1966 F-SN----ripicola (Kiesenwetter, 1844) FKSND---carbonaria (Mannerheim, 1830) FKSNDE--rubrior Tottenham, 1939 F-S-DE--

Ischnopoda Stephens, 1835Tachyusa Erichson, 1837

constricta (Erichson, 1837) FKSNDEAIcoarctata (Erichson, 1837) FKS-DEAIbalteata (Erichson, 1839) -K------atra (Gravenhorst, 1806) FKSNDEAIleucopus (Marsham, 1802) FKSNDEAIumbratica (Erichson, 1837) FKS-DE-Iscitula (Erichson, 1837) FKSNDEAI

Brachyusa Mulsant & Rey, 1874concolor (Erichson, 1839) FKSND-AI

Meoticina Seevers, 1978Meotica Mulsant & Rey, 1873pallens (Redtenbacher, 1849) FKSND---

lohsei Benick, 1953hanseni Scheerpeltz, 1954strandi Scheerpeltz, 1958

stockmanni Muona, 1978 F--N----finnmarchica Benick, 1953 F-SN----

angulata Benick, 1953

Athetini Casey, 1910Trichomicra Brundin, 1945sahlbergiana (Bernhauer, 1910) F-SN----

Dochmonota Thomson, 1859clancula (Erichson, 1837) FKSNDE--rudiventris (Eppelsheim, 1886) FKSN----

Dacrila Mulsant & Rey, 1874fallax (Kraatz, 1856) F-SNDE--

Brundinia Tottenham, 1949meridionalis (Mulsant & Rey, 1853) --S-DE--marina (Mulsant & Rey, 1853) --S-DE--

Hydrosmecta Thomson, 1858Hydrosmectina Ganglbauer, 1895

longula (Heer, 1839) FKSNDE--thinobioides (Kraatz, 1854)

delicatula (Sharp, 1869) F-SN----tenella (Mannerheim, 1830) FK------


subtilissima (Kraatz, 1854) F-SN-E--obscurior (Benick, 1969)

septentrionum (Benick, 1969) ----D---

Dilacra Thomson, 1858luteipes (Erichson, 1837) FKSNDE--vilis (Erichson, 1837) F-SNDE-I

Ousipalia Des Gozis, 1886caesula (Erichson, 1839) --S-DE--

Tomoglossa Kraatz, 1856luteicornis (Erichson, 1837) F-S-DE--

Schistoglossa Kraatz, 1856viduata (Erichson, 1837) FKSNDEAIdrusilloides (J.Sahlberg, 1876) FKS-D---gemina (Erichson, 1837) FKSND-AIpseudogemina Benick, 1981 ----D---curtipennis (Sharp, 1869) FKSNDE-Ibergvalli Palm, 1968 --S-D---

benicki Lohse, 1981aubei (Brisout de Barneville, 1860) FKSNDE--

Aloconota Thomson, 1858eichhoffi (Scriba, 1867) ---N----planifrons (Waterhouse, 1864) F-SNDE--strandi (Benick, 1939) FKSN----currax (Kraatz, 1856) FKSN----sulcifrons (Stephens, 1832) FKSNDE-Iinsecta (Thomson, 1856) FKSNDE--subgrandis (Brundin, 1954) F-SND---gregaria (Erichson, 1839) FKSNDEAI

Geostiba Thomson, 1858Sipalia auct., nec Mulsant & Rey, 1853Evanystes auct., nec Gistel, 1856

circellaris (Gravenhorst, 1802) FKSNDEAI

Callicerus Gravenhorst, 1802obscurus Gravenhorst, 1802 F-SND---rigidicornis (Erichson, 1839) ----D---

Dadobia Thomson, 1858immersa (Erichson, 1837) FKSNDEAI

Paranopleta Brundin, 1954inhabilis (Kraatz, 1856) FKSNDE--

Lundbergia Muona, 1975trybomi (J.Sahlberg, 1880) F-SN----

Cadaverota Yosii & Sawada, 1976cadaverina (Brisout de Barneville, 1860) FKSNDEAIhansseni (Strand, 1943) --SND---

Disopora Thomson, 1859languida (Erichson, 1837) F-S-DEAIultima (Benick & Lohse, 1959) -----E--longicollis (Mulsant & Rey, 1852) F----E--coulsoni (Last, 1952) --S-D---

Liogluta Thomson, 1858pagana (Erichson, 1839) F-SNDE--granigera (Kiesenwetter, 1850) FKSNDE-Imicans (Mulsant & Rey, 1852) FKSNDE--

letzneri (Eppelsheim, 1880)longiuscula (Gravenhorst, 1802) FKSNDEAImicroptera Thomson, 1867 FKSNDEAI

oblongiuscula (Sharp, 1869)oblonga (Erichson, 1839) nec (Lacordaire, 1835)

alpestris (Heer, 1839) FKSNDE-Initidula (Kraatz, 1856)

Philhygra Mulsant & Rey, 1873pseudoelongatula (Bernhauer, 1907) F-------arctica (Thomson, 1856) FKSNDE-Ibotanicarum (Muona, 1983) F-------polaris (Bernhauer, 1900) FKSN-E--elongatula (Gravenhorst, 1802) FKSNDEAIhygrobia (Thomson, 1856) FKSNDEAI

magniceps (J.Sahlberg, 1876)hygrotopora (Kraatz, 1856) F-SNDE--pinegensis (Muona, 1983) F--N----luridipennis (Mannerheim, 1830) FKSNDEAIterminalis (Gravenhorst, 1806) FKSNDEA-grisea (Thomson, 1852) F-S-DE--botildae (Brundin, 1954) --S-DE--tmolosensis (Bernhauer, 1940) --S-D---

dentifera (Brundin, 1943)gyllenhalii (Thomson, 1856) FKSNDE-Imelanocera (Thomson, 1856) FKSNDEAI

?vaga (Heer, 1839)malleus (Joy, 1913) FKSNDE-I

hygrobia auct. nec (Thomson, 1856)volans (Scriba, 1859) FKSNDE--

halophila (Thomson, 1861)tomlini (Joy, 1913)

obtusangula (Joy, 1913) F-SND---palustris (Kiesenwetter, 1844) FKSNDE-Idebilis (Erichson, 1837) FKSNDEAIdebiloides (Strand, 1962) F-SN----scotica (Elliman, 1909) F-SND---


parca (Mulsant & Rey, 1874) F-S-DEA-nannion (Joy, 1931)

deformis (Kraatz, 1856) FKSND---kaiseriana (Brundin, 1943) --S-D---britteni (Joy, 1913) FKSN-EA-mahleri Muona, 1995 ----D---ripicola (Hanssen, 1932) FKSNDE-I

flavithorax (Benick, 1976)fallaciosa (Sharp, 1869) FKSNDE-I

celata (Erichson, 1837) FKSNDEAIarenicola Thomson, 1868

sg. Xenota Mulsant & Rey, 1874myrmecobia (Kraatz, 1856) FKSNDE-I

sg. Mycetota Ádám, 1987laticollis (Stephens, 1832) FKSNDE-I

sg. Mocyta Mulsant & Rey, 1874clientula (Erichson, 1839) F-SND---orphana (Erichson, 1837) FKSNDEA-orbata (Erichson, 1837) FKSNDE-Inegligens (Mulsant & Rey, 1873) --SNDE--fungi (Gravenhorst, 1806) FKSNDEAIamplicollis (Mulsant & Rey, 1873) FKSND---amblystegii Brundin, 1952 F-SN----

sg. Lypoglossa Fenyes, 1918lateralis (Mannerheim, 1830) FKSN-E--

sg. Alaobia Thomson, 1858scapularis (Sahlberg, 1831) FKSNDE--sodalis (Erichson, 1837) FKSNDEAIgagatina (Baudi, 1848) FKSNDEAIpandionis Scheerpeltz, 1958 F-SN-E--pallidicornis (Thomson, 1856) FKSNDE-Ihybrida (Sharp, 1869) --S-D---trinotata (Kraatz, 1856) FKSNDEAIsparreschneideri Munster, 1923 FKSN----serrata Benick, 1938 --S-----taxiceroides Munster, 1935 F-SN----subglabra (Sharp, 1869) F-SND--I

sg. Notothecta Thomson, 1858flavipes (Gravenhorst, 1806) FKSND-AIconfusa (Märkel, 1844) FKSNDE-I

sg. Neohilara Lohse, 1971subterranea (Mulsant & Rey, 1853) FKSNDE--

sg. Badura Mulsant & Rey, 1873macrocera (Thomson, 1856) FKSNDE-Iehnstroemi Baranowski, 1982 --S-----puncticollis Benick, 1938 F-SNDE--

Atheta Thomson, 1858

sg. Parameotica Ganglbauer, 1895laticeps (Thomson, 1856) F-S-D---

sg. Amidobia Thomson, 1858talpa (Heer, 1841) FKSND-AI

sg. Microdota Mulsant & Rey, 1873palleola (Erichson, 1837) F-SN----benickiella Brundin, 1948 FKSNDE--wireni Brundin, 1948 F-S-----amicula (Stephens, 1832) FKSNDE-Ipittionii Scheerpeltz, 1950 F-------spatuloides Benick, 1939 FKSN----inquinula (Gravenhorst, 1802) FKS-DE-Iganglbaueri Brundin, 1948 -----E--excelsa Bernhauer, 1911 FKSND---subtilis (Scriba, 1866) FKSNDEAInesslingi Bernhauer, 1928 FKSND---atomaria (Kraatz, 1856) F-SND---kerstensi Benick, 1968 ----D---minuscula (Brisout de Barneville, 1859) --SND---

perexigua (Sharp, 1869))glabricula Thomson, 1867 F-SND---glabriculoides Strand, 1958 --SND---sundti Strand, 1971 ---N----indubia (Sharp, 1869) F-SND---liliputana (Brisout de Barneville, 1860) FKSND---

alpina Benick, 1940boreella Brundin, 1948 FKSND---platonoffi Brundin, 1948 FKS-----

sg. Datomicra Mulsant & Rey, 1874Datostiba Yosii & Sawada

zosterae (Thomson, 1856) FKSNDEA-nigra (Kraatz, 1856) FKSNDEAIdadopora Thomson, 1867 FKSND--Icanescens (Sharp, 1869) FKSNDE-Isordidula (Erichson, 1837) FKSNDE-I

sg. Chaetida Mulsant & Rey, 1874longicornis (Gravenhorst, 1802) FKSNDEAI

sg. Trochanterella Brundin, 1954cribripennis J.Sahlberg, 1890 FKSN----

sg. Rhagocneme Munster, 1923subsinuata (Erichson, 1839) FKSND-AIdwinensis Poppius, 1908 FKSN----


sg. Boreophilia Benick, 1973munsteri Bernhauer, 1902 FKSN----subplana J.Sahlberg, 1880 F-SN----islandica (Kraatz, 1857) FKSN-E--eremita (Rye, 1866) FKSNDE--

hercynica Renkonen, 1936hyperborea Brundin, 1940 FKSN----fusca (Sahlberg, 1831) FKSN----insecuta Eppelsheim, 1893 F-------latifemorata Brundin, 1940 F-SN----

sg. Dimetrota Mulsant & Rey, 1873nigripes (Thomson, 1856) FKSNDE-Imarcida (Erichson, 1837) F-SNDE--putrida (Kraatz, 1856) -------Ieuropaea Likovský, 1984 FKS--E--

livida (Mulsant & Rey, 1852) nec (Erichson, 1837)cinnamoptera (Thomson, 1856) FKSNDE--aeneipennis (Thomson, 1856) FKSNDE-I

picipennis auct. nec (Mannerheim, 1843)parapicipennis Brundin, 1954 FKSN----picipennoides Hanssen, 1932 FKSN-E--lapponica J.Sahlberg, 1876 FKSN----altaica Bernhauer, 1901 FKSN----intermedia (Thomson, 1852) FKSNDE-I

ravilla (Erichson, 1839) FKSND-AIangusticollis (Thomson, 1856)

procera (Kraatz, 1856) FKSN-E--basicornis (Mulsant & Rey, 1852) FKSNDEAInidicola (Johansen, 1914) FKSNDE-Iallocera Eppelsheim, 1893 FKSN----oblita (Erichson, 1839) FKSNDE--autumnalis (Erichson, 1839) FKSN-E-Iboletophila (Thomson, 1856) FKSNDE-Idiversa (Sharp, 1869) FKSN---Istrandiella Brundin, 1954 FKSNDEAIpilicornis (Thomson, 1852) FKSNDEAIacutangula Hanssen, 1936 F-SN----boleticola J.Sahlberg, 1876 FKSN-E-Ifungicola (Thomson, 1852) F-SNDE--britanniae Bernhauer & Scheerpeltz, 1926 --SND--Icrassicornis (Fabricius, 1793) FKSNDEAI

repanda Mulsant & Rey, 1873paracrassicornis Brundin, 1954 FKSN-E-Ipfaundleri Benick, 1940 --S-----pachycera (Eppelsheim, 1893) F-S-----contristata (Kraatz, 1856) F-------euryptera (Stephens, 1832) FKSNDEAIdivisa (Märkel, 1844) FKSNDE-Iliturata (Stephens, 1832) --S-DE-Inigricornis (Thomson, 1852) FKSNDE-Iharwoodi Williams, 1930 FKSNDE-Icoriaria (Kraatz, 1856) FKSNDE-I

sg. Boreostiba Lohse, 1990sibirica (Mäklin, 1880) FKSN----

frigida J.Sahlberg, 1880piligera J.Sahlberg, 1876 FKSN----thulea Poppius, 1909 F-------

sg. Plataraea Thomson, 1858nigrifrons (Erichson, 1839) --S-D---dubiosa Benick, 1934 FK---E--brunnea (Fabricius, 1798) F-SNDEA-nigritula (Gravenhorst, 1802) FKSNDEAI

sg. Traumoecia Mulsant & Rey, 1874picipes (Thomson, 1856) FKSNDEAI

sg. Bessobia Thomson, 1858occulta (Erichson, 1837) FKSND-AIfungivora (Thomson, 1867) FKS-D---monticola (Thomson, 1852) FKSND--Iexcellens (Kraatz, 1856) FKSND---

sg. Atheta s.str.Coproceramius Gistel, 1857

cauta (Erichson, 1837) FKSNDE-Iischnocera Thomson, 1870 FKSNDE-Isetigera (Sharp, 1869) FKSNDE-Ilaevana (Mulsant & Rey, 1852) FKSNDEAIatramentaria (Gyllenhal, 1810) FKSNDE-Ihypnorum (Kiesenwetter, 1850) FKSNDEA-laevicauda J.Sahlberg, 1876 FKSN---Ibrunneipennis (Thomson, 1852) FKSN-E--castanoptera (Mannerheim, 1830) FKSNDEA-

pertyi (Heer, 1839)triangulum (Kraatz, 1856) F-S-D---xanthopus (Thomson, 1856) F-SNDE--graminicola (Gravenhorst, 1806) FKSNDEAIebenina (Mulsant & Rey, 1873) FKSNDE--heymesi Hubenthal, 1913 F-SN----aquatilis (Thomson, 1867) --S-D---incognita (Sharp, 1869) FKSNDE--aquatica (Thomson, 1852) F-SNDE-Iaeneicollis (Sharp, 1869) --SND-A-

pertyi auct. nec (Heer, 1839)


sg. Enalodroma Thomson, 1859hepatica (Erichson, 1839) --SND---

sg. Thinobaena Thomson, 1859vestita (Gravenhorst, 1806) FKSNDE--

Anopleta Mulsant & Rey, 1874corvina (Thomson, 1856) FKSNDE-Isodermani (Bernhauer, 1931) F-SND---depressicollis (Fauvel, 1872) FKSN-E-Ijanssoni (Bernhauer, 1923) F-S-----nitella (Brundin, 1948) --SND---

Alevonota Thomson, 1858rufotestacea (Kraatz, 1856) --SND---gracilenta (Erichson, 1839) FKSND--I

Alianta Thomson, 1858incana (Erichson, 1837) FKSNDE-I

Trichiusa Casey, 1893immigrata Lohse, 1984 F-SND---

Pachnida Mulsant & Rey, 1874nigella (Erichson, 1837) F-S-DE--

Dinaraea Thomson, 1858angustula (Gyllenhal, 1810) FKSNDEAIaequata (Erichson, 1837) FKSNDEAIlinearis (Gravenhorst, 1802) FKSNDEAIarcana (Erichson, 1839) FKSN-E-I

Lyprocorrhe Thomson, 1859anceps (Erichson, 1837) FKSNDEAI

Nehemitropia Lohse, 1971lividipennis (Mannerheim, 1830) FKSNDEAI

curvipes (Stephens, 1832)sordida (Marsham, 1802) nec (Gravenhorst, 1802)

aterrima (Gravenhorst, 1802) FKSNDEAIbenicki (Allen, 1940) F-SND--I

pusilla (Brundin, 1952)parvula (Mannerheim, 1830) FKSNDEAI

Actophylla Bernhauer, 1908varendorffiana (Bernhauer & Scheerpeltz, 1926) ----D---

Coprothassa Thomson, 1859melanaria (Mannerheim, 1830) FKSNDEAI

Halobrecta Thomson, 1858puncticeps (Thomson, 1852) FKSNDE--flavipes Thomson, 1861 F-SNDE--

Amischa Thomson, 1858decipiens (Sharp, 1869) F-SNDEAIandreasi Muona, 1990 F-------nigrofusca (Stephens, 1832) FKSNDEAI

soror (Kraatz, 1856)simillima (Sharp, 1869)sarsi Munster, 1927

analis (Gravenhorst, 1802) FKSNDEAIbifoveolata (Mannerheim, 1830) FKSND---

cavifrons (Sharp, 1869)

Pycnota Mulsant & Rey, 1874paradoxa (Mulsant & Rey, 1861) F-SNDE--

nidorum (Thomson, 1868)

Pachyatheta Munster, 1930mortuorum (Thomson, 1867) FKSN----cribrata (Kraatz, 1856) FKSNDE--

Thamiaraea Thomson, 1858cinnamomea (Gravenhorst, 1802) FKSNDEAIhospita (Märkel, 1844) F-SND---

Falagriini Mulsant & Rey, 1873Falagria Samouelle, 1819caesa Erichson, 1837 FKSNDEAI

sulcata (Paykull, 1789) nec (Müller, 1776)sulcatula (Gravenhorst, 1806) F-SNDEAI

Myrmecopora Saulcy, 1865sulcata (Kiesenwetter, 1850) --SND---

simillima (Wollaston, 1864)lohmanderi Bernhauer, 1927

Acrotona Thomson, 1859Colpodota Mulsant & Rey, 1873

exigua (Erichson, 1837) F-SNDE-Ipilosicollis (Brundin, 1952) -K-----Isylvicola (Kraatz, 1856) FKSNDEAIpygmaea (Gravenhorst, 1802) FKSNDEAIconvergens (Strand, 1958) FKSND---obfuscata (Gravenhorst, 1802) F-SNDE-Itroglodytes (Motschulsky, 1858) F-SNDE--

consanguinea (Eppelsheim, 1875)pseudotenera (Cameron, 1933) F-SND---

rassii Muona, 1993parens (Mulsant & Rey, 1852) F-SNDE-Imuscorum (Brisout de Barneville, 1860) FKSNDE-I

Borboropora Kraatz, 1862Aneurota Casey, 1893

kraatzi Fuss, 1862 --S-----


Bohemiellina Machulka, 1941flavipennis (Cameron, 1921) F-SND---

paradoxa Machulka, 1941

Cordalia Jacobs, 1925obscura (Gravenhorst, 1802) FKSNDEAI

Myrmecocephalus MacLeay, 1871concinnus (Erichson, 1839) F-SND---

longipes (Wollaston, 1871)

Falagrioma Casey, 1906Falagriusa Ádám, 1987Stenagria auct., nec Sharp, 1883

thoracica (Stephens, 1832) FKSND---

Anaulacaspis Ganglbauer, 1895nigra (Gravenhorst, 1802) FKSNDEA-

Lomechusini Fleming, 1821Myrmedoniini Thomson, 1867

Drusilla Leach, 1819Astilbus Dillwyn, 1829

canaliculata (Fabricius, 1787) FKSNDEAI

Zyras Stephens, 1835collaris (Paykull, 1800) FKSNDEAIlimbatus (Paykull, 1789) FKSNDEAIfunestus (Gravenhorst, 1806) FKSNDEA-humeralis (Gravenhorst, 1802) FKSNDEAIcognatus (Märkel, 1842) FKSNDEAIlugens (Gravenhorst, 1802) FKSNDEAIlaticollis (Märkel, 1842) FKSNDEAI

Lomechusoides Tottenham, 1939strumosus (Fabricius, 1793) FKSNDE--wellenii (Palm, 1949) FKS-----inflatus (Zetterstedt, 1828) FKS-----

teres (Eppelsheim, 1886)mariae (Palm, 1949)

Lomechusa Gravenhorst, 1806Atemeles Dillwyn, 1829

emarginata (Paykull, 1789) FKSNDEA-paradoxa Gravenhorst, 1806 --SND-A-pubicollis Brisout de Barneville, 1860 FKSN----

Homalotini Heer, 1839Gyrophaenina Kraatz, 1856Gyrophaena Mannerheim, 1830nitidula (Gyllenhal, 1810) -KS-----pulchella Heer, 1839 FKSNDE--obsoleta Ganglbauer, 1895 --SN----affinis Mannerheim, 1830 FKSNDEAI

kangasi Rutanen, 1994 F-------rousi Dvoáák, 1966 --S-----nana (Paykull, 1800) FKSNDEAIpseudonana Strand, 1839 FKSND---rugipennis Mulsant & Rey, 1861 --S-----gentilis Erichson, 1839 FKSNDEAIpoweri Crotch, 1866 FKSNDEAIminima Erichson, 1837 FKSNDEAImunsteri Strand, 1935 F-SNDE-Ifasciata (Marsham, 1802) FKSNDEAI

laevipennis Kraatz, 1856congrua Erichson, 1837 FKSNDEA-hanseni Strand, 1946 ----D---williamsi Strand, 1935 FKSND---orientalis Strand, 1938 FKS-----bihamata Thomson, 1867 FKSNDEAIlucidula Erichson, 1837 F-S-DE--joyi Wendeler, 1924 FKSNDEAIjoyioides Wüsthoff, 1937 FKSNDE-Iangustata (Stephens, 1832) FKSNDEAI

manca Erichson, 1839strictula Erichson, 1839 FKSNDE-Iboleti (Linnaeus, 1758) FKSNDEAI

Agaricochara Kraatz, 1856latissima (Stephens, 1832) --SND---

laevicollis (Kraatz, 1854)

Encephalus Stephens, 1832complicans Stephens, 1832 FKSNDEAI

Bolitocharina Thomson, 1859Bolitochara Mannerheim, 1830obliqua Erichson, 1837 F-S-DEAImulsanti Sharp, 1875 FKSNDEAIpulchra (Gravenhorst, 1806) FKSNDEAI

lunulata (Paykull, 1789) nec (Linnaeus, 1761)bella Märkel, 1844 ----D---lucida (Gravenhorst, 1802) --SND-AI

Phymatura J.Sahlberg, 1876brevicollis (Kraatz, 1856) FKSN-E-I

Leptusa Kraatz, 1856Sipalia auct., nec Mulsant & Rey, 1853Pachygluta Thomson, 1858

pulchella (Mannerheim, 1830) FKSNDEAIsaalasi Kangas, 1952

fuliginosa (Aubé, 1850) -----E--norvegica Strand, 1941 FKSND-A-fumida (Erichson, 1839) F-SNDEAI


ruficollis (Erichson, 1839) F-SNDE--

Tachyusida Mulsant & Rey, 1872gracilis (Erichson, 1837) --S-D--I

Euryusa Erichson, 1837castanoptera Kraatz, 1856 FKSNDEAIoptabilis Heer, 1839 F-S-----sinuata Erichson, 1837 --S-D---coarctata Märkel, 1844 --S-----

Heterota Mulsant & Rey, 1874plumbea (Waterhouse, 1858) --S-----

Silusina Fenyes, 1918Silusa Erichson, 1837rubiginosa Erichson, 1837 F-SNDE-Irubra Erichson, 1839 -K---E--

Homalotina Heer, 1839Anomognathus Solier, 1849Theetura Thomson, 1858

cuspidatus (Erichson, 1839) FKSNDEAI

Thecturota Casey, 1893Pragensiella Machulka, 1941

marchii (Dodero, 1922) F-SNDE--

Homalota Mannerheim, 1830plana (Gyllenhal, 1810) FKSNDEAI

Pseudomicrodota Machulka, 1935jelineki (Krása, 1914) --SND---

flavicollis (Brundin, 1948)

Rhopalocerina Reitter, 1909clavigera (Scriba, 1859) --S-----

Cyphea Fauvel, 1863curtula (Erichson, 1837) FKSNDE--latiuscula Sjöberg, 1934 FKSN-E--

Placusini Mulsant & Rey, 1871Placusa Erichson, 1837complanata Erichson, 1839 FKS--E-Idepressa Mäklin, 1845 FKSNDE-Itachyporoides (Waltl, 1838) FKSNDEAIincompleta Sjöberg, 1934 FKSNDEAIcribrata Johnson & Lundberg, 1977 FKSN----suecica Johnson & Lundberg, 1977 F-SN----atrata (Mannerheim, 1830) FKSNDEAIpumilio (Gravenhorst, 1802) --S-D---adscita Erichson, 1839 F-------

Autaliini Thomson, 1859Autalia Leach, 1819impressa (Olivier, 1795) FKSNDEA-longicornis Scheerpeltz, 1947 F-SND---rivularis (Gravenhorst, 1802) FKSNDEAIpuncticollis Sharp, 1864 FKSNDEAI

Phytosini Thomson, 1867Phytosus Curtis, 1838spinifer Curtis, 1838 --SND---balticus Kraatz, 1859 --SND--I

Arena Fauvel, 1862tabida (Kiesenwetter, 1850) ----D---

Diglottini Jacobson, 1909Diglotta Champion, 1887sinuaticollis (Mulsant & Rey, 1870) --SND---

submarina auct. nec (Fairmaire & Laboulbène, 1856)mersa (Haliday, 1837) ----D---

submarina (Fairmaire & Laboulbène, 1856)

Hygronomini Thomson, 1859Hygronoma Erichson, 1837dimidiata (Gravenhorst, 1806) FKSNDEAI

Hypocyphtini Laporte de Castelnau, 1835Oligotini Thomson, 1859

Holobus Solier, 1849apicatus (Erichson, 1837) FKSNDE--flavicornis (Lacordaire, 1835) F-SNDEAI

Oligota Mannerheim, 1830granaria Erichson, 1837 --SNDE-Iparva Kraatz, 1862 F-SNDE--inflata (Mannerheim, 1830) FKSNDEA-muensteri Bernhauer, 1923 FKSN----tantilla Mannerheim, 1843 FK-N----punctulata Heer, 1839 ----D-A-

ruficornis Sharp, 1870pusillima (Gravenhorst, 1806) FKSNDEAI

atomaria Erichson, 1837intermedia Kangas, 1938

inexpectata Williams, 1994 F-------picipes (Stephens, 1832) ----D---pumilio Kiesenwetter, 1858 --S-D-A-uralensicola Kangas, 1982 F-------

Cypha Samouelle, 1819Hypocyphtus Gyllenhal, 1827

longicornis (Paykull, 1800) --S-DEA-imitator (Luze, 1902) F-SNDEA-


tarsalis (Luze, 1902) FKSNDE--laeviuscula (Mannerheim, 1830) FKSND-AIovulum (Heer, 1839) FKSN----discoidea (Erichson, 1839) F-S-D---suecica (Palm, 1936) FKSND---nitida (Palm, 1936) --SND---pulicaria (Erichson, 1839) F-SND-AIhanseni (Palm, 1949) -KS-D---seminulum (Erichson, 1839) F-S-D-AIpunctum (Motschulsky, 1857) --SND---

Myllaenini Ganglbauer, 1895Myllaena Erichson, 1837dubia (Gravenhorst, 1806) FKSNDEAIintermedia Erichson, 1837 FKSNDEAIkraatzi Sharp, 1871 FKSNDEAIhyperborea Strand, 1943 F-SN----elongata (Matthews, 1838) ---NDE--brevicornis (Matthews, 1838) FKSNDE-Igracilis (Matthews, 1838) FKSNDE--minuta (Gravenhorst, 1806) FKSNDEAIinfuscata Kraatz, 1853 --S-DE-Imasoni A.Matthews, 1883 ---ND---

Gymnusini Heer, 1839Gymnusa Gravenhorst, 1806brevicollis (Paykull, 1800) FKSNDEAIvariegata Kiesenwetter, 1845 FKSN----

Deinopsini Sharp, 1883Deinopsis Matthews, 1838erosa (Stephens, 1832) FKSNDEAI

Scaphidiinae Latreille, 1807Scaphidiini Latreille, 1807Scaphidium Olivier, 1790quadrimaculatum Olivier, 1790 FKSNDEAI

Scaphisomatini Casey, 1894Scaphisoma Leach, 1815agaricinum (Linnaeus, 1758) FKSNDEAIinopinatum Löbl, 1967 FKSN--AIboleti (Panzer, 1793) FKSNDEAIsubalpinum Reitter, 1881 FKSN-EA-balcanicum Tamanini, 1954 F-SN--A-boreale Lundblad, 1952 FKSN--A-assimile Erichson, 1845 FKSNDEAI

Oxytelinae Fleming, 1821Deleasterini Reitter, 1909Syntomium Curtis, 1828aeneum (Müller, 1821) FKSNDEAI

Deleaster Erichson, 1839dichrous (Gravenhorst, 1802) FKSNDEAI

Coprophilini Heer, 1839Coprophilus Latreille, 1829Elonium Samouelle, 1819 (Suppr.)

striatulus (Fabricius, 1793) FKSNDEAI

Thinobiini J.Sahlberg, 1876Manda Blackwelder, 1952Acrognathus Erichson, 1839, nec Agassiz, 1826

mandibularis (Gyllenhal, 1827) --S-DEAI

Planeustomus Jacquelin du Val, 1857palpalis (Erichson, 1839) --S-D---

Ochthephilus Mulsant & Ray, 1856Ancyrophorus Kraatz, 1857

omalinus (Erichson, 1840) FKSN-E--strandi (Scheerpeltz, 1950) FKSN----praepositus Mulsant & Rey, 1878 ----D---

longipennis auct. nec (Fairmaire & Laboulbène, 1856)

Thinobius Kiesenwetter, 1844brundini Scheerpeltz, 1959 --SN----munsteri Scheerpeltz, 1959 F-SN----longicornis J.Sahlberg, 1874 -K-N----ciliatus Kiesenwetter, 1844 FKSND---

praetor Smetana, 1959crinifer Smetana, 1959ssp. crinifer Smetana, 1959 ----D---ssp. strandi Smetana, 1960 FKSN----flagellatus Lohse, 1984 FKS-----

pusillimus auct. nec (Heer, 1839)brevipennis Kiesenwetter, 1850 F-SNDE--

Thinodromus Kraatz, 1857hirticollis Mulsant & Rey, 1879 -----E--arcuatus (Stephens, 1834) FKSNDE--Carpelimus Leach, 1819Trogophloeus Mannerheim, 1830

bilineatus Stephens, 1834 FKSNDEAIerichsoni (Sharp, 1871) ------A-

augustae (Bernhauer, 1901)rivularis (Motschulsky, 1860) FKSNDEAIobesus (Kiesenwetter, 1844) FKSND-AIfuliginosus (Gravenhorst, 1802) F-SNDE-Initidus (Baudi, 1848) -------Ilindrothi (Palm, 1943) F-S-DEA-impressus (Lacordaire, 1835) FKSNDEA-heidenreichi (L.Benick, 1934) --S---A-corticinus (Gravenhorst, 1806) FKSNDEAI


subtilicornis (Roubal, 1946) FKSN-EA-strandi (Scheerpeltz, 1950)

foveolatus (Sahlberg, 1832) FKSNDEA-halophilus (Kiesenwetter, 1844) FKS-DE--despectus (Baudi, 1870) FKSNDEA-zealandicus (Sharp, 1900) --s-----pusillus (Gravenhorst, 1802) FKSNDEAI

lindbergi (Scheerpeltz, 1937)gracilis (Mannerheim, 1830) FKSNDEAI

graciliformis Konzelmann & Lohse, 1981subtilis (Erichson, 1839) -KSNDE-Ielongatulus (Erichson, 1839) FKSNDEAIsubterraneus (Smetana, 1960) ------A-schneideri (Ganglbauer, 1895) --S-D---

Aploderus Stephens, 1833caelatus (Gravenhorst, 1802) FKSNDEAIcaesus (Erichson, 1839) -KS-DEA-

bernhaueri Poppius, 1909 FKSN----kutsae Kangas, 1937 -KS-----arcticus J.Sahlberg, 1890 FKSN----terebrans (Schiödte, 1866) FKSNDEA-filipes Sharp, 1911 -KS-DEA-poppiusi Bernhauer, 1902 FKSN----fennicus Kangas, 1937 FK------vilis Mäklin, 1876 FK-N-EAIlongulus Erichson, 1839 FKSNDEAIlativentris Jansson, 1928 F-SN----denticollis Fauvel, 1872 FKSN-EA-opacus (Block, 1799) FKSNDEAI

subsinuatus Mulsant & Rey, 1878baudii Fauvel, 1872 --S-D---pygmaeus Erichson, 1839 ----D---

agricultor Heer, 1841pusillus Erichson, 1839 ----D---erraticus Erichson, 1839 FKSND-AI

bosnicus auct. nec Bernhauer, 1902fontinalis auct. nec Bernhauer, 1929

fergussoni Joy, 1912 FKSNDEAIarenarius (Paykull, 1800) nec (Geoffroy, 1785)

subniger Schneider, 1898 ----D---frater Kraatz, 1857 ------A-tibialis Heer, 1839 F-SND---

Oxytelini Fleming, 1821Oxytelus Gravenhorst, 1802sculptus Gravenhorst, 1806 FKSNDEAImigrator Fauvel, 1904 F-S-D--Ifulvipes Erichson, 1839 FKSNDEAIpiceus (Linnaeus, 1767) FKS-DEAIlaqueatus (Marsham, 1802) FKSNDEAI

Bledius Leach, 1819tricornis (Herbst, 1784) FKSNDEAIlimicola Tottenham, 1940 F-S-DE--

germanicus Wagner, 1935, nec (Gravenhorst, 1806)spectabilis Kraatz, 1857 ----D---furcatus (Olivier, 1811) --S-D---bicornis (Germar, 1823) ----D-A-

dama Motschulsky, 1857jutlandensis Herman, 1986

diota Schiödte, 1866 F-S-DE--dissimilis Erichson, 1840 -K--DEAIoccidentalis Bondroit, 1907 F-SND--I

crassicollis auct. nec Lacordaire, 1835cribricollis Heer, 1839 ----D-A-femoralis (Gyllenhal, 1827) --SND-AIgallicus (Gravenhorst, 1806) FKSNDEAI

fracticornis (Paykull, 1790) nec (Müller, 1776)procerulus Erichson, 1840 --S----Iatricapillus (Germar, 1825) --S-D-A-

praetermissus Williams, 1929nanus Erichson, 1840 --S-D-A-talpa (Gyllenhal, 1810) FKSNDEAIsubterraneus Erichson, 1839 FKSNDEAIlittoralis Heer, 1839 -KSN-EA-defensus Fauvel, 1872 FK--DE-Ipallipes (Gravenhorst, 1806) FKSNDEAI

larseni Hansen, 1940rossicus Bernhauer & Schubert, 1911 -------Ifuscipes Rye, 1865 FKSNDE--condensus Kangas, 1938 -K------

Anotylus Thomson, 1859insecatus (Gravenhorst, 1806) F-SNDEA-rugifrons (Hochhuth, 1849) --S-DEAIrugosus (Fabricius, 1775) FKSNDEAImaritimus Thomson, 1861 --SND-A-

perrisii (Fauvel, 1862)inustus (Gravenhorst, 1806) --S-D-AIsculpturatus (Gravenhorst, 1806) FKS-DEAImutator (Lohse, 1963) --S-D---politus (Erichson, 1840) ------A-nitidulus (Gravenhorst, 1802) FKSNDEAIpetzi (Bernhauer, 1914) --S-----complanatus (Erichson, 1839) F-SNDEAIintricatus (Erichson, 1840) ------A-clypeonitens (Pandellé, 1867) ----DEA-


pumilus (Erichson, 1839) FKS-----affinis (Czwalina, 1870) -----E--hamatus (Fairmaire & Laboulbène, 1856) FKSN-EA-saulcyi (Pandellé, 1867) --S-D---fairmairei (Pandellé, 1867) FKSNDEAIclavatus (Strand, 1946) FKSN----tetracarinatus (Block, 1799) FKSNDEAItetratoma (Czwalina, 1870) FK-N----

Platystethus Mannerheim, 1830arenarius (Geoffroy, 1785) FKSNDEAIcornutus (Gravenhorst, 1802) F-S-DEAIalutaceus Thomson, 1861 FKSNDEAIcapito Heer, 1839 FK---E--nodifrons Mannerheim, 1830 FKSNDEAInitens (Sahlberg, 1832) FKS-DEAI

Oxyporinae Fleming, 1821Oxyporus Fabricius, 1775rufus (Linnaeus, 1758) FKSNDEAImannerheimii Gyllenhal, 1827 FK---EAImaxillosus Fabricius, 1793 FK--DEAI

Steninae MaLeay, 1825Stenus Latreille, 1797biguttatus (Linnaeus, 1758) FKSNDEAIcomma LeConte, 1863 FKSNDEAI

bipunctatus Erichson, 1839, nec Ljungh, 1804longipes Heer, 1839 ------AIguttula Müller, 1821 --S-D-A-fossulatus Erichson, 1840 FKSNDE--aterrimus Erichson, 1839 FKSND--Ijuno (Paykull, 1789) FKSNDEAIcalcaratus Scriba, 1864 -KS---AIater Mannerheim, 1830 FKSNDEAIlongitarsis Thomson, 1851 FKSND-A-bilineatus J.Sahlberg, 1871 FKSN----fasciculatus J.Sahlberg, 1871 FKSN----lustrator Erichson, 1839 FKSNDEAIproditor Erichson, 1839 FKSND-AIgallicus Fauvel, 1873 --S-D---sylvester Erichson, 1839 FKSNDEAIclavicornis (Scopoli, 1763) FKSNDEAIprovidus Erichson, 1839 FKSNDEAI

rogeri Kraatz, 1857scrutator Erichson, 1840 FKSNDEAIbimaculatus Gyllenhal, 1810 FKSNDEAIpalposus Zetterstedt, 1828 FKSN-EAIruralis Erichson, 1840 FKSN--A-strandi L.Benick, 1937 FKSN-E--

boops Ljungh, 1810 FKSNDEAIincrassatus Erichson, 1839 FKSNDEAImelanarius Stephens, 1833 FKSNDEAIsubdepressus Mulsant & Rey, 1861 ----D---morio Gravenhorst, 1806 FKSNDEAIexcubitor Erichson, 1839 FKS-DE--

rossicus Bernhauer, 1903ampliventris J.Sahlberg, 1890 FK------atratulus Erichson, 1839 FKSNDEAImelanopus (Marsham, 1802) F-S-D-A-canaliculatus Gyllenhal, 1827 FKSNDEAInitens Stephens, 1833 FKSNDEA-latipennis J.Sahlberg, 1880 FK------confusus J.Sahlberg, 1876 FKSN-E--vinnulus Casey, 1884 FKSN----

confusoides Renkonen, 1935labilis Erichson, 1840 FKSN----fuscipes Gravenhorst, 1802 FKSNDEAIkongsbergensis Munster, 1912 FKSN-E--argus Gravenhorst, 1806 FKSNDEAIageus Casey, 1884 FKSN-EA-

gerhardti L. Benick, 1915europaeus Puthz, 1966 FKSNDE-Icautus Erichson, 1839 FKS-DEAI

vafellus Erichson, 1839mendicus Erichson, 1840 ------A-incanus Erichson, 1839 -----EA-subarcticus Poppius, 1907 FKSN----pusillus Stephens, 1833 FKSNDEAInanus Stephens, 1833 FKSNDEAIassequens Rey, 1884 FKSNDE--

pusio Casey, 1884simillimus L.Benick, 1949

circularis Gravenhorst, 1802 FKSNDEAIpumilio Erichson, 1839 FKSND-A-humilis Erichson, 1839 FKSNDEAIcarbonarius Gyllenhal, 1827 FKSNDEAIglabellus Thomson, 1870 --S-D--Ihyperboreus J.Sahlberg, 1876 FKSN----eumerus Kiesenwetter, 1850 ------A-scabriculus J.Sahlberg, 1876 FKSN--A-opticus Gravenhorst, 1806 FKSNDEAIcrassus Stephens, 1833 FKSNDEAIformicetorum Mannerheim, 1843 FKSNDE-Iintermedius Rey, 1884 FKSND--I

problematicus Kevan & Allen, 1962brunnipes Stephens, 1833 F-SNDEAInigritulus Gyllenhal, 1827 FKSNDEAI


latifrons Erichson, 1839 FKSNDEAIfulvicornis Stephens, 1833 FKSND-AItarsalis Ljungh, 1804 FKSNDEAIbohemicus Machulka, 1947 FKS-DEAIoscillator Rye, 1870 --SND---similis (Herbst, 1784) FKSNDEAIsolutus Erichson, 1840 --S-D--Icicindeloides (Schaller, 1783) FKSNDEAIkiesenwetteri Rosenhauer, 1856 -K------fornicatus Stephens, 1833 FKSND-AIpallitarsis Stephens, 1833 FKSNDEAIniveus Fauvel, 1865 FKSNDE--binotatus Ljungh, 1804 FKSNDEAIpubescens Stephens, 1833 FKS-DEAIumbratilis Casey, 1884 FKSNDEA-

pseudopubescens Strand, 1940flavipes Stephens, 1833 FKS-DEAIaudax J.Sahlberg, 1900 FKSN----nitidiusculus Stephens, 1833 --SND---picipennis Erichson, 1840 ----DE--bifoveolatus Gyllenhal, 1827 FKSNDEAIpicipes Stephens, 1833 -KSNDE-I

brevipennis Thomson, 1851foveicollis Kraatz, 1857

impressus Germar, 1824 FKSNDEAIludyi Fauvel, 1886 FKSNDE--ochropus Kiesenwetter, 1858 --SNDE-Ifuscicornis Erichson, 1840 -------Ipalustris Erichson, 1839 FKSNDE-Iflavipalpis Thomson, 1860 FKSNDEAIgeniculatus Gravenhorst, 1806 FKSNDEAIkolbei Gerhardt, 1893 -----E-Ipallipes Gravenhorst, 1802 FKS-DEAInoctivagus Casey, 1884 FKSN----

sibiricus auct. nec J.Sahlberg, 1880

Paederinae Fleming, 1821Paederidus Mulsant & Rey, 1877ruficollis (Fabricius, 1781) --S-D---

Paederus Fabricius, 1775riparius (Linnaeus, 1758) FKSNDEAIlittoralis Gravenhorst, 1802 --S-DEAIfuscipes Curtis, 1826 F-SNDEAIbrevipennis Lacordaire, 1835 ------AI

Astenus Stephens, 1833procerus (Gravenhorst, 1806) F-SND-AI

filiformis (Latreille, 1806) nec (Fabricius, 1793)pulchellus (Heer, 1839) FKSNDEAIgracilis (Paykull, 1789) FKSNDEAI

angustatus (Paykull, 1789) nec (Schrank, 1781)lyonessius (Joy, 1908) F-S-DEAI

longelytratus Palm, 1936immaculatus Stephens, 1833 --S-D---

Rugilus Leach, 1819Stilicus Berthold, 1827

fragilis (Gravenhorst, 1806) FKS--EAIangustatus (Geoffroy, 1785) nec (Schrank, 1781)scutellatus (Motschulsky, 1858)

rufipes Germar, 1836 FKSNDEAIorbiculatus (Paykull, 1789) FKSNDEAIerichsonii (Fauvel, 1867) FKSNDEAIsimilis (Erichson, 1839) FKSNDEAIgeniculatus (Erichson, 1839) ----DEAI

Medon Stephens, 1833apicalis (Kraatz, 1857) F-SND---rufiventris (Nordmann, 1837) --S-----dilutus (Erichson, 1839) --S-----brunneus (Erichson, 1839) --S-D---castaneus (Gravenhorst, 1802) --S-D--Iripicola (Kraatz, 1854) --S-D--Ifusculus (Mannerheim, 1830) --SN---I

Sunius Stephens, 1829Hypomedon Mulsant & Rey, 1877

melanocephalus (Fabricius, 1793) --S-D-A-bicolor (Olivier, 1795) --S-D---

Pseudomedon Mulsant & Rey, 1877obsoletus (Nordmann, 1837) FKS-D---obscurellus (Erichson, 1840) FKSND-A-

Lithocharis Dejean, 1833nigriceps Kraatz, 1859 FKSNDEA-ochracea (Gravenhorst, 1802) FKSND-A-

tricolor auct. nec (Fabricius, 1787)

Dianous Samouelle, 1819coerulescens (Gyllenhal, 1810) FKSNDEA-

Euaesthetinae Thomson, 1859Edaphus Motschulsky, 1857beszedesi Reitter, 1913 -----ea-

bluehweissi Scheerpeltz, 1936

Euaesthetus Gravenhorst, 1806ruficapillus (Lacordaire, 1835) FKSNDEAIlaeviusculus Mannerheim, 1844 FKSNDE-Ibipunctatus (Ljungh, 1804) FKSNDEAI


Chloecharis Lynch, 1884debilicornis (Wollaston, 1857) --S-----

Scopaeus Erichson, 1840laevigatus (Gyllenhal, 1827) FKSNDEAIsulcicollis (Stephens, 1833) --SNDE--

cognatus Mulsant & Rey, 1855minutus Erichson, 1840 FKS-D---pusillus Kiesenwetter, 1843 F-SN--A-ryei Wollaston, 1872 ----D---

minimus auct, nec (Erichson, 1839)

Lathrobium Gravenhorst, 1802Tetartopeus Czwalina, 1888

multipunctum Gravenhorst, 1802 --SNDEAIzetterstedti Rye, 1872 FKSN-EA-

punctatum Zetterstedt, 1828, nec (Goeze, 1785)rufonitidum Reitter, 1909 F-S-DE--

fennicum Renkonen, 1938sphagnetorum Muona, 1977 FKSN----

gracile Hampe, 1866, nec Gravenhorst, 1802terminatum Gravenhorst, 1802 FKSNDEAIquadratum (Paykull, 1789) FKSNDEAIscutellare Nordmann, 1837 -----EAIelongatum (Linnaeus, 1767) FKSNDEAIrufipenne Gyllenhal, 1813 FKSNDEAIgeminum Kraatz, 1857 FKSNDEAI

boreale Hochhuth, 1851volgense Hochhuth, 1851

pallidipenne Hochhuth, 1851 -K--D-A-ripicola Czwalina, 1888

castaneipenne Kolenati, 1846 --S-DEA-fulvipenne Gravenhorst, 1806 FKSNDEAIbrunnipes (Fabricius, 1793) FKSNDEAIimpressum Heer, 1841 FKSNDEAI

filiforme Gravenhorst, 1806, nec (Fabricius, 1793)fovulum Stephens, 1833 FKSNDEAIlongulum Gravenhorst, 1802 FKSNDEAI

patris Benick, 1950pallidum Nordmann, 1837 --SNDE--dilutum Erichson, 1839 F-SNDEAI

Achenium Samouelle, 1819humile (Nicolai, 1822) --S-DE--

Ochthephilum Stephens, 1829Cryptobium Mannerheim, 1830

fracticorne (Paykull, 1800) FKSNDEAIcollare (Reitter, 1884) F-S-D---

Staphylininae Latreille, 1802Xantholinini Erichson, 1839Leptacinus Erichson, 1839pusillus (Stephens, 1833) FKSNDE-I

linearis (Gravenhorst, 1802) nec (Olivier, 1793)formicetorum Märkel, 1841 FKSNDE-Isulcifrons (Stephens, 1833) F-SN-EAI

othioides auct. nec Baudi, 1870intermedius Donisthorpe, 1936 FKSNDEAIbatychrus (Gyllenhal, 1827) FKSNDEAI

Phacophallus Coiffait, 1956parumpunctatus (Gyllenhal, 1827) FKSNDEA-

Gauropterus Thomson, 1860fulgidus (Fabricius, 1787) FKSNDEA-

Gyrohypnus Leach, 1819punctulatus (Paykull, 1789) FKSNDEAI

ater auct. nec (Stephens, 1833)fracticornis (Müller, 1776) FKSNDEAIangustatus Stephens, 1833 FKSNDEAI

liebei (Scheerpeltz, 1926)scoticus (Joy, 1913)

atratus (Heer, 1839) FKSNDE-I

Nudobius Thomson, 1860lentus (Gravenhorst, 1806) FKSNDEAI

Megalinus Mulsant & Rey, 1877glabratus (Gravenhorst, 1802) --S-D--I

Hypnogyra Casey, 1906Phalacrolinus Coiffait, 1972

angularis (Ganglbauer, 1895) F-S--EAIglabra (Gravenhorst, 1806) nec (Müller, 1776)

Leptophius Coiffait, 1983flavocinctus (Hochhuth, 1849) ------AI

relucens auct. nec (Gravenhorst, 1806)Xantholinus Dejean, 1821linearis (Olivier, 1794) FKSNDEAIlongiventris Heer, 1839 FKS-DEAIaudrasi Coiffait, 1956 --SN-E--

strandi Coiffait, 1958gallicus Coiffait, 1956 ----DEA-

rhenanus Coiffait, 1962dissimilis Coiffait, 1956 ----D---roubali Coiffait, 1956 -KS-DE--tricolor (Fabricius, 1787) FKSNDEAI

meyeri Drugmand, 1994


laevigatus Jacobsen, 1847 FKSNDEAIclairei Coiffait, 1956

distans Mulsant & Rey, 1853 -----EAI

Othiini Thomson, 1859Othius Stephens, 1829punctulatus (Goeze, 1777) FKSNDEAIlapidicola Märkel & Kiesenwetter, 1847FKSN-E-Ilaeviusculus Stephens, 1832 -------Iangustus Stephens, 1833 FKSNDEA-

melanocephalus (Gravenhorst, 1806) nec (Geoffroy, 1785)volans J.Sahlberg, 1876 FKSN-EA-subuliformis Stephens, 1833 FKSNDEAI

myrmecophilus Kiesenwetter, 1843

Atrecus Jacquelin du Val, 1856Baptolinus Kraatz, 1858

pilicornis (Paykull, 1790) FKSN-E-Iaffinis (Paykull, 1789) FKSNDEAIlongiceps (Fauvel, 1872) FKSNDEAI

Staphylinini Latreille, 1802Philonthina Kirby, 1837Erichsonius Fauvel, 1874Actobius Fauvel, 1876

cinerascens (Gravenhorst, 1802) FKSNDEAI

Gabrius Stephens, 1829nigritulus (Gravenhorst, 1802) FKSNDEAIsubnigritulus Joy, 1913 -----E--

subnigrituloides (Scheerpeltz, 1933)appendiculatus Sharp, 1910 FKSNDEAI

subnigritulus (Reitter, 1909) (infrasubspec.)toxotes Joy, 1913 FKS-DE--sphagnicola (Sjöberg, 1950) FKSN-E--austriacus Scheerpeltz, 1947 FKSNDEAI

velox auct. nec (Sharp, 1910)keysianus Sharp, 1910 ----D---breviventer (Sperk, 1835) FKSNDEAI

coxalus (Hochhuth, 1871)pennatus Sharp, 1910

splendidulus (Gravenhorst, 1802) FKSNDEAIexpectatus Smetana, 1952 FKSN-E--bescidicus Smetana, 1954 F-S-----osseticus (Kolenati, 1846) FKSNDEAI

vernalis (Gravenhorst, 1802) nec (Müller, 1776)dufbergi Gillerfors, 1997 --S-----exiguus (Nordmann, 1837) --S-DE--astutoides (Strand, 1946) F-SN----

lividipes (Baudi, 1848) FKS----Itirolensis (Luze, 1903) --SN---piliger Mulsant & Rey, 1876 --S-DE--trossulus (Nordmann, 1837) FKSNDEAI

Bisnius Stephens, 1829nitidulus (Gravenhorst, 1802) FKSNDEAIscoticus (Joy & Tomlin, 1910) FKSN----diversipennis (Bernhauer, 1899) FK------fimetarius (Gravenhorst, 1802) FKSNDEAI

rigidicornis (Gravenhorst, 1802)puella (Nordmann, 1837) FKSNDEAIsubuliformis (Gravenhorst, 1802) FKSNDEAI

fuscus (Gravenhorst, 1802) nec (Gmelin, 1790)spermophili (Ganglbauer, 1897) -----E--nigriventris (Thomson, 1867) FKSND-A-sordidus (Gravenhorst, 1802) FKSNDEAI

pachycephalus (Nordmann, 1837)varipennis (Scriba, 1864) -------I

scribai (Fauvel, 1867)parcus (Sharp, 1874) F-SND---pseudoparcus (Brunne, 1976) --SND---cephalotes (Gravenhorst, 1802) FKSNDEAI

Rabigus Mulsant & Rey, 1876tenuis (Fabricius, 1793) FK---EAI

Gabronthus Tottenham, 1955thermarum (Aubé, 1850) F-SNDE--sulcifrons (Sharp, 1889) F-------

Philonthus Stephens, 1829Spatulonthus Tottenham, 1955Paragabrius Coiffait, 1963

lepidus (Gravenhorst, 1802) FKSNDEAIrectangulus Sharp, 1874 FKSNDEAIdiscoideus (Gravenhorst, 1802) F-SND-AIsalinus Kiesenwetter, 1844 --S-D---quisquiliarius (Gyllenhal, 1810) FKSNDEAIcorvinus Erichson, 1839 FKSNDEAIsanguinolentus (Gravenhorst, 1802) FKSNDEAIbinotatus (Gravenhorst, 1802) --S-DE-Iventralis (Gravenhorst, 1802) FKSNDEAIintermedius (Lacordaire, 1835) --S-DEAIlaminatus (Creutzer, 1799) FKSNDEAIpolitus (Linnaeus, 1758) FKSNDEAIsuccicola Thomson, 1860 FKSNDEAI

chalceus Ganglbauer, 1895, nec Stephens, 1832addendus Sharp, 1867 FKSNDEAIcyanipennis (Fabricius, 1793) ------A-


caerulescens (Lacordaire, 1835) ------A-nitidus (Fabricius, 1787) FKSNDEAIspinipes Sharp, 1874 F-S-D--Irotundicollis (Ménétriés, 1832) FKSN-EAItemporalis Mulsant & Rey, 1853 -----EA-tenuicornis Mulsant & Rey, 1853 FKSNDEAI

carbonarius (Gyllenhal, 1810) nec (Gravenhorst, 1802)decorus (Gravenhorst, 1802) FKSNDEAIcognatus Stephens, 1832 FKSNDEAI

fuscipennis (Mannerheim, 1830) nec (Block, 1799)mannerheimi Fauvel, 1868 --SNDEAIcruentatus (Gmelin, 1790) FKSNDEAIconfinis Strand, 1941 FKSNDE--

isthmius Kangas, 1979varians (Paykull, 1789) FKSNDEAIpseudovarians Strand, 1941 F--NDE--jurgans Tottenham, 1937 --S-D---marginatus (Müller, 1764) FKSNDEAIlederi Eppelsheim, 1893 FKS-----atratus (Gravenhorst, 1802) FKSNDEAIsubvirescens Thomson, 1884 FKSN-E-Initidicollis (Lacordaire, 1835) --S-DE-I

bimaculatus (Gravenhorst, 1802) nec (Schrank, 1798)carbonarius (Gravenhorst, 1802) FKSNDEAI

varius (Gyllenhal, 1810)alpinus Eppelsheim, 1875 F-SNDE-Ialbipes (Gravenhorst, 1802) FKSNDEAIcaucasicus Nordmann, 1837 FK------

dimidiatus (Sahlberg, 1830) nec (Panzer, 1796)corruscus (Gravenhorst, 1802) --S-DE-Iebeninus (Gravenhorst, 1802) FKSNDEAIconcinnus (Gravenhorst, 1802) FKSNDEAI

ochropus (Gravenhorst, 1802)rufipes (Stephens, 1832) --S-D-AI

immundus (Gyllenhal, 1810) nec (Gravenhorst, 1806)debilis (Gravenhorst, 1802) FKSNDEAIsplendens (Fabricius, 1793) FKSNDEAIpunctus (Gravenhorst, 1802) FKS-DEAIlinki Solsky, 1866 -------Ifumarius (Gravenhorst, 1806) FKSNDEAIumbratilis (Gravenhorst, 1802) FKSNDEAInigrita (Gravenhorst, 1806) FKSNDEAIlaevicollis (Lacordaire, 1835) -----EAImontivagus Heer, 1839 ------AIlongicornis Stephens, 1832 FKSNDEAIcoprophilus Jarrige, 1949 ------AIparvicornis (Gravenhorst, 1802) FKSNDEAI

agilis (Gravenhorst, 1806)

rubripennis Stephens, 1832 FKSNDEAIfulvipes (Fabricius, 1792) nec (Scopoli, 1763)

furcifer Renkonen, 1937 FKSNDE-Imicans (Gravenhorst, 1802) FKSNDEAImicantoides Benick & Lohse, 1956 F--NDEAI

Neobisnius Ganglbauer, 1895villosulus (Stephens, 1833) FKSNDE-Ilathrobioides (Baudi, 1848) F-SNDE--

cerrutii Gridelli, 1943prolixus (Erichson, 1840) F-------procerulus (Gravenhorst, 1806) F-S-DE--

Remus Holme, 1837sericeus Holme, 1837 --SND---

Cafius Stephens, 1826xantholoma (Gravenhorst, 1806) F-SNDEAI

Staphylinina Latreille, 1802Dinothenarus Thomson, 1858pubescens (DeGeer, 1774) FKSNDEAI

Ontholestes Ganglbauer, 1895tessellatus (Geoffroy, 1785) FKSNDEAImurinus (Linnaeus, 1758) FKSNDEAI

Emus Leach, 1819hirtus (Linnaeus, 1758) --S-DEAI

Platydracus Thomson, 1858fulvipes (Scopoli, 1763) FKSNDEAIstercorarius (Olivier, 1795) FKSNDEAIchalcocephalus (Fabricius, 1801) ----DEA-latebricola (Gravenhorst, 1806) FKSNDEAI

Staphylinus Linnaeus, 1758erythropterus Linnaeus, 1758 FKSNDEAIcaesareus Cederhjelm, 1798 FKSNDEAIdimidiaticornis Gemminger, 1851 -KSNDEAI

Ocypus Leach, 1819olens (Müller, 1764) --S-DE--ophthalmicus (Scopoli, 1763) FKSNDEAInitens (Schrank, 1781) FKS-DEAI

nero (Faldermann, 1835)similis (Fabricius, 1792) nec (Herbst, 1784)

brunnipes (Fabricius, 1781) FKSNDEAIfuscatus (Gravenhorst, 1802) FKSNDEAIpicipennis (Fabricius, 1793) FKSNDEAIfulvipennis Erichson, 1840 FK---EAIaeneocephalus (DeGeer, 1774) FKSNDEAI


Tasgius Stephens, 1829Alapsodus Tottenham, 1939

pedator (Gravenhorst, 1802) --S-DEA-ater (Gravenhorst, 1802) FKSNDEAImorsitans (Rossi, 1790) --SND--I

compressus (Marsham, 1802) nec (Geoffroy, 1785)winkleri (Bernhauer, 1906) --S-D---melanarius (Heer, 1839) F-SNDEAIglobulifer (Geoffroy, 1785) --S-DEA-

Creophilus Leach, 1819maxillosus (Linnaeus, 1758) FKSNDEAI

Quediina Kraatz, 1857Heterothops Stephens, 1829binotatus (Gravenhorst, 1802) F-SNDE--stiglundbergi Israelson, 1979 FKSND--Ipraevius Erichson, 1839 FKSNDE-Iniger Kraatz, 1868 F-S-D--Iquadripunctulus (Gravenhorst, 1806) FKSNDEAIdissimilis (Gravenhorst, 1802) FKS-DEAIminutus Wollaston, 1860 --S-D---

brunneipennis auct. nec Kiesenwetter, 1858

Euryporus Erichson, 1839picipes (Paykull, 1800) FKSNDEAI

Velleius Leach, 1819dilatatus (Fabricius, 1787) FKS-DEA-

Quedius Stephens, 1829longicornis Kraatz, 1857 FKS-DEAIfulgidus (Fabricius, 1793) FKSND-AI

assimilis (Nordmann, 1837)nigrocaeruleus Fauvel, 1874 -KS-D--Imesomelinus (Marsham, 1802) FKSNDEAImaurus (Sahlberg, 1830) FKSNDEAIcruentus (Olivier, 1795) FKSND-AIinvreai Gridelli, 1924 --SND---ochripennis (Ménétriés, 1832) -----EAIbrevicornis (Thomson, 1860) FKSNDEAIpuncticollis (Thomson, 1867) FKS-D---brevis Erichson, 1840 FKSNDEAIvexans Eppelsheim, 1881 F-S---A-microps Gravenhorst, 1847 FKSNDE-Ilundbergi Palm, 1972 F-S-----infuscatus Erichson, 1840 ----DE--truncicola Fairmaire & Laboulbène, 1856 --S-D--I

ventralis (Aragona, 1830) nec (Gravenhorst, 1802)tenellus (Gravenhorst, 1806) FKSN-E-Iscitus (Gravenhorst, 1806) FKSNDEAI

xanthopus Erichson, 1839 FKSNDEAIlateralis (Gravenhorst, 1802) --S-DE--cinctus (Paykull, 1790) FKSNDEAIplagiatus Mannerheim, 1843 FKSN-EAI

laevigatus (Gyllenhal, 1810) nec (Marsham, 1802)fuliginosus (Gravenhorst, 1802) FKSNDEAIcurtipennis Bernhauer, 1908 FKSNDEAIlevicollis (Brullé, 1832) --SNDEAI

tristis (Gravenhorst, 1802) nec (Fabricius, 1793)balticus Korge, 1960 --S-DE-Isubunicolor Korge, 1961 FKSN---Imolochinus (Gravenhorst, 1806) FKSNDEAI

picipennis (Paykull, 1800) nec (Fabricius, 1793)simplicifrons Fairmaire, 1861 ----D---

hispanicus Bernhauer, 1898auricomus Kiesenwetter, 1850 ----DE--scintillans (Gravenhorst, 1806) -S--DE-Iplancus Erichson, 1840 ------A-lucidulus Erichson, 1839 FKSNDEAIpicipes (Mannerheim, 1830) FKSNDEAIhumeralis Stephens, 1832 -----E-I

oblitteratus Erichson, 1840nemoralis Baudi, 1848 FKS-DE-Iumbrinus Erichson, 1839 FKSNDEAI

maritimus (J.Sahlberg, 1876)pseudoumbrinus Lohse, 1958

suturalis Kiesenwetter, 1845 --S-DE-Ihumeralis auct. nec (Stephens, 1832)

fumatus (Stephens, 1833) F-SNDE-Isublimbatus Mäklin, 1853 FKSN----

arcticus Munster, 1920nigriceps Kraatz, 1857 F-SNDEAImaurorufus (Gravenhorst, 1806) F-S-DE-Ipseudolimbatus Strand, 1938 --SN----limbatus (Heer, 1834) FKSN-EAI

limbatoides Coiffait, 1963fellmani (Zetterstedt, 1838) FKSN-E--nitipennis (Stephens, 1833) FKSNDEAI

attenuatus (Gyllenhal, 1808) nec (Gravenhorst, 1806)semiaeneus (Stephens, 1833) --S-D---semiobscurus (Marsham, 1802) --S-D---

rufipes (Gravenhorst, 1802) nec (Linnaeus, 1758)paradisianus (Heer, 1839) ------A-persimilis Mulsant & Rey, 1876 --SND---

aridulus Jansson, 1939fulvicollis (Stephens, 1833) FKSNDE-Iboopoides Munster, 1923 FKSNDE-Iboops (Gravenhorst, 1802) FKSNDEAI


Acylophorus Nordmann, 1837wagenschieberi Kiesenwetter, 1850 FKSND--Iglaberrimus (Herbst, 1784) --S-D-A-

Tanygnathinina Reitter, 1909Atanygnathus Jacobson, 1909terminalis (Erichson, 1839) F-S--E--

SCARABAEOIDEALucanidae Latreille, 1806Aesalinae MacLeay, 1819Aesalus Fabricius, 1801scarabaeoides (Panzer, 1794) --S-----

Syndesinae MacLeay, 1819Ceruchus MacLeay, 1819chrysomelinus (Hochenwarth, 1785) FKSN-EAI

Sinodendron Schneider, 1791cylindricum (Linnaeus, 1758) FKSNDEAI

Lucaninae Latreille, 1806Platycerus Geoffroy, 1762caprea (DeGeer, 1774) FKSN-EAIcaraboides (Linnaeus, 1758) FKSNDEAI

Dorcus MacLeay, 1819parallelepipedus (Linnaeus, 1758) --S-DEAI

Lucanus Scopoli, 1763cervus (Linnaeus, 1758) --S-D-AI

Trogidae MacLeay, 1819Trox Fabricius, 1775sabulosus (Linnaeus, 1758) FKSNDEAIhispidus (Pontoppidan, 1763) --S-D---scaber (Linnaeus, 1767) FKSNDEAI

Geotrupidae Latreille, 1806Bolboceratinae Mulsant, 1842Bolboceras Kirby, 1819Odonteus Samouelle, 1819

armiger (Scopoli, 1772) --S-DEAI

Geotrupinae Latreille, 1806Typhaeus Leach, 1815typhoeus (Linnaeus, 1758) --S-D---

Geotrupes Latreille, 1796mutator (Marsham, 1802) --S---AIspiniger (Marsham, 1802) --SNDEAIstercorarius (Linnaeus, 1758) FKSNDEAIstercorosus (Scriba, 1791) FKSNDEAIvernalis (Linnaeus, 1758) FKSNDEAI

Scarabaeidae Latreille, 1802Aegialiinae Lacordaire, 1856Aegialia Latreille, 1807arenaria (Fabricius, 1787) F-SNDEAIrufa (Fabricius, 1792) --SND-A-

spissipes LeConte, 1878rufina Silfverberg, 1977

sabuleti (Panzer, 1797) FKSNDEAI

Aphodiinae Leach, 1815Aphodiini Leach, 1815Aphodius Illiger, 1798erraticus (Linnaeus, 1758) FKSNDEAIsubterraneus (Linnaeus, 1758) FKSNDEAIfossor (Linnaeus, 1758) FKSNDEAIhaemorrhoidalis (Linnaeus, 1758) FKSNDEAIbrevis Erichson, 1848 F-SNDEA-putridus (Geoffroy, 1785) F-S-DEAI

arenarius (Olivier, 1789) nec (Fabricius, 1787)rhododactylus (Marsham, 1802)

rufipes (Linnaeus, 1758) FKSNDEAIdepressus (Kugelann, 1792) FKSNDEAIluridus (Fabricius, 1775) FKSNDEAIbimaculatus (Laxman, 1770) -----EA-zenkeri Germar, 1813 --S-D---pusillus (Herbst, 1789) FKSNDEAIcoenosus (Panzer, 1798) FKSNDEAI

tristis (Zenker, 1801)quadriguttatus (Herbst, 1783) --S-D-AIquadrimaculatus (Linnaeus, 1761) -K------obliteratus Panzer, 1823 ----D---contaminatus (Herbst, 1783) F-SND--Iequestris (Panzer, 1798) --SNDEAI

sticticus (Panzer, 1798) nec (Linnaeus, 1767)conspurcatus (Linnaeus, 1758) FKSNDEAIpaykulli Bedel, 1908 FKSNDEAI

tessulatus (Paykull, 1798) nec (Moll, 1782)pictus Sturm, 1805 --S-D-A-melanostictus Schmidt, 1840 ------AIdistinctus (Müller, 1776) FKSNDEAIpunctatosulcatus Sturm, 1805 FKSN-EA-

hirtipes Fischer v. Waldheim, 1844sabulicola Thomson, 1868

sphacelatus (Panzer, 1798) FKSNDEAIprodromus (Brahm, 1790) FKSNDEAIserotinus (Panzer, 1799) FK------tomentosus (Müller, 1776) FKS-DEA-obscurus (Fabricius, 1792) ------A-


porcus (Fabricius, 1792) --SNDE-Iscrofa (Fabricius, 1787) -KS-DEAImerdarius (Fabricius, 1775) FKSNDEAIfrater Mulsant & Rey, 1872 --s-----foetidus (Herbst, 1783) --S-DEAI

scybalarius auct. nec (Fabricius, 1781)fimetarius (Linnaeus, 1758) FKSNDEAIfoetens (Fabricius, 1787) FKSNDEAI

aestivalis Stephens, 1839uliginosus Hardy, 1847 FKSNDEAI

putridus (Herbst, 1789) nec (Geoffroy, 1785)fasciatus (Olivier, 1789) nec (Linnaeus, 1758)tenellus auct. nec Say, 1823

ater (DeGeer, 1774) FKSNDEAIborealis Gyllenhal, 1827 FKSNDEAInemoralis Erichson, 1848 F-SNDEAI

ardoei Landin, 1954lapponum Gyllenhal, 1806 FKSN----piceus Gyllenhal, 1808 FKSN-E-Isordidus (Fabricius, 1775) FKSNDEAIimmundus Creutzer, 1799 --S-DEAIrufus (Moll, 1782) FKSNDEAI

scybalarius (Fabricius, 1781)lugens Creutzer, 1799 -----EA-ictericus (Laicharting, 1781) FKSNDEAI

nitidulus (Fabricius, 1792)punctipennis Erichson, 1848 -------Iniger (Panzer, 1797) FKSN-EAIvarians Duftschmid, 1805 --S-DEAIplagiatus (Linnaeus, 1767) FKSNDEAIlividus (Olivier, 1789) FKS--E-Igranarius (Linnaeus, 1767) FKSNDEAI

Heptaulacus Mulsant, 1842sus (Herbst, 1783) -KS-DEAIvillosus (Gyllenhal, 1806) F-SNDEA-testudinarius (Fabricius, 1775) --S-DEAI

Oxyomus Dejean, 1833sylvestris (Scopoli, 1763) F-S-DEAI

Pleurophorus Mulsant, 1842caesus (Creutzer, 1796) --S---A-

Rhyssemus Mulsant, 1842germanus (Linnaeus, 1767) --S-D-AI

asper (Fabricius, 1775)

Scarabaeinae Latreille, 1802Copris Geoffroy, 1762lunaris (Linnaeus, 1758) --SNDEAI

Oniticellus Dejean, 1821fulvus (Goeze, 1777) -------I

Caccobius Thomson, 1859schreberi (Linnaeus, 1767) --S-DEAI

Onthophagus Latreille, 1802taurus (Schreber, 1759) -------Iillyricus (Scopoli, 1763) --S-----ovatus (Linnaeus, 1767) --S-DEAIjoannae Goljan, 1953 --SND--Inuchicornis (Linnaeus, 1758) FKSNDEAIvacca (Linnaeus, 1767) --S-D-AIgibbulus (Pallas, 1781) FKS--EAIfracticornis (Preyssler, 1790) FKSNDEAIsimilis (Scriba, 1790) FKS-D--Icoenobita (Herbst, 1787) --S-D-AI

Sericinae Hope, 1837Serica MacLeay, 1819brunnea (Linnaeus, 1758) FKSNDEAI

Maladera Mulsant, 1871holosericea (Scopoli, 1772) --S-D-AI

Omaloplia Schönherr, 1817nigromarginata (Herbst, 1785) ----DEAI

occidentalis Baraud, 1965ruricola auct. nec (Fabricius, 1775)

Psammodiini Mulsant, 1842Psammodius Fallén, 1807sulcicollis (Illiger, 1802) FKSNDEAI

asper auct. nec (Fabricius, 1775)

Diastictus Mulsant, 1842vulneratus (Sturm, 1805) --S-DEAI

Melolonthinae Samouelle, 1819Amphimallon Berthold, 1827solstitiale (Linnaeus, 1758) FKSNDEAIfallenii (Gyllenhal, 1817) --S-D---

?ochraceum (Knoch, 1801)Melolontha Fabricius, 1775hippocastani Fabricius, 1801 FKSNDEAImelolontha (Linnaeus, 1758) f-S-DEAI

Polyphylla Harris, 1842fullo (Linnaeus, 1758) --S-D-AI


Rutelinae MacLeay, 1819Anomala Leach, 1819dubia (Scopoli, 1763) FKSNDEAI

Anisoplia Dejean, 1821segetum (Herbst, 1783) -------Iagricola (Poda, 1761) -------I

Phyllopertha Stephens, 1830horticola (Linnaeus, 1758) FKSNDEAI

Hopliinae Latreille, 1829Hoplia Illiger, 1803farinosa (Linnaeus, 1761) --S-D---

philanthus (Herbst, 1783) nec (Fuessly, 1775)parvula Krynicki, 1832 -----EAIzaitzevi Jacobson, 1914 -----EA-praticola Duftschmid, 1805 ------A-graminicola (Fabricius, 1792) ----DEAI

Dynastinae MacLeay, 1819Oryctes Illiger, 1798nasicornis (Linnaeus, 1758) FKSNDEAI

Cetoniinae Leach, 1815Oxythyrea Mulsant, 1842funesta (Poda, 1761) F---dEAI

Tropinota Mulsant, 1842hirta (Poda, 1761) --s---AI

Cetonia Fabricius, 1775aurata (Linnaeus, 1758) FKSNDEAI

Liocola Thomson, 1859marmorata (Fabricius, 1792) FKSNDEAI

lugubris (Herbst, 1786) nec (Fabricius, 1775)

Potosia Mulsant & Rey, 1871aeruginosa (Drury, 1770) -------Ifieberi (Kraatz, 1880) -------Icuprea (Fabricius, 1775)ssp. metallica (Herbst, 1786) FKSNDEAI

Valginae Mulsant, 1842Valgus Scriba, 1790hemipterus (Linnaeus, 1758) ----D-AI

Trichiinae Fleming, 1821Osmoderma LePeletier & Audinet-Serville, 1828eremita (Scopoli, 1763) F-SNDEAI

Gnorimus LePeletier & Audinet-Serville, 1828nobilis (Linnaeus, 1758) --SNDEA-variabilis (Linnaeus, 1758) --S-DEAI

octopunctatus (Fabricius, 1775)

Trichius Fabricius, 1787fasciatus (Linnaeus, 1758) FKSNDEAIzonatus Germar, 1794 ----D---

SCIRTOIDEAEucinetidae Lacordaire, 1857Eucinetus Germar, 1818haemorrhoidalis (Germar, 1818) FKS-DEAI

haemorrhous (Duftschmid, 1825)

Eucilodes Vit, 1985caucasicus (Reitter, 1885) FK------

Clambidae Fischer v. Waldheim, 1821Calyptomerus Redtenbacher, 1849dubius (Marsham, 1802) --SND---

Clambus Fischer v. Waldheim, 1821pubescens Redtenbacher, 1849 FKSNDEAIpunctulum (Beck, 1817) FKSNDEAI

borealis Strand, 1946gibbulus (LeConte, 1850) F-SND-AI

radula Endrödy-Younga, 1960pallidulus Reitter, 1911 ----D---simsoni Blackburn, 1902 --S-D---armadillo (DeGeer, 1774) FKSNDEAInigrellus Reitter, 1914 F-SND---minutus (Sturm, 1807) -------I

Scirtidae Fleming, 1821Elodes Latreille, 1796minuta (Linnaeus, 1767) FKSNDEAI

pseudominuta Klausnitzer, 1971marginata (Fabricius, 1798) --SND---elongata Tournier, 1868 FKSNDE--

tricuspis Nyholm, 1985

Microcara Thomson, 1859testacea (Linnaeus, 1767) FKSNDEAI

bohemanni (Mannerheim, 1844)

Cyphon Paykull, 1799coarctatus Paykull, 1799 FKSNDEAIpalustris Thomson, 1855 FKSNDEAIruficeps Tournier, 1868 ------A-kongsbergensis Munster, 1924 FKSND-AIochraceus Stephens, 1830 FKSNDEAIvariabilis (Thunberg, 1787) FKSNDEAIlaevipennis Tournier, 1868 F-SNDEAI

phragmiteticola Nyholm, 1955pubescens (Fabricius, 1792) FKSNDEAIpunctipennis Sharp, 1873 FKSNDEAI


padi (Linnaeus, 1758) FKSNDEAIhilaris Nyholm, 1944 F-S-D-AI

Prionocyphon Redtenbacher, 1858serricornis (Müller, 1821) --SND-AI

Scirtes Illiger, 1807hemisphaericus (Linnaeus, 1758) FKSNDEAIorbicularis (Panzer, 1793) --S-D-AI

DASCILLOIDEADascillidae Guérin-Ménéville, 1843 (1836)Dascillus Latreille, 1796cervinus (Linnaeus, 1758) FKSNDEAI

BUPRESTOIDEABuprestidae Leach, 1815Buprestinae Leach, 1815Buprestini Leach, 1815Chalcophora Dejean, 1833mariana (Linnaeus, 1758) FKSNdEAI

Buprestis Linnaeus, 1758splendens Fabricius, 1775 -KS-d---rustica Linnaeus, 1758 FKSNdEAIhaemorrhoidalis Herbst, 1780 FKSNdEAInovemmaculata Linnaeus, 1767 FKS?DEAIoctoguttata Linnaeus, 1758 FKSN-EAI

Dicerca Eschscholtz, 1829aenea (Linnaeus, 1761) --SN-EAIalni (Fischer v. Waldheim, 1823) FKS--EAIfurcata (Thunberg, 1787) FKSN-EAI

acuminata (Pallas, 1782) nec (DeGeer, 1774)moesta (Fabricius, 1792) FKSNdEAI

Poecilonota Eschscholtz, 1829Descarpentriesina Leraut, 1983

variolosa (Paykull, 1799) FKSN-EAI

Ovalisia Kerremans, 1900Scintillatrix Obenberger, 1956Lampra Lacordaire, 1835, nec Hübner, 1821

rutilans (Fabricius, 1777) F-N----Idives (Guillebeau, 1889) ------A-

decipiens auct., nec (Gebler, 1847)

formaneki Jacobson, 1913 FKS-----

Anthaxia Eschscholtz, 1829manca (Linnaeus, 1767) -----EAIsalicis (Fabricius, 1777) -----EA-nitidula (Linnaeus, 1758) -----EAIsimilis Saunders, 1871 --SN-EAI

morio (Herbst, 1801) nec (Fabricius, 1792)helvetica Stierlin, 1868 -----EA-quadripunctata (Linnaeus, 1758) FKSNDEAIgodeti Gory, 1841 F-SN--A-

submontana Obenberger, 1930

Chrysobothrini Laporte de Castelnau & Gory, 1839Chrysobothris Eschscholtz, 1829chrysostigma (Linnaeus, 1758) FKSNdEAIaffinis (Fabricius, 1794) --SNDEAI

Agrilinae Laporte de Castelnau, 1835Agrilini Laporte de Castelnau, 1835Agrilus Curtis, 1825guerini Lacordaire, 1835 --S-----ater (Linnaeus, 1767) FK---EAIbiguttatus (Fabricius, 1777) --SN-EAI

pannonicus (Piller & Mitterpacher, 1783)subauratus (Gebler, 1833) FKS--EAIconvexicollis Redtenbacher, 1849 --S-D-A-cyanescens Ratzeburg, 1837 ----DEAI

coeruleus (Rossi, 1790) nec (Thunberg, 1789)laticornis (Illiger, 1803) F-SNDEAI

scaberrimus Ratzeburg, 1837angustulus (Illiger, 1803) --SNDEAIsulcicollis Lacordaire, 1835 FKSNDEAImendax Mannerheim, 1837 FKS--EAIsinuatus (Olivier, 1790) -----E--roberti Chevrolat, 1837 FKSN-EAI

pratensis (Ratzeburg, 1839)integerrimus (Ratzeburg, 1839) FK---EAIauricollis Kiesenwetter, 1857 ------A-delphinensis Abeille de Perrin, 1897 FK---EAI

pseudocyaneus auct. nec Kiesenwetter, 1857betuleti (Ratzeburg, 1837) FKSNDEAI

foveicollis Marseul, 1869viridis (Linnaeus, 1758) FKSNDEAIpopulneus Schaefer, 1946 F-SN--A-

suvorovi auct. nec Obenberger, 1935cuprescens Ménétriés, 1832 FKSN-EAI

aurichalceus Redtenbacher, 1849communis Obenberger, 1924

Anthaxiini Laporte de Castelnau & Gory, 1839Melanophila Eschscholtz, 1829Oxypteris Kirby, 1837

acuminata (DeGeer, 1774) FKSNDEAI

Phaenops Dejean, 1833cyanea (Fabricius, 1775) FKSNdEAI


paludicola Krogerus, 1922 FKSN-E?-olivicolor Kiesenwetter, 1857 -KSN----kaluganus Obenberger, 1940 ------AIhyperici (Creutzer, 1799) -----EAI

Coraebini Bedel, 1921Coraebus Laporte de Castelnau & Gory, 1839undatus (Fabricius, 1787) ------A-elatus (Fabricius, 1787) -----EAI

Aphanisticini Jacquelin du Val, 1859Aphanisticus Latreille, 1810emarginatus (Olivier, 1790) -------Ipusillus (Olivier, 1790) --S-DEAI

Trachyini Laporte de Castelnau, 1835Trachys Fabricius, 1801minutus (Linnaeus, 1758) FKSNDEAItroglodytes Gyllenhal, 1817 --S-DEAIscrobiculatus Kiesenwetter, 1857 ---ND--I

pumilus auct. nec Illiger, 1803fragariae Brisout de Barneville, 1874 -------I

Habroloma Thomson, 1864nana (Paykull, 1799) FKSNDE--

geranii (Silfverberg, 1977)

BYRRHOIDEAByrrhidae Latreille, 1804Byrrhinae Latreille, 1804Simplocaria Stephens, 1830semistriata (Fabricius, 1794) FKSNDEAIelongata J.Sahlberg, 1903 ---N----arctica Poppius, 1904 ---N----metallica (Sturm, 1807) FKSN--A-

Morychus Erichson, 1846aeneus (Fabricius, 1775) FKSNDEAI

Arctobyrrhus Münster, 1902dovrensis Münster, 1902 F-SN----

Lamprobyrrhulus Ganglbauer, 1902nitidus (Schaller, 1783) FK---EAI

Cytilus Erichson, 1846sericeus (Forster, 1771) FKSNDEAIauricomus (Duftschmid, 1825) FKSNDEAI

Byrrhus Linnaeus, 1767fasciatus (Forster, 1771) FKSNDEAIarietinus Steffahny, 1842 FKSND-AIpustulatus (Forster, 1771) FKSNDEAIpilula (Linnaeus, 1758) FKSNDEAI

Porcinolus Mulsant & Rey, 1869murinus (Fabricius, 1794) FKSND-AI

Syncalyptinae Mulsant & Rey, 1869Chaetophora Kirby & Spence, 1823Syncalypta Dillwyn, 1829

spinosa (Rossi, 1794) FKS-D---

Curimopsis Ganglbauer, 1902paleata (Erichson, 1846) FKSN-EAIsetigera (Illiger, 1798) FKS-DEAInigrita (Palm, 1934) -KS-D---cyclolepidia (Münster, 1902) FKSN----

DRYOPOIDEAElmidae Curtis, 1830Stenelmis Dufour, 1835canaliculata (Gyllenhal, 1808) F-SN----

Elmis Latreille, 1798maugetii Latreille, 1798 ------A-aenea (Müller, 1806) FKSNDEAIobscura (Müller, 1806) ------A-

Esolus Mulsant & Rey, 1872pygmaeus (Müller, 1806) -------Iangustatus (Müller, 1821) --S---A-

Oulimnius Des Gozis, 1886tuberculatus (Müller, 1806) FKSNDEAItroglodytes (Gyllenhal, 1827) --S-D--I

Limnius Illiger, 1802Latelmis Reitter, 1883

volckmari (Panzer, 1793) FKSNDEAImuelleri (Erichson, 1847) ------AIintermedius Fairmaire, 1881 ----D---opacus Müller, 1806 ------A-

Normandia Pic, 1900nitens (Müller, 1817) FKS--EAI

Riolus Mulsant & Rey, 1872cupreus (Müller, 1806) --S-DEAI

Macronychus Müller, 1806quadrituberculatus Müller, 1806 ------AI

Dryopidae Billberg, 1820Helichus Erichson, 1847substriatus (Müller, 1806) -------I

Dryops Olivier, 1791auriculatus (Geoffroy, 1785) FKSNDEAIanglicanus Edwards, 1909 FKS-DE--griseus (Erichson, 1847) FKSNDEA-


similaris Bollow, 1936 F-SNDE--ernesti Des Gozis, 1886 FKSNDEAIluridus (Erichson, 1847) --SNDEAInitidulus (Heer, 1841) --SNDE-Iviennensis (Laporte de Castelnau, 1840) -------Ilutulentus (Erichson, 1847) -----E--

Limnichidae Erichson, 1846Limnichus Dejean, 1821pygmaeus (Sturm, 1807) -KS-D-AIsericeus (Duftschmid, 1825) FK---EA-

Heteroceridae MacLeay, 1825Heterocerus Fabricius, 1792fossor Kiesenwetter, 1843 --S-----flexuosus Stephens, 1828 FKSNDE-Iobsoletus Curtis, 1828 FKSNDEAImarginatus (Fabricius, 1787) FK--DEAIfenestratus (Thunberg, 1784) F-SND-AIfusculus Kiesenwetter, 1843 FKSNDEAIhispidulus Kiesenwetter, 1843 FKSNDEAIintermedius Kiesenwetter, 1843 FKSND-AIsenescens Kiesenwetter, 1865 --S-----aureolus (Schiödte, 1866) ----D---

Psephenidae Lacordaire, 1857Eubria Germar, 1818palustris Germar, 1818 FK--D-AI

ELATEROIDEAEucnemidae Eschscholtz, 1829Melasinae Fleming, 1821Melasini Fleming, 1821Isorhipis Lacordaire, 1835marmottani Bonvouloir, 1871 --SN-E--

Melasis Olivier, 1790buprestoides (Linnaeus, 1761) F-SND---

Hylocharini Jacquelin du Val, 1859Hylochares Latreille, 1834cruentatus (Gyllenhal, 1808) FK---EA-

Calyptocerini Muona, 1993Otho Lacordaire, 1857sphondyloides (Germar, 1818) ------A-

Xylobiini Reitter, 1911Xylophilus Mannerheim, 1823Xylobius Latreille, 1834

corticalis (Paykull, 1800) F-SNDEA-

Epiphanini Muona, 1993Hylis Des Gozis, 1886Hypocaelus Guérin-Ménéville, 1843 , nec Dejean, 1833

procerulus (Mannerheim, 1823) FKSN-EAIcariniceps (Reitter, 1902) F-SN----foveicollis (Thomson, 1874) FKS-D-A-

fleischeri (Olexa, 1954)olexai (Palm, 1955) --S-D-A-

Dirhagini Reitter, 1911Microrhagini Fleutiaux, 1919

Clypeorhagus Olexa, 1975clypeatus (Hampe, 1850) ------A-

Microrhagus Dejean, 1833Dirhagus Latreille, 1834

lepidus Rosenhauer, 1847 FKSN--AIlindbergi (Palm, 1958)

pygmaeus (Fabricius, 1793) FKSNDEA-emyi (Rouget, 1855) --S-----

Dirrhagofarsus Fleutiaux, 1935Attenuorhagus Olexa, 1975

attenuatus (Mäklin, 1845) FK------

Rhacopus Hampe, 1855sahlbergi (Mannerheim, 1823) FKSN-EA-

Eucneminae Eschscholtz, 1829Eucnemis Ahrens, 1812capucina Ahrens, 1812 FKSND-AI

Macraulacinae Fleutiaux, 1922Macraulacini Fleutiaux, 1922

Dromaeolini Leiler, 1976Dromaeolus Kiesenwetter, 1858barnabita (Villa, 1838) --S-----

Throscidae Laporte de Castelnau, 1840Aulonothroscus Horn, 1890laticollis (Rybinski, 1897) FK------brevicollis (Bonvouloir, 1859) F-S---A-

Trixagus Kugelann, 1794Throscus Latreille, 1796

dermestoides (Linnaeus, 1767) F-SNDEAIcarinifrons (Bonvouloir, 1859) FKSNDEAIleseigneuri Muona, 2002 F-S-----elateroides (Heer, 1841) ------A-exul (Bonvouloir, 1859) F-S-D---

duvalii auct. nec (Bonvouloir, 1859)atticus Reitter, 1921 F-S-D---

caucasicus Reitter, 1921


Elateridae Leach, 1815Lissominae Laporte de Castelnau, 1840Drapetes Dejean, 1821mordelloides (Host, 1789) FKS-dEAI

cinctus (Panzer, 1796)biguttatus (Piller & Mitterpacher, 1783) nec (Fabricius, 1777)

Agrypninae Lacordaire, 1857Pyrophorinae Candeze, 1863

Agrypnini Lacordaire, 1857Agrypnus Eschscholtz, 1829murinus (Linnaeus, 1758) FKSNDEAI

Adelocerini Buysson, 1893Lacon Laporte de Castelnau, 1838Danosoma Thomson, 1859

punctatus (Herbst, 1779) --S-----lepidopterus (Panzer, 1801) F-S-D-A-querceus (Herbst, 1784) --S----Iconspersus (Gyllenhal, 1808) FKSN-EA-fasciatus (Linnaeus, 1758) FKSN-EAI

Denticollinae Stein & Weise, 1877 (1856)Athoinae Candeze, 1859

Denticollini Stein & Weise, 1877 (1856)Hemicrepidius Germar, 1839Pseudathous Méquignon, 1930

hirtus (Herbst, 1784) --SNDEAIniger (Linnaeus, 1758) FKSNDEAI

Crepidophorus Mulsant & Guillebeau, 1853mutilatus (Rosenhauer, 1847) F-SND-AI

Athous Eschscholtz, 1829vittatus (Fabricius, 1792) --SNDEAIhaemorrhoidalis (Fabricius, 1801) F-SNDEAIsubfuscus (Müller, 1764) FKSNDEAI

Limonius Eschscholtz, 1829Pheletes Kiesenwetter, 1858

aeneoniger (DeGeer, 1774) FKSNDE--

Limoniscus Reitter, 1905violaceus (Müller, 1821) ----DE--

Harminius Fairmaire, 1851Diacanthous Reitter, 1905

undulatus (DeGeer, 1774) FKSN-EAI

Stenagostus Thomson, 1859rufus (DeGeer, 1774) --SN-EAIrhombeus (Olivier, 1790) --S-D---

villosus auct. nec (Geoffroy, 1785)

Denticollis Piller & Mitterpacher, 1783rubens Piller & Mitterpacher, 1783 --SNDEA-linearis (Linnaeus, 1758) FKSNDEAIborealis (Paykull, 1800) FKSN-EA-

Cidnopus Thomson, 1859pilosus (Leske, 1785) --SNDEAIaeruginosus (Olivier, 1790) FKSNDEAIminutus (Linnaeus, 1758) F-S-DEAIparvulus (Panzer, 1799) -----EA-

Hypnoidus Dillwyn, 1829riparius (Fabricius, 1792) FKSNDEAIrivularius (Gyllenhal, 1808) FKSN-EA-

consobrinus (Mulsant & Guillebeau, 1855)

Ascoliocerus Mequignon, 1930Berninelsonius Leseigneur, 1972

hyperboreus (Gyllenhal, 1827) FKSN----

Prosternini Gistel, 1856Ctenicerini Fleutiaux, 1936 (1861)

Ctenicera Latreille, 1829Corymbites Latreille, 1834

virens (Schrank, 1781) -----E--pectinicornis (Linnaeus, 1758) FKSNDEAIcuprea (Fabricius, 1775) FKSN-E-I

Liotrichus Kiesenwetter, 1858affinis (Paykull, 1800) FKSN-EAI

Orithales Kiesenwetter, 1858serraticornis (Paykull, 1800) FKSN-EAI

Actenicerus Kiesenwetter, 1858sjaelandicus (Müller, 1764) FKSNDEAI

Prosternon Latreille, 1834tessellatum (Linnaeus, 1758) FKSNDEAI

Anostirus Thomson, 1859castaneus (Linnaeus, 1758) FKSNDEAIpurpureus (Poda, 1761) ----DEAI

Pseudanostirus Dolin, 1964globicollis (Germar, 1843) F-------

Aplotarsus Stephens, 1830incanus (Gyllenhal, 1827) FKSNDEA-

Selatosomus Stephens, 1830Mosotalesus Kishii, 1977

impressus (Fabricius, 1792) FKSNDEAInigricornis (Panzer, 1799) FKSNDEAIcruciatus (Linnaeus, 1758) FKSNDEAImelancholicus (Fabricius, 1798) FKSN----


aeneus (Linnaeus, 1758) FKSNDEAIgravidus (Germar, 1843) -----EAI

latus (Fabricius, 1801) nec (Fuessly, 1775)

Hypoganus Kiesenwetter, 1858inunctus (Lacordaire, 1835) --SND---

cinctus (Paykull, 1800) nec (Panzer, 1796)

Calambus Thomson, 1859bipustulatus (Linnaeus, 1767) --SND-AI

Eanus LeConte, 1861Paranomus Kiesenwetter, 1858, nec Chaudoir, 1842

costalis (Paykull, 1800) FKSN----

Negastriinae Nakane, 1953Negastrius Thomson, 1859pulchellus (Linnaeus, 1761) FKSNDEAIarenicola (Boheman, 1853) F-SND---sabulicola (Boheman, 1853) --S-D-AI

Zorochros Thomson, 1859minimus (Lacordaire, 1835) FKSNDEA-

dermestoides (Herbst, 1806) nec (Linnaeus, 1767)

Fleutiauxellus Méquignon, 1930maritimus (Curtis, 1840) F-SN----algidus (J.Sahlberg, 1883) FKSN----

Oedostethus LeConte, 1853tenuicornis (Germar, 1824) -K---EAIquadripustulatus (Fabricius, 1792) -KS-DEAI

Neohypdonus Stibick, 1971arcticus (Candeze, 1860) FKSN----

Elaterinae Leach, 1815Megapenthini Gurjeva, 1973Procraerus Reitter, 1905tibialis (Lacordaire, 1835) --SNDEAI

hjorti (Rye, 1905) --SND-A-triangularis (Dorn, 1925) --S-----

nemoralis Bouwer, 1980elongatulus (Fabricius, 1787) -----EAIelegantulus (Schönherr, 1817) --S-D-A-suecicus Palm, 1976 FKSN--A-

borealis (Palm, 1947) nec (Paykull, 1800)balteatus (Linnaeus, 1758) FKSNDEAIlepidus (Mäklin, 1876) F-s-----

sorsakoskii (Hellén, 1934)praeustus (Fabricius, 1792) FKSNDEAIcardinalis (Schiödte, 1865) --SND-A-tristis (Linnaeus, 1758) FKSN-EAIerythrogonus (Müller, 1821) FKS-DEAInigrinus (Herbst, 1784) FKSNDEAI

Brachygonus Buysson, 1912Reitterelater Platia & Cate, 1990

dubius Platia & Cate, 1990 --S-----megerlei auct. nec (Lacordaire, 1835)

Ischnodes Germar, 1844sanguinicollis (Panzer, 1793) --S-D---

Elaterini Leach, 1815Elater Linnaeus, 1758Ludius Berthold, 1827

ferrugineus Linnaeus, 1758 --S-D---

Sericus Eschscholtz, 1829brunneus (Linnaeus, 1758) FKSNDEAI

Melanotini Candeze, 1859 (1856)Cratonychini Gistel, 1856

Melanotus Eschscholtz, 1829villosus (Geoffroy, 1785) FKSNDEAI

erythropus (Gmelin, 1790)rufipes (Herbst, 1784) nec (Goeze, 1777)

castanipes (Paykull, 1800) FKSNDEAIpunctolineatus (Pelerin, 1829) --S-D-A-

niger (Fabricius, 1792) nec (Linnaeus, 1758)crassicollis (Erichson, 1841) -----EA-

Ampedini Gistel, 1856Ampedus Dejean, 1833Elater auct., nec Linnaeus, 1758

rufipennis (Stephens, 1830) --S-D---cinnabarinus (Eschscholtz, 1829) FKSNDEAIsanguineus (Linnaeus, 1758) FKSNDEAIpomonae (Stephens, 1830) FKSNDEAIquercicola (Buysson, 1887) ----D---sanguinolentus (Schrank, 1776) FKSNDEAInigerrimus (Lacordaire, 1835) --S-D---nigroflavus (Goeze, 1777) FKSNDEAIpomorum (Herbst, 1784) FKSNDEAI

ferrugatus (Lacordaire, 1835)

Agriotini Laporte de Castelnau, 1840Agriotes Eschscholtz, 1829gallicus Lacordaire, 1835 ------A-ustulatus (Schaller, 1783) F---DE-Ipilosellus (Schönherr, 1817) --S-D---

elongatus auct. nec (Marsham, 1802)acuminatus (Stephens, 1830) ----D--Isputator (Linnaeus, 1758) -KSNDEAIlineatus (Linnaeus, 1767) FKSNDEAI


obscurus (Linnaeus, 1758) FKSNDEAI

Ectinus Eschscholtz, 1829aterrimus (Linnaeus, 1761) FKSNDEAI

Dalopius Eschscholtz, 1829marginatus (Linnaeus, 1758) FKSNDEAI

Adrastus Eschscholtz, 1829limbatus (Fabricius, 1777) -------Ipallens (Fabricius, 1792) FKSNDEAI

nitidulus (Marsham, 1802) nec (Fabricius, 1787)rachifer (Geoffroy, 1785) -----E-Imontanus (Scopoli, 1763) -K------

Synaptus Eschscholtz, 1829filiformis (Fabricius, 1781) -----EAI

Cardiophorinae Candeze, 1859Cardiophorus Eschscholtz, 1829gramineus (Scopoli, 1763) --S---A-ruficollis (Linnaeus, 1758) FKSNDEAIrufipes (Goeze, 1777) --S---A-widenfalki Leiler, 1967 --S-----atramentarius Erichson, 1840 FKSN----ebeninus (Germar, 1824) -----EA-asellus Erichson, 1840 FKS-DEA-

Dicronychus Brullé, 1832cinereus (Herbst, 1784) --S-D-AIequiseti (Herbst, 1784) ---N-EAIequisetioides Lohse, 1976 -KS-D-A-

Paracardiophorus Schwarz, 1895musculus (Erichson, 1840) -------I

CANTHAROIDEADrilidae Blanchard, 1845Drilus Olivier, 1790concolor Ahrens, 1812 F-SNDE-Iflavescens (Geoffroy, 1785) --S-----

Lycidae Laporte de Castelnau, 1836Erotinae LeConte, 1881Dictyoptera Latreille, 1829aurora (Herbst, 1784) FKSNDEAI

Pyropterus Mulsant, 1838nigroruber (DeGeer, 1774) FKSNDEAI

affinis (Paykull, 1799)

Platycis Thomson, 1859Glabroplatycis Pic, 1914

minuta (Fabricius, 1787) FKSNDEAIcosnardi (Chevrolat, 1829) --S-D---

Lopheros LeConte, 1884rubens (Gyllenhal, 1817) FKSN-EAI

Metriorrhynchinae Kleine, 1926Xylobanellus Kleine, 1930Rossioptera Kasantsev, 1988

erythropterus (Baudi, 1871) ------A-

Calochrominae Lacordaire, 1857Lygistopterus Dejean, 1833sanguineus (Linnaeus, 1758) FKSNDEAI

Lampyridae Latreille, 1817Lampyris Geoffroy, 1762noctiluca (Linnaeus, 1758) FKSNDEAI

Phosphaenus Laporte de Castelnau, 1833hemipterus (Goeze, 1777) FKS-DEAI

Cantharidae Imhoff, 1856 (1815)Cantharinae Imhoff, 1856 (1815)Podabrus Westwood, 1838alpinus (Paykull, 1798) FKSNDEAIlapponicus (Gyllenhal, 1810) FKSN----obscuripes J.Sahlberg, 1871 FKSN----

Ancistronycha Märkel, 1852cyanipennis (Faldermann, 1835) FKSNDEAI

violacea (Paykull, 1798) nec (Thunberg, 1784)

Cantharis Linnaeus, 1758annularis Ménétriés, 1836 ----D---fusca Linnaeus, 1758 FKSNDEAIrustica Fallén, 1807 FKSNDEAIobscura Linnaeus, 1758 FKSNDEAIpulicaria Fabricius, 1781 F----EA-nigricans (Müller, 1776) FKSNDEAIpellucida Fabricius, 1792 FKSNDEAIlivida Linnaeus, 1758 FKSNDEAIfigurata Mannerheim, 1843 FKSNDEAIdecipiens Baudi, 1871 --SND-A-quadripunctata (Müller, 1776) FKSNDEAIrufa Linnaeus, 1758 FKSNDEAIpallida Goeze, 1777 --S-D-AIcryptica Ashe, 1947 --S-D---flavilabris Fallén, 1807 FKSNDEAI

fulvicollis Fabricius, 1792, nec Scopoli, 1763nigra (DeGeer, 1774) FKSNDEAI

thoracica (Olivier, 1790)bicolor Herbst, 1784 nec Linnaeus, 1763

paludosa Fallén, 1807 FKSNDEAIlateralis Linnaeus, 1758 FKS-DEAI


Metacantharis Bourgeois, 1886clypeata (Illiger, 1798) -----E-I

haemorrhoidalis (Fabricius, 1792) nec (Gmelin, 1790)

Armidia Mulsant, 1863signata (Germar, 1814) -K------

Rhagonycha Eschscholtz, 1830lutea (Müller, 1764) --SNDE--fulva (Scopoli, 1763) -KSNDEAItestacea (Linnaeus, 1758) FKSNDEAIlimbata Thomson, 1864 FKSNDEAIlignosa (Müller, 1764) FKSNDEAIelongata (Fallén, 1807) FKSNDEAIatra (Linnaeus, 1767) FKSNDEAIfugax Mannerheim, 1843 FK---EA-nigripes Redtenbacher, 1842 ------AI

Absidia Mulsant, 1863rufotestacea (Letzner, 1845) FKSNDEA-schoenherri (Dejean, 1837) FKSNDEAI

pilosa (Paykull, 1798) nec (Scopoli, 1763)

Silinae Mulsant, 1862Silis Charpentier, 1825Crudosilis Kasantsev, 1994

ruficollis (Fabricius, 1775) F-S-D-AInitidula (Fabricius, 1792) -----EAI

Malthininae Kiesenwetter, 1852Malthodinae Böving & Craighead, 1931

Malthinus Latreille, 1806biguttatus (Linnaeus, 1758) FKSNDEAI

biguttulus (Paykull, 1800)punctatus (Geoffroy, 1785) FKSNDEAI

flaveolus (Herbst, 1786)facialis Thomson, 1864 --SND--Iseriepunctatus Kiesenwetter, 1851 --S-----balteatus Suffrian, 1851 --SN--A-frontalis (Marsham, 1802) FKSNDEA-

guttifer Kiesenwetter, 1852 FKSNDEAImarginatus (Latreille, 1806) FKSNDEAImysticus Kiesenwetter, 1852 FKSNDEA-pumilus (Brébisson, 1835) FKSNDEAIspathifer Kiesenwetter, 1852 FKSNDEA-crassicornis (Mäklin, 1846) FKSND-A-dimidiatocollis (Rosenhauer, 1847) ------A-brevicollis (Paykull, 1798) FKSNDEAI

Malthodes Kiesenwetter, 1852debilis Kiesenwetter, 1852 ------A-flavoguttatus Kiesenwetter, 1852 FKSNDEAIdispar (Germar, 1824) --S-D---europaeus Wittmer, 1970 ------A-fibulatus Kiesenwetter, 1852 FKSNDEA-maurus (Laporte de Castelnau, 1840) FKSNDEA-misellus Kiesenwetter, 1852 F-------fuscus (Waltl, 1838) FKSNDEA-minimus (Linnaeus, 1758) F-SNDEAI

DERODONTOIDEADerodontidae LeConte, 1861Laricobiinae Mulsant & Rey, 1864Laricobius Rosenhauer, 1846erichsonii Rosenhauer, 1846 --S-D---

BOSTRICHOIDEANosodendridae Erichson, 1846Nosodendron Latreille, 1804fasciculare (Olivier, 1790) --S-D--I

Dermestidae Latreille, 1804Dermestinae Latreille, 1804Dermestes Linnaeus, 1758maculatus DeGeer, 1774 F-SNdEAI

vulpinus Fabricius, 1781frischii Kugelann, 1792 f-snDeAImurinus Linnaeus, 1758 FKSNDEAIlaniarius Illiger, 1801 F-SNDEAImustelinus Erichson, 1846 -----EA-gyllenhalii Laporte de Castelnau, 1840 FKS-DEAI

atomarius Erichson, 1846szekessyi Kalik, 1950 FKSND---erichsoni Ganglbauer, 1904 ------A-

tessellatus Erichson, 1846, nec Fabricius, 1775ater DeGeer, 1774 Fksndeai

cadaverinus Fabricius, 1775haemorrhoidalis Küster, 1852 f-snD---peruvianus Laporte de Castelnau, 1840f-s-----lardarius Linnaeus, 1758 FKSNDEAIpalmi Sjöberg, 1950 FKS-----

Thorictinae Wollaston, 1854Thaumaphrastini Anderson, 1949Thorictodes Reitter, 1875heydeni Reitter, 1875 --s-----

Trinodinae Casey, 1900Trinodes Dejean, 1821hirtus (Fabricius, 1781) --S-D-A-


Thylodriinae Semenov, 1912Thylodrias Motschulsky, 1839contractus Motschulsky, 1839 FKSND---

Attageninae Laporte de Castelnau, 1840Attagenus Latreille, 1802schaeferi (Herbst, 1792) F-----AIunicolor (Brahm, 1791) f-sn-eai

piceus (Olivier, 1790) nec (Thunberg, 1781)brunneus Faldermann, 1835 ------A-augustatus Ballion, 1871 --s-----smirnovi Zhantiev, 1973 FKSnD-a-pellio (Linnaeus, 1758) FKSNDEAIpunctatus (Scopoli, 1772) --S-----bifasciatus (Olivier, 1790) ----d---fasciatus (Thunberg, 1795) f-------woodroffei Halstead & Green, 1979 F-SND---

fasciatus auct. nec (Thunberg, 1795)gloriosae auct. nec (Fabricius, 1798)

trifasciatus (Fabricius, 1787) f-------

Megatominae Leach, 1815Megatomini Leach, 1815Trogoderma Dejean, 1821variabile Ballion, 1878 f-s----iangustum (Solier, 1849) f-snD-A-versicolor (Creutzer, 1799) f-s-d-aiglabrum (Herbst, 1783) FK--d-AI

nigrum (Herbst, 1797)granarium Everts, 1898 --snD---

Reesa Beal, 1967vespulae (Milliron, 1939) F-SND---

Globicornis Latreille, 1829emarginata (Gyllenhal, 1808) FKSNDEAI

marginata (Paykull, 1798) nec (Thunberg, 1781)corticalis (Eichhoff, 1863) --S-D---nigripes (Fabricius, 1792) --S-----

rufitarsis (Panzer, 1796)fasciata (Fairmaire, 1859) ----D---

Megatoma Herbst, 1792undata (Linnaeus, 1758) FKSNDEAIpubescens (Zetterstedt, 1828) F-SN--AI

Ctesias Stephens, 1830serra (Fabricius, 1792) F-SNDEAI

Anthrenini Gistel, 1856Anthrenus Geoffroy, 1762pimpinellae Fabricius, 1775 --S-DEAIscrophulariae (Linnaeus, 1758) F-SN-EAI

flavipes LeConte, 1854 f---D---verbasci (Linnaeus, 1767) f-snD-AImuseorum (Linnaeus, 1761) FKSNDEAIolgae Kalik, 1946 F-s---A-fuscus Olivier, 1789 --SnDEAIpolonicus Mroczkowski, 1951 -----EAI

Bostrichidae Latreille, 1802Bostrichinae Latreille, 1802Bostrichus Geoffroy, 1762capucinus (Linnaeus, 1758) f-SNd-A-

Dinoderinae Thomson, 1863Stephanopachys Waterhouse, 1888substriatus (Paykull, 1800) FKSN-EA-linearis (Kugelann, 1792) FKSN-EAI

Rhyzopertha Stephens, 1830dominica (Fabricius, 1793) fkSnD-A-

Dinoderus Stephens, 1830brevis Horn, 1878 fks-----minutus (Fabricius, 1775) f-snd---

Lyctinae Billberg, 1820Lyctus Fabricius, 1792brunneus (Stephens, 1830) fksnD---africanus Lesne, 1907 f-------linearis (Goeze, 1777) F-SND-AI

Minthea Pascoe, 1863rugicollis (Walker, 1858) f-s-d---

Trogoxylon LeConte, 1862impressum (Comolli, 1837) f-snd---

Anobiidae Fleming, 1821Eucradinae LeConte, 1861Hedobiini Mulsant & Rey, 1868Hedobia Dejean, 1821Ptinomorphus Mulsant & Rey, 1868

imperialis (Linnaeus, 1767) F-SNDEA-

Ptininae Latreille, 1802Gibbiini Jacquelin du Val, 1860Gibbium Scopoli, 1777psylloides (Czempinski, 1778) --s-de--

Mezium Curtis, 1828affine Boieldieu, 1856 f-s-----sulcatum (Fabricius, 1781) --s-d---

Sphaericini Bellés, 1982Trigonogenius Solier, 1849globulus Solier, 1849 fksnd---


Sphaericus Wollaston, 1854gibboides (Boieldieu, 1854) ----D---

Ptinini Latreille, 1802Niptus Boieldieu, 1856hololeucus (Faldermann, 1836) FKSNDEAI

Epauloecus Mulsant & Rey, 1868Tipnus Thomson, 1859, nec Boieldieu, 1856

unicolor (Piller & Mitterpacher, 1783) FKSNDEAI

Pseudeurostus Heyden, 1906hilleri (Reitter, 1877) f-S-d---

Ptinus Linnaeus, 1767tectus Boieldieu, 1856 F-SND---dubius Sturm, 1837 FKSNDEA-sexpunctatus Panzer, 1795 FKSND-A-rufipes Olivier, 1790 F-SNDEAIlichenum Marsham, 1802 --S-D---clavipes Panzer, 1792 f---Deai

testaceus Olivier, 1790, nec Thunberg, 1784pusillus Sturm, 1837 ----D---fur (Linnaeus, 1758) FKSNDEAIbicinctus Sturm, 1837 FKSNdEA-villiger Reitter, 1884 FKSND-AI

balticus Khnzorian & Karapetian, 1991subpillosus Sturm, 1837 FKSNDEAIpilosus Müller, 1821 ------AIraptor Sturm, 1837 FKSNDEAI

explanatus (Mannerheim, 1843) FKSN-E--gigas (Mulsant & Rey, 1863) -------Imollis (Linnaeus, 1758) FKSNDEAIangusticollis (Ratzeburg, 1847) F-SND-AI

tabidus Kiesenwetter, 1877abietinus (Gyllenhal, 1808) F-SNDEA-abietis (Fabricius, 1792) F-SNDEAIpini (Sturm, 1837) F-SNDEA-

Anobiinae Fleming, 1821Oligomerus Redtenbacher, 1849brunneus (Olivier, 1790) --S----I

Stegobium Motschulsky, 1860Sitodrepa Thomson, 1863

paniceum (Linnaeus, 1758) FKSNDEAI

Gastrallus Jacquelin du Val, 1860immarginatus (Müller, 1821) --SND-A-

Anobium Fabricius, 1775Hemicoelus LeConte, 1861

punctatum (DeGeer, 1774) FKSNDEAInitidum Fabricius, 1792 FKSNDEA-costatum Aragona, 1830 --S-D---fulvicorne Sturm, 1837 F-SND--Irufipes Fabricius, 1792 FKSN-EAIthomsoni (Kraatz, 1881) FKSN-EAI

Microbregma Seidlitz, 1889emarginata (Duftschmid, 1825) FKSN--AI

Hadrobregmus Thomson, 1859denticollis (Creutzer, 1796) ----D-AIpertinax (Linnaeus, 1758) FKSNDEAIconfusus (Kraatz, 1881) FKS---A-

Priobium Motschulsky, 1845carpini (Herbst, 1793) FKS-DEAI

Ptilininae Shuckard, 1840Ptilinus Geoffroy, 1762pectinicornis (Linnaeus, 1758) --SND-AIfuscus Geoffroy, 1785 FKSN-EA-

Pseudoptilinus Leiler, 1963fissicollis (Reitter, 1877) --S-----

Xyletininae Gistel, 1856Xyletinus Latreille, 1809laticollis (Duftschmid, 1825) --SNDEAIater (Creutzer, 1796) FKSNDEAIpectinatus (Fabricius, 1792) FKSN-EA-vaederoeensis Lundberg, 1969 --S-----

Dryophilinae LeConte, 1861Grynobius Thomson, 1859planus (Fabricius, 1787) --SND---

tricolor (Olivier, 1790)

Dryophilus Chevrolat, 1832pusillus (Gyllenhal, 1808) F-SNDEAI

Ernobiinae Pic, 1912Ochina Sturm, 1826ptinoides (Marsham, 1802) ----D--I

Xestobium Motschulsky, 1845plumbeum (Illiger, 1801) ----D---rufovillosum (DeGeer, 1774) F-SND-A-

Episernus Thomson, 1863angulicollis Thomson, 1863 FKSN----

striatulus Leinberg, 1904

Ernobius Thomson, 1859longicornis (Sturm, 1837) FKSNDEAInigrinus (Sturm, 1837) FKSNDEAIkiesenwetteri Schilsky, 1898 ------AI


fibyensis Lundblad, 1949 F-S---A-gronblomi Y.Kangas, 1955

tremulicola Y.Kangas, 1958 F-S-----longitarsis Jansson, 1942 F-SN--A-hanseni Jansson, 1947 FKSND---planicollis Lohse, 1957 F-S---A-

suecicus Lundberg, 1972

Lasioderma Stephens, 1835serricorne (Fabricius, 1792) f-snDeA-

Dorcatominae Thomson, 1859Stagetus Wollaston, 1861Theca Mulsant & Rey, 1860, nec Morris, 1845

borealis Israelson, 1971 FKSN--A-pilulus auct. nec (Aubé, 1861)

Dorcatoma Herbst, 1792flavicornis (Fabricius, 1793) F-SNDEAIambjoerni Baranowski, 1985 --S-----chrysomelina Sturm, 1837 F-SNDEAIjanssoni Büche & Lundberg, 2002 --S---A-substriata Hummel, 1829 F-S-DEA-

serra (Panzer, 1795) nec (Fabricius, 1792)minor Zahradnik, 1993 --S-----punctulata Mulsant & Rey, 1864 FKSN-EA-dresdensis Herbst, 1792 FKSNDEAIrobusta Strand, 1938 FKSNDEA-lomnickii Reitter, 1903 -----E--

Caenocara Thomson, 1859bovistae (Hoffmann, 1803) FKSND-AIaffinis (Sturm, 1837) ------AI

Anitys Thomson, 1863rubens (Hoffmann, 1803) --SND---

LYMEXYLOIDEALymexylidae Fleming, 1821Hylecoetinae Gistel, 1856Hylecoetus Latreille, 1806Elateroides Houlbert & Bétis, 1905

dermestoides (Linnaeus, 1761) FKSNDEAIflabellicornis (Schneider, 1791) FK---EAI

Lymexylinae Fleming, 1821Lymexylon Fabricius, 1775navale (Linnaeus, 1758) --SNDEAI

CLEROIDEAPhloiophilidae Kiesenwetter, 1863Phloiophilus Stephens, 1830edwardsii Stephens, 1830 --S-D---

Trogossitidae Latreille, 1802Peltinae Kirby, 1837Peltis Kugelann, 1792grossa (Linnaeus, 1758) FKSNdEAI

Ostoma Laicharting, 1781ferruginea (Linnaeus, 1758) FKSNDEAI

Thymalus Latreille, 1802limbatus (Fabricius, 1787) FKSNDEAIsubtilis Reitter, 1889 FKSN----

Lophocaterini Crowson, 1964Grynocharis Thomson, 1859oblonga (Linnaeus, 1758) FKSN-EAI

Lophocateres Olliff, 1883pusillus (Klug, 1832) ----d---

Calitinae Reitter, 1922Calitys Thomson, 1859scabra (Thunberg, 1784) FKSN-EAI

Trogossitinae Latreille, 1802Nemozoma Latreille, 1804elongatum (Linnaeus, 1761) F-SNDEAI

Temnoscheila Westwood, 1835caerulea (Olivier, 1790) --S-----

Tenebroides Piller & Mitterpacher, 1783Trogossita Olivier, 1790

mauritanicus (Linnaeus, 1758) FKSNDEAICleridae Latreille, 1802Tillinae Leach, 1815Tillus Olivier, 1790elongatus (Linnaeus, 1758) FKSNDEAI

Clerinae Latreille, 1802Opilo Latreille, 1802mollis (Linnaeus, 1758) -KSNDEA-domesticus (Sturm, 1837) f-SND---

Thanasimus Latreille, 1806formicarius (Linnaeus, 1758) FKSNDEAIfemoralis (Zetterstedt, 1828) FKSNDEAI

rufipes (Brahm, 1797) nec (DeGeer, 1775)

Trichodes Herbst, 1792apiarius (Linnaeus, 1758) ----dEAI

Enopliinae Gistel, 1856Dermestoides Houlbert & Bétis, 1905Aporthopleura Barr, 1977Dermestoides Schäffer, 1771 – not binominal

sanguinicollis (Fabricius, 1787) --S-----


Tarsosteninae Jacquelin du Val, 1860Tarsostenus Spinola, 1844univittatus (Rossi, 1792) fk------

Korynetinae Laporte de Castelnau, 1836Korynetes Herbst, 1792caeruleus (DeGeer, 1775) F-SND-AIruficornis Sturm, 1837 --S-----

obenbergeri Jansson, 1936

Necrobia Olivier, 1795ruficollis (Fabricius, 1775) FKSNDEAIviolacea (Linnaeus, 1758) FKSNDEAIrufipes (DeGeer, 1775) F-SNDEAI

Opetiopalpus Spinola, 1844scutellaris (Panzer, 1797) -----E-I

Melyridae Leach, 1815Rhadalinae LeConte, 1861Aplocnemus Stephens, 1830impressus (Marsham, 1802) F-SNDEA-

pini (Redtenbacher, 1849)nigricornis (Fabricius, 1793) FKSNDEAItarsalis (Sahlberg, 1822) FKSN-EAI

Trichoceble Thomson, 1859floralis (Olivier, 1790) --SN---Imemnonia (Kiesenwetter, 1861) --SND--I

Dasytinae Laporte de Castelnau, 1840Danacaeini Thomson, 1859Danacea Laporte de Castelnau, 1836pallipes (Panzer, 1793) --S-DE--nigritarsis (Küster, 1850) --S-----

Psilothrix Redtenbacher, 1858viridicoeruleus (Geoffroy, 1785) --S-D---

cyaneus auct. nec (Fabricius, 1775)

Dolichosoma Stephens, 1830lineare (Rossi, 1792) FKSNDEAI

Malachiinae Fleming, 1821Colotini Abeille de Perrin, 1890Hypebaeus Kiesenwetter, 1863flavipes (Fabricius, 1787) --SN----

Charopus Erichson, 1840graminicola (Dejean, 1833) FKSNDEAI

flavipes (Paykull, 1798) nec (Fabricius, 1787)

Attalini Abeille de Perrin, 1890Ebaeus Erichson, 1840lapplandicus Evers, 1993 F-SND-AI

pedicularius auct. nec (Linnaeus, 1758)

Nepachys Thomson, 1859cardiacae (Linnaeus, 1761) FKSN-EA-

Axinotarsus Motschulsky, 1854ruficollis (Olivier, 1790) --S-D---pulicarius (Fabricius, 1775) --SND-AImarginalis (Laporte de Castelnau, 1840) --S-D-AI

Dasytini Laporte de Castelnau, 1840Dasytes Paykull, 1799niger (Linnaeus, 1761) FKSNDEAIalpigradus Kiesenwetter, 1863 -----E--obscurus Gyllenhal, 1813 FKSN-EA-cyaneus (Fabricius, 1775) F-SNDEAI

caeruleus (DeGeer, 1774) nec (Linnaeus, 1758)nigrocyaneus Mulsant & Rey, 1868 --S-D---virens (Marsham, 1802) -----EA-

flavipes (Olivier, 1790) nec (Fabricius, 1781)aeratus Stephens, 1830 --SNDE--

aerosus Kiesenwetter, 1867plumbeus (Müller, 1776) FKSNDEAIsubaeneus Schönherr, 1817 ----d-A-fusculus (Illiger, 1801) FKSNDEAI

Malachiini Fleming, 1821Cordylepherus Evers, 1985viridis (Fabricius, 1787) FKSNDEAI

Malachius Fabricius, 1775scutellaris Erichson, 1840 -------Irubidus Erichson, 1840 -------Iaeneus (Linnaeus, 1758) FKSNDEAIbipustulatus (Linnaeus, 1758) FKSNDEAI

Clanoptilus Motschulsky, 1854marginellus (Olivier, 1790) --S-D-AIbarnevillei (Puton, 1865) --S-D---

Anthocomus Erichson, 1840rufus (Herbst, 1784) F-S-DEAI

coccineus (Schaller, 1783) nec (Linnaeus, 1761)fasciatus (Linnaeus, 1758) F-SNDEAIequestris (Fabricius, 1781) ---N-EAI

bipunctatus (Harrer, 1784)

Cerapheles Mulsant & Rey, 1867terminatus (Ménétriés, 1832) ----d---


Laiini Jacobson, 1913Paratinus Abeille de Perrin, 1891femoralis (Erichson, 1840) FKS-DEAI

CUCUJOIDEASphindidae Jacquelin du Val, 1861Sphindus Megerle, 1821dubius (Gyllenhal, 1808) FKSNDE-I

Aspidiphorus Ziegler, 1821orbiculatus (Gyllenhal, 1808) FKSNDEAI

Kateretidae Erichson, 1846Brachypteridae Erichson, 1845

Kateretes Herbst, 1793pedicularius (Linnaeus, 1758) FKSNDEAIpusillus (Thunberg, 1794) FKSN-EAI

bipustulatus (Paykull, 1798) nec (Thunberg, 1781)rufilabris (Latreille, 1807) --S-D-AI

Heterhelus Jacquelin du Val, 1858scutellaris (Heer, 1841) FKSNDEA-

Brachypterus Kugelann, 1794urticae (Fabricius, 1792) FKSNDEAI

erythropus (Marsham, 1802)fulvipes Erichson, 1843 --S--EAIglaber (Stephens, 1835) F-SNDEAI

Brachypterolus Grouvelle, 1913pulicarius (Linnaeus, 1758) FKSNDEAIlinariae (Stephens, 1830) FKSNDEA-

cornelii Spornraft, 1966antirrhini (Murray, 1864) -------I

villiger (Reitter, 1885)

Nitidulidae Latreille, 1802Carpophilinae Erichson, 1842Carpophilini Erichson, 1842Carpophilus Stephens, 1830ligneus Murray, 1864 f-snd---dimidiatus (Fabricius, 1792) f-snde--nepos Murray, 1864 ----d---

freemani Dobson, 1956marginellus Motschulsky, 1858 F-SND---obsoletus Erichson, 1843 ----d---hemipterus (Linnaeus, 1758) F-snD-aimutilatus Erichson, 1843 --s-d---

Epuraeini Kirejtshuk, 1986Epuraea Erichson, 1843melanocephala (Marsham, 1802) --SNDE--guttata (Olivier, 1811) F-SNDEAIfuscicollis (Stephens, 1832) --S-D---

hilleri Reitter, 1877 ----DEA-fageticola Audisio, 1991

concurrens Sjöberg, 1939 FK------castanea (Duftschmid, 1825) nec (Sahlberg, 1820)

neglecta (Heer, 1841) FKSNDEAIpallescens (Stephens, 1832) FKSNDEAI

florea Erichson, 1845abietina J.Sahlberg, 1889

laeviuscula (Gyllenhal, 1827) FKSNDE-Irufobrunnea Sjöberg, 1939 FK------deubeli Reitter, 1898 FKSNDE--thoracica Tournier, 1872 FKSNDE--angustula Sturm, 1844 FKSNDE-Ioblonga (Herbst, 1793) FKSNDEAI

danica Sjöberg, 1939longipennis Sjöberg, 1939 FKSN----

rugulosa auct. nec J.Sahlberg, 1889fussii Reitter, 1875 FKSN-E--

interjecta Sjöberg, 1939boreella (Zetterstedt, 1828) FKSNdEAIopalizans J.Sahlberg, 1889 FKSNDE--marseuli Reitter, 1872 FKSNDEAI

bickhardti Saint-Claire Deville, 1906acuta Biström, 1977pusilla (Illiger, 1798) nec (Thunberg, 1794)

pygmaea (Gyllenhal, 1808) FKSNDEAIlongula Erichson, 1845 FKS-D-A-binotata Reitter, 1872 FKSNDEA-placida Mäklin, 1853 FKSNDEA-terminalis (Mannerheim, 1843) FKSNDEAI

adumbrata Mannerheim, 1852tenenbaumi Sjöberg, 1939

longiclavis Sjöberg, 1939 FKSN----distincta (Grimmer, 1841) F-----AIexcisicollis Reitter, 1872 --S-D---biguttata (Thunberg, 1784) FKSNDEA-unicolor (Olivier, 1790) FKSNDEAI

x-rubrum J.Sahlberg, 1911variegata (Herbst, 1793) FKSNDEA-muehli Reitter, 1908 FKSNDEA-silacea (Herbst, 1784) FKSNDEA-

deleta Erichson, 1843aestiva (Linnaeus, 1758) FKSNDEAI

depressa (Illiger, 1798)melina Erichson, 1843 FKSNDEA-rufomarginata (Stephens, 1830) FKSNDEAIcontractula J.Sahlberg, 1889 FKSN----

helleni Sjöberg, 1939


silesiaca Reitter, 1872 FKS--E--limbata (Fabricius, 1787) FKSNDEA-

Meligethinae Thomson, 1859Pria Stephens, 1830Laria Scopoli, 1763 – suppr.

dulcamarae (Scopoli, 1763) --SND-AI

Meligethes Stephens, 1830solidus (Kugelann, 1794) --S---A-denticulatus (Heer, 1841) FKSN-EAI

hebes Erichson, 1845atratus (Olivier, 1790) FKSN-EA-flavimanus Stephens, 1830 FKSNDEA-

lumbaris Sturm, 1845subrugosus (Gyllenhal, 1808) FKSNDEA-corvinus Erichson, 1845 F-SNDEAIcaudatus Guillebeau, 1897 --SND---anthracinus Brisout de Barneville, 1863 -----E--subaeneus Sturm, 1845 F-SN-E--matronalis Audisio & Spornraft, 1990 --S-D---coracinus Sturm, 1845 FKSNDEAIcoeruleivirens Förster, 1849 FKSNDEA-aeneus (Fabricius, 1775) FKSNDEAIczwalinai Reitter, 1871 --S---A-viridescens (Fabricius, 1787) FKSNDEAIbidens Brisout de Barneville, 1863 FKSNDEAIsulcatus Brisout de Barneville, 1863 F---DEA-atramentarius Förster, 1849 --S-D-A-difficilis (Heer, 1841) F-S-DEAIkunzei Erichson, 1845 ------A-ochropus Sturm, 1845 F-SNDEAImorosus Erichson, 1845 FKSNDEAIbrunnicornis Sturm, 1845 FKSNDEA-haemorrhoidalis Förster, 1849 --S-D-AIbuyssoni Brisout de Barneville, 1882 ------A-persicus Faldermann, 1837 FKSNDEAI

pedicularius auct. nec (Gyllenhal, 1808)pedicularius (Gyllenhal, 1808) --S-DEAI

viduatus (Heer, 1841)serripes (Gyllenhal, 1827) FKS-----nanus Erichson, 1845 --S-----ruficornis (Marsham, 1802) F-S-D---

flavipes Sturm, 1845ovatus Sturm, 1845 FKSND-A-maurus Sturm, 1845 --SNDEA-incanus Sturm, 1845 -----E--umbrosus Sturm, 1845 FKSNDEA-distinctus Sturm, 1845 --S-----

obscurus auct. nec Erichson, 1845

hoffmanni Reitter, 1871 --S-----exilis Sturm, 1845 FKS-DEA-lugubris Sturm, 1845 FKS-DEA-gagathinus Erichson, 1845 F-S-DE--norvegicus Easton, 1959 ---N----egenus Erichson, 1845 FKS---AIcarinulatus Förster, 1849 FKSNDEA-

erythropus auct. nec (Marsham, 1802)bidentatus Brisout de Barneville, 1863 ----D---nigrescens Stephens, 1830 FKSNDEAIsymphyti (Heer, 1841) --S-DEA-planiusculus (Heer, 1841) --S-D---tristis Sturm, 1845 F-SNDEA-

Nitidulinae Latreille, 1802Omosita Erichson, 1843depressa (Linnaeus, 1758) FKSNDEAIdiscoidea (Fabricius, 1775) --SNDEAIcolon (Linnaeus, 1758) FKSNDEAI

Nitidula Fabricius, 1775bipunctata (Linnaeus, 1758) FKSNDEAIrufipes (Linnaeus, 1767) FKSNDEAIcarnaria (Schaller, 1783) --S-DEAI

Amphotis Erichson, 1843marginata (Fabricius, 1781) --S-DEAI

Soronia Erichson, 1843punctatissima (Illiger, 1794) FKSNDEAIgrisea (Linnaeus, 1758) FKSNDEAI

Ipidia Erichson, 1843binotata Reitter, 1875 FKSN-EAI

quadriplagiata Biström, 1978quadrimaculata (Quensel, 1790) nec (Scopoli, 1772)quadrinotata (Fabricius, 1798) nec (Scriba, 1793)

sexguttata (F.Sahlberg, 1834) FK-----I

Pocadius Erichson, 1843ferrugineus (Fabricius, 1775) FKSNDEAIadustus Reitter, 1888 --S-D---

lanuginosus Franz, 1969

Physoronia Reitter, 1884Pocadiodes Ganglbauer, 1899

wajdelota (Wankowicz, 1869) -------I

Thalycra Erichson, 1843fervida (Olivier, 1790) FKSNDEAI

Cyllodes Erichson, 1843ater (Herbst, 1792) FKS-DEAI


Cychramus Kugelann, 1794variegatus (Herbst, 1792) FKSNDEAI

quadripunctatus (Herbst, 1792)luteus (Fabricius, 1787) FKSNDEAI

Cryptarchinae Thomson, 1859Cryptarcha Shuckard, 1839strigata (Fabricius, 1787) FKSNDEAIundata (Olivier, 1790) F-SND-AI

Glischrochilus Reitter, 1873Librodor Reitter, 1874

quadriguttatus (Fabricius, 1777) F-SNDEAIgrandis (Tournier, 1872) ------A-hortensis (Geoffroy, 1785) FKSNDEAIquadrisignatus (Say, 1835) ----D---quadripunctatus (Linnaeus, 1758) FKSNDEAI

Pityophagus Shuckard, 1839ferrugineus (Linnaeus, 1761) FKSNDEAI

Cybocephalinae Jacquelin du Val, 1858Cybocephalus Erichson, 1844politus (Gyllenhal, 1813) FKSNDEA-

Monotomidae Laporte de Castelnau, 1840Rhizophaginae Redtenbacher, 1845Cyanostolus Ganglbauer, 1899aeneus (Richter, 1820) FKSN-EAI

Rhizophagus Herbst, 1793grandis Gyllenhal, 1827 FKSNDEAIdepressus (Fabricius, 1793) FKSNDEAIferrugineus (Paykull, 1800) FKSNDEAIparallelocollis Gyllenhal, 1827 FKSNDE-Iperforatus Erichson, 1845 F-SND-A-picipes (Olivier, 1790) FKSNDEAIdispar (Paykull, 1800) FKSNDEAIbipustulatus (Fabricius, 1793) FKSNDEAInitidulus (Fabricius, 1798) FKSNDEAIbrancsiki Reitter, 1905 --S-----parvulus (Paykull, 1800) FKSNDEAIcribratus Gyllenhal, 1827 FKSNDEAIpuncticollis Sahlberg, 1837 FK------

Monotominae Laporte de Castelnau, 1840Monotoma Herbst, 1793quadrifoveolata Aubé, 1837 --S-D---conicicollis Aubé, 1837 FKSNDEAIangusticollis (Gyllenhal, 1827) FKSNDEAIspinicollis Aubé, 1837 FKSNDE-Ipicipes Herbst, 1793 FKSNDEAI

brevipennis Kunze, 1839

brevicollis Aubé, 1837 F-SNDEA-bicolor Villa, 1835 FKSNDEA-quadricollis Aubé, 1837 F-S-D-gotzi Holzschuh & Lohse, 1981 ----D---testacea Motschulsky, 1845 F-SNDE--longicollis (Gyllenhal, 1827) FKSNDE--

Phloeostichidae Reitter, 1911Phloeostichus Redtenbacher, 1842denticollis Redtenbacher, 1842 ----D---

Silvanidae Kirby, 1837Brontinae Erichson, 1845

Uleiotinae Gistel, 1856Brontini Erichson, 1845Uleiota Latreille, 1796planata (Linnaeus, 1761) FKS-D-AI

Dendrophagus Schönherr, 1809crenatus (Paykull, 1799) FKSN-EAI

Cryptamorphini Casey, 1884Psammoecus Latreille, 1829desjardinsi Guérin-Ménéville, 1844 f-s-d---bipunctatus (Fabricius, 1792) F-SNDEAI

Silvaninae Kirby, 1837Airaphilus Redtenbacher, 1858elongatus (Gyllenhal, 1813) F-SN-EA-perangustus Lindberg, 1943 F-------

Nausibius Redtenbacher, 1858clavicornis (Kugelann, 1794) --snd---

Ahasverus Des Gozis, 1881advena (Waltl, 1834) FkSnDEA-

Oryzaephilus Ganglbauer, 1899surinamensis (Linnaeus, 1758) F-SNDEAImercator (Fauvel, 1889) F-SNDEAI

Silvanus Latreille, 1807bidentatus (Fabricius, 1792) FKSNDEAIunidentatus (Olivier, 1790) FKSNDEAI

Silvanoprus Reitter, 1911fagi (Guérin-Ménéville, 1844) FKSNDEAI

Cucujidae Latreille, 1802Cucujus Fabricius, 1775cinnaberinus (Scopoli, 1763) FKSN-EAIhaematodes Erichson, 1845 F----E-I

Pediacus Shuckard, 1839depressus (Herbst, 1797) FKSND--Idermestoides (Fabricius, 1793) ----D---fuscus Erichson, 1845 FKSN-EA-


Laemophloeidae Ganglbauer, 1899Laemophloeus Dejean, 1835monilis (Fabricius, 1787) --SNd---muticus (Fabricius, 1781) FKSNdEA-

Notolaemus Lefkovitch, 1959castaneus (Erichson, 1845) --S-----unifasciatus (Latreille, 1804) --S-d---

bimaculatus (Paykull, 1801) nec (Olivier, 1791)

Cryptolestes Ganglbauer, 1899duplicatus (Waltl, 1839) f-S-d---ferrugineus (Stephens, 1831) F-SnD-AIabietis (Wankowicz, 1865) FKSN-EAIweisei (Reitter, 1879) FK------alternans (Erichson, 1846) FKSND--Icorticinus (Erichson, 1846) FKSNd-AIspartii (Curtis, 1834) f-s--e--

ater (Olivier, 1795) nec (Geoffroy, 1785)capensis (Waltl, 1832) ----d---pusillus (Schönherr, 1817) f-snd---

minutus (Olivier, 1791) nec (Geoffroy, 1785)pusilloides (Steel & Howe, 1952) ----d---turcicus (Grouvelle, 1876) f-snD---

Placonotus MacLeay, 1871testaceus (Fabricius, 1787) ----D---

Phalacridae Leach, 1815Phalacrus Paykull, 1800coruscus (Panzer, 1797) FKSNDEAI

fimetarius auct. nec (Fabricius, 1775)dieckmanni Vogt, 1967 --S-D---

grossus auct. nec Erichson, 1845championi Guillebeau, 1892 --S--EA-

suecicus Palm, 1947substriatus Gyllenhal, 1813 FKSNDEAInigrinus (Marsham, 1802) FKSNDEAI

caricis Sturm, 1807fimetarius (Fabricius, 1775) --S-D-A-

brisouti Rye, 1872

Olibrus Erichson, 1845aeneus (Fabricius, 1792) FKSNDEAIbaudueri Flach, 1888 FKS-D---millefolii (Paykull, 1800) FKSNDEAIcorticalis (Panzer, 1797) --SND-AIpygmaeus (Sturm, 1807) --S-D-A-flavicornis (Sturm, 1807) FKSN-EA-norvegicus Munster, 1901 FKSNDEA-affinis (Sturm, 1807) --S-DEAI

liquidus Erichson, 1845 ------A-bicolor (Fabricius, 1792) FKS-DEAIbimaculatus Küster, 1848 FKSN-EA-

Stilbus Seidlitz, 1872testaceus (Panzer, 1797) F-SNDEAIatomarius (Linnaeus, 1767) FKS-DEA-oblongus (Erichson, 1845) FKSNDEAI

Cryptophagidae Kirby, 1837Hypocoprinae Reitter, 1880Hypocoprus Motschulsky, 1839latridioides Motschulsky, 1839 FKSNDE-I

quadricollis Reitter, 1877

Cryptophaginae Kirby, 1837Telmatophilini Jacquelin du Val, 1858Telmatophilus Heer, 1841caricis (Olivier, 1790) FKSNDEAIbrevicollis Aubé, 1862 ----D---typhae (Fallén, 1802) FKSNDEAIschoenherrii (Gyllenhal, 1808) FKSND-A-

Cryptophagini Kirby, 1837Paramecosoma Curtis, 1833melanocephalum (Herbst, 1793) FKSNDE-I

Henoticus Thomson, 1868serratus (Gyllenhal, 1808) FKSNDEAIcalifornicus (Mannerheim, 1843) f-s-d---

Pteryngium Reitter, 1887crenatum (Fabricius, 1798) FKSNDEAI

Micrambe Thomson, 1863bimaculata (Panzer, 1798) FKSNDEA-lindbergorum (Bruce, 1934) FK---E--villosa (Heer, 1841) -KSND---abietis (Paykull, 1798) FKSNDEAIlongitarsis (J.Sahlberg, 1900) FKSN----

Cryptophagus Herbst, 1792acutangulus Gyllenhal, 1827 FKSNDEAIparallelus Brisout de Barneville, 1863 FKSNDEA-

angustus Ganglbauer, 1899cylindrus Kiesenwetter, 1858 --S-DE--fallax Balfour-Browne, 1953 FKSNDE--

fumatus auct. nec Marsham, 1802quercinus Kraatz, 1852 F-SN-EAIbadius Sturm, 1845 FKSNDEAIpopuli Paykull, 1800 F-SNDEA-lysholmi Munster, 1932 F-SN----quadrihamatus Mäklin, 1853 FKSN----

plagiatus Poppius, 1900


confertus Casey, 1900 FKSN----archangelicus J.Sahlberg, 1926

lapponicus Gyllenhal, 1827 FKSN-E--subdepressus Gyllenhal, 1827 FKSNDEA-tuberculosus Mäklin, 1852 FKSN----

instabilis Bruce, 1936subfumatus Kraatz, 1856 FKSNDE-Ipubescens Sturm, 1845 F-SND--Imicaceus Rey, 1889 --SND---saginatus Sturm, 1845 FKSNDEA-fuscicornis Sturm, 1845 F-SN---Ilabilis Erichson, 1846 F-SNDEAIconfusus Bruce, 1934 FKSN-E--intermedius Bruce, 1934 --SNDE--dentatus (Herbst, 1793) FKSNDE-Ipseudodentatus Bruce, 1934 FKSNDEA-dorsalis Sahlberg, 1819 FKSNDEAIsubvittatus Reitter, 1887 FK------distinguendus Sturm, 1845 FKSNDEAIquadrimaculatus Reitter, 1877 F-------falcozi Roubal, 1927 ----DE--

westi Bruce, 1943corticinus Thomson, 1863 FKSND-A-scanicus (Linnaeus, 1758) FKSNDEAIpallidus Sturm, 1845 FKSNDE-Iimmixtus Rey, 1889 --S-D---

postpositus J.Sahlberg, 1903obsoletus Reitter, 1879 --S-----scutellatus Newman, 1834 FKSNDEAIlycoperdi (Scopoli, 1763) --SND-A-pilosus Gyllenhal, 1827 FKSNDEA-cellaris (Scopoli, 1763) FKSND-A-laticollis Lucas, 1846 F-SND-AI

affinis Sturm, 1845, nec F.Sahlberg, 1834setulosus Sturm, 1845 FKSNDE-Ischmidti Sturm, 1845 ------A-

Spavius Motschulsky, 1844Emphylus Erichson, 1846

glaber (Gyllenhal, 1808) FKSNDEAI

Antherophagus Dejean, 1821nigricornis (Fabricius, 1787) FKSNDEAIcanescens Grouvelle, 1916 --S-DE--pallens (Linnaeus, 1758) FKSNDEAI

Caenoscelini Falcoz, 1929Caenoscelis Thomson, 1863subdeplanata Brisout de Barneville, 1882 FKSNDE--ferruginea (Sahlberg, 1820) FKSNDE-I

sibirica Reitter, 1889 F-SNDE--fleischeri Reitter, 1889grandis Thomson, 1892

Atomariinae LeConte, 1861Atomariini LeConte, 1861Atomaria Stephens, 1830impressa Erichson, 1846 FKSN-E--munda Erichson, 1846 F-SNDE--nigripennis (Kugelann, 1794) F-SNDE-Imorio Kolenati, 1846 FKSND-A-ornata Heer, 1841 FKSND---

contaminata Erichson, 1846peltata Kraatz, 1853 FKSNDE--peltataeformis Sjöberg, 1947 FKSN-E--fuscipes (Gyllenhal, 1808) FKSNDEAIpusilla (Paykull, 1798) FKSNDEAIsodermani Sjöberg, 1947 F-SNDE--lundbergi Johnson, 1978 F-S-----clavigera Ganglbauer, 1899 FKSND---fuscata (Schönherr, 1808) FKSNDEAI

agnita Kangas, 1961zetterstedti (Zetterstedt, 1838) FKSNDEA-lewisi Reitter, 1877 F-SNDEA-pseudatra Reitter, 1887 FKS-DE--

reitteri Lövendal, 1892atra (Herbst, 1793) F-----A-gutta Newman, 1834 F-S-DEA-rhenonum Kraatz, 1853 F-SND---

rhenana Dohrn, 1853basalis Erichson, 1846 F-S-DEA-mesomela (Herbst, 1792) FKSNDEAIscutellaris Motschulsky, 1849 --S---A-rubella Heer, 1841 FKSND-A-

berolinensis Kraatz, 1853atricapilla Stephens, 1830 FKSNDEA-mongolica Johnson, 1970 F-------gravidula Erichson, 1846 -----E--nitidula (Marsham, 1802) FKSND-A-

borealis Sjöberg, 1947analis Erichson, 1846 --S-DEA-semitestacea Reitter, 1887 FKSN----rubida Reitter, 1875 F-S-----rubricollis Brisout de Barneville, 1863 FKS-DE--hislopi Wollaston, 1857 FKSN-E--attila Reitter, 1878 ----D---turgida Erichson, 1846 FKSNDEAIapicalis Erichson, 1846 FKSNDEA-


testacea Stephens, 1830 FKSNDEAIruficornis (Marsham, 1802) nec (Gmelin, 1790)

fimetarii (Fabricius, 1792) FKSND---umbrina (Gyllenhal, 1827) FKSNDE-I

sjobergi Palm, 1949diluta Erichson, 1846 F-SND--Ibarani Brisout de Barneville, 1863 F-S-DE--lapponica Johnson, 1978 FKS-----sparreschneideri Munster, 1927 FKSN----nigriventris Stephens, 1830 FKSNDE--punctithorax Reitter, 1887 --S-D---

consanguinea Johnson, 1976puncticollis Thomson, 1868 FKSND---nigrirostris Stephens, 1830 FKSNDEA-

fuscicollis Mannerheim, 1852linearis Stephens, 1830 --SNDEA-alpina Heer, 1841 FKSN----subangulata J.Sahlberg, 1926 FKSND---abietina Reitter, 1887 F-S-----

norica Ganglbauer, 1899bescidica Reitter, 1887 FKSN----grossepunctata Reitter, 1896 F-------

peltatula auct. nec Reitter, 1887badia Erichson, 1846 FKSN----

sahlbergi Sjöberg, 1947elongatula Erichson, 1846 F-S-----affinis (F.Sahlberg, 1834) FKSN---Ipseudaffinis Johnson & Strand, 1968 F-SN----bella Reitter, 1875 F-SNDE--lohsei Johnson & Strand, 1968 ----D---wollastoni Sharp, 1867 FKSND---pulchra Erichson, 1846 FKSN-EA-atrata Reitter, 1875 FKSN-EA-gracilicornis Reitter, 1887 F-------strandi Johnson, 1967 F-SND---procerula Erichson, 1846 FKSND---ihsseni Johnson, 1978 F-------

Ephistemini Leng, 1920Ootypus Ganglbauer, 1899globosus (Waltl, 1838) FKSNDEAI

Ephistemus Stephens, 1829globulus (Paykull, 1798) FKSNDEAI

Curelius Casey, 1900exiguus (Erichson, 1846) -----EA-

Erotylidae Latreille, 1802Xenoscelinae Ganglbauer, 1899Xenoscelini Ganglbauer, 1899Zavaljus Reitter, 1880Eicolyctus J.Sahlberg, 1919

brunneus (Gyllenhal, 1808) FKS---A-fausti Reitter, 1880

Cryptophilini Casey, 1900Cryptophilus Reitter, 1874integer (Heer, 1838) ----D---obliteratus Reitter, 1878 ----D---

Erotylinae Latreille, 1802Dacnini Gistel, 1856Dacne Latreille, 1796notata (Gmelin, 1790) -----E--rufifrons (Fabricius, 1775) --S-D---bipustulata (Thunberg, 1781) FKSNDEAI

Combocerus Bedel, 1867glaber (Schaller, 1783) FKSNDE-I

Tritomini Curtis, 1834Tritoma Fabricius, 1775bipustulata Fabricius, 1775 FKSNDEAIsubbasalis (Reitter, 1896) -----EAI

jakowlewi (Semenow, 1898)

Triplax Herbst, 1793aenea (Schaller, 1783) FKSNDEAIrussica (Linnaeus, 1758) FKSNDEAIscutellaris Charpentier, 1825 FKSNDEAIrufipes (Fabricius, 1781) FKSNDEAI

Byturidae Jacquelin du Val, 1858Byturus Latreille, 1796tomentosus (DeGeer, 1774) FKSNDEAIochraceus (Scriba, 1791) FKSNDEAI

aestivus auct. nec (Linnaeus, 1758)

Biphyllidae Sharp, 1900Biphyllus Dejean, 1821lunatus (Fabricius, 1792) F-SN?---

Diplocoelus Guérin-Ménéville, 1844fagi Guérin-Ménéville, 1844 --SND---

Bothrideridae Erichson, 1845Teredinae Seidlitz, 1888Teredus Dejean, 1835cylindricus (Olivier, 1790) --S-D---

Oxylaemus Erichson, 1845variolosus (Dufour, 1843) --S-----


Anommatinae Ganglbauer, 1899Anommatus Wesmael, 1835duodecimstriatus (Müller, 1821) --S-D---diecki Reitter, 1875 --S-D---

Bothriderinae Erichson, 1845Bothrideres Dejean, 1835contractus (Geoffroy, 1785) FKSNd-AI

bipunctatus (Gmelin, 1790)

Cerylonidae Billberg, 1820Cerylon Latreille, 1802fagi Brisout de Barneville, 1867 F-SNDEAIhisteroides (Fabricius, 1793) FKSNDEAIferrugineum Stephens, 1830 FKSNDEAIimpressum Erichson, 1845 FKSN-EAIdeplanatum Gyllenhal, 1827 FKSNDEAI

Alexiidae Imhoff, 1856Sphaerosomatidae Ganglbauer, 1899

Sphaerosoma Stephens, 1832pilosum (Panzer, 1793) F-SNDE--

Endomychidae Leach, 1815Holoparamecinae Seidlitz, 1888Holoparamecus Curtis, 1833caularum (Aubé, 1843) --S-D---

Leiestinae Thomson, 1863Leiestes Chevrolat, 1837seminigra (Gyllenhal, 1808) FKSN-EAI

Endomychinae Leach, 1815Endomychus Panzer, 1795coccineus (Linnaeus, 1758) FKSNDEAI

Lycoperdininae Redtenbacher, 1844Lycoperdina Latreille, 1807bovistae (Fabricius, 1793) --S-D---succincta (Linnaeus, 1767) F-SNDEAI

Mycetina Mulsant, 1846cruciata (Schaller, 1783) F-SNDEAI

Mycetaeinae Jacquelin du Val, 1858Mycetaea Stephens, 1830subterranea (Fabricius, 1801) FKSNDEA-

hirta (Marsham, 1802) nec (Schaeffer, 1769)

Anamorphinae Strohecker, 1953Symbiotes Redtenbacher, 1849latus Redtenbacher, 1849 ----D---gibberosus (Lucas, 1849) ----D---

Coccinellidae Latreille, 1807Coccidulinae Mulsant, 1846Coccidula Kugelann, 1798scutellata (Herbst, 1783) F-SNDEAIrufa (Herbst, 1783) FKSNDEAI

Rhyzobius Stephens, 1832litura (Fabricius, 1787) --S-D---chrysomeloides (Herbst, 1792) ----D---

Scymninae Mulsant, 1846Stethorini Dobzhansky, 1924Stethorus Weise, 1885punctillum (Weise, 1891) FKSNDEAI

Scymnini Mulsant, 1846Cryptolaemus Mulsant, 1850montrouzieri Mulsant, 1853 --s-----

Scymnus Kugelann, 1794Pullus Mulsant, 1846

nigrinus Kugelann, 1794 FKSNDEAIpallipediformis Günther, 1958ssp. apetzoides Capra & Fürsch, 1967 FKSN--A-

apetzii auct., nec Mulsant, 1846frontalis (Fabricius, 1787) FKSNDEAImimulus Capra & Fürsch, 1967 FKSND--I

rufipes auct. nec (Fabricius, 1798)jakowlewi Weise, 1892 F-SN----

triangularis J.Sahlberg, 1914rubromaculatus (Goeze, 1777) F-S-DEAIfemoralis (Gyllenhal, 1827) FKSND-A-abietis (Paykull, 1798) FKSNDEAIferrugatus (Moll, 1785) FKS--EAIauritus Thunberg, 1795 FKSNDEAIimpexus Mulsant, 1850 --s-----suturalis Thunberg, 1795 FKSNDEAIater Kugelann, 1794 --SN-EA-haemorrhoidalis Herbst, 1797 FKSNDEA-limbatus Stephens, 1832 FKSNDEA-silesiacus Weise, 1902 --S-----

stiglundbergi Fürsch, 1969fennicus J.Sahlberg, 1886 FKSN----

Nephus Mulsant, 1846redtenbacheri (Mulsant, 1846) FKSNDEAIlimonii (Donisthorpe, 1903) F-SND---rutaneni Fürsch, 1986 F-------quadrimaculatus (Herbst, 1783) --S---AIbipunctatus (Kugelann, 1794) FKSNDEAIbisignatus (Boheman, 1850) FKSND---


Hyperaspini Mulsant, 1846Hyperaspis Chevrolat, 1837inexpectata Günther, 1959 FK------pseudopustulata Mulsant, 1853 FKSNDEAI

reppensis auct. nec (Herbst, 1783)campestris (Herbst, 1783) -----EAIconcolor Suffrian, 1843 --S-----

Chilocorinae Mulsant, 1846Platynaspidini Mulsant, 1846Platynaspis Redtenbacher, 1843luteorubra (Goeze, 1777) FKSNDEAI

Chilocorini Mulsant, 1846Chilocorus Leach, 1815bipustulatus (Linnaeus, 1758) FKSNDEAIrenipustulatus (Scriba, 1790) FKSNDEAI

Exochomus Redtenbacher, 1843quadripustulatus (Linnaeus, 1758) FKSNDEAInigromaculatus (Goeze, 1777) FKS-DEAI

Coccinellinae Latreille, 1807Coccinellini Latreille, 1807Coccinula Dobzhansky, 1925quatuordecimpustulata (Linnaeus, 1758) FKSNDEAI

Anisosticta Chevrolat, 1837novemdecimpunctata (Linnaeus, 1758) FKSNDEAIstrigata (Thunberg, 1795) FKSN----

Sospita Mulsant, 1846vigintiguttata (Linnaeus, 1758) FKSNDEAI

Myzia Mulsant, 1846Neomysia Casey, 1899

oblongoguttata (Linnaeus, 1758) FKSNDEAI

Myrrha Mulsant, 1846octodecimguttata (Linnaeus, 1758) FKSNDEAI

Propylea Mulsant, 1846quatuordecimpunctata (Linnaeus, 1758) FKSNDEAI

Calvia Mulsant, 1846decemguttata (Linnaeus, 1767) F-S--EAIquatuordecimguttata (Linnaeus, 1758) FKSNDEAIquindecimguttata (Fabricius, 1777) FK---EAI

Hippodamia Chevrolat, 1837Adonia Mulsant, 1846Semiadalia Crotch, 1874

tredecimpunctata (Linnaeus, 1758) FKSNDEAIseptemmaculata (DeGeer, 1775) FKSNDEAIarctica (Schneider, 1792) FKSN----variegata (Goeze, 1777) FKSNDEAInotata (Laicharting, 1781) FK-N-EAIundecimnotata (Schneider, 1792) ------A-

Coccinella Linnaeus, 1758undecimpunctata Linnaeus, 1758 FKSNDEAIsaucerottii Mulsant, 1850ssp. lutshniki Dobzhansky, 1917 ------A-trifasciata Linnaeus, 1758 FKSN-EA-septempunctata Linnaeus, 1758 FKSNDEAImagnifica Redtenbacher, 1843 FKSNDEAI

divaricata auct. nec Olivier, 1808distincta Faldermann, 1837, nec Herbst, 1793

hieroglyphica Linnaeus, 1758 FKSNDEAIquinquepunctata Linnaeus, 1758 FKSNDEAI

Oenopia Mulsant, 1850Synharmonia Ganglbauer, 1899

conglobata Linnaeus, 1758 FKS-?EAI

Adalia Mulsant, 1850conglomerata (Linnaeus, 1758) FKSN-EAIbipunctata (Linnaeus, 1758)ssp. bipunctata (Linnaeus, 1758) FKSNDEAIssp. frigida (Schneider, 1792) FKSN----decempunctata (Linnaeus, 1758) FKSNDEAI

Harmonia Mulsant, 1846quadripunctata (Pontoppidan, 1763) F-S-DEAIoctomaculata (Fabricius, 1781) --S-----

arcuata (Fabricius, 1787)axyridis (Pallas, 1773) F-------

Anatis Mulsant, 1846ocellata (Linnaeus, 1758) FKSNDEAI

Aphidecta Weise, 1893obliterata (Linnaeus, 1758) FKSNDEAI

Psylloborini Casey, 1899Vibidia Mulsant, 1846duodecimguttata (Poda, 1761) --S-DEAI

Halyzia Mulsant, 1846sedecimguttata (Linnaeus, 1758) FKSNDEAI

Psyllobora Chevrolat, 1837Thea Mulsant, 1846

vigintiduopunctata (Linnaeus, 1758) FKSNDEAI

Tytthaspidini Crotch, 1874Tytthaspis Crotch, 1874sedecimpunctata (Linnaeus, 1761) --SNDEAI


Epilachninae Mulsant, 1846Epilachnini Mulsant, 1846Subcoccinella Agassiz, 1846vigintiquatuorpunctata (Linnaeus, 1758) FKSNDEAI

Cynegetini Thomson, 1866Madaini Gordon, 1975

Cynegetis Chevrolat, 1837impunctata (Linnaeus, 1767) F-SND-A-

Corylophidae LeConte, 1852Corylophinae LeConte, 1852Corylophus Stephens, 1835cassidoides (Marsham, 1802) F-S-DEAI

Rypobiinae Paulian, 1950Rypobius LeConte, 1852praetermissus Bowestead, 1999 --S-----

ruficollis auct. nec (Jacquelin du Val, 1854)

Sericoderinae Matthews, 1888Sericoderus Stephens, 1829lateralis (Gyllenhal, 1827) F-SNDEAI

Parmulinae Poey, 1854Clypastraea Haldeman, 1842Sacium LeConte, 1852

pusilla (Gyllenhal, 1810) FKS---AI

Arthrolips Wollaston, 1854obscura (Sahlberg, 1833) FKS----I

Orthoperinae Jacquelin du Val, 1859Orthoperus Stephens, 1829brunnipes (Gyllenhal, 1808) FKSNDEA-nigrescens Stephens, 1829 F-S-D-A-

mundus Matthews, 1885coriaceus auct. nec Mulsant & Rey, 1861

punctatus Wankowicz, 1865 FKSN--AIrogeri Kraatz, 1874 F-SND-AI

punctulatus Reitter, 1876atomus (Gyllenhal, 1808) FKSNDEA-corticalis (Redtenbacher, 1849) FKSND-A-

improvisus Bruce, 1946

Latridiidae Erichson, 1842Latridiinae Erichson, 1842Latridiini Erichson, 1842Latridius Herbst, 1793hirtus Gyllenhal, 1827 FKSNDEAIconsimilis Mannerheim, 1844 FKSND---anthracinus Mannerheim, 1844 F-SND-A-minutus (Linnaeus, 1767) FKSNDEAIpseudominutus (Strand, 1958) FKSND-A-

nidicola (Palm, 1944) F-SND-A-brevicollis (Thomson, 1868) F-SND-A-

Enicmus Thomson, 1859brevicornis (Mannerheim, 1844) --SN--A-lundbladi Palm, 1956 F-SN----fungicola Thomson, 1868 FKSNDEA-planipennis Strand, 1940 FKSN--A-apicalis J.Sahlberg, 1926 FKSN----rugosus (Herbst, 1793) FKSNDEAItestaceus (Stephens, 1830) --SND-A-atriceps Hansen, 1962 --S-D---transversus (Olivier, 1790) FKSNDEAIhistrio Joy & Tomlin, 1910 FKSNDEA-

Dienerella Reitter, 1911Cartodere Thomson, 1863, nec Thomson, 1859

elongata (Curtis, 1830) F-SNDEA-clathrata (Mannerheim, 1844) F-SND---

separanda auct. nec (Reitter, 1887)ruficollis (Marsham, 1802) F-SND--Ifiliformis (Gyllenhal, 1827) F-SNDEA-argus (Reitter, 1884) F-----a-filum (Aubé, 1850) FKSNDEA-costulata (Reitter, 1877) ----d-A-elegans (Aubé, 1850) --S-----

Adistemia Fall, 1899watsoni (Wollaston, 1871) f-snD---

Stephostethus LeConte, 1878Latridius auct., nec Herbst, 1793

lardarius (DeGeer, 1775) FKSNDEAIattenuatus (Mannerheim, 1844) F-------angusticollis (Gyllenhal, 1827) FKS?DEAI

kokujewi (Semenow, 1898)pandellei (Brisout de Barneville, 1863) FKSN-EAIcinnamopterus (Mannerheim, 1853) FKSN----

sahlbergi (Reitter, 1894)variolosus (Mannerheim, 1844) FKSN----alternans (Mannerheim, 1844) F-SND---rugicollis (Olivier, 1790) FKSNDE--

Thes Semenov-Tian-Shansky, 1910bergrothi (Reitter, 1880) FKSNDEAI

Aridius Motschulsky, 1866norvegicus (Strand, 1940) ---n----bifasciatus (Reitter, 1877) --S-D---nodifer (Westwood, 1839) F-SNDEA-

Cartodere Thomson, 1859constricta (Gyllenhal, 1827) FKSNDEAI


Corticariini Curtis, 1829Corticaria Marsham, 1802pubescens (Gyllenhal, 1827) FKSNDEAIpunctulata Marsham, 1802 ----D---dentiventris Poppius, 1904 F-SN----pineti Lohse, 1960 F-SND---crenulata (Gyllenhal, 1827) FKSNDEA-fulva (Comolli, 1837) FKSNDEAIumbilicata (Beck, 1817) FKSNDEA-longicornis (Herbst, 1793) FKS--EA-impressa (Olivier, 1790) FKSNDEAIlapponica (Zetterstedt, 1838) FKSNDE--saginata Mannerheim, 1844 FKSND---serrata (Paykull, 1798) FKSNDEAIplanula Fall, 1899 F-S-----

strandi Palm, 1949, nec Roubal, 1934orbicollis Mannerheim, 1853 FKSN----

munsteri Strand, 1937abietorum Motschulsky, 1867 FKSNDEA-interstitialis Mannerheim, 1844 FKSN----foveola (Beck, 1817) FKSND--Irubripes Mannerheim, 1844 FKSNDEA-

linearis (Paykull, 1798) nec (Thunberg, 1784)polypori J.Sahlberg, 1900 FKSN--A-

eppelsheimi Reitter, 1886, nec Reitter, 1875alleni Johnson, 1974 F-S-----crenicollis Mannerheim, 1844 FKSN----fennica Johnson, 1974 F-S-----lateritia Mannerheim, 1844 FKSN--A-longicollis (Zetterstedt, 1838) FKSNDEAIinconspicua Wollaston, 1860 F-S-D---elongata (Gyllenhal, 1827) FKSNDEAIobsoleta Strand, 1940 FKSN----fagi Wollaston, 1854 F-SND---ferruginea Marsham, 1802 FKSNDEAI

Cortinicara Johnson, 1975gibbosa (Herbst, 1793) FKSNDEAI

Corticarina Reitter, 1880similata (Gyllenhal, 1827) FKSNDEAIlambiana (Sharp, 1910) F-SND-A-obfuscata Strand, 1937 F-SN--A-truncatella (Mannerheim, 1844) F-SNDEAIfuscula (Gyllenhal, 1827) FKSNDEAIlatipennis (J.Sahlberg, 1871) FKSN-EA-

Melanophthalma Motschulsky, 1866curticollis (Mannerheim, 1844) FKSNDEAI

transversalis auct. nec (Gyllenhal, 1827)

suturalis (Mannerheim, 1844) F-S-D---distinguenda (Comolli, 1837) --S-D--I

Migneauxia Jacquelin du Val, 1859orientalis Reitter, 1877 --S-D---

TENEBRIONOIDEAMycetophagidae Leach, 1815Triphyllus Dejean, 1821bicolor (Fabricius, 1777) --S-D-A-

Litargus Erichson, 1846connexus (Geoffroy, 1785) FKSNDEAI

Mycetophagus Hellwig, 1792quadripustulatus (Linnaeus, 1761) FKSNDEAIater (Reitter, 1879) ------AIpiceus (Fabricius, 1777) FKSNDEAIsalicis Brisout de Barneville, 1862 ---N--A-decempunctatus Fabricius, 1801 FKSN-EAIatomarius (Fabricius, 1787) FKSNDE-Iquadriguttatus Müller, 1821 FKS-D-AImultipunctatus Fabricius, 1792 FKSNDEA-fulvicollis Fabricius, 1792 FKSNDEAIpopuli Fabricius, 1798 FKSNDEAI

Typhaea Stephens, 1829stercorea (Linnaeus, 1758) FKSNDEAIdecipiens Lohse, 1989 F-S-D---

Ciidae Leach, 1819Cis Latreille, 1796lineatocribratus Mellié, 1848 FKSND-A-alter Silfverberg, 1991 FKSNDEAI

nitidus auct. nec (Fabricius, 1792)jacquemartii Mellié, 1848 FKSNDEAIhanseni Strand, 1965 FKS-D-A-glabratus Mellié, 1848 FKSNDEA-comptus Gyllenhal, 1827 FKSNDEAIstriatulus Mellié, 1848 -K------hispidus (Paykull, 1798) FKSNDEAIsetiger Mellié, 1848 -SND--AI

petropolitanus Jacobson, 1895micans (Fabricius, 1792) FKSNDEAIboleti (Scopoli, 1763) FKSNDEAIrugulosus Mellié, 1848 FKS-D-A-quadridens Mellié, 1848 FKSN--AIpunctulatus Gyllenhal, 1827 FKSNDEA-quadridentulus Perris, 1874 F-------fagi Waltl, 1839 FKSND--Icastaneus Mellié, 1848 --S-D-A-

fusciclavis Nyholm, 1954


dentatus Mellié, 1848 FKSN--A-bidentatus (Olivier, 1790) FKSNDEAIfissicornis Mellié, 1848 FK---E-I

Ennearthron Mellié, 1847cornutum (Gyllenhal, 1827) FKSNDEAIpruinosulum (Perris, 1864) --S-----laricinum (Mellié, 1848) FKSN-EAIpalmi Lohse, 1966 FKS-----

filum auct. nec Abeille de Perrin, 1874

Orthocis Casey, 1898alni (Gyllenhal, 1813) FKSNDEAIlinearis (J.Sahlberg, 1900) FKSN--A-

perrisi auct. nec (Abeille de Perrin, 1874)pygmaeus (Marsham, 1802) --SND-A-

rhododactylus (Marsham, 1802)vestitus (Mellié, 1848) F-SND-A-

pygmaeus auct. nec (Marsham, 1802)festivus (Panzer, 1793) FKSND-AI

Hadreule Thomson, 1859elongatula (Gyllenhal, 1827) F-SN--AI

Sulcacis Dury, 1917affinis (Gyllenhal, 1827) FKSNDEAIbidentulus (Rosenhauer, 1847) -K------fronticornis (Panzer, 1809) FKSND-A-

Wagaicis Lohse, 1964wagai (Wankowicz, 1869) FK-----I

Ropalodontus Mellié, 1847perforatus (Gyllenhal, 1813) --SND-A-strandi Lohse, 1969 FKSN----baudueri (Abeille de Perrin, 1874) ----D---

Xylographus Mellié, 1847bostrychoides (Dufour, 1843) ----d---

Octotemnus Mellié, 1847glabriculus (Gyllenhal, 1827) FKSNDEAImandibularis (Gyllenhal, 1813) F-SN--AI

Tetratomidae Billberg, 1820Tetratoma Fabricius, 1790fungorum Fabricius, 1790 F-SNDEAIdesmarestii Latreille, 1807 --S-D---ancora Fabricius, 1790 FKSNDEA-

Melandryidae Leach, 1815Hallomeninae Mulsant, 1856Hallomenus Panzer, 1794binotatus (Quensel, 1790) FKSNDEAIaxillaris (Illiger, 1807) FKSNDEA-

Eustrophinae Gistel, 1856Eustrophus Latreille, 1809dermestoides (Fabricius, 1793) F----EA-

Melandryinae Leach, 1815Orchesiini Lacordaire, 1859Orchesia Latreille, 1807micans (Panzer, 1794) FKSNDEAIfusiformis Solsky, 1871 ------A-luteipalpis Mulsant & Guillebeau, 1857 --SND-A-minor Walker, 1837 FKSNDEAIfasciata (Illiger, 1798) FKSNDEAIundulata Kraatz, 1853 F-SNDEA-

Serropalpini Latreille, 1829Anisoxya Mulsant, 1856fuscula (Illiger, 1798) --SND---

Abdera Stephens, 1832affinis (Paykull, 1799) FKSNDEAIflexuosa (Paykull, 1799) FKSNDEA-biflexuosa (Curtis, 1829) ----DE--triguttata (Gyllenhal, 1810) FKSNDEA-

Dircaea Fabricius, 1798quadriguttata (Paykull, 1798) FKS--EA-australis Fairmaire, 1856 --S-----

Phloiotrya Stephens, 1832rufipes (Gyllenhal, 1810) F-SN--A-

Rushia Forel, 1893parreyssii (Mulsant, 1856) --S-D-A-

Xylita Paykull, 1798laevigata (Hellenius, 1786) FKSNdEAIlivida (Sahlberg, 1833) FKS--EA-

Serropalpus Hellenius, 1786barbatus (Schaller, 1783) FKSNDEAI

Hypulini Seidlitz, 1875Hypulus Paykull, 1798quercinus (Quensel, 1790) --SNDEA-bifasciatus (Fabricius, 1792) F-S-D--I

Zilora Mulsant, 1856obscura (Fabricius, 1807) -----EAI

sericea (Sturm, 1807)ferruginea (Paykull, 1798) FKSN-EA-elongata J.Sahlberg, 1881 F----E--

Melandryini Leach, 1815Melandrya Fabricius, 1801caraboides (Linnaeus, 1761) --SND-A-barbata (Fabricius, 1787) FKSND---


dubia (Schaller, 1783) FKSNDEAI

Phryganophilus Sahlberg, 1833ruficollis (Fabricius, 1798) FKSN-EA-auritus Motschulsky, 1860 -----E--

Osphyinae Mulsant, 1856 (1840)Nothinae Shuckard, 1840

Conopalpus Gyllenhal, 1810testaceus (Olivier, 1790) --SND-A-

Osphya Illiger, 1807bipunctata (Fabricius, 1775) --SND---

Mordellidae Latreille, 1802Mordellinae Latreille, 1802Mordellini Latreille, 1802Tomoxia Costa, 1854bucephala Costa, 1854 FKSNDEAI

biguttata (Gyllenhal, 1827) nec (Rossi, 1794)

Variimorda Méquignon, 1946villosa (Schrank, 1781) FKS--EAI

fasciata (Fabricius, 1775) nec (Forster, 1771)briantea (Comolli, 1837) -----EA-basalis (Costa, 1854) F-------

Mordella Linnaeus, 1758huetheri Ermisch, 1956 --SN----leucaspis Küster, 1849 -----E--aculeata Linnaeus, 1758 FKSNDEAIholomelaena Apfelbeck, 1914 FKSNDEAIbrachyura Mulsant, 1856 F-SN----

Hoshihananomia Kono, 1935perlata (Sulzer, 1776) FK---EAI

Curtimorda Méquignon, 1946maculosa (Naezen, 1794) FKSNDEAI

Conalia Mulsant & Rey, 1858baudii Mulsant & Rey, 1858 F-------

Mordellistenini Ermisch, 1941Mordellistenula Stshegoleva-Barovskaja, 1930perrisi (Mulsant, 1856) --SND---

rectangula (Thomson, 1868)

Mordellistena Costa, 1854parvula (Gyllenhal, 1827) FKSNDE-Ifalsoparvula Ermisch, 1956 -----E--connata Ermisch, 1969 FKS-----thurepalmi Ermisch, 1965 FKSN----

palmi Ermisch, 1963, nec Liljeblad, 1945rhenana Ermisch, 1956 F----E--carinthiaca Ermisch, 1966 F-S-----

gemellata Schilsky, 1899 -----E--brevicauda (Boheman, 1849) --S-?-AIpseudobrevicauda Ermisch, 1963 -----E--stenidea Mulsant, 1856 ------I-reitteri Schilsky, 1894 F-------purpureonigrans Ermisch, 1963 F-SNDE--dieckmanni Ermisch, 1963 F-S-----pumila (Gyllenhal, 1810) FKSNDEAIpseudopumila Ermisch, 1963 FKS-D---koelleri Ermisch, 1956 --S-----pygmaeola Ermisch, 1956 F-SN----praesagita Kangas, 1988 FKS-----

dalmatica auct. nec Ermisch, 1956multicicatrix Kangas, 1986 F-------secreta Horák, 1983 --SN----neuwaldeggiana (Panzer, 1796) --S-D---variegata (Fabricius, 1798) F-SNDEAIhumeralis (Linnaeus, 1758) FKSNDEAIrufifrons Schilsky, 1894 ------A-pseudonana Ermisch, 1956 ------AI

nana auct. nec Motschulsky, 1860

Mordellochroa Emery, 1876abdominalis (Fabricius, 1775) FKSNDEAItournieri (Emery, 1876) FKS-----

Rhipiphoridae Laporte de Castelnau, 1840Pelecotominae Seidlitz, 1875Pelecotoma Fischer v. Waldheim, 1809fennica (Paykull, 1799) FK---EAI

Ripidiinae Gerstaecker, 1855Ripidius Thunberg, 1806quadriceps Abeille de Perrin, 1872 F-S-----

apicipennis Kraatz, 1892

Rhipiphorinae Laporte de Castelnau, 1840Metoecus Dejean, 1834paradoxus (Linnaeus, 1761) F-SNDEAI

Zopheridae Solier, 1834Colydiinae Erichson, 1842Colydiini Erichson, 1842Colydium Fabricius, 1792elongatum (Fabricius, 1787) --SND--Ifiliforme Fabricius, 1792 --SN--A-

Aulonium Erichson, 1845trisulcum (Geoffroy, 1785) ----D--I


Synchitini Erichson, 1845Diodesmini Reitter, 1911

Synchita Hellwig, 1792Cicones Curtis, 1827

humeralis (Fabricius, 1793) FKSNDEAIseparanda (Reitter, 1881) --S-----variegata Hellwig, 1792 --S-D---

Bitoma Herbst, 1793Ditoma Illiger, 1807

crenata (Fabricius, 1775) FKSNDEAI

Lasconotus Erichson, 1845Lado Wankowicz, 1867

jelskii (Wankowicz, 1867) FKS----I

Xylolaemus Dejean, 1835fasciculosus (Gyllenhal, 1827) --S-----

Orthocerini Blanchard, 1845 (1820)Orthocerus Latreille, 1796clavicornis (Linnaeus, 1758) FKSNDEAIcrassicornis (Erichson, 1845) -----E--

Zopherinae Solier, 1834Pycnomerini Erichson, 1845Pycnomerus Erichson, 1842Penthelispa Pascoe, 1860

inexpectus Jacquelin du Val, 1859 -----e--

Tenebrionidae Latreille, 1802Lagriinae Latreille, 1825 (1820)Lagria Fabricius, 1775hirta (Linnaeus, 1758) FKSNDEAI

Tenebrioninae Latreille, 1802Blaptini Leach, 1815Blaps Fabricius, 1775mortisaga (Linnaeus, 1758) FKSNDEAIlethifera Marsham, 1802 f-SNDE--sinuatocollis Solier, 1848ssp. suecica Ferrer & Picka, 1990 --S-----mucronata Latreille, 1804 f-Snd---

obtusa Gyllenhal, 1813

Platyscelidini Lacordaire, 1859Oodescelis Motschulsky, 1845polita (Sturm, 1807) -------I

Dendarini Español, 1945Phylan Dejean, 1821gibbus (Fabricius, 1775) --SND-AI

Pedinini Eschscholtz, 1829Pedinus Latreille, 1796femoralis (Linnaeus, 1767) -------I

Opatrini Hope, 1840Opatrum Fabricius, 1775riparium Scriba, 1865 FKSNDEAIsabulosum (Linnaeus, 1761) FKSNDEAI

Melanimon Steven, 1829tibialis (Fabricius, 1781) FKSNDEAI

Ulomini Blanchard, 1845Uloma Dejean, 1821culinaris (Linnaeus, 1758) --SNDEAIrufa (Piller & Mitterpacher, 1783) FKSN-EAI

perroudii Mulsant & Guillebeau, 1855

Triboliini Mulsant, 1854Tribolium MacLeay, 1825madens (Charpentier, 1825) fkS-deaicastaneum (Herbst, 1797) FKSNDEAIdestructor Uyttenboogart, 1934 FKSNDEAIconfusum Jacquelin du Val, 1863 FKSNDEAI

Palorus Mulsant, 1854subdepressus (Wollaston, 1864) f-s-d---depressus (Fabricius, 1790) FKSN-EAIratzeburgii (Wissmann, 1848) -ksnD--i

Latheticus Waterhouse, 1880oryzae Waterhouse, 1880 fksnd-a-

Martianus Fairmaire, 1893dermestoides (Chevrolat, 1878) --S-----

Tenebrionini Latreille, 1802Tenebrio Linnaeus, 1758opacus Duftschmid, 1812 --S-D---obscurus Fabricius, 1792 FKSndE-Imolitor Linnaeus, 1758 FKSNDEAI

Neatus LeConte, 1862picipes (Herbst, 1797) -----EAI

Bius Dejean, 1834thoracicus (Fabricius, 1792) FKSN-E--

Alphitobiini Reitter, 1917Alphitobius Stephens, 1829diaperinus (Panzer, 1797) FkSnDeailaevigatus (Fabricius, 1781) fkSndea-

Helopini Latreille, 1802Stenomax Allard, 1876aeneus (Scopoli, 1763) ------A-

lanipes (Linnaeus, 1771)

Nalassus Mulsant, 1854dermestoides (Illiger, 1798) f-S---AI


Alleculinae Laporte de Castelnau, 1840Alleculini Laporte de Castelnau, 1840Allecula Fabricius, 1801morio (Fabricius, 1787) F-S-D-A-rhenana Bach, 1856 --S-D---

Hymenophorus Mulsant, 1851doublieri Mulsant, 1851 F-S-----

Prionychus Solier, 1835ater (Fabricius, 1775) FKSNDEAImelanarius (Germar, 1813) FKSN----

Hymenalia Mulsant, 1856rufipes (Fabricius, 1793) --SND-A-

Pseudocistela Crotch, 1873ceramboides (Linnaeus, 1758) F-SNDEAI

Gonodera Mulsant, 1856luperus (Herbst, 1783) --S-DE--

Isomira Mulsant, 1856murina (Linnaeus, 1758) FKSNDEAI

Mycetochara Berthold, 1827flavipes (Fabricius, 1793) FKSNDEAIaxillaris (Paykull, 1799) FKSNDEAIhumeralis (Fabricius, 1787) FKSNDEAIobscura (Zetterstedt, 1838) FKSN--A-linearis (Illiger, 1794) --SNDEA-

Cteniopini Solier, 1835Omophlini Mulsant, 1856

Cteniopus Solier, 1835sulphureus (Linnaeus, 1758) F-SND-AI

flavus (Scopoli, 1763)

Omophlus Dejean, 1834betulae (Herbst, 1783) -KS---AI

rufitarsis (Leske, 1785)

Bolitophagini Kirby, 1837Bolitophagus Illiger, 1798reticulatus (Linnaeus, 1767) FKSNDEAI

Eledonoprius Reitter, 1911armatus (Panzer, 1799) --S-D---

Eledona Latreille, 1796agricola (Herbst, 1783) FKSNDEAI

Phaleriini Blanchard, 1845Phaleria Latreille, 1802cadaverina (Fabricius, 1792) --S-D---

Crypticini Mulsant, 1854Crypticus Latreille, 1817quisquilius (Linnaeus, 1761) FKSNDEAI

Adelinini LeConte, 1862Gnatocerus Thunberg, 1814cornutus (Fabricius, 1798) F-SNDEA-maxillosus (Fabricius, 1801) f-------

Cynaeus LeConte, 1862angustus (LeConte, 1851) F-S-----depressus Horn, 1870 --S-----

opacus Champion, 1886

Pentaphyllini Mulsant, 1854Neomida Dahl, 1823Oplocephala Laporte de Castelnau & Brullé, 1831

haemorrhoidalis (Fabricius, 1787) FKSNDEAI

Scaphidema Redtenbacher, 1849metallicum (Fabricius, 1793) FKSNDEAI

Platydema Laporte de Castelnau & Brullé, 1831violaceum (Fabricius, 1790) FKS-DEAI

Alphitophagus Stephens, 1832bifasciatus (Say, 1823) F-SNDEAI

Pentaphyllus Dejean, 1821testaceus (Hellwig, 1792) F-S-DEA-

Diaperini Latreille, 1802Diaperis Geoffroy, 1762boleti (Linnaeus, 1758) FKSNDEAI

Myrmechixenini Jacquelin du Val, 1858Myrmechixenus Chevrolat, 1835subterraneus Chevrolat, 1835 FKSNDEAIvaporariorum Guérin-Ménéville, 1843 F-S-D---

Coelometopinae Lacordaire, 1859Menephilus Mulsant, 1854cylindricus (Herbst, 1784) --S-----

Diaperinae Latreille, 1802Hypophlaeini Billberg, 1820Corticeus Piller & Mitterpacher, 1783Hypophlaeus Fabricius, 1790

unicolor Piller & Mitterpacher, 1783 --SNDEAIlongulus (Gyllenhal, 1827) FKSN-EAIfraxini (Kugelann, 1794) FKSN-EAIpini (Panzer, 1799) --S--EAIsuturalis (Paykull, 1800) FKSN-EAIbicolor (Olivier, 1790) FKSNDEAIfasciatus (Fabricius, 1790) --S-DE-Ilinearis (Fabricius, 1790) FKSNDEAI


Upis Fabricius, 1792ceramboides (Linnaeus, 1758) FKSNdEAI

Prostomidae Thomson, 1859Prostomis Latreille, 1829mandibularis (Fabricius, 1801) --S-D---

Oedemeridae Latreille, 1810Calopodinae Costa, 1852Calopus Fabricius, 1775serraticornis (Linnaeus, 1758) FKSN-EAI

Oedemerinae Latreille, 1810Nacerdini Mulsant, 1858Nacerdes Dejean, 1834Xanthochroa Schmidt, 1844

melanura (Linnaeus, 1758) FKSNDEAIcarniolica (Gistl, 1832) --S-----

Anogcodes Dejean, 1834rufiventris (Scopoli, 1763) -KS--EAIustulatus (Scopoli, 1763) -----EAIferrugineus (Schrank, 1776) --S--E-I

Ditylini Mulsant, 1858Ditylus Fischer v. Waldheim, 1817laevis (Fabricius, 1787) FK---EAI

Chrysanthia Schmidt, 1844viridissima (Linnaeus, 1758) FKSN-EAI

viridis (DeGeer, 1775)geniculata Heyden, 1877 FKSNDEAI

nigricornis (Westhoff, 1881)viridis Schmidt, 1846, nec (DeGeer, 1775)

Asclerini Gistel, 1856Ischnomera Stephens, 1832Asclera Dejean, 1834

sanguinicollis (Fabricius, 1787) --SND-AIcaerulea (Linnaeus, 1758) --SN----cyanea (Fabricius, 1792) --S-D---cinerascens (Pandellé, 1867) --SND---

Oedemerini Latreille, 1810Oedemera Olivier, 1789flavipes (Fabricius, 1792) --SN--AIcroceicollis (Gyllenhal, 1827) F-S-DE--podagrariae (Linnaeus, 1767) --s---AIlurida (Marsham, 1802) FKSNDEAIvirescens (Linnaeus, 1767) FKSNDEAIpthysica (Scopoli, 1763) --S-d---

subulata Olivier, 1795nobilis (Scopoli, 1763) ----D---

femorata (Scopoli, 1763) FKSNDEAIflavescens (Linnaeus, 1767)

Stenotrachelidae Thomson, 1859Cephaloidae LeConte, 1862

Stenotrachelus Berthold, 1827aeneus (Paykull, 1799) FKSN----

Scotodes Eschscholtz, 1818annulatus Eschscholtz, 1818 FK---EA-

Meloidae Gyllenhal, 1810Meloinae Gyllenhal, 1810Cerocomini Leach, 1815Cerocoma Geoffroy, 1762schaefferi (Linnaeus, 1758) ------A-

Lyttini Thomson, 1859Lytta Fabricius, 1775vesicatoria (Linnaeus, 1758) --S-DEAI

Meloini Gyllenhal, 1810Meloe Linnaeus, 1758proscarabaeus Linnaeus, 1758 FKSNDEAIviolaceus Marsham, 1802 FKSNDEAIvariegatus Donovan, 1793 --S-D--Ibrevicollis Panzer, 1793 FKSNDEAI

Nemognathinae Laporte de Castelnau, 1840Zonitidinae Lacordaire, 1857

Apalus Fabricius, 1775bimaculatus (Linnaeus, 1761) FKSNDEA-

Mycteridae Blanchard, 1845Mycterus Clairville & Schellenberg, 1798curculioides (Fabricius, 1781) ------A-

Boridae Thomson, 1859Boros Herbst, 1797schneideri (Panzer, 1795) FKS--EAI

Pythidae Solier, 1834Pytho Latreille, 1796depressus (Linnaeus, 1767) FKSNDEAIkolwensis Sahlberg, 1833 FKS--E--abieticola J.Sahlberg, 1875 FKSN----

Pyrochroidae Latreille, 1807Pyrochroa Geoffroy, 1762coccinea (Linnaeus, 1761) FKSNDEAIserraticornis (Scopoli, 1763) ----DEAI

Schizotus Newman, 1838pectinicornis (Linnaeus, 1758) FKSNDEAI


Salpingidae Leach, 1815Agleninae Horn, 1878Aglenus Erichson, 1845brunneus (Gyllenhal, 1813) --S-D---

Salpinginae Leach, 1815Lissodemini Seidlitz, 1916Lissodema Curtis, 1833cursor (Gyllenhal, 1813) F-SND---denticolle (Gyllenhal, 1813) --S-D---

quadripustulatum (Marsham, 1802) nec (Fabricius, 1775)

Salpingini Leach, 1815Colposis Mulsant, 1859mutilatus (Beck, 1817) ------AI

Rabocerus Mulsant, 1859foveolatus (Ljungh, 1823) FKSNDEAIgabrieli (Gerhardt, 1901) FKSNDEAI

Sphaeriestes Stephens, 1829Salpingus auct., nec Illiger, 1801

castaneus (Panzer, 1796) F-SNDEAIbimaculatus (Gyllenhal, 1810) FKSN-EAIstockmanni (Biström, 1977) FKSNDEAI

ater (Paykull, 1798) nec (DeGeer, 1774)reyi (Abeille de Perrin, 1874) F-SND---

Vincenzellus Reitter, 1911ruficollis (Panzer, 1794) --S-D---

viridipennis (Latreille, 1804)

Salpingus Illiger, 1801Rhinosimus Latreille, 1802

aeneus (Olivier, 1807) -----EA-planirostris (Fabricius, 1787) FKSNDEAIruficollis (Linnaeus, 1761) FKSNDEAI

Anthicidae Latreille, 1819Anthicinae Latreille, 1819Notoxini Sturm, 1826Notoxus Geoffroy, 1762monoceros (Linnaeus, 1761) FKSNDEAI

Formicomini Bonadona, 1974Anthelephila Hope, 1833Formicomus LaFerté-Sénectère, 1849

pedestris (Rossi, 1790) ------A-

Anthicini Latreille, 1819Cyclodinus Mulsant & Rey, 1866humilis (Germar, 1824) --S-D---

sibiricus auct. nec (Pic, 1893)

Omonadus Mulsant & Rey, 1866floralis (Linnaeus, 1758) FKSNDEAIformicarius (Goeze, 1777) FKSNDEAI

quisquilius (Thomson, 1864)kolenatii (Kolenati, 1846) --S-D---

bifasciatus (Rossi, 1792) nec (Fabricius, 1792)

Cordicomus Pic, 1894instabilis (Schmidt, 1842) --SND---gracilis (Panzer, 1797) -KSN---Isellatus (Panzer, 1797) FKSNDEAI

Stricticomus Pic, 1894tobias (Marseul, 1879) F-S-DEAI

Hirticomus Pic, 1894hispidus (Rossi, 1792) --S---A-

Anthicus Paykull, 1798flavipes (Panzer, 1797) FKSNDEAIantherinus (Linnaeus, 1761) FKSNDEAIaxillaris Schmidt, 1842 -KS--EAIater (Panzer, 1796) FKSNDEAIumbrinus LaFerté-Sénectère, 1848 FKS---A-bimaculatus (Illiger, 1801) FKSNDEAI

Aderidae Winkler, 1927Phytobaenini Báguena, 1948Phytobaenus F.Sahlberg, 1834amabilis F.Sahlberg, 1834 F-S--EA-

Aderini Winkler, 1927Euglenesini Seidlitz, 1875

Aderus Stephens, 1829Hylophilus Berthold, 1827, nec Temminck, 1822

populneus (Creutzer, 1796) F-SNDEAI

Euglenes Westwood, 1830pygmaeus (DeGeer, 1775) FKSN-EAI

lokvenci (Roubal, 1938)oculatus (Paykull, 1798) --SND-A-

Anidorus Mulsant & Rey, 1866nigrinus (Germar, 1831) FKSNDEAI

Vanonus Casey, 1895brevicornis (Perris, 1869) --S-D---

Pseudeuglenes Pic, 1897pentatomus (Thomson, 1864) FKSN----

Scraptiidae Mulsant, 1856Scraptiinae Mulsant, 1856Scraptia Latreille, 1807fuscula Müller, 1821 F-SND-A-

innata (Kangas, 1959)


Anaspidinae Mulsant, 1856Cyrtanaspis Emery, 1876phalerata (Germar, 1831) FKS--EA-

Anaspis Geoffroy, 1762fasciata (Forster, 1771) ----D---

humeralis (Fabricius, 1775) nec (Linnaeus, 1758)bohemica Schilsky, 1898 FKSND-A-

norvegica Munster, 1924lurida Stephens, 1832 F-----A-viennensis Schilsky, 1895 -----E--frontalis (Linnaeus, 1758) FKSNDEAImaculata Geoffroy, 1785 --S-D---marginicollis Lindberg, 1925 FKSNDEA-

schilskyana Csiki, 1915 – infrasubsp.thoracica (Linnaeus, 1758) FKSNDEAIruficollis (Fabricius, 1793) ----D-AIpulicaria Costa, 1854 -----E-Iregimbarti Schilsky, 1895 --S-D---arctica Zetterstedt, 1828 FKSN-EA-garneysi Fowler, 1889 --S-D-A-rufilabris (Gyllenhal, 1827) FKSNDEAImelanostoma Costa, 1854 ----D--Icostai Emery, 1876 ----D---flava (Linnaeus, 1758) F-SND-AIbrunnipes (Mulsant, 1856) FKS--EAIvarians (Mulsant, 1856) -----EA-

CHRYSOMELOIDEACerambycidae Latreille, 1802Prioninae Latreille, 1802Ergates Audinet-Serville, 1832faber (Linnaeus, 1761) --S-d-AI

Prionus Geoffroy, 1762coriarius (Linnaeus, 1758) FKSNDEAI

Tragosoma Audinet-Serville, 1832depsarius (Linnaeus, 1767) FKSN-EAI

Spondylidinae Audinet-Serville, 1832Spondylis Fabricius, 1775buprestoides (Linnaeus, 1758) FKSNDEAI

Aseminae J.Thomson, 1860Nothorhina Redtenbacher, 1845punctata (Fabricius, 1798) FKSN-EAI

muricata (Dalman, 1817)

Arhopalus Audinet-Serville, 1834Criocephalus Dejean, 1835

rusticus (Linnaeus, 1758) FKSNDEAI

tristis (Fabricius, 1787) FKS-DEAIferus (Mulsant, 1839)

Asemum Eschscholtz, 1830striatum (Linnaeus, 1758) FKSNDEAItenuicorne Kraatz, 1879 --S-----

Tetropium Kirby, 1837castaneum (Linnaeus, 1758) FKSNDEAIfuscum (Fabricius, 1787) FKSNDEAIgabrieli Weise, 1905 f-s-D---aquilonium Plavistshikov, 1940 FKS-----

Lepturinae Latreille, 1802Rhagiini Kirby, 1837

Stenocorini J.Thomson, 1860Rhagium Fabricius, 1775bifasciatum Fabricius, 1775 ---NDE-Isycophanta (Schranck, 1781) --S-D-AImordax (DeGeer, 1775) FKSNDEAIinquisitor (Linnaeus, 1758) FKSNDEAI

Rhamnusium Latreille, 1829bicolor (Schranck, 1781) F---dEAI

gracilicorne Théry, 1895

Oxymirus Mulsant, 1863cursor (Linnaeus, 1758) FKSNDEAI

Stenocorus Geoffroy, 1762meridianus (Linnaeus, 1758) F-SNDEAI

Pachyta Dejean, 1821quadrimaculata (Linnaeus, 1758) FK--DEAIlamed (Linnaeus, 1758) FKSNDEAI

Brachyta Fairmaire, 1864interrogationis (Linnaeus, 1758) FKSNdEAI

Evodinus LeConte, 1850borealis (Gyllenhal, 1827) FKSN-EAI

Gaurotes LeConte, 1850Carilia Mulsant, 1863

virginea (Linnaeus, 1758) FKSN-EAI

Acmaeops LeConte, 1850Dinoptera Mulsant, 1863Gnathacmaeops Linsley & Chemsak, 1972

septentrionis (Thomson, 1866) FKSN-E--marginata (Fabricius, 1781) FKSN-EAIsmaragdula (Fabricius, 1792) FKSN---Ipratensis (Laicharting, 1784) FKSN-EAIcollaris (Linnaeus, 1758) --SNDEAI

Pidonia Mulsant, 1863lurida (Fabricius, 1787) ------A-


Lepturini Latreille, 1802Cortodera Mulsant, 1863femorata (Fabricius, 1787) FKSN-EAI

Grammoptera Audinet-Serville, 1835ustulata (Schaller, 1783) --SND---ruficornis (Fabricius, 1781) --SNDEAIabdominalis (Stephens, 1831) --S-D---

Alosterna Mulsant, 1863tabacicolor (DeGeer, 1775) FKSNDEAIerythropus (Gebler, 1841)ssp. ingrica Baeckmann, 1902 -----E-I

Nivellia Mulsant, 1863sanguinosa (Gyllenhal, 1827) FKSN----extensa (Gebler, 1841) -K------

Anoplodera Mulsant, 1839Corymbia Des Gozis, 1886

rufipes (Schaller, 1783) --S---A-sexguttata (Fabricius, 1775) F-SNDEAIlivida (Fabricius, 1777) -K--DEAIfulva (DeGeer, 1775) -------Imaculicornis (DeGeer, 1775) FKSNDEAIrubra (Linnaeus, 1758) FKSNDEAIvariicornis (Dalman, 1817) -K---EAIscutellata (Fabricius, 1781) --S-D-A-sanguinolenta (Linnaeus, 1761) FKSNDEAIdubia (Scopoli, 1763) -------Ireyi (Heyden, 1889) FKSNdEAI

inexpectata (Jansson & Sjöberg, 1928)virens (Linnaeus, 1758) FKSNdEAI

Judolia Mulsant, 1863sexmaculata (Linnaeus, 1758) FKSNDEAIcerambyciformis (Schrank, 1781) ----D?AI

Leptura Linnaeus, 1758Pedostrangalia Sokolow, 1897

revestita Linnaeus, 1767 --S-D---pubescens Fabricius, 1787 FKSN-EAInigripes DeGeer, 1775 FKSN-EAIthoracica Creutzer, 1799 FK---EAIaurulenta Fabricius, 1792 -------Iquadrifasciata Linnaeus, 1758 FKSNDEAIannularis Fabricius, 1801 -----EAI

mimica Bates, 1884arcuata Panzer, 1793, nec Linnaeus,1758

aethiops Poda, 1761 ----DEAImaculata Poda, 1761 F-SNDEAImelanura Linnaeus, 1758 FKSNDEAI

bifasciata Müller, 1776 -----EAInigra Linnaeus, 1758 --SNDEAI

Strangalia Audinet-Serville, 1835attenuata (Linnaeus, 1758) FKSNDEAI

Necydalini Latreille, 1825Necydalis Linnaeus, 1758major Linnaeus, 1758 FKSNDEAIulmi Chevrolat, 1836 ------AI

Cerambycinae Latreille, 1802Cerambycini Latreille, 1802Cerambyx Linnaeus, 1758cerdo Linnaeus, 1758 --S-d-AIscopoli Fuessli, 1775 --SND?-I

Graciliini Mulsant, 1839Gracilia Audinet-Serville, 1834minuta (Fabricius, 1781) f-snd--i

Axinopalpis Duponchel & Chevrolat, 1842gracilis (Krynicki, 1832) -------I

Obriini Mulsant, 1839Obrium Dejean, 1821cantharinum (Linnaeus, 1767) FKS--EAIbrunneum (Fabricius, 1792) -KS-DEA-

Psebiini Lacordaire, 1869Nathrius Brèthes, 1916brevipennis (Mulsant, 1839) f-snd---

Molorchini Mulsant, 1862Molorchus Fabricius, 1792Glaphyra Newman, 1840

minor (Linnaeus, 1758) FKSNDEAIumbellatarum (Schreber, 1759) --SNDEAI

Callichromatini Blanchard, 1845Aromia Audinet-Serville, 1833moschata (Linnaeus, 1758) FKSNDEAI

Compsocerini J.Thomson, 1864Rosaliini Fairmaire, 1864

Rosalia Audinet-Serville, 1833alpina (Linnaeus, 1758) --S-d---

Callidiini Kirby, 1837Hylotrupes Audinet-Serville, 1834bajulus (Linnaeus, 1758) F-SNDEAI

Ropalopus Mulsant, 1839femoratus (Linnaeus, 1758) --S---A-macropus (Germar, 1824) --S----Iclavipes (Fabricius, 1775) ----D-AI


Leioderes Redtenbacher, 1849kollari Redtenbacher, 1849 --SN--AI

Semanotus Mulsant, 1839undatus (Linnaeus, 1758) FKSNdEAI

Callidium Fabricius, 1775coriaceum Paykull, 1800 FKSN-EAIviolaceum (Linnaeus, 1758) FKSNDEAIaeneum (DeGeer, 1775) FKSN-EAI

Callidiellum Linsley, 1940rufipenne (Motschulsky, 1860) ----d---

Pyrrhidium Fairmaire, 1864sanguineum (Linnaeus, 1758) F-SND--I

Phymatodes Mulsant, 1839testaceus (Linnaeus, 1758) F-SNDEAIpusillus (Fabricius, 1787) --S-----

Poecilium Fairmaire, 1864alni (Linnaeus, 1767) F-SNDEAI

Clytini Mulsant, 1839Xylotrechus Chevrolat, 1860Rusticoclytus Vives Noguera, 1977

rusticus (Linnaeus, 1758) FKSNDEAIpantherinus (Savenius, 1825) FKSN----antilope (Schönherr, 1817) --SN----ibex (Gebler, 1825) F------Iarvicola (Olivier, 1795) -----EAI

Clytus Laicharting, 1784arietis (Linnaeus, 1758) F-SNDEAIrhamni Germar, 1817 -------I

Cyrtoclytus Ganglbauer, 1881capra (Germar, 1824) -----EAI

Plagionotus Mulsant, 1842detritus (Linnaeus, 1758) --S-DEAIarcuatus (Linnaeus, 1758) F-SND?AIfloralis (Pallas, 1773) ------AI

Chlorophorus Chevrolat, 1863varius (Müller, 1766) --S----Iherbstii (Brahm, 1790) F-S--EAIfiguratus (Scopoli, 1763) ------AI

Anaglyptus Mulsant, 1839mysticus (Linnaeus, 1758) --SNDEAI

Lamiinae Latreille, 1825Dorcadiini Latreille, 1825Dorcadion Dalman, 1817Iberodorcadion Breuning, 1943

fuliginator (Linnaeus, 1758) ------A-

Lamiini Latreille, 1825Lamia Fabricius, 1775textor (Linnaeus, 1758) FKSNDEAI

Monochamus Dejean, 1821sartor (Fabricius, 1787) ----d--Iurussovii (Fischer v. Waldheim, 1806) FKSNdEAIsutor (Linnaeus, 1758) FKSNDEAIgalloprovincialis (Olivier, 1795) FKSN-EAIsaltuarius Gebler, 1830 -------I

Mesosini Mulsant, 1839Mesosa Latreille, 1829Aplocnemia Stephens, 1831

curculionoides (Linnaeus, 1761) --SNd--Imyops (Dalman, 1817) F-S---A-nebulosa (Fabricius, 1781) --SND---

Apodasyini Lacordaire, 1872Rhodopini Lacordaire, 1872

Anaesthetis Dejean, 1835testacea (Fabricius, 1781) --S--EA-

Pogonocherini Mulsant, 1839Pogonocherus Dejean, 1821Pityphilus Mulsant, 1863Eupogonocherus Linsley, 1935

hispidulus (Piller & Mitterpacher, 1781) --SNDEAIhispidus (Linnaeus, 1758) F-SNDEAIcaroli Mulsant, 1863 --S-----fasciculatus (DeGeer, 1775) FKSNDEAIdecoratus Fairmaire, 1855 FKSNDEAIovatus (Goeze, 1777) ------AI

Acanthoderini J.Thomson, 1860Oplosia Mulsant, 1863cinerea (Mulsant, 1839) F-SNDEAI

fennica (Paykull, 1800) nec (Linnaeus, 1758)

Aegomorphus Haldeman, 1847Acanthoderes auct., nec Audinet-Serville, 1835

clavipes (Schrank, 1781) FKSNDEAI

Acanthocinini J.Thomson, 1860Leiopus Audinet-Serville, 1835nebulosus (Linnaeus, 1758) F-SNDEAIpunctulatus (Paykull, 1800) FKS--EAI

Acanthocinus Dejean, 1821aedilis (Linnaeus, 1758) FKSNDEAIgriseus (Fabricius, 1792) FKSNdEAI

Exocentrus Dejean, 1835adspersus Mulsant, 1846 --S-----


lusitanus (Linnaeus, 1767) FKSNDEAI

Agapanthiini Mulsant, 1839Agapanthia Audinet-Serville, 1835villosoviridescens (DeGeer, 1775) FKSNDEAI

Saperdini Mulsant, 1839Saperda Fabricius, 1775Anaerea Mulsant, 1839Compsidia Mulsant, 1839

carcharias (Linnaeus, 1758) FKSNDEAIsimilis Laicharting, 1784 FKSN-EAIpopulnea (Linnaeus, 1758) FKSNDEAIscalaris (Linnaeus, 1758) FKSNDEAIperforata (Pallas, 1773) FKSN-EAIpunctata (Linnaeus, 1767) ------A-octopunctata (Scopoli, 1772) --S----I

Menesia Mulsant, 1856bipunctata (Zoubkoff, 1829) -----EAI

Phytoeciini Mulsant, 1839Oberea Dejean, 1835pupillata (Gyllenhal, 1817) -----EAIoculata (Linnaeus, 1758) FKSNDEAIlinearis (Linnaeus, 1761) --SND-A-erythrocephala (Schrank, 1776) -------I

Stenostola Dejean, 1835dubia (Laicharting, 1784) F-SND-A-ferrea (Schrank, 1776) --SNDEAI

Phytoecia Dejean, 1835affinis (Harrer, 1784) -------I

nigripes (Panzer, 1794) nec (DeGeer, 1775)nigricornis (Fabricius, 1781) FKS--EAIcylindrica (Linnaeus, 1758) FKSNDEAIpustulata (Schrank, 1776) -------Ivirgula (Charpentier, 1825) ------AIcaerulea (Scopoli, 1772) -------Icoerulescens (Scopoli, 1763) -------I

Tetraopini J.Thomson, 1860Tetrops Kirby, 1826praeusta (Linnaeus, 1758) FKSNDEAIstarkii Chevrolat, 1859 --SND---

frontalis Suffrian, 1840 --S-DEAIflavicollis auct. nec (Marsham, 1802)

Orsodacnidae Thomson, 1859Orsodacne Latreille, 1802cerasi (Linnaeus, 1758) F-SNDEAI

Megalopodidae Latreille, 1802Zeugophorinae Böving & Craighead, 1931Zeugophora Kunze, 1818scutellaris Suffrian, 1840 FKSND-AIsubspinosa (Fabricius, 1781) FKSNDEAIturneri Power, 1863 F-SNDEAI

Chrysomelidae Latreille, 1802Donaciinae Kirby, 1837Macroplea Samouelle, 1819Haemonia Dejean, 1821

appendiculata (Panzer, 1794) FKSNDEAImutica (Fabricius, 1792) FKSNDEA-pubipennis (Reuter, 1875) F-------

Donacia Fabricius, 1775clavipes Fabricius, 1792 FKSNDEAIcrassipes Fabricius, 1775 FKSNDEAIdentata Hoppe, 1795 FKSNDEAIversicolorea (Brahm, 1790) FKSNDEAIsemicuprea Panzer, 1796 FKSNDEAImalinovskyi Ahrens, 1810 -------Ifennica (Paykull, 1800) FK---EAIsparganii Ahrens, 1810 FKSNDEAIaquatica (Linnaeus, 1758) FKSNDEAIimpressa Paykull, 1799 FKSNDEAIspringeri Müller, 1916 --S-----brevitarsis Thomson, 1884 FKS-D-A-brevicornis Ahrens, 1810 FKSNDE-Imarginata Hoppe, 1795 FKS?DEAIbicolor Zschach, 1788 FKSNDEAIobscura Gyllenhal, 1813 FKSNDEA-aureocincta J.Sahlberg, 1921 F-S-----antiqua Kunze, 1818 FK---EAIthalassina Germar, 1811 FKSNDEAIvulgaris Zschach, 1788 FKSNDEAIsimplex Fabricius, 1775 FKSNDEAItomentosa Ahrens, 1810 FKS--EAIcinerea Herbst, 1784 FKSNDEAI

Plateumaris Thomson, 1859discolor (Herbst, 1795) FKSNDEAIsericea (Linnaeus, 1758) FKSNDEAIbraccata (Scopoli, 1772) F-SNDEAIconsimilis (Schrank, 1781) --S-D--Iweisei (Duvivier, 1885) FKS-----affinis (Kunze, 1818) FKSNDEAIrustica (Kunze, 1818) ----DEAI


Criocerinae Latreille, 1807Criocerini Latreille, 1807Crioceris Geoffroy, 1762duodecimpunctata (Linnaeus, 1758) --S-DEAIasparagi (Linnaeus, 1758) f-S-D-AI

Lilioceris Reitter, 1912lilii (Scopoli, 1763) F-SNDEAImerdigera (Linnaeus, 1758) FKSNDEAI

Lemini Gyllenhal, 1813Lema Fabricius, 1798cyanella (Linnaeus, 1758) FKSNDEAI

puncticollis (Curtis, 1830)

Oulema Des Gozis, 1886erichsonii (Suffrian, 1841) FKSNDEAI

septentrionis (Weise, 1880)gallaeciana (Heyden, 1870) FKSNDEAI

lichenis (Weise, 1881)melanopus (Linnaeus, 1758) FKSNDEAIduftschmidi (Redtenbacher, 1874) --S-D---

Clytrinae Kirby, 1837Labidostomis Germar, 1817tridentata (Linnaeus, 1758) FKSNDEAIhumeralis (Schneider, 1792) --SN--AIlucida (Germar, 1823) ------A-longimana (Linnaeus, 1761) --SNDEAIlepida Lefèvre, 1872 ------A-

Clytra Laicharting, 1781quadripunctata (Linnaeus, 1758) FKSNDEAIlaeviuscula Ratzeburg, 1837 -----EA-

Smaragdina Chevrolat, 1837Gynandrophthalma Lacordaire, 1848

salicina (Scopoli, 1763) FKS-DEAIcyanea (Fabricius, 1775)

flavicollis (Charpentier, 1825) FK---EAIaurita (Linnaeus, 1767) --S-D---affinis (Illiger, 1794) FK----A-

Coptocephala Chevrolat, 1837unifasciata (Scopoli, 1763) -----EAI

Cryptocephalinae Gyllenhal, 1813Pachybrachini Chapuis, 1874Pachybrachis Chevrolat, 1837hieroglyphicus (Laicharting, 1781) FK---EAI

Cryptocephalini Gyllenhal, 1813Cryptocephalus Geoffroy, 1762coryli (Linnaeus, 1758) FKSNDEAI

cordiger (Linnaeus, 1758) FKSNDEAIoctopunctatus (Scopoli, 1763) FKS--EAIsexpunctatus (Linnaeus, 1758) FKSNDEAIdistinguendus Schneider, 1792 FKSNDEAIcruciger Hellén, 1922 F-S-----bipunctatus (Linnaeus, 1758) FKSNDEAIbiguttatus (Scopoli, 1763) FKS-DEAIaureolus Suffrian, 1847 FKS-DEAIsericeus (Linnaeus, 1758) --SNDEAIhypochoeridis (Linnaeus, 1758) FKSNDEAIviolaceus Laicharting, 1781 -------Ielongatus Germar, 1824 --S-----nitidus (Linnaeus, 1758) FKSNDEAInitidulus Fabricius, 1787 FKSN-EAIparvulus Müller, 1776 FKSNDEAIcaerulescens Sahlberg, 1838 FKS--EAImarginatus Fabricius, 1781 F-------decemmaculatus (Linnaeus, 1758) FKSNDEAImoraei (Linnaeus, 1758) FKSNDEAIquadripustulatus Fabricius, 1775 FKSN-E-Iflavipes Fabricius, 1781 F-S?DEAIsignatifrons Suffrian, 1847 ------A-vittatus Fabricius, 1775 --S-D-AIpini (Linnaeus, 1758) FKSN-EAIfrenatus Laicharting, 1781 -------Ipunctiger Paykull, 1799 FKSNDEA-pallifrons Gyllenhal, 1813 FKS-DEAIjanthinus Germar, 1824 -------Ibilineatus (Linnaeus, 1767) FKSNDEAIelegantulus Gravenhorst, 1807 ------A-chrysopus Gmelin, 1790 ------A-frontalis Marsham, 1802 FKSNDEAIsaliceti Zebe, 1855 F-------ocellatus Drapiez, 1819 -----EAIquerceti Suffrian, 1848 FKSND-AIlabiatus (Linnaeus, 1761) FKSNDEAIexiguus Schneider, 1792 FKSN-EAIpygmaeus Fabricius, 1792 ------A-fulvus (Goeze, 1777) FKSNDEAIpopuli Suffrian, 1848 ------A-pusillus Fabricius, 1777 FKSNDEAIrufipes (Goeze, 1777) ----D-A-

Lamprosomatinae Lacordaire, 1849Oomorphus Curtis, 1831concolor (Sturm, 1807) --S-D-A-


Synetinae Edwards, 1953Syneta Dejean, 1837betulae (Fabricius, 1792) FKSN-EA-

Eumolpinae Hope, 1840Eumolpini Hope, 1840Chrysochus Chevrolat, 1837asclepiadeus (Pallas, 1776) ------AI

Adoxini Baly, 1863Pachnephorus Chevrolat, 1837pilosus (Rossi, 1790) FK---EAItessellatus (Duftschmid, 1825) -----EAI

Bromius Chevrolat, 1837Adoxus Kirby, 1837

obscurus (Linnaeus, 1758) FKSNDEAI

Chrysomelinae Latreille, 1802Chrysomelini Latreille, 1802Leptinotarsa Chevrolat, 1837decemlineata (Say, 1824) FKS-DEAI

Chrysolina Motschulsky, 1860Dlochrysa Motschulsky, 1860

herbacea (Duftschmid, 1825) ----D---menthastri (Suffrian, 1851)

caerulans (Scriba, 1791) -----E--graminis (Linnaeus, 1758) FKSNDEAIpolita (Linnaeus, 1758) FKSNDEAIreitteri (Weise, 1884)ssp. saxonica Silfverberg, 1977 -----E-I

lurida (Linnaeus, 1767) nec (Scopoli, 1763)staphylaea (Linnaeus, 1758) FKSNDEAIcerealis (Linnaeus, 1767) FK--DEAIrelucens (Rosenhauer, 1847) -K------rufa (Duftschmid, 1825) -----EA-oricalcia (Müller, 1776) --SND--Ihaemoptera (Linnaeus, 1758) F-S-DEAIfuliginosa (Olivier, 1807) ------A-sturmi (Westhoff, 1882) --SNDEAI

diversipes (Bedel, 1892)violacea auct. nec (Müller, 1776)

sanguinolenta (Linnaeus, 1758) FKSNDEAIgypsophilae (Küster, 1845) FKSNDEAIlatecincta (Demaison, 1896)ssp. hellieseni Silfverberg, 1977 ---N----

crassicornis (Helliesen, 1912) nec (Fabricius, 1775)kuesteri (Helliesen, 1911) ------A-varians (Schaller, 1783) FKSNDEAIhyperici (Forster, 1771) F-SNDEAI

americana (Linnaeus, 1758) ------A-brunsvicensis (Gravenhorst, 1807) ----D---geminata (Paykull, 1799) -KSNDEAIquadrigemina (Suffrian, 1851) ----D-A-carnifex (Fabricius, 1792) ----DEAIanalis (Linnaeus, 1767) FKSNDEAImarginata (Linnaeus, 1758) FKSNDEAIlimbata (Fabricius, 1775) --S-DEAIfastuosa (Scopoli, 1763)ssp. speciosa (Linnaeus, 1767) FKSNDEAI

Oreina Chevrolat, 1837Chrysochloa Hope, 1840

luctuosa (Olivier, 1807) -K--D---rugulosa (Suffrian, 1851) nec (Gebler, 1841)suffriani Bontems, 1984

Colaphellus Weise, 1916sophiae (Schaller, 1783) ----D-AI

Gastrophysa Chevrolat, 1837polygoni (Linnaeus, 1758) FKSNDEAIviridula (DeGeer, 1775) FKSNDEAI

Phaedon Megerle, 1823laevigatus (Duftschmid, 1825) -----EAIcochleariae (Fabricius, 1792) FKSNDEAIarmoraciae (Linnaeus, 1758) FKSNDEAIconcinnus (Stephens, 1831) FKSNDE--

Hydrothassa Thomson, 1859glabra (Herbst, 1783) FKSNDEAImarginella (Linnaeus, 1758) FKSNDEAIhannoveriana (Fabricius, 1775) FKS-DEAI

Prasocuris Latreille, 1802junci (Brahm, 1790) --S-D-AIphellandrii (Linnaeus, 1758) FKSNDEAI

Plagiodera Chevrolat, 1837versicolora (Laicharting, 1781) FKSNDEAI

Chrysomela Linnaeus, 1758cuprea Fabricius, 1775 FKSN-EAIlapponica Linnaeus, 1758 FKSN-EAIcollaris Linnaeus, 1758 FKSNDEAIvigintipunctata (Scopoli, 1763) FKS--EAIpopuli Linnaeus, 1758 FKSNDEAItremula Fabricius, 1787 FKSNDEAIsaliceti (Weise, 1884) FKS-DEAI

Plagiosterna Motschulsky, 1860Linaeidea Motschulsky, 1860

aenea (Linnaeus, 1758) FKSNDEAI


Gonioctena Chevrolat, 1837Phytodecta Kirby, 1837

flavicornis (Suffrian, 1851) FK-N--AIviminalis (Linnaeus, 1758) FKSNDEAIdecemnotata (Marsham, 1802) FKSNDEAI

rufipes (DeGeer, 1775) nec (Linnaeus,1758)linnaeana (Schranck, 1781) FKSN-EAIarctica Mannerheim, 1853 FKSN----

affinis (Gyllenhal, 1808) nec (Fabricius, 1787)decaspilota (Achard, 1924)

sundmani (Jacobson, 1901) -K------norvegica (Strand, 1936) FKSN----olivacea (Forster, 1771) ----D---quinquepunctata (Fabricius, 1787) FKSNDEAIintermedia (Helliesen, 1913) FKSN-EA-pallida (Linnaeus, 1758) FKSNDEAI

Phratora Chevrolat, 1837Phyllodecta Kirby, 1837

vulgatissima (Linnaeus, 1758) FKSNDEAItibialis (Suffrian, 1851) ----DEAIpolaris (Sparre Schneider, 1886) FKSN--A-laticollis (Suffrian, 1851) FKSNDEAIvitellinae (Linnaeus, 1758) FKSNDEAIatrovirens (Cornelius, 1857) FKSNDEAI

Entomoscelis Chevrolat, 1837adonidis (Pallas, 1771) -k----AI

Timarchini Motschulsky, 1860Timarcha Dejean, 1821tenebricosa (Fabricius, 1775) ------A-goettingensis (Linnaeus, 1758) ------AI

Galerucinae Latreille, 1802Galerucini Latreille, 1802Galerucella Crotch, 1873nymphaeae (Linnaeus, 1758) FKSNDEAIaquatica (Geoffroy, 1785) F---D---sagittariae (Gyllenhal, 1813) FKSNDEAI

kerstensi Lohse, 1989grisescens (Joannis, 1865) FK---EAIlineola (Fabricius, 1781) FKSNDEAIcalmariensis (Linnaeus, 1767) FKSNDEAIpusilla (Duftschmid, 1825) FKSNDEAItenella (Linnaeus, 1761) FKSNDEAI

Pyrrhalta Joannis, 1865viburni (Paykull, 1799) FKSNDEAI

Xanthogaleruca Laboissière, 1934luteola (Müller, 1766) --S-D-AI

Lochmaea Weise, 1883caprea (Linnaeus, 1758) FKSNDEAIsuturalis (Thomson, 1866) FKSNDEAIcrataegi (Forster, 1771) --S-D-AI

Galeruca Geoffroy, 1762tanaceti (Linnaeus, 1758) FKSNDEAIinterrupta Illiger, 1802 --S-D-AI

circumdata Duftschmid, 1825, nec Say, 1824pomonae (Scopoli, 1763) FKSNDEAIlaticollis Sahlberg, 1838 FKS-DEAIdahlii (Joannis, 1865) ------A-melanocephala Ponza, 1805 FKS-D---

Hylaspini Chapuis, 1875Sermylassa Reitter, 1912halensis (Linnaeus, 1767) --S-D-AI

Agelastica Chevrolat, 1837alni (Linnaeus, 1758) FKSNDEAI

Luperini Gistel, 1856Phyllobrotica Chevrolat, 1837quadrimaculata (Linnaeus, 1758) FKSNDEAI

Calomicrus Dillwyn, 1829pinicola (Duftschmid, 1825) FKS--EAI

Luperus Geoffroy, 1762luperus (Sulzer, 1776) ------AIlongicornis (Fabricius, 1781) FKSNDEAIflavipes (Linnaeus, 1758) FKSNDEAI

Alticinae Newman, 1834Phyllotreta Chevrolat, 1837vittula (Redtenbacher, 1849) FKSNDEAIarmoraciae (Koch, 1803) F-S-DEAInemorum (Linnaeus, 1758) FKSNDEAIundulata Kutschera, 1860 FKSNDEAItetrastigma (Comolli, 1837) FKSNDEAIdilatata Thomson, 1866 --S-D---flexuosa (Illiger, 1794) FKSNDEAIzimmermanni (Crotch, 1873) F-S-----striolata (Fabricius, 1803) FKSNDEAI

vittata auct. nec (Fabricius, 1775)ochripes (Curtis, 1837) FKSND-AIexclamationis (Thunberg, 1784) FKSNDEAIatra (Fabricius, 1775) FKSNDEAIdiademata Foudras, 1860 --S-----cruciferae (Goeze, 1777) --S-D-AInigripes (Fabricius, 1775) FKS-DEAI


Aphthona Chevrolat, 1837cyparissiae (Koch, 1803) ------A-abdominalis (Duftschmid, 1825) ------AIpallida (Bach, 1856) --SN--AIlutescens (Gyllenhal, 1813) FKSNDEAInigriceps (Redtenbacher, 1842) ------A-violacea (Koch, 1803) --S--EAIpygmaea Kutschera, 1861 FKS--EAIatrocaerulea (Stephens, 1831) --SND--I

cyanella (Redtenbacher, 1849)venustula Kutschera, 1861 --S--E-Ieuphorbiae (Schrank, 1781) FKSNDEAInonstriata (Goeze, 1777) FKSNDEAI

coerulea (Geoffroy, 1785)erichsoni (Zetterstedt, 1838) FKSN-EAI

Longitarsus Berthold, 1827pellucidus (Foudras, 1860) --SNDEAIochroleucus (Marsham, 1802) F-SND-A-jacobaeae (Waterhouse, 1858) --SNDEAIsymphyti Heikertinger, 1912 ------AIsuccineus (Foudras, 1860) FKSNDEAIaeruginosus (Foudras, 1860) ------A-noricus Leonardi, 1976 ------A-rubiginosus (Foudras, 1860) FKS-DEAItabidus (Fabricius, 1775) FKSNDEAInigrofasciatus (Goeze, 1777) ---N--A-lycopi (Foudras, 1860) FKS-DEAImembranaceus (Foudras, 1860) ------A-ferrugineus (Foudras, 1860) --S-D-AInanus (Foudras, 1860) ------A-melanocephalus (DeGeer, 1775) FKSNDEAIplantagomaritimus Dollman, 1912 -KS-D---kutscherai (Rye, 1872) FKSND--Icurtus (Allard, 1860) FKSNDEAIexsoletus (Linnaeus, 1758) --SNDEAIpulmonariae Weise, 1893 ----D---longipennis Kutschera, 1864 ------A-lewisii Baly, 1874 FK---EA-

scutellaris auct. nec (Rey, 1874)pratensis (Panzer, 1794) F-SNDEAIreichei (Allard, 1860) --SND---ballotae (Marsham, 1802) ------A-gracilis Kutschera, 1864 --S-DEAIlongiseta Weise, 1889 FKSN-EAIganglbaueri Heikertinger, 1912 FKSN--AIaeneicollis (Faldermann, 1837) -----EAI

suturalis (Marsham, 1802) nec (Fabricius, 1775)

atricillus (Linnaeus, 1761) FKSNDEAIsuturellus (Duftschmid, 1825) FKSNDEAInasturtii (Fabricius, 1792) FKSNDEAIquadriguttatus (Pontoppidan, 1765) --S-DEA-apicalis (Beck, 1817) FKSNDEA-holsaticus (Linnaeus, 1758) FKSNDEAIluridus (Scopoli, 1763) FKSNDEAIfulgens (Foudras, 1860) F-----A-brunneus (Duftschmid, 1825) FKSNDEAIminusculus (Foudras, 1860) ------A-fuscoaeneus Redtenbacher, 1849 ------A-echii (Koch, 1803) -------Iniger (Koch, 1803) --S-DEAInigerrimus (Gyllenhal, 1827) FKSNDEAIparvulus (Paykull, 1799) FKSNDEAImedvedevi Shapiro, 1956 --S-----anchusae (Paykull, 1799) --S-DEAIabsynthii Kutschera, 1862 F-S---A-obliteratus (Rosenhauer, 1847) ------AI

Altica Geoffroy, 1762aenescens Weise, 1888 FKS--EAIampelophaga (Guérin-Ménéville, 1858)------A-lythri Aubé, 1843 F---D--Icarduorum (Guérin-Ménéville, 1858) F-S---A-

cirsii Israelson, 1956brevicollis Foudras, 1860 --S-DEA-quercetorum Foudras, 1860ssp. saliceti Weise, 1888 F-SN--AItamaricis Schrank, 1785 F---DEAIdeserticola Weise, 1889 ------A-engstromi J.Sahlberg, 1893 FKSN-EAIoleracea (Linnaeus, 1758) FKSNDEAIchamaenerii Håkan Lindberg, 1926 FKSN-EAIpalustris Weise, 1888 FKSNDEAIhelianthemi (Allard, 1859) -----EAI

pusilla Duftschmid, 1825, nec Gyllenhal, 1813longicollis (Allard, 1860) FKSNDEA-

britteni Sharp, 1914sandini Kemner, 1919

opacifrons Lindberg, 1938 FKS-----carinthiaca Weise, 1888 F--N--A-

Hermaeophaga Foudras, 1860mercurialis (Fabricius, 1792) --S-DE-I

Batophila Foudras, 1860rubi (Paykull, 1799) FKSNDEAIfallax Weise, 1888 ------A-


Lythraria Bedel, 1897salicariae (Paykull, 1800) FKSNDEAI

Ochrosis Foudras, 1860ventralis (Illiger, 1807) ------A-

Neocrepidodera Heikertinger, 1911Asiorestia Jacobson, 1925Crepidodera auct., nec Chevrolat, 1837

brevicollis (Daniel, 1904) --S-D---transversa (Marsham, 1802) --S-DEAIimpressa (Fabricius, 1801) --S-----interpunctata (Motschulsky, 1859) FK-ND-AImotschulskii (Konstantinov, 1991) FKSNDEAI

sublaevis auct. nec (Motschulsky, 1859)crassicornis (Faldermann, 1837) ------A-ferruginea (Scopoli, 1763) FKSNDEAIfemorata (Gyllenhal, 1813) FKSNDE-Inigritula (Gyllenhal, 1813) FKS--EAI

Derocrepis Weise, 1886rufipes (Linnaeus, 1758) -KS-DEAI

Hippuriphila Foudras, 1860modeeri (Linnaeus, 1761) FKSNDEAI

laevicollis Hellén, 1933

Crepidodera Chevrolat, 1837Chalcoides Foudras, 1860

aurea (Geoffroy, 1785) ----DEAIfulvicornis (Fabricius, 1792) FKSNDEAI

lapponica (Heikertinger, 1950)aurata (Marsham, 1802) FKSNDEAIplutus (Latreille, 1804) --s-D-A-lamina (Bedel, 1901) ---N--A-nitidula (Linnaeus, 1758) FKSNDEAI

Epitrix Foudras, 1860pubescens (Koch, 1803) FKS-DEAI

Podagrica Chevrolat, 1837fuscicornis (Linnaeus, 1767) ----D---

Mantura Stephens, 1831chrysanthemi (Koch, 1803) FKSNDEAIobtusata (Gyllenhal, 1813) F-SNDEA-

ambigua Kutschera, 1862rustica (Linnaeus, 1767) FKSNDEAI

Chaetocnema Stephens, 1831semicoerulea (Koch, 1803) -K---EAIconcinna (Marsham, 1802) FKSNDEAIlaevicollis (Thomson, 1866) FKSNDEAI

heikertingeri Lubischev, 1963

tibialis (Illiger, 1807) F-----A-obesa (Boieldieu, 1859) ------A-breviuscula (Faldermann, 1837) ------AIaerosa (Letzner, 1846) FKSN-EA-aridula (Gyllenhal, 1827) FKSNDEAIconfusa (Boheman, 1851) --S-D-A-mannerheimii (Gyllenhal, 1827) FKS--EAIarida Foudras, 1860 ------AIsubcoerulea (Kutschera, 1864) F-S-D-A-hortensis (Geoffroy, 1785) FKSNDEAIsahlbergii (Gyllenhal, 1827) FKSNDEAIcompressa (Letzner, 1846) ------A-

Dibolia Latreille, 1829depressiuscula Letzner, 1846 FKS-DEAIoccultans (Koch, 1803) --S-DEA-timida (Illiger, 1794) ------A-cryptocephala (Koch, 1803) -----EAIcynoglossi (Koch, 1803) ----DEA-

Sphaeroderma Stephens, 1831testaceum (Fabricius, 1775) F-SNDEAIrubidum (Graëlls, 1858) --S-D-AI

Argopus Fischer v. Waldheim, 1824ahrensi (Germar, 1817) ------A-

Apteropeda Chevrolat, 1837splendida Allard, 1860 ----D---globosa (Illiger, 1794) ----D---orbiculata (Marsham, 1802) ----D---

Mniophila Stephens, 1831muscorum (Koch, 1803) --SND--I

Psylliodes Berthold, 1827affinis (Paykull, 1799) FKSNDEAImarcida (Illiger, 1807) FKSNDEAIluteola (Müller, 1776) -----EA-picina (Marsham, 1802) FKSND-A-attenuata (Koch, 1803) FKS---AIchrysocephala (Linnaeus, 1758) FKSNDEAIsophiae Heikertinger, 1914 FKSNDE-Inapi (Fabricius, 1792) FKSNDEAIbrisouti Bedel, 1898 --S-----cuprea (Koch, 1803) --S-DEAIisatidis Heikertinger, 1912 F-SN----crambicola Lohse, 1954 F-S-D---cupreata (Duftschmid, 1825) ------AIhyoscyami (Linnaeus, 1758) FKSNDEAIchalcomera (Illiger, 1807) --S-DEAIdulcamarae (Koch, 1803) FKS-DEAI


cucullata (Illiger, 1807) FKSNDEAIreitteri Weise, 1888 ------A-

Hispinae Gyllenhal, 1813Hispa Linnaeus, 1767atra Linnaeus, 1767 --S-D--I

Cassidinae Stephens, 1831Pilemostoma Desbrochers des Loges, 1891fastuosa (Schaller, 1783) --S-D-AI

Hypocassida Weise, 1893subferruginea (Schrank, 1776) FK--DEAI

Cassida Linnaeus, 1758viridis Linnaeus, 1758 FKSNDEAIhemisphaerica Herbst, 1799 FKSNDEAImurraea Linnaeus, 1767 F-S-DEAInebulosa Linnaeus, 1758 FKSNDEAIflaveola Thunberg, 1794 FKSNDEAIberolinensis Suffrian, 1844 --S-----lineola Creutzer, 1799 ------A-atrata Fabricius, 1787 ------A-seladonia Gyllenhal, 1827 --S-D-AIvibex Linnaeus, 1767 FKSNDEAIpannonica Suffrian, 1844 ------AIpanzeri Weise, 1907 FKSN-EAIferruginea Goeze, 1777 --S--EA-rubiginosa Müller, 1776 FKSNDEAIstigmatica Suffrian, 1844 F---D-AIsanguinosa Suffrian, 1844 FKSNDEAIdenticollis Suffrian, 1844 FKSNDEAIsanguinolenta Müller, 1776 FKSNDEAIprasina Illiger, 1798 FKS---AIsubreticulata Suffrian, 1844 ------A-margaritacea Schaller, 1783 ----D-AInobilis Linnaeus, 1758 FKSNDEAIvittata Villers, 1789 --S-DEAI

Bruchinae Latreille, 1802Bruchini Latreille, 1802Bruchus Linnaeus, 1767loti Paykull, 1800 FKSNDEAIatomarius (Linnaeus, 1761) FKSNDEAIluteicornis Illiger, 1794 --S-----rufimanus Boheman, 1833 F-S-DEAIaffinis Frölich, 1799 F-SND-AIviciae Olivier, 1795 --S-D---pisorum (Linnaeus, 1758) FKSNdEAI

Acanthoscelidini Bridwell, 1946Acanthoscelides Schilsky, 1905obtectus (Say, 1831) fksnd-ai

obsoletus auct. nec (Say, 1831)

Bruchidius Schilsky, 1905marginalis (Fabricius, 1777) ------AIunicolor (Olivier, 1795) -------Ilividimanus (Gyllenhal, 1833) f-------villosus (Fabricius, 1792) --S-D---

fasciatus (Olivier, 1795)

Callosobruchus Pic, 1902chinensis (Linnaeus, 1758) f-snd---maculatus (Fabricius, 1775) ---nd---

Amblycerini Bridwell, 1932Spermophagus Schönherr, 1833sericeus (Geoffroy, 1785) f-S-D--I

CURCULIONOIDEANemonychidae Bedel, 1882Cimberidinae Des Gozis, 1882Cimberidini Des Gozis, 1882Cimberis Des Gozis, 1881Rhinomacer auct., nec Müller, 1764

attelaboides (Fabricius, 1787) FKSNDEAI

Doydirhynchini Pierce, 1916Doydirhynchus Dejean, 1821austriacus (Olivier, 1807) --SN--A-

Anthribidae Billberg, 1820Anthribinae Billberg, 1820Tropiderini Lacordaire, 1866Tropideres Schönherr, 1823albirostris (Herbst, 1783) --S----Idorsalis (Thunberg, 1796) FKSN-EAI

Stenocerini Kolbe, 1895Allandrus LeConte, 1876Tropiderinus auct., nec Reitter, 1916

undulatus (Panzer, 1795) FKSNDE--

Enedreytes Schönherr, 1839sepicola (Fabricius, 1792) --SND---

Zygaenodini Lacordaire, 1866Rhaphitropis Reitter, 1916marchicus (Herbst, 1797) ------A-

Dissoleucas Jordan, 1925niveirostris (Fabricius, 1798) FKSNDEAI


Platyrhinini Bedel, 1882Platyrhinus Clairville & Schellenberg, 1798resinosus (Scopoli, 1763) FKSNDEAI

Platystomini Pierce, 1916Platystomos Schneider, 1791albinus (Linnaeus, 1758) FKSNDEAI

Anthribini Billberg, 1820Anthribus Geoffroy, 1762Brachytarsus Schönherr, 1823

fasciatus Forster, 1770 F-S?DEA-scapularis (Gebler, 1833) FKSNDEAInebulosus Forster, 1770 FKSNDEAI

Choraginae Kirby, 1819Araecerus Schönherr, 1823coffeae (Fabricius, 1801) f-snd---

fasciculatus auct., nec (DeGeer, 1775)

Choragus Kirby, 1819sheppardi Kirby, 1819 FKS-D---horni Wolfrum, 1930 --SND-A-

Urodontinae Thomson, 1859Bruchela Dejean, 1821Urodon Schönherr, 1823

suturalis (Fabricius, 1793) -------I

Attelabidae Billberg, 1820Rhynchitinae Gistel, 1848Rhynchitini Gistel, 1848Lasiorhynchites Jekel, 1860sericeus (Herbst, 1797) ----D---cavifrons (Gyllenhal, 1833) ----D---olivaceus (Gyllenhal, 1833) --S-D---coeruleocephalus (Schaller, 1783) -------I

Temnocerus Thunberg, 1815Pselaphorhynchites Schilsky, 1903

nanus (Paykull, 1792) FKSNDEAItomentosus (Gyllenhal, 1839) FKSNDEAIlongiceps (Thomson, 1888) FKSND-AI

Neocoenorrhinus Voss, 1952Caenorhinus auct., nec Thomson, 1859

germanicus (Herbst, 1797) FKSNDEAIaeneovirens (Marsham, 1802) --SNDE-Iinterpunctatus (Stephens, 1831) --S-DE--pauxillus (Germar, 1824) --S-D--Iaequatus (Linnaeus, 1767) F-SNDEAI

Rhynchites Schneider, 1791cupreus (Linnaeus, 1758) FKSNDEAI

auratus (Scopoli, 1763) -----EAIbacchus (Linnaeus, 1758) -------I

Byctiscini Voss, 1923Byctiscus Thomson, 1859betulae (Linnaeus, 1758) FKSNDEAIpopuli (Linnaeus, 1758) FKSNDEAI

Deporaini Voss, 1929Deporaus Leach, 1819Caenorhinus Thomson, 1859

mannerheimii (Hummel, 1823) FKSNDEAIbetulae (Linnaeus, 1758) FKSNDEAI

Attelabinae Billberg, 1820Attelabus Linnaeus, 1758nitens (Scopoli, 1763) --S-DEAI

Apoderinae Jekel, 1860Apoderus Olivier, 1807coryli (Linnaeus, 1758) FKSNDEAIerythropterus (Gmelin, 1790) ------AI

Apionidae Schönherr, 1823Apioninae Schönherr, 1823Apion Herbst, 1797laevigatum (Paykull, 1792) FKSNDE--

sorbi (Fabricius, 1792)dispar Germar, 1817 --S-DEA-hookerorum Kirby, 1808 FKSNDEAIonopordi Kirby, 1808 FKSNDEAIpenetrans Germar, 1817 --S-DEAIbasicorne Illiger, 1807 -KS-D---

alliariae auct. nec (Linnaeus, 1758)carduorum Kirby, 1808 --S-D---gibbirostre Gyllenhal, 1813 FKSNDEAI

carduorum auct. nec Kirby, 1808armatum Gerstaecker, 1854 --S-DE--austriacum Wagner, 1904 ----DEA-confluens Kirby, 1808 FKSNDEAIstolidum Germar, 1817 FKSNDEAIdetritum Mulsant & Rey, 1859 ----D---sulcifrons Herbst, 1797 --S-DEAIaeneum (Fabricius, 1775) --S-D--Iradiolus (Marsham, 1802) FKSNDEAIholosericeum Gyllenhal, 1833 ------A-minimum Herbst, 1797 FKSNDEAIelongatum Germar, 1817 -----EAIcineraceum Wencker, 1864 FKS--E-Iflavimanum Gyllenhal, 1833 --S-DE--vicinum Kirby, 1808 FKSNDE--


origani Planet, 1918 --S-----atomarium Kirby, 1808 FKS-DEA-oblivium Schilsky, 1902 --S-D---hoffmanni Wagner, 1930 F-S-----pallipes Kirby, 1808 --S-DEA-urticarium (Herbst, 1784) FKSNDEAIrufirostre (Fabricius, 1775) --S-D---compactum Desbrochers des Loges, 1888 ----D---difficile Herbst, 1797 ----D---elongatulum Desbrochers des Loges, 1891 -------Ifuscirostre (Fabricius, 1775) --S-D---astragali (Paykull, 1800) --SNDEAIgracilipes Dietrich, 1857 F-S----Ifulvipes (Geoffroy, 1785) FKSNDEAI

dichroum Bedel, 1885flavipes (Paykull, 1792) nec (DeGeer, 1775)

nigritarse Kirby, 1808 --S-DEA-filirostre Kirby, 1808 --S-DEAItrifolii (Linnaeus, 1768) FKSNDEAI

aestivum Germar, 1817ruficrus Germar, 1817 -----EA-interjectum Desbrochers des Loges, 1895 FKSND---

boreum Gönget, 1997apricans Herbst, 1797 FKSNDEAIvaripes Germar, 1817 -KSNDE-Iassimile Kirby, 1808 FKSNDEAIononidis Gyllenhal, 1827 -KSNDEA-

ononicola Bach, 1854dissimile Germar, 1817 --S-DEA-brevirostre Herbst, 1797 ----D---simum Germar, 1817 FKS-DEAIviolaceum Kirby, 1808 FKSNDEAIhydrolapathi (Marsham, 1802) --S-D---laticeps Desbrochers des Loges, 1870 FK------

lindbergi Wagner, 1943marchicum Herbst, 1797 FKSNDEAIaffine Kirby, 1808 FKSNDEA-sedi Germar, 1818 FKSNDEAIcurtirostre Germar, 1817 FKSNDEAIoblongum Gyllenhal, 1827 ------A-frumentarium (Linnaeus, 1758) --SNDEAI

sanguineum (DeGeer, 1775)miniatum Germar, 1833

rubiginosum Grill, 1893 FKSNDEAIsanguineum auct. nec (DeGeer, 1775)

rubens Stephens, 1839 FKSNDEAIseniculus Kirby, 1808 FKSNDEAImeieri (Desbrochers des Loges, 1901) FKSNDE--pubescens Kirby, 1811 --S-DE--simile Kirby, 1811 FKSNDEAItenue Kirby, 1808 --SNDEAImeliloti Kirby, 1808 F-SNDEA-loti Kirby, 1808 FKSNDEAImodestum Germar, 1817 --S-D---

plumbeomicans auct. nec Rosenhauer, 1856sicardi auct. nec Desbrochers des Loges, 1893

virens Herbst, 1797 FKSNDEAIatratulum Kirby, 1808 ----D---ebeninum Kirby, 1808 FKSNDEAIpisi (Fabricius, 1801) F-S-DEAIaestimatum Faust, 1890 ------AIaethiops Herbst, 1797 FKSNDEAIononis Kirby, 1808 --S-D--Ipavidum Germar, 1817 -----EAIimmune Kirby, 1808 ----D---punctirostre Gyllenhal, 1839 -------Iscandinavicum Dieckmann, 1977 F-SN----brundini Wagner, 1943 FKSN----columbinum Germar, 1817 F-SNDE--spencii Kirby, 1808 FKSNDEAIafer Gyllenhal, 1833 FKSN-EA-

platalea auct. nec Germar, 1817gnarum Faust, 1890 -K------gyllenhali Kirby, 1808 FKSNDEAIsubulatum Kirby, 1808 FKSNDEAIopeticum Bach, 1854 FKS-DEAIcraccae (Linnaeus, 1767) --SNDEAIcerdo Gerstaecker, 1854 FKSNDEAIpomonae (Fabricius, 1798) --S-D-AIflavifemoratum Herbst, 1797 ----D---viciae (Paykull, 1800) FKSNDEAI

voisini (Ehret, 1997)ervi Kirby, 1808 FKSNDEAImelancholicum Wencker, 1864 F-SND-A-vorax Herbst, 1797 --S-D-AIpunctigerum (Paykull, 1792) F-SNDE-Ifacetum Gyllenhal, 1839 FKSNDEA-

Nanophyinae Gistel, 1848Nanophyes Schönherr, 1838Nanodes Schönherr, 1825

circumscriptus Aubé, 1864 F-S--EA-

haematodes Kirby, 1808 FKSNDEAIfrumentarium (Paykull, 1792) nec (Linnaeus, 1758)

cruentatum Walton, 1844 F-SNDEAI


marmoratus (Goeze, 1777) FKSNDEAIglobulus (Germar, 1821) --S-----sahlbergi (Sahlberg, 1835) FKS-DE--

Curculionidae Latreille, 1802Entiminae Schönherr, 1823Otiorhynchini Schönherr, 1826Otiorhynchus Germar, 1822lugdunensis Boheman, 1843 --S-DEA-fuscipes (Olivier, 1807) ----dE--arcticus (O.Fabricius, 1780) FKSNd---politus Gyllenhal, 1834 FK------raucus (Fabricius, 1777) FKSNDEAInodosus (Müller, 1764) FKSNDEAI

dubius (Ström, 1783)rugosostriatus (Goeze, 1777) --SND---porcatus (Herbst, 1795) F-SNDE--scaber (Linnaeus, 1758) FKSNDEAIuncinatus Germar, 1824 ----D---ligneus (Olivier, 1807) --SND---singularis (Linnaeus, 1767) FKSNDEAIlepidopterus (Fabricius, 1794) --SN----

salicis (Ström, 1788) nec (Linnaeus, 1758)sulcatus (Fabricius, 1775) F-SNDEA-rugifrons (Gyllenhal, 1813) FKSND-A-smreczynskii Cmoluch, 1968 ----D---rotundus Marseul, 1872 -------I

rotundatus Siebold, 1837, nec (Gmelin, 1790)desertus Rosenhauer, 1847 FKSNDEA-crataegi Germar, 1824 ----D---ovatus (Linnaeus, 1758) FKSNDEAItristis (Scopoli, 1763) F-S-DEAIaurosparsus Germar, 1824 ------A-ligustici (Linnaeus, 1758) FKSNDEAIatroapterus (DeGeer, 1775) --SND--Idieckmanni Magnano, 1979 --S-----conspersus (Herbst, 1795) ------A-orbicularis (Herbst, 1795) -------I

Peritelini Lacordaire, 1863Simo Dejean, 1821Homorhythmus Bedel, 1883Peritelus auct., nec Germar, 1824

hirticornis (Herbst, 1795) --SND---variegatus (Boheman, 1843) ----D---

Trachyphloeini Lacordaire, 1863Trachyphloeus Germar, 1817alternans Gyllenhal, 1834 --S-D---

rectus Thomson, 1865 F-SND---laticollis auct. nec Boheman, 1843

spinimanus Germar, 1824 --S-D-A-digitalis (Gyllenhal, 1827) --S-DEA-heymesi Hybenthal, 1934 --S-D---scabriculus (Linnaeus, 1771) -KS-DE-Ibifoveolatus (Beck, 1817) FKSNDEAIangustisetulus Hansen, 1915 -KS-D-A-aristatus (Gyllenhal, 1827) FKSNDEAI

Omiini Shuckard, 1840Mylacini Reitter, 1913

Omiamima Silfverberg, 1977Omias auct., nec Germar, 1817

mollina (Boheman, 1834) --SND---

Phyllobiini Schönherr, 1826Phyllobius Germar, 1824cinerascens (Fabricius, 1792) -------Iviridicollis (Fabricius, 1792) FKSNDEA-brevis Gyllenhal, 1834 -------Iroboretanus Gredler, 1882 -----E-I

parvulus (Olivier, 1807) nec (Fabricius, 1792)virideaeris (Laicharting, 1781) FKSNDEAIoblongus (Linnaeus, 1758) FKSNDEAIpyri (Linnaeus, 1758) FKSNDEAIvespertinus (Fabricius, 1792) --S-DEA-maculicornis Germar, 1824 FKSNDEAIargentatus (Linnaeus, 1758) FKSNDEAIbetulinus (Bechstein & Scharfenberg, 1805) ------AI

betulae (Fabricius, 1801) nec (Linnaeus, 1758)arborator (Herbst, 1797) FK---EAIscutellaris Redtenbacher, 1849 -K------glaucus (Scopoli, 1763) FKSNDEAI

calcaratus (Fabricius, 1792)pomaceus Gyllenhal, 1834 FKSNDEAI

urticae (DeGeer, 1775) nec (Scopoli, 1763)

Polydrosini Schönherr, 1823Polydrusus Germar, 1817marginatus Stephens, 1831 ---N----pallidus (Gyllenhal, 1834) ----DEAI

atomarius (Olivier, 1807) nec (Linnaeus, 1761)pterygomalis Boheman, 1840 -KS-DEA-corruscus Germar, 1824 -----EAIflavipes (DeGeer, 1775) FKSNDEAIimpressifrons Gyllenhal, 1834 --S---A-cervinus (Linnaeus, 1758) F-SNDEAIpilosus Gredler, 1866 FKSN-EAIpulchellus Stephens, 1831 ----D---


confluens Stephens, 1831 ----D---undatus (Fabricius, 1781) FKSNDEAIfulvicornis (Fabricius, 1792) FKSN-EAI

ruficornis (Bonsdorff, 1785) nec (Linnaeus, 1758)picus (Fabricius, 1792) -------Iformosus (Mayer, 1779) --S-D--I

splendidus (Herbst, 1784)sericeus (Schaller, 1783) nec (Goeze, 1777)

mollis (Ström, 1768) FKSNDEAI

Sciaphilini Sharp, 1891Sciaphobus Daniel, 1904rubi (Gyllenhal, 1813) --S-----

Sciaphilus Schönherr, 1823asperatus (Bonsdorff, 1785) FKSNDEAI

Brachysomus Schönherr, 1823echinatus (Bonsdorff, 1785) FKSNDEAI

Foucartia Jacquelin du Val, 1854squamulata (Herbst, 1795) --S-----

Barypeithes Jacquelin du Val, 1854chevrolati (Boheman, 1843) --S-----pellucidus (Boheman, 1834) FKSNDEA-trichopterus (Gautier des Cottes, 1863) --S-DEAImollicomus (Ahrens, 1812) FKSNDE--araneiformis (Schrank, 1781) --sNDE--

Brachyderini Schönherr, 1826Brachyderes Schönherr, 1823incanus (Linnaeus, 1758) FKSNDEAI

Strophosoma Billberg, 1820Neliocarus Thomson, 1859

fulvicorne Walton, 1848 F-SND-A-curvipes Thomson, 1865

melanogrammum (Forster, 1771) F-SNDEAIseptentrionale Pelletier, 1995

capitatum (DeGeer, 1775) FKSNDEAIrufipes Stephens, 1831

faber (Herbst, 1785) --SND-AIsus Stephens, 1831 --SND---

laterale (Paykull, 1792) nec (Panzer, 1789)

Cneorhinini Lacordaire, 1863Liophloeus Germar, 1817tessulatus (Müller, 1776) F-SNDEAI

Attactagenus Tournier, 1876Cneorhinus auct., nec Schönherr, 1823

plumbeus (Marsham, 1802) --S-D---exaratus (Marsham, 1802) nec (Gmelin, 1790)

Philopedon Schönherr, 1826plagiatus (Schaller, 1783) FKSNDEAI

Geonemini Gistel, 1856Barynotus Germar, 1817obscurus (Fabricius, 1775) FKSNDEAIsquamosus Germar, 1824 --SND---moerens (Fabricius, 1792) FKSNDEA-

Tropiphorini Marseul, 1863Tropiphorus Schönherr, 1842elevatus (Herbst, 1795) FKSNDEAI

carinatus (Müller, 1776) nec (Linnaeus, 1767)terricola (Newman, 1838) F-S-D---

tomentosus (Marsham, 1802) nec (Olivier, 1790)obtusus (Bonsdorff, 1785) -KSNDE-I

Tanymecini Lacordaire, 1863Cycloderes Sahlberg, 1823pilosulus (Herbst, 1795) -------I

Chlorophanus Sahlberg, 1823viridis (Linnaeus, 1758) FK--DEAIgraminicola (Schönherr, 1832) -----EAI

Tanymecus Germar, 1817palliatus (Fabricius, 1787) FKS-DEAI

Sitonini Gistel, 1856Sitona Germar, 1817gressorius (Fabricius, 1792) --S----Igriseus (Fabricius, 1775) --SNDEAIcambricus Stephens, 1831 ----DE--cinerascens (Fåhraeus, 1840) --S-D---regensteinensis (Herbst, 1797) F-S-D---striatellus Gyllenhal, 1834 -K--DEAI

tibialis (Herbst, 1795) nec (Sparrman, 1787)lineatus (Linnaeus, 1758) FKSNDEAIinops Schönherr, 1832 -----EA-suturalis Stephens, 1831 FKSNDEAIlateralis Gyllenhal, 1834 FKSND-A-

ononidis Sharp, 1866sulcifrons (Thunberg, 1798) FKSNDEAIpuncticollis Stephens, 1831 FKSNDEAIlepidus Gyllenhal, 1834 FKSNDEAI

flavescens (Marsham, 1802) nec (Fabricius, 1787)longulus Gyllenhal, 1834 -----EA-lineellus (Bonsdorff, 1785) FKSNDEAI

decipiens Lindberg, 1933ambiguus Gyllenhal, 1834 FKSNDEAImacularius (Marsham, 1802) FK--DEAI

crinitus (Herbst, 1795) nec (Gmelin, 1790)


waterhousei Walton, 1846 ----D---hispidulus (Fabricius, 1777) FKSNDEAIcylindricollis (Fåhraeus, 1840) F-SNDEAIhumeralis Stephens, 1831 F-SNDEAI

Lixinae Schönherr, 1823Lixini Schönherr, 1823Larinus Dejean, 1821planus (Fabricius, 1792) ---NDEAIsturnus (Schaller, 1783) FKS--EAIjaceae (Fabricius, 1775) -----EA-

Lixus Fabricius, 1801paraplecticus (Linnaeus, 1758) FKS-DEAIiridis Olivier, 1807 FKSNDEAIalbomarginatus Boheman, 1843 -------I

ascanii auct. nec Linnaeus, 1767myagri Olivier, 1807 ----D---punctiventris Boheman, 1836 ------?Ibardanae (Fabricius, 1787) --SN-EAI

Rhinocyllini Lacordaire, 1863Rhinocyllus Germar, 1819conicus (Frölich, 1792) -----E-I

Cleonini Schönherr, 1826Temnorhinus Chevrolat, 1873hololeucus (Pallas, 1781) -------I

Coniocleonus Motschulsky, 1860hollbergi (Fåhraeus, 1842) FKSNDEAI

turbatus (Fåhraeus, 1842)glaucus (Fabricius, 1787) nec (Scopoli, 1763)

nebulosus (Linnaeus, 1758) FKSNDEAI

Stephanocleonus Motschulsky, 1860tetragrammus (Pallas, 1781) ------A-

Bothrynoderes Schönherr, 1823Chromoderus Motschulsky, 1860

affinis (Schrank, 1781) FKSNDEAIfasciatus (Müller, 1776) nec (Scopoli, 1763)

Pseudocleonus Chevrolat, 1873grammicus (Panzer, 1789) --S-----cinereus (Schrank, 1781) -------I

Cyphocleonus Motschulsky, 1860dealbatus (Gmelin, 1790) ----DEAI

tigrinus (Panzer, 1789) nec (Geoffroy, 1785)trisulcatus (Herbst, 1795) FKS-DE-I

Cleonis Dejean, 1821pigra (Scopoli, 1763) FKSNDEAI

Cyclominae Schönherr, 1826Rhytirrhinini Lacordaire, 1863Gronops Schönherr, 1823lunatus (Fabricius, 1775) --S-DE--inaequalis Boheman, 1842 FKS-DEAI

Hyperinae Marseul, 1863Hypera Germar, 1817Phytonomus Schönherr, 1823

zoilus (Scopoli, 1763) FKSNDEAIpunctata (Fabricius, 1775)

vidua Gené, 1837 --SN----dauci (Olivier, 1807) -KS-DEAI

fasciculata (Herbst, 1795) nec (DeGeer, 1775)arundinis (Paykull, 1792) --S-DEAIpollux (Fabricius, 1796) FKSNDEAI

adspersa (Fabricius, 1793) nec (Fabricius, 1775)rumicis (Linnaeus, 1758) FKSNDEAIcontaminata (Herbst, 1795) ------A-meles (Fabricius, 1792) FKSNDEAIfornicata (Penecke, 1928) F-S-----arator (Linnaeus, 1758) FKSNDEAIdiversipunctata (Schrank, 1798) FKSNDEAI

elongata (Paykull, 1792) nec (Fabricius, 1775)suspiciosa (Herbst, 1795) FKSNDEAI

pedestris (Paykull, 1792) nec (Poda, 1761)obovata (Csiki, 1934) FKSN-E--denominanda (Capiomont, 1868) FKSND---postica (Gyllenhal, 1813) FKSNDEAI

variabilis (Herbst, 1795) nec (Fabricius, 1777)fuscocinerea (Marsham, 1802) -KS-DEAI

murina (Fabricius, 1793) nec (Müller, 1764)plantaginis (DeGeer, 1775) F-SNDEAInigrirostris (Fabricius, 1775) FKSNDEAIvenusta (Fabricius, 1781) --SNDEAI

trilineata (Marsham, 1802)viciae (Gyllenhal, 1813) FKSNDEAI

Limobius Schönherr, 1847borealis (Paykull, 1792) FKSNDE--

Erirhininae Schönherr, 1825Erirhinini Schönherr, 1825

Notarini Zumpt, 1926Grypus Germar, 1817equiseti (Fabricius, 1775) FKSNDEAIbrunnirostris (Fabricius, 1792) --S-DEAI

Thryogenes Bedel, 1884fiorii Zumpt, 1928 ----D---

atrirostris Lohse, 1992


festucae (Herbst, 1795) FKSNDEAInereis (Paykull, 1800) FKSNDEAIscirrhosus (Gyllenhal, 1836) FKSNDEA-

Notaris Germar, 1817bimaculatus (Fabricius, 1787) -KSNDEAIscirpi (Fabricius, 1792) FKSNDEAIacridulus (Linnaeus, 1758) FKSNDEAIaethiops (Fabricius, 1792) FKSNDEAImaerkeli (Boheman, 1843) -----EAI

Tanysphyrinae Gistel, 1856Tanysphyrus Germar, 1817lemnae (Paykull, 1792) FKSNDEAI

callae Voss, 1943ater Blatchley, 1928 FKS-----

makolskii Smreczynski, 1957

Curculioninae Latreille, 1802Ellescini Thomson, 1859Ellescus Dejean, 1821scanicus (Paykull, 1792) FKSNDEAIbipunctatus (Linnaeus, 1758) FKSNDEAIinfirmus (Herbst, 1795) -----EAI

Dorytomus Germar, 1817longimanus (Forster, 1771) --S-D-AIschoenherri Faust, 1882 -----EAInordenskioldi Faust, 1882 -K---E-Itremulae (Fabricius, 1787) FKSNDEAItortrix (Linnaeus, 1761) FKSNDEAIfilirostris (Gyllenhal, 1836) --S-DE--minutus (Gyllenhal, 1836) -----EA-suratus (Gyllenhal, 1836) ------AI

flavipes (Panzer, 1797) nec (DeGeer, 1775)lapponicus J.Sahlberg, 1892 F-SN----ictor (Herbst, 1795) F-S-D-AI

validirostris (Gyllenhal, 1836)hirtipennis (Bedel, 1884) F-SNDE-Iedoughensis Desbrochers des Loges, 1875 FKSNDEAI

affinis (Paykull, 1800) nec (Schrank, 1781)reussi Formanek, 1908 ------A-dejeani Faust, 1882 F-SNDEAItaeniatus (Fabricius, 1781) FKSNDEAIoccallescens (Gyllenhal, 1836) ------A-melanophthalmus (Paykull, 1792) FKSNDEAIrufatus (Bedel, 1886) FKSNDEAI

rufulus (Bedel, 1884) nec (Mannerheim, 1853)salicinus (Gyllenhal, 1827) FKSNDEA-salicis Walton, 1851 FKSNDEAI

majalis (Paykull, 1792) FKSNDEA-dorsalis (Linnaeus, 1758) FKSNDEAI

Styphlini Jekel, 1861Comasinus Dejean, 1821Orthochaetes Germar, 1824

setiger (Beck, 1817) --S-D---

Pseudostyphlus Tournier, 1874pillumus (Gyllenhal, 1836) FKS-DEA-

Smicronychini Seidlitz, 1891Smicronyx Schönherr, 1843reichii (Gyllenhal, 1836) --S-----jungermanniae (Reich, 1797) --SND---smreczynskii Solari, 1952 FKS-D---

politus Thomson, 1865, nec Boheman, 1843coecus (Reich, 1797) ----DE--

Cionini Schönherr, 1825Cionus Clairville & Schellenberg, 1798alauda (Herbst, 1784) ---N----tuberculosus (Scopoli, 1763) FKSNDEAIscrophulariae (Linnaeus, 1758) FKSNDEAIhortulanus (Geoffroy, 1785) FKSNDEAIlongicollis Brisout de Barneville, 1863 FKSN-EAI

montanus Wingelmüller, 1914olivieri Rosenschoeld, 1838 -----EA-nigritarsis Reitter, 1904 FKSNDEA-

Stereonychus Suffrian, 1854fraxini (DeGeer, 1775) FKS-DEAI

Cleopus Dejean, 1821pulchellus (Herbst, 1795) F-SNDEA-solani (Fabricius, 1792) ------A-

Tychiini Gistel, 1848Tychius Germar, 1817quinquepunctatus (Linnaeus, 1758) FKSNDEAIparallelus (Panzer, 1794) --S-D--I

venustus (Fabricius, 1787) nec (Fabricius, 1781)polylineatus (Germar, 1824) FKSNDEAIschneideri (Herbst, 1795) -KS-D-AIsquamulatus Gyllenhal, 1836 --SND-AI

flavicollis auct. nec Stephens, 1831medicaginis Brisout de Barneville, 1862------A-aureolus Kiesenwetter, 1851 ------A-trivialis Boheman, 1843 ----DE--

kiesenwetteri Tournier, 1873crassirostris Kirsch, 1871 ----D---junceus (Reich, 1797) --S-D-AI


breviusculus Desbrochers des Loges, 1873 F-S--EAImicaceus Rey, 1895haematopus auct. nec Gyllenhal, 1836

meliloti Stephens, 1831 F-SNDEAIstephensi Gyllenhal, 1836 FKSNDEAI

tomentosus (Herbst, 1795) nec (Olivier, 1790)sharpi Tournier, 1873 -----EA-picirostris (Fabricius, 1787) FKSNDEAIlineatulus Stephens, 1831 --S-DEA-

Sibinia Germar, 1817sodalis Germar, 1824 ----D---phalerata (Gyllenhal, 1836) --S-DE-Iprimita (Herbst, 1795) FKSND-A-pyrrhodactyla (Marsham, 1802) FKSND-AI

potentillae Germar, 1824pellucens (Scopoli, 1772) FK---EAIviscariae (Linnaeus, 1761) FKSN-EAI

Acalyptini Thomson, 1859Acalyptus Schönherr, 1833carpini (Fabricius, 1792) FKSNDEAIsericeus Gyllenhal, 1836 FKS-D---

Anthonomini Thomson, 1859Anthonomus Germar, 1817Furcipus Desbrochers des Loges, 1868

pomorum (Linnaeus, 1758) -KSNDEAIhumeralis (Panzer, 1795) FKSNDEA-piri Kollar, 1837 --S---AIulmi (DeGeer, 1775) --S-DEAIbituberculatus Thomson, 1868 --SNDE--spilotus Redtenbacher, 1849 --S-----undulatus Gyllenhal, 1836 FKSN-E--pedicularius (Linnaeus, 1758) FKSNDEAIconspersus Desbrochers des Loges, 1868 F-SNDEA-rufus Gyllenhal, 1836 --S-D--Isorbi Germar, 1821 F-S-DEA-rubi (Herbst, 1795) FKSNDEAIbrunnipennis Curtis, 1840 FKSNDEAIphyllocola (Herbst, 1795) FKSNDEAI

varians (Paykull, 1792) nec (Gmelin, 1790)pinivorax Silfverberg, 1977 FKS--EAI

pubescens (Paykull, 1792) nec (Fabricius, 1775)rectirostris (Linnaeus, 1758) FKSNDEAI

Brachonyx Schönherr, 1825pineti (Paykull, 1792) FKSNDEAI

Bradybatus Germar, 1824kellneri Bach, 1854 --SNDEAI

Curculionini Latreille, 1802Curculio Linnaeus, 1758venosus (Gravenhorst, 1807) --S-D-AInucum Linnaeus, 1758 F-SNDEAIglandium Marsham, 1802 ----DEAIvillosus Fabricius, 1781 --SNDEAIbetulae (Stephens, 1831) FKSNDEAI

cerasorum Paykull, 1792, nec Fabricius, 1775rubidus (Gyllenhal, 1836) -------I

Archarius Gistel, 1848crux (Fabricius, 1777) FKSND-AIsalicivorus (Paykull, 1792) FKSNDEAIpyrrhoceras (Marsham, 1802) --SNDEAI

Rhamphini Rafinesque, 1815Rhynchaenus Clairville & Schellenberg, 1798Orchestes Illiger, 1804

rufus (Schrank, 1781) --S-D-AIalni (Linnaeus, 1758) -K--DEAI

saltator (Geoffroy, 1785)pilosus (Fabricius, 1781) --SNDEAIquercus (Linnaeus, 1758) FKSND-AIsignifer (Creutzer, 1799) --SND-AI

avellanae (Donovan, 1797) nec (Paykull, 1792)iota (Fabricius, 1787) FKSNDEAIlonicerae (Herbst, 1795) FK---EAIfagi (Linnaeus, 1758) F-SNDEA-testaceus (Müller, 1776) FKSNDEAIcalceatus (Germar, 1821) FKSNDE--rusci (Herbst, 1795) FKSNDEAIpratensis (Germar, 1821) ------AIermischi Dieckmann, 1958 -----E--cinereus Fåhraeus, 1843 -------I

Tachyerges Schönherr, 1826decoratus (Germar, 1821) FKSNDEAIrufitarsis (Germar, 1821) F-SND-AIsalicis (Linnaeus, 1758) FKSNDEAIstigma (Germar, 1821) FKSNDEAIpseudostigma (Tempère, 1982) F-SND-A-

Isochnus Thomson, 1859populicola (Silfverberg, 1977) FKSNDEAI

populi (Fabricius, 1793) nec (Linnaeus, 1758)foliorum (Müller, 1764) FKSNDEAI

saliceti (Paykull, 1792)flagellum (Ericson, 1902) FKSN----angustifrons (West, 1917) ---ND---


Rhamphus Clairville & Schellenberg, 1798pulicarius (Herbst, 1795) FKSNDEAIoxyacanthae (Marsham, 1802) FKSND-A-subaeneus (Illiger, 1807) -------I

Mecinini Gistel, 1848Gymnetron Schönherr, 1825rostellum (Herbst, 1795) FKS-DEA-stimulosum (Germar, 1821) ------A-melanarium (Germar, 1821) F-SNDEAIvillosulum Gyllenhal, 1838 --S-DEAIbeccabungae (Linnaeus, 1761) FKSNDEAIveronicae (Germar, 1821) FKSNDEA-

Rhinusa Stephens, 1829tetrum (Fabricius, 1792) -------Iantirrhini (Paykull, 1800) FKSNDEAIhispida (Brullé, 1832) --S-D---netum (Germar, 1821) -------Ithapsicola (Germar, 1821) --S-D--Icollina (Gyllenhal, 1813) -KSNDEA-linariae (Panzer, 1793) -KSNDEAI

Mecinus Germar, 1821collaris Germar, 1821 FKSNDEAIianthinus Germar, 1821 --S---A-heydeni Wencker, 1866 --S-D-A-pyraster (Herbst, 1795) --SNDEAI

Miarini Pierce, 1919Miarus Schönherr, 1826labilis (Herbst, 1795) --SNDEAIpascuorum (Gyllenhal, 1813) FKS-DEAIportai Solari, 1947 --SN----

confusus Kangas, 1976thuleus Kangas, 1980 FK------campanulae (Linnaeus, 1767) FKSNDEAI

frigidus Franz, 1947fennicus Kangas, 1978

Cleopomiarus Pierce, 1919Miaromimus Solari, 1947

micros (Germar, 1821) --S-DEA-distinctus (Boheman, 1845) FK---EAI

longirostris auct. nec Gyllenhal, 1813graminis (Gyllenhal, 1813) FKSNDEAIdulcinasutus (Kangas, 1976) FK------graminoides (Kangas, 1976) F-------

Anoplini Bedel, 1884Anoplus Germar, 1820plantaris (Naezen, 1794) FKSNDEAI

roboris Suffrian, 1840 FKSNDEAIsetulosus Kirsch, 1870 -------I

Bagoinae Thomson, 1859Dicranthus Motschulsky, 1845elegans (Fabricius, 1801) --S-DE-I

Hydronomus Schönherr, 1825alismatis (Marsham, 1802) FKSNDEA-

Bagous Germar, 1817Ephimeropus Hochhuth, 1847

petro (Herbst, 1795) F-S-DEA-tubulus Caldara & O'Brien, 1994 --S-D-AI

scanicus Silfverberg, 1997angustus Silfverberg, 1977, nec Tanner, 1954cylindrus (Paykull, 1800) nec (Fabricius, 1781)

friwaldszkyi Tournier, 1874 -K---E--binodulus (Herbst, 1795) -KS-D-AInodulosus Gyllenhal, 1836 --S-DE--argillaceus Gyllenhal, 1836 --S-DE--limosus (Gyllenhal, 1827) F-SNDE--subcarinatus Gyllenhal, 1836 --S-DE--longitarsis Thomson, 1868 FKS-DE--collignensis (Herbst, 1797) F-S-D---claudicans Boheman, 1845 --S-----frit (Herbst, 1795) FKSNDE--lutulosus (Gyllenhal, 1827) FKSNDEA-brevis Gyllenhal, 1836 F-SND---diglyptus Boheman, 1845 --SNDE--

curtus Gyllenhal, 1845brevitarsis Hansen, 1917

tempestivus (Herbst, 1795) FKS-DEA-thomsoni Bruce, 1968sjobergi Bruce, 1968

czwalinai Seidlitz, 1891 F-S-D---lutosus (Gyllenhal, 1813) FKSNDEAIpuncticollis Boheman, 1845 F-S-D---robustus Brisout de Barneville, 1863 F-S-D---lutulentus (Gyllenhal, 1813) FKSNDEAIglabrirostris (Herbst, 1795) FKSNDEA-

Dryophthorinae Schönherr, 1825Dryophthorus Germar, 1824corticalis (Paykull, 1792) FKSNDEAI

Cossoninae Schönherr, 1825Pentarthrini Lacordaire, 1866Pentarthrum Wollaston, 1854huttoni Wollaston, 1854 ----D---


Euophryum Broun, 1909rufum (Broun, 1880) ---nD---confine (Broun, 1881) ---nD---

Cossonini Schönherr, 1825Rhopalomesites Wollaston, 1873tardii (Curtis, 1825) ---N----

Cossonus Clairville & Schellenberg, 1798linearis (Fabricius, 1775) --S-D--Iparallelepipedus (Herbst, 1795) FKSNDEAIcylindricus Sahlberg, 1835 FKS--EAI

Rhyncolini Gistel, 1848Rhyncolus Germar, 1817Eremotes Wollaston, 1861

reflexus Boheman, 1838 -----EA-elongatus (Gyllenhal, 1827) FKSN-EAIater (Linnaeus, 1758) FKSNDEAI

chloropus auct. nec (Linnaeus, 1758)punctatulus Boheman, 1838 --S-----sculpturatus Waltl, 1839 FKSNdEA-

nitidipennis Thomson, 1868

Phloeophagus Schönherr, 1838lignarius (Marsham, 1802) F-SNDEA-turbatus Schönherr, 1845 FKSN-EA-thomsoni (Grill, 1896) --S-DE--

Onycholipini Wollaston, 1873Brachytemnus Wollaston, 1873porcatus (Germar, 1824) --S-d-AI

Stereocorynes Wollaston, 1873truncorum (Germar, 1824) --S-D-AI

Pselactus Broun, 1886spadix (Herbst, 1795) -KS-DEAI

Pseudophloeophagus Wollaston, 1873Caulotrupodes Voss, 1955

aeneopiceus (Boheman, 1845) -------I

Mesoptiliinae Lacordaire, 1863Magdalidini Pascoe, 1870Magdalis Germar, 1817phlegmatica (Herbst, 1797) FKSNDE-Irufa Germar, 1824 ------A-nitida (Gyllenhal, 1827) FKSN-EAImemnonia (Gyllenhal, 1837) ----DEAIlinearis (Gyllenhal, 1827) FKSNDEAIduplicata Germar, 1819 FKSNDEAI

weisei Schreiner, 1882frontalis (Gyllenhal, 1827) FKSNDEAI

violacea (Linnaeus, 17587) FKSNDEAIarmigera (Geoffroy, 1785) --SND-AIcarbonaria (Linnaeus, 1758) FKSNDEAIcaucasica Tournier, 1872 ------A-cerasi (Linnaeus, 1758) --SNDE-Iexarata (Brisout de Barneville, 1862) ----D---fuscicornis (Desbrochers des Loges, 1870) -------Ibarbicornis (Latreille, 1804) F-SNDEA-

mixta (Desbrochers des Loges, 1870)flavicornis (Gyllenhal, 1836) --S-D--Iruficornis (Linnaeus, 1758) FKSNDEAInitidipennis (Boheman, 1843) -------I

Molytinae Schönherr, 1823Lepyrini Kirby, 1837Lepyrus Germar, 1817palustris (Scopoli, 1763) ----DEAIcapucinus (Schaller, 1783) --S-DEAIarcticus (Paykull, 1792) FKSN----

Hylobiini Kirby, 1837Hylobius Germar, 1817piceus (DeGeer, 1775) FKSN-EAIabietis (Linnaeus, 1758) FKSNDEAIpinastri (Gyllenhal, 1813) FKSNDEAItransversovittatus (Goeze, 1777) FKSND---

Molytini Schönherr, 1823Liparini Latreille, 1828

Liparus Olivier, 1807coronatus (Goeze, 1777) --S-DEAIglabrirostris Küster, 1849 ----d--I

Leiosoma Stephens, 1829deflexum (Panzer, 1795) --SND---

Pissodini Gistel, 1848Pissodes Germar, 1817piceae (Illiger, 1807) ------AIcastaneus (DeGeer, 1775) FKSNDEAI

notatus (Fabricius, 1787) nec (Bonsdorff, 1785)pini (Linnaeus, 1758) FKSNDEAIgyllenhalii (Sahlberg, 1834) FKSN-E--validirostris (Sahlberg, 1834) FKSNDEAIharcyniae (Herbst, 1795) FKSN-EAIpiniphilus (Herbst, 1797) FKSNdEAI

Trachodini Gistel, 1848Trachodes Germar, 1824hispidus (Linnaeus, 1758) FKSNDEAI


Cryptorhynchinae Schönherr, 1825Cryptorhynchus Illiger, 1807lapathi (Linnaeus, 1758) FKSNDEAI

Acalles Schönherr, 1825roboris Curtis, 1835 --SND--Icamelus (Fabricius, 1792) --S-DEAIechinatus (Germar, 1824) FKSNDEAIparvulus Boheman, 1837 --SN---I

turbatus Boheman, 1844misellus Boheman, 1844 ----D---ptinoides (Marsham, 1802) --SND---

Baridinae Schönherr, 1836Baridini Schönherr, 1836Baris Germar, 1817artemisiae (Herbst, 1795) F-SN-EAIspitzyi Motschulsky, 1847 -----E--dalmatina Brisout de Barneville, 1870 ------A-laticollis (Marsham, 1802) --S-D-AIcarbonaria (Boheman, 1836) ------AIlepidii Germar, 1824 --S-D-AIcoerulescens (Scopoli, 1763) -------Itimida (Rossi, 1792) ------A-

Madopterini Lacordaire, 1866Limnobaris Bedel, 1885t-album (Linnaeus, 1758) FKSNDEAIpilistriata (Stephens, 1831) FKSNDEAIreitteri Munster, 1928 F-SNDEA-

Ceutorhynchinae Gistel, 1848Mononychini LeConte, 1876Mononychus Germar, 1824punctumalbum (Herbst, 1784) ------AI

Phytobiini Gistel, 1848Eubrychius Thomson, 1859velutus (Beck, 1817) FKSNDEA-

Phytobius Schönherr, 1833Litodactylus Redtenbacher, 1845

leucogaster (Marsham, 1802) FKSNDE--

Pelenomus Thomson, 1859Phytobius Dejean, 1835

comari (Herbst, 1795) FKSNDEAIwaltoni (Boheman, 1843) FKSND-AIcanaliculatus (Fåhraeus, 1843) FKSNDEAIquadrituberculatus (Fabricius, 1787) FKSNDE-Izumpti (Wagner, 1939) ----D---olssoni (Israelson, 1972) --S-D---velaris (Gyllenhal, 1827) FKSN-EA-

quadricorniger (Colonnelli, 1986) -KS-DEAIquadricornis (Gyllenhal, 1813) nec (Paykull, 1792)

Neophytobius Wagner, 1936muricatus (Brisout de Barneville, 1867) FKSNDE-Iquadrinodosus (Gyllenhal, 1813) FKSNDEA-

Rhinoncus Schönherr, 1825albicinctus Gyllenhal, 1837 ------A-perpendicularis (Reich, 1797) FKSNDEAIinconspectus (Herbst, 1795) FKSNDEAI

gramineus (Fabricius, 1793) nec (Gmelin, 1790)pericarpius (Linnaeus, 1758) FKSNDEAIbruchoides (Herbst, 1784) FKSNDEAIcastor (Fabricius, 1792) FKSNDEAI

Cnemogonini Colonnelli, 1979Auleutes Dietz, 1896epilobii (Paykull, 1800) FKSNDEA-

Scleropterini Schultze, 1902Rutidosoma Stephens, 1831globulus (Herbst, 1795) FKSNDEAIfallax (Otto, 1897) --S-D-A-

Scleropterus Schönherr, 1825serratus (Germar, 1824) F-S--EAI

Ceutorhynchini Gistel, 1848Amalus Schönherr, 1825scortillum (Herbst, 1795) FKSNDEAI

haemorrhous (Herbst, 1795) nec (Gmelin, 1790)

Amalorrhynchus Reitter, 1913melanarius (Stephens, 1831) --S-D---

Poophagus Schönherr, 1837sisymbrii (Fabricius, 1777) F-S-DEAI

Tapeinotus Schönherr, 1826sellatus (Fabricius, 1794) FKSNDEAI

Coeliodes Schönherr, 1837rubicundus (Herbst, 1795) FKSNDEAInigritarsis Hartmann, 1895 FKSNDE-Irana (Fabricius, 1787) F-SNDEAI

dryados (Gmelin, 1790)quercus (Fabricius,1787) nec (Linnaeus, 1758)

trifasciatus Bach, 1854 -----EA-ruber (Marsham, 1802) F-SNDE-Itransversealbofasciatus (Goeze, 1777) --SNDE-I

erythroleucos (Gmelin, 1790)cinctus (Geoffroy, 1785) nec (Drury, 1782)


Thamiocolus Thomson, 1859viduatus (Gyllenhal, 1813) FKSNDEA-sahlbergi (Sahlberg, 1845) FK--DEA-

Micrelus Thomson, 1859ericae (Gyllenhal, 1813) FKSNDEAI

Zacladus Reitter, 1913geranii (Paykull, 1800) FKSNDEAI

affinis (Paykull, 1792) nec (Schrank, 1781)

Ceutorhynchus Germar, 1824Ceuthorhynchidius Jacquelin du Val, 1854Neosirocalus Wagner, 1944

scapularis Gyllenhal, 1837 FKS-DE--pallipes Crotch, 1866 FKSNDEAI

minutus (Reich, 1797) nec (Drury, 1773)contractus (Marsham, 1802) nec (Geoffroy, 1785)

erysimi (Fabricius, 1787) FKSNDEAIviridanus Gyllenhal, 1837 ---N----ignitus Germar, 1824 FKSNDEA-pervicax Weise, 1883 FKS-DEAIbarbareae Suffrian, 1847 FK---EA-chalybaeus Germar, 1824 F-SNDEAIthomsoni Kolbe, 1900 F-SND---pectoralis Weise, 1895 -K--DEA-hirtulus Germar, 1824 F-SNDEAIsulcicollis (Paykull, 1800) FKSNDEAIpallidactylus (Marsham, 1802) FKSNDEAI

quadridens (Panzer, 1795) nec (Fabricius, 1775)atomus Boheman, 1845 F-SND---cochleariae (Gyllenhal, 1813) FKSNDEAIconstrictus (Marsham, 1802) F-S-D---plumbeus Brisout de Barneville, 1869 -----EA-unguicularis Thomson, 1871 F-SNDE--

curvirostris Schultze, 1898pleurostigma (Marsham, 1802) FKS-DEAI

assimilis (Paykull, 1792) nec (Fabricius, 1775)dubius Brisout de Barneville, 1883 ------A-sophiae Gyllenhal, 1837 -KS----Iroberti Gyllenhal, 1837 FKSN----alliariae Brisout de Barneville, 1860 --S-D---puncticollis Boheman, 1845 F-S--E-Inapi Gyllenhal, 1837 -K--DE-Irapae Gyllenhal, 1837 FKSNDEAIobstrictus (Marsham, 1802) FKSNDEAI

assimilis auct. nec (Paykull, 1792), nec (Fabricius, 1775)gallorhenanus Hoffmann, 1954 F-S--EA-griseus Brisout de Barneville, 1869 F-S-D---parvulus Brisout de Barneville, 1869 ----D---

inaffectatus Gyllenhal, 1837 F-SNDEA-syrites Germar, 1824 --S-D-AIgerhardti Schultze, 1899 F-S-E---

granulicollis Thomson, 1865, nec (Schönherr, 1837)querceti (Gyllenhal, 1813) FKSNDE--typhae (Herbst, 1795) FKSNDEAI

floralis (Paykull, 1792) nec (Olivier, 1790)cakilis (Hansen, 1917) F-SNDE--posthumus Germar, 1824 --S-D-A-pumilio (Gyllenhal, 1827) --S-D---hampei Brisout de Barneville, 1869 FKSNDEAIpulvinatus Gyllenhal, 1837 FKSNDEAIrhenanus Schultze, 1895 --SND-A-pyrrhorhynchus (Marsham, 1802) --SNDE-Isuturalis (Fabricius, 1775) ------AIdenticulatus (Schrank, 1781) -------Ipollinarius (Forster, 1771) F-SND-AIdistinctus Brisout de Barneville, 1870 FKS-DEAI

simillimus Edwards, 1911marginatus (Paykull, 1792) nec (Olivier, 1790)

molleri Thomson, 1868 FKS-DE--punctiger (Sahlberg, 1835) FKSNDEAIfennicus Faust, 1895 FKSNDE--arquatus (Herbst, 1795) F----E-Iangulosus Boheman, 1845 --S-DE--melanostictus (Marsham, 1802) -KS-DE-Iurticae Boheman, 1845 ----D---rugulosus (Herbst, 1795) FKSNDEAI

figuratus Gyllenhal, 1837triangulum Boheman, 1845 FKS-DEA-millefolii Schultze, 1897 --S-DE-Icampestris Gyllenhal, 1837 FK--DEAIlitura (Fabricius, 1775) FKSNDEAIeuphorbiae Brisout de Barneville, 1866--SNDEA-abbreviatulus (Fabricius, 1792) -------Ipallidicornis Gougelet & Brisout de

Barneville, 1860 F-S-D---asperifoliarum (Gyllenhal, 1813) FKSNDEAIcrucifer (Pallas, 1781) --SND-AI

cruciger (Herbst, 1784)javetii Gerhardt, 1867 --S-DE--larvatus Schultze, 1897 F---D--Igeographicus (Goeze, 1777) --S-D--I

Sirocalodes Voss, 1958depressicornis (Gyllenhal, 1813) FKSND-AI

nigrinus (Marsham, 1802) nec (Herbst, 1795)quercicola (Paykull, 1792) FKSNDEAI


Calosirus Thomson, 1859terminatus (Herbst, 1795) --S-DEA-apicalis (Gyllenhal, 1827) F-SNDE--

Trichosirocalus Colonnelli, 1979Ceuthorhynchidius auct., nec Jacquelin du Val, 1854

troglodytes (Fabricius, 1787) F-SNDEAIthalhammeri (Schultze, 1906) --S-D---barnevillei (Grenier, 1866) FKSNDEAI

Stenocarus Thomson, 1859umbrinus (Gyllenhal, 1827) --S-DE--

fuliginosus (Marsham, 1802) nec (Gmelin, 1790)cardui (Herbst, 1784) --S-D---

Nedyus Schönherr, 1825Cidnorhinus Thomson, 1859

quadrimaculatus (Linnaeus, 1758) FKSNDEAI

Coeliastes Weise, 1883lamii (Fabricius, 1792) ----DEAI

Conoderinae Schönherr, 1833Coryssomerini Thomson, 1859Coryssomerus Schönherr, 1825capucinus (Beck, 1817) --S-DEAI

Euryommatus Roger, 1857mariae Roger, 1857 ------A-

Orobitidinae Thomson, 1859Orobitis Germar, 1817cyaneus (Linnaeus, 1758) FKSNDEAI

Rhynchophorinae Schönherr, 1833Sitophilus Schönherr, 1838granarius (Linnaeus, 1758) F-SNDEAIoryzae (Linnaeus, 1763) f-snDeA-zeamais Motschulsky, 1855 fksnD---

Scolytinae Latreille, 1807Hylastini LeConte, 1876Hylurgops LeConte, 1876glabratus (Zetterstedt, 1828) FKSN-EA-palliatus (Gyllenhal, 1813) FKSNDEAI

Hylastes Erichson, 1836ater (Fabricius, 1792) F-S-D-AIbrunneus Erichson, 1836 FKSNDEA-

aterrimus Eggers, 1933cunicularius Erichson, 1836 FKSNDEAI

starki (Eggers, 1933)attenuatus Erichson, 1836 --SN-EAI

scandinavicus Lekander, 1955angustatus (Herbst, 1793) FKSNDEAI

opacus Erichson, 1836 FKSNDEAIobscurus Chapuis, 1875 -K------

septentrionalis Eggers, 1923

Hylesinini Erichson, 1836Hylastinus Bedel, 1888obscurus (Marsham, 1802) --S-D-A-

Pteleobius Bedel, 1888vittatus (Fabricius, 1787) ------AI

Hylesinus Fabricius, 1801Leperisinus Reitter, 1913

crenatus (Fabricius, 1787) F-SNDEAIoleiperda (Fabricius, 1792) --SND-A-fraxini (Panzer, 1799) FKSNDEAIvarius (Fabricius, 1775) F-SNDE--

orni Fuchs, 1906

Tomicini Shuckard, 1840Xylechinus Chapuis, 1869pilosus (Ratzeburg, 1837) FKSNDEA-

Hylurgus Latreille, 1807ligniperda (Fabricius, 1787) --S-DEAI

Tomicus Latreille, 1802Blastophagus Eichhoff, 1864

minor (Hartig, 1834) FKSNDEAIpiniperda (Linnaeus, 1758) FKSNDEAI

Dendroctonus Erichson, 1836micans (Kugelann, 1794) FKSNDEAI

Phloeotribini Chapuis, 1869Phloeotribus Latreille, 1796rhododactylus (Marsham, 1802) --S-D-A-spinulosus (Rey, 1883) FKSNDEAI

Polygraphini Chapuis, 1869Polygraphus Erichson, 1836subopacus Thomson, 1871 FKSN-EA-poligraphus (Linnaeus, 1758) FKSNDEAI

griseus Eggers, 1923punctifrons Thomson, 1886 FKSN-E--

Carphoborus Eichhoff, 1864cholodkovskyi Spessivstev, 1916 FKS--E--rossicus Semenov, 1902 FKS-----minimus (Fabricius, 1798) F-------teplouchovi Spessivtsev, 1916 -KS-----

Scolytini Latreille, 1807Scolytus Geoffroy, 1762multistriatus (Marsham, 1802) --S-DEAIpygmaeus (Fabricius, 1787) --S----I


scolytus (Fabricius, 1775) ----DEAItriarmatus (Eggers, 1912) --SND---ratzeburgi Janson, 1856 FKSNDEAIlaevis Chapuis, 1873 --SNDEA-mali (Bechstein & Scharfenberg, 1805) F-S-DEAIintricatus (Ratzeburg, 1837) FKSNDEAIcarpini (Ratzeburg, 1837) --S--EA-morawitzi Semenov, 1902 -K------rugulosus (Ratzeburg, 1837) FKSNDEAI

Ipini Latreille, 1807Pityogenes Bedel, 1888chalcographus (Linnaeus, 1761) FKSNDEAImonacensis Fuchs, 1911 --S----Iirkutensis Eggers, 1910 FKS-----saalasi Eggers, 1914 FKS-----trepanatus (Nördlinger, 1848) FKSNDEA-quadridens (Hartig, 1834) FKSNDEAIbidentatus (Herbst, 1783) FKSNDEAI

Pityokteines Fuchs, 1911curvidens (Germar, 1824) ------AI

Orthotomicus Ferrari, 1867longicollis (Gyllenhal, 1827) FKSNdEAIstarki Spessivtseff, 1926 -K-----Iproximus (Eichhoff, 1868) FKSNDEAIsuturalis (Gyllenhal, 1827) FKSNDEAIlaricis (Fabricius, 1792) FKSNDEAI

Ips DeGeer, 1775cembrae (Heer, 1836) -Ks-D---

subelongatus Motschulsky, 1860acuminatus (Gyllenhal, 1827) FKSNDEAIsexdentatus (Börner, 1776) FKSNdEAIduplicatus (Sahlberg, 1836) FKSN-EAItypographus (Linnaeus, 1758) FKSNDEAIamitinus (Eichhoff, 1872) FKs--EAI

Dryocoetini Lindemann, 1876Lymantor Lövendal, 1889coryli (Perris, 1855) FKSNDEAIaceris (Lindemann, 1875) ------AI

Xylocleptes Ferrari, 1867bispinus (Duftschmid, 1825) ---D---

Taphrorychus Eichhoff, 1878bicolor (Herbst, 1793) --SND--I

Dryocoetes Eichhoff, 1864baikalicus Reitter, 1900 -K------villosus (Fabricius, 1792) --SND-AIalni (Georg, 1856) FKSNDEAI

autographus (Ratzeburg, 1837) FKSNDEAIsuecicus Eggers, 1923

hectographus Reitter, 1913 FKSN-EAI

Crypturgini LeConte, 1876Crypturgus Erichson, 1836subcribrosus Eggers, 1933 FKSNDEAIcinereus (Herbst, 1793) FKSN-EAIpusillus (Gyllenhal, 1813) FKSNDEAIhispidulus Thomson, 1870 FKSNDEAI

Xyloterini Lindemann, 1876Trypodendron Stephens, 1830Xyloterus Erichson, 1836

domesticum (Linnaeus, 1758) F-SNDEAIlaeve Eggers, 1939 FKSN-E--

piceum Strand, 1946proximum auct. nec (Niijima, 1909)

lineatum (Olivier, 1795) FKSNDEAIsignatum (Fabricius, 1792) FKSNDEAI

Xyleborini LeConte, 1876Xyleborus Eichhoff, 1864Anisandrus Ferrari, 1867

dispar (Fabricius, 1792) FKSNDEAIcryptographus (Ratzeburg, 1837) FKSNDEAImonographus (Fabricius, 1792) --S-DEAI

Xyleborinus Reitter, 1913saxesenii (Ratzeburg, 1837) --SND--I

Cryphalini Lindemann, 1876Trypophloeus Fairmaire, 1868alni (Lindemann, 1875) FK-N-E--bispinulus Eggers, 1927 FKSN--A-granulatus (Ratzeburg, 1837)ssp. granulatus (Ratzeburg, 1837) -------Issp. grothii (Hagedorn, 1904) --SND---asperatus (Gyllenhal, 1813) FKSN-EA-discedens Palm, 1950 F-S--EA-

palmi Hansen, 1955

Ernoporicus Berger, 1917Ernopocerus Wood, 1954

caucasicus (Lindemann, 1876) F-SND---fagi (Fabricius, 1798) --S-D---

Ernoporus Thomson, 1859tiliae (Panzer, 1793) F-SNDEAI

Cryphalus Erichson, 1836piceae (Ratzeburg, 1837) ------AIabietis (Ratzeburg, 1837) FKSNDEAIsaltuarius Weise, 1891 FKSN-E--


Corthylini LeConte, 1876Pityophthorus Eichhoff, 1864micrographus (Linnaeus, 1758) FKSN-EAIlichtensteinii (Ratzeburg, 1837) FKSNDEAIpubescens (Marsham, 1802) --SND---morosovi Spessivtseff, 1926 FKS---A-glabratus Eichhoff, 1878 FKSND--Itragardhi Spessivtseff, 1921 FKSN-E--lapponicus Stark, 1952 -K------

Gnathotrichus Eichhoff, 1868materiarius (Fitch, 1855) F-s-----

Platypodidae Shuckard, 1840Platypus Herbst, 1793cylindrus (Fabricius, 1792) --S-d-A-

Appendix 1. The following species have been reportedfrom Finland, but the information has not yet been pub-lished. The details will be provided by the discoverers inforthcoming reports.

Microderes brachypus (Dej.) - S. KarjalainenBradycellus csikii Laczo - H. LappalainenBradycellus verbasci (Duft.) - P. RassiHydrochus elongatus (Schall.) - S. KarjalainenMargarinotus terricola (Germ.) - P. MartikainenAloeochara haematodes Kr. - T. ClayhillsPhilhygra pseudoelongatula (Bernh.) - P. RassiAtheta (Microdota) indubia (Sharp) – J. MuonaBolitochara obliqua (Er.) - M. LandvikTachinus schneideri Luze - I. RutanenGabronthus sulcifrons (Sharp) - I. RutanenPhilonthus alpinus Epp. - P. RassiQuedius vexans Epp. - P. RassiAttagenus trifasciatus (F.) - J. MuonaDorcatoma flavicornis (F.) - J. Mattila,

P. Ahlroth & M. v.NumersCryptarcha undata (Ol.) - J. Mattila & P. AhlrothMonotoma quadricollis Aubé - I. RutanenCryptolestes weisei (Rtt.) - I. RutanenAtomaria pseudatra Rtt. - I. RutanenCalvia decemguttata (L.) - S. KarjalainenDienerella ruficollis (Marsh.) - I. RutanenCorticarina truncatella (Mannh.) -

P. MartikainenStricticomus tobias (Mars.) - S. KarjalainenOtiorhynchus lugdunensis Boh. -

I. MannerkoskiCeutorhynchus arquatus (Hbst.) - P. Rassi


Hakemisto — Index

Abax ................................. 11Abdera ............................. 70Ablattaria ......................... 23Abraeus ............................ 17Absidia ............................. 55Acalles .............................. 95Acalyptus ......................... 92Acanthocinus ................... 78Acanthoderes .................... 78Acanthoscelides ............... 85Achenium ......................... 42Acidota ............................. 25Acilius ................................ 8Aclypea ............................ 23Acmaeops ......................... 76Acritus .............................. 17Acrognathus ...................... 38Acrolocha ......................... 24Acrostiba .......................... 30Acrotona .......................... 35Acrotrichis ........................ 20Acrulia ............................. 24Actenicerus ...................... 52Actidium ........................... 19Actinopteryx .................... 20Actobius ............................ 43Actophylla ........................ 35Acupalpus ........................ 14Acylophorus ..................... 46Adalia ............................... 67Aderus .............................. 75Adistemia ......................... 68Adonia .............................. 67Adoxus ............................. 81Adrastus ........................... 54Aegialia ............................ 46Aegomorphus ................... 78Aeletes .............................. 17Aepus ............................... 10Aesalus ............................. 46Agabus ............................... 7Agapanthia ...................... 79

Agaricochara ................... 36Agaricophagus ................. 21Agathidium ...................... 21Agelastica ........................ 82Aglenus ............................ 75Agonum ............................ 12Agrilus .............................. 49Agriotes ............................ 53Agrypnus .......................... 52Agyrtes ............................. 20Ahasverus ........................ 62Airaphilus ......................... 62Alaobia, subg. ................... 33Alapsodus ......................... 45Aleochara ......................... 29Alevonota ......................... 35Alianta ............................. 35Allandrus .......................... 85Allecula ............................ 73Aloconota ......................... 32Alosterna .......................... 77Alphitobius ....................... 72Alphitophagus .................. 73Altica ................................ 83Amalorrhynchus ............... 95Amalus ............................. 95Amara .............................. 12Amarochara ..................... 31Amauronyx ....................... 26Amidobia, subg. ................ 33Amischa ........................... 35Ampedus .......................... 53Amphicyllis ....................... 21Amphimallon .................... 47Amphotis .......................... 61Anacaena ......................... 16Anaerea ............................ 79Anaesthetis ....................... 78Anaglyptus ....................... 78Anaspis ............................. 76Anatis ............................... 67Anaulacaspis .................... 36Anchomenus ..................... 12Ancistronycha .................. 54

Ancyrophorus .................... 38Aneurota ........................... 35Anidorus ........................... 75Anisandrus ........................ 98Anisodactylus ................... 14Anisoplia .......................... 48Anisosticta ....................... 67Anisotoma ........................ 21Anisoxya .......................... 70Anitys ............................... 58Anobium ........................... 57Anogcodes ........................ 74Anomala ........................... 48Anommatus ...................... 66Anomognathus ................. 37Anopleta ........................... 35Anoplodera ...................... 77Anoplus ............................ 93Anostirus .......................... 52Anotylus ........................... 39Anthaxia .......................... 49Anthelephila ..................... 75Antherophagus ................. 64Anthicus ........................... 75Anthobium ......................... 23Anthobium ....................... 25Anthocomus ..................... 59Anthonomus ..................... 92Anthophagus .................... 25Anthracus ......................... 14Anthrenus ......................... 56Anthribus ......................... 86Apalus .............................. 74Aphanisticus ..................... 50Aphidecta ......................... 67Aphodius .......................... 46Aphthona ......................... 83Apion ................................ 86Aplocnemia ....................... 78Aplocnemus ...................... 59Aploderus ......................... 39Aplotarsus ........................ 52Apoderus .......................... 86Aporthopleura ................... 58


Apteropeda ....................... 84Araecerus ......................... 86Archarius .......................... 92Arctobyrrhus .................... 50Arena ................................ 37Argopus ............................ 84Arhopalus ......................... 76Aridius .............................. 68Armidia ............................ 55Aromia ............................. 77Arpedium ......................... 25Arrhenopeplus .................. 26Arthrolips ......................... 68Asaphidion ....................... 10Asclera .............................. 74Ascoliocerus ..................... 52Asemum ........................... 76Asiorestia .......................... 84Aspidiphorus .................... 60Astenus ............................. 41Astilbus ............................. 36Atanygnathus ................... 46Atemeles ........................... 36Atheta .............................. 33Atheta, subg. .................... 34Atholus ............................. 18Athous .............................. 52Atomaria .......................... 64Atrecus ............................. 43Attactagenus .................... 89Attagenus ......................... 56Attelabus .......................... 86Attenuorhagus ................... 51Auleutes ........................... 95Aulonium .......................... 71Aulonothroscus ................ 51Autalia ............................. 37Axinopalpis ...................... 77Axinotarsus ...................... 59Badister ............................ 13Badura, subg. ................... 33Baeckmanniolus ................. 17Baeocrara ........................ 20Bagous ............................. 93

Baptolinus ......................... 43Baranowskiella ................ 19Baris ................................. 95Barynotus ......................... 89Barypeithes ...................... 89Batophila .......................... 83Batrisodes ........................ 26Batrisus ............................ 26Bembidion ........................ 10Berninelsonius .................... 52Berosus ............................. 15Bessobia, subg. ................. 34Bibloplectus ...................... 26Bibloporus ........................ 26Bidessus .............................. 5Biphyllus ........................... 65Bisnius .............................. 43Bitoma .............................. 72Bius .................................. 72Blaps ................................ 72Blastophagus ..................... 97Bledius .............................. 39Blemus .............................. 10Blethisa .............................. 9Blitophaga ......................... 23Bohemiellina .................... 36Bolboceras ....................... 46Bolitobius .......................... 28Bolitobius ......................... 28Bolitochara ...................... 36Bolitophagus .................... 73Borboropora ..................... 35Boreaphilus ...................... 25Boreophilia, subg. ............. 34Boreostiba, subg. .............. 34Boros ................................ 74Bostrichus ........................ 56Bothrideres ....................... 66Bothrynoderes .................. 90Brachinus ......................... 15Brachonyx ........................ 92Brachyderes ...................... 89Brachygluta ...................... 27Brachygonus .................... 53

Brachypterolus ................. 60Brachypterus .................... 60Brachysomus .................... 89Brachyta ........................... 76Brachytarsus ...................... 86Brachytemnus .................. 94Brachyusa ........................ 31Bradybatus ....................... 92Bradycellus ....................... 14Bromius ............................ 81Broscus ............................... 9Bruchela ........................... 86Bruchidius ........................ 85Bruchus ............................ 85Brundinia ......................... 31Bryaxis ............................. 26Bryaxis .............................. 27Brychius ............................. 5Bryocharis ......................... 28Bryophacis ....................... 27Bryoporus ......................... 27Buprestis ........................... 49Byctiscus .......................... 86Byrrhus ............................. 50Bythinus ........................... 26Byturus ............................. 65Caccobius ........................ 47Cadaverota ...................... 32Caenocara ....................... 58Caenorhinus ...................... 86Caenorhinus ...................... 86Caenoscelis ...................... 64Cafius ............................... 44Calambus ......................... 53Calathus ........................... 11Calitys .............................. 58Callicerus ......................... 32Callidiellum ...................... 78Callidium ......................... 78Callosobruchus ................ 85Calodera .......................... 30Calodromius ..................... 15Calomicrus ....................... 82Calopus ............................ 74


Calosirus .......................... 97Calosoma ........................... 8Calvia .............................. 67Calyptomerus ................... 48Cantharis ......................... 54Carabus ............................. 8Carcinops ......................... 17Cardiophorus ................... 54Carilia ............................... 76Carpelimus ....................... 38Carphacis ......................... 28Carphoborus .................... 97Carpophilus ..................... 60Cartodere .......................... 68Cartodere ......................... 68Cassida ............................ 85Catops .............................. 22Caulotrupodes ................... 94Centrotoma ...................... 27Cephalocousya ................. 30Cephennium ..................... 22Cerambyx ........................ 77Cerapheles ....................... 59Cercyon ............................ 16Cerocoma ......................... 74Ceruchus .......................... 46Cerylon ............................ 66Cetonia ............................ 48Ceuthorhynchidius ............. 96Ceuthorhynchidius ............. 97Ceutorhynchus ................. 96Chaetabraeus ................... 17Chaetarthria .................... 16Chaetida, subg. ................ 33Chaetocnema ................... 84Chaetophora .................... 50Chalcionellus ................... 17Chalcoides ........................ 84Chalcophora .................... 49Chanoma ......................... 30Charopus .......................... 59Chennium ......................... 27Chilocorus ........................ 67Chilomorpha .................... 30

Chilopora .......................... 30Chlaenius ......................... 13Chloecharis ...................... 42Chlorophanus ................... 89Chlorophorus ................... 78Choleva ............................ 22Choragus ......................... 86Chromoderus .................... 90Chrysanthia ..................... 74Chrysobothris .................. 49Chrysochloa ...................... 81Chrysochus ...................... 81Chrysolina ........................ 81Chrysomela ...................... 81Cicindela ............................ 9Cicones ............................. 72Cidnopus .......................... 52Cidnorhinus ....................... 97Cilea ................................. 29Cillenus ............................ 10Cimberis ........................... 85Cionus .............................. 91Cis .................................... 69Clambus ........................... 48Clanoptilus ....................... 59Claviger ........................... 27Cleonis ............................. 90Cleopomiarus ................... 93Cleopus ............................ 91Clivina ............................... 9Clypastraea ...................... 68Clypeorhagus ................... 51Clytra ............................... 80Clytus ............................... 78Cneorhinus ........................ 89Coccidula ......................... 66Coccinella ........................ 67Coccinula ......................... 67Coelambus ......................... 6Coeliastes ......................... 97Coeliodes ......................... 95Coelostoma ...................... 16Colaphellus ...................... 81Colenis ............................. 21

Colon ............................... 21Colpodota ......................... 35Colposis ........................... 75Colydium .......................... 71Colymbetes ........................ 8Comasinus ....................... 91Combocerus ..................... 65Compsidia ......................... 79Conalia ............................ 71Coniocleonus ................... 90Conopalpus ...................... 71Conosoma ......................... 28Copelatus ........................... 5Copris .............................. 47Coproceramius .................. 34Coprophilus ...................... 38Coproporus ...................... 28Coprothassa ..................... 35Coptocephala ................... 80Coraebus .......................... 50Cordalia ........................... 36Cordicomus ...................... 75Cordylepherus .................. 59Corticaria ........................ 69Corticarina ...................... 69Corticeus .......................... 73Cortinicara ...................... 69Cortodera ........................ 77Corylophus ....................... 68Corymbia .......................... 77Corymbites ........................ 52Coryphiomorphus ............ 25Coryphium ....................... 25Coryssomerus .................. 97Cossonus .......................... 94Crataraea ........................ 30Creophilus ........................ 45Crepidodera ...................... 84Crepidodera ..................... 84Crepidophorus ................. 52Criocephalus ..................... 76Crioceris .......................... 80Crudosilis .......................... 55Cryphalus ......................... 98


Cryptarcha ....................... 62Crypticus .......................... 73Cryptobium ....................... 42Cryptocephalus ................ 80Cryptolaemus ................... 66Cryptolestes ..................... 63Cryptophagus .................. 63Cryptophilus .................... 65Cryptopleurum ................. 16Cryptorhynchus ............... 95Crypturgus ....................... 98Ctenicera ......................... 52Cteniopus ......................... 73Ctesias ............................. 56Cucujus ............................ 62Curculio ........................... 92Curelius ............................ 65Curimopsis ....................... 50Curtimorda ...................... 71Cyanostolus ..................... 62Cybister .............................. 8Cybocephalus ................... 62Cychramus ....................... 62Cychrus .............................. 9Cycloderes ........................ 89Cyclodinus ....................... 75Cylindera ........................... 9Cylister .............................. 18Cylletron .......................... 24Cyllodes ........................... 61Cymbiodyta ...................... 16Cymindis .......................... 15Cynaeus ........................... 73Cynegetis ......................... 68Cypha ............................... 37Cyphea ............................. 37Cyphocleonus ................... 90Cyphon ............................. 48Cyrtanaspis ...................... 76Cyrtoclytus ....................... 78Cyrtusa ............................ 21Cytilus .............................. 50Dacne ............................... 65Dacrila ............................. 31

Dadobia ........................... 32Dalopius ........................... 54Danacea ........................... 59Danosoma ......................... 52Dascillus ........................... 49Dasyglossa ........................ 30Dasygnypeta .................... 31Dasytes ............................. 59Datomicra, subg. .............. 33Datostiba ........................... 33Deinopsis .......................... 38Deleaster .......................... 38Deliphrosoma ................... 25Deliphrum ........................ 25Demetrias ......................... 14Dendroctonus ................... 97Dendrophagus .................. 62Dendrophilus .................... 17Dendroxena ...................... 23Denticollis ........................ 52Deporaus .......................... 86Dermestes ........................ 55Dermestoides ................... 58Dermestoides .................... 58Derocrepis ........................ 84Deronectes ......................... 7Descarpentriesina .............. 49Deubelia ............................ 30Devia ................................ 30Dexiogyia ......................... 30Diacanthous ....................... 52Diacheila ............................ 9Diachromus ...................... 14Dianous ............................ 41Diaperis ............................ 73Diastictus ......................... 47Dibolia ............................. 84Dicerca ............................. 49Dicheirotrichus ................. 14Dicranthus ....................... 93Dicronychus ..................... 54Dictyoptera ...................... 54Dienerella ......................... 68Diglotta ............................ 37

Dilacra ............................. 32Dimetrota, subg. ............... 34Dinaraea .......................... 35Dinarda ............................ 31Dinoderus ......................... 56Dinoptera .......................... 76Dinothenarus ................... 44Diplocoelus ...................... 65Dircaea ............................. 70Dirhagus ............................ 51Dirrhagofarsus ................. 51Disopora .......................... 32Dissoleucas ...................... 85Ditoma .............................. 72Ditylus .............................. 74Dlochrysa .......................... 81Dochmonota .................... 31Dolichosoma .................... 59Dolichus ........................... 12Donacia ............................ 79Dorcadion ........................ 78Dorcatoma ....................... 58Dorcus .............................. 46Dorytomus ....................... 91Doydirhynchus ................. 85Drapetes ........................... 52Dreposcia ......................... 22Drilus ............................... 54Dromaeolus ...................... 51Dromius ............................ 15Drusilla ............................ 36Dryocoetes ....................... 98Dryophilus ........................ 57Dryophthorus ................... 93Dryops .............................. 50Dyschirius .......................... 9Dytiscus ............................. 8Eanus .............................. 53Ebaeus ............................ 59Ecanus ............................ 20Ectinus ............................ 54Edaphus .......................... 41Eicolyctus ......................... 65Elaphrus ............................ 9


Elater ............................... 53Elater .............................. 53Elateroides ....................... 58Eledona ........................... 73Eledonoprius ................... 73Ellescus ........................... 91Elmis ............................... 50Elodes ............................. 48Elonium ............................ 38Emphylus .......................... 64Emus ............................... 44Enalodroma, subg. .......... 35Encephalus ...................... 36Endomychus .................... 66Enedreytes ....................... 85Enicmus .......................... 68Ennearthron .................... 70Enochrus ......................... 16Entomoscelis ................... 82Eocatops ......................... 22Epauloecus ...................... 57Ephimeropus .................... 93Ephistemus ...................... 65Episernus ........................ 57Epitrix ............................. 84Epuraea .......................... 60Erchomus ......................... 28Eremotes .......................... 94Ergates ............................ 76Erichsonius ..................... 43Ernobius .......................... 57Ernopocerus ..................... 98Ernoporicus .................... 98Ernoporus ....................... 98Esolus .............................. 50Euaesthetus ..................... 41Eubria ............................. 51Eubrychius ...................... 95Eucilodes ......................... 48Eucinetus ........................ 48Eucnecosum .................... 25Eucnemis ......................... 51Euconnus ........................ 23Eudectus ......................... 25

Euglenes .......................... 75Euophryum ..................... 94Euplectus ........................ 26Eupogonocherus ............... 78Eurylister .......................... 18Eurymniusa ....................... 31Euryommatus .................. 97Euryporus ....................... 45Euryptilium ..................... 19Euryusa ........................... 37Eusphalerum ................... 23Eustrophus ...................... 70Eutheia ............................ 22Euthiconus ...................... 22Evanystes ......................... 32Evodinus ......................... 76Exocentrus ...................... 78Exochomus ..................... 67Falagria .......................... 35Falagrioma ..................... 36Falagriusa ......................... 36Fleutiauxellus ................. 53Formicomus ..................... 75Foucartia ........................ 89Furcipus ........................... 92Gabrius ........................... 43Gabronthus ..................... 43Galeruca ......................... 82Galerucella ..................... 82Gastrallus ....................... 57Gastrophysa .................... 81Gauropterus .................... 42Gaurotes ......................... 76Geodromicus ................... 25Georissus ........................ 15Geostiba .......................... 32Geotrupes ....................... 46Gibbium .......................... 56Glabroplatycis .................. 54Glaphyra .......................... 77Glischrochilus ................. 62Globicornis ..................... 56Gnathacmaeops ................ 76Gnathoncus ..................... 17

Gnathotrichus ................. 99Gnatocerus ..................... 73Gnorimus ........................ 48Gnypeta .......................... 31Gonioctena ..................... 82Gonodera ........................ 73Gracilia ........................... 77Grammoplectus ................ 26Grammoptera ................. 77Grammostethus ................. 18Graphoderus ..................... 8Graptodytes ...................... 6Gronops .......................... 90Grynobius ....................... 57Grynocharis .................... 58Grypus ............................ 90Guignotus ........................... 6Gymnetron ...................... 93Gymnusa ......................... 38Gynandrophthalma ............ 80Gyrinus ............................. 5Gyrohypnus ..................... 42Gyrophaena .................... 36Habrocerus ...................... 29Habroloma ...................... 50Hadreule .......................... 70Hadrobregmus ................. 57Haemonia ......................... 79Haliplus ............................. 5Hallomenus ..................... 70Halobrecta ...................... 35Halyzia ............................ 67Hapalaraea ..................... 24Haploglossa .................... 30Harminius ....................... 52Harmonia ........................ 67Harpalus ......................... 13Hedobia ........................... 56Helichus .......................... 50Helochares ...................... 16Helophorus ...................... 15Hemicoelus ....................... 57Hemicrepidius ................. 52Henoticus ........................ 63


Heptaulacus .................... 47Hermaeophaga ............... 83Hetaerius ......................... 18Heterhelus ....................... 60Heterocerus ..................... 51Heterota .......................... 37Heterothops ..................... 45Hippodamia .................... 67Hippuriphila .................... 84Hirticomus ...................... 75Hispa ............................... 85Hister .............................. 18Holobus ........................... 37Hololepta ........................ 18Holoparamecus ............... 66Homalota ........................ 37Homorhythmus ................. 88Hoplia ............................. 48Hoshihananomia ............. 71Hydaticus .......................... 8Hydnobius ....................... 20Hydraena ........................ 18Hydrobius ........................ 16Hydrochara ..................... 16Hydrochus ....................... 15Hydrocolus ......................... 6Hydroglyphus .................... 6Hydronomus .................... 93Hydrophilus ..................... 16Hydroporus ....................... 6Hydrosmecta ................... 31Hydrosmectina ................. 31Hydrothassa .................... 81Hydrous ........................... 16Hydrovatus ........................ 5Hygronoma ..................... 37Hygropora ....................... 30Hygrotus ............................ 6Hylastes ........................... 97Hylastinus ....................... 97Hylecoetus ....................... 58Hylesinus ......................... 97Hylis ................................ 51Hylobius .......................... 94

Hylochares ...................... 51Hylophilus ........................ 75Hylotrupes ....................... 77Hylurgops ........................ 97Hylurgus .......................... 97Hymenalia ....................... 73Hymenophorus ................ 73Hypebaeus ....................... 59Hypera ............................ 90Hyperaspis ...................... 67Hyphydrus ......................... 6Hypnogyra ...................... 42Hypnoidus ....................... 52Hypocacculus .................. 17Hypocaccus ..................... 17Hypocaelus ...................... 51Hypocassida .................... 85Hypocoprus ..................... 63Hypocyphtus .................... 37Hypoganus ...................... 53Hypomedon ..................... 41Hypophlaeus .................... 73Hypopycna ...................... 24Hypulus ........................... 70Iberodorcadion ................. 78Ilybius ................................ 7Ilyobates .......................... 31Ipidia ............................... 61Ips ................................... 98Ischnodes ........................ 53Ischnoglossa .................... 30Ischnomera ..................... 74Ischnopoda ...................... 31Ischnosoma ..................... 27Isochnus .......................... 92Isomira ............................ 73Isorhipis .......................... 51Ityocara ............................ 30Judolia ............................ 77Kateretes ......................... 60Korynetes ........................ 59Labidostomis ................... 80Laccobius ........................ 16Laccophilus ....................... 5

Laccornis .......................... 5Lacon .............................. 52Lado ................................ 72Laemophloeus ................. 63Laemostenus ................... 12Lagria ............................. 72Lamia .............................. 78Lampra ............................ 49Lamprinodes ................... 28Lamprinus ....................... 28Lamprobyrrhulus ............ 50Lampyris ......................... 54Laria ................................ 61Laricobius ....................... 55Larinus ............................ 90Lasconotus ...................... 72Lasioderma ..................... 58Lasiorhynchites ............... 86Latelmis ............................ 50Latheticus ....................... 72Lathrimaeum ..................... 25Lathrobium ..................... 42Latridius .......................... 68Latridius ........................... 68Lebia ............................... 14Leiestes ........................... 66Leioderes ......................... 78Leiodes ............................ 20Leiopus ............................ 78Leiosoma ......................... 94Leistus ............................... 8Lema ............................... 80Leperisinus ....................... 97Leptacinus ....................... 42Leptinotarsa .................... 81Leptinus .......................... 22Leptophius ...................... 42Leptoplectus .................... 26Leptura ........................... 77Leptusa ........................... 36Lepyrus ........................... 94Lesteva ............................ 25Leucoparyphus ................. 29Librodor ........................... 62


Licinus ............................. 13Lilioceris ......................... 80Limnebius ........................ 18Limnichus ........................ 51Limnius ........................... 50Limnobaris ...................... 95Limnoxenus ..................... 16Limobius ......................... 90Limoniscus ...................... 52Limonius ......................... 52Linaeidea .......................... 81Liocola ............................ 48Liocyrtusa ........................ 21Liodopria ........................ 21Liogluta ........................... 32Lionychus ........................ 15Liophloeus ....................... 89Liotrichus ........................ 52Liparus ............................ 94Lissodema ....................... 75Litargus ........................... 69Lithocharis ...................... 41Litodactylus ...................... 95Lixus ............................... 90Lochmaea ....................... 82Lomechusa ...................... 36Lomechusoides ................ 36Longitarsus ..................... 83Lopheros ......................... 54Lophocateres ................... 58Lordithon ........................ 28Loricera ............................ 9Lucanus ........................... 46Ludius .............................. 53Lundbergia ...................... 32Luperus ........................... 82Lycoperdina .................... 66Lyctus .............................. 56Lygistopterus .................. 54Lymantor ........................ 98Lymexylon ....................... 58Lypoglossa, subg. ............ 33Lyprocorrhe .................... 35Lythraria ......................... 84

Lytta ................................ 74Macronychus ................... 50Macroplea ....................... 79Magdalis ......................... 94Malachius ........................ 59Maladera ......................... 47Malthinus ........................ 55Malthodes ....................... 55Manda ............................. 38Mannerheimia ................. 24Mantura .......................... 84Margarinotus .................. 18Martianus ........................ 72Masoreus ......................... 14Mecinus ........................... 93Medon ............................. 41Megalinus ........................ 42Megarthrus ..................... 25Megasternum .................. 16Megatoma ....................... 56Melandrya ....................... 70Melanimon ...................... 72Melanophila .................... 49Melanophthalma ............. 69Melanotus ....................... 53Melasis ............................ 51Meliceria ......................... 26Meligethes ....................... 61Meloe .............................. 74Melolontha ...................... 47Menephilus ...................... 73Menesia ........................... 79Meotica ........................... 31Mesosa ............................ 78Metacantharis ................. 55Metoecus ......................... 71Metopsia ......................... 25Mezium ............................ 56Miaromimus ..................... 93Miarus ............................. 93Micralymma .................... 24Micrambe ........................ 63Micrelus ........................... 96Micridium ........................ 19

Microbregma ................... 57Microcara ........................ 48Microderes ....................... 14Microdota, subg. .............. 33Microglotta ....................... 30Microlestes ...................... 15Microlomalus .................... 18Micropeplus ..................... 26Microptilium .................... 19Microrhagus .................... 51Microscydmus ................. 23Microsporus ....................... 5Migneauxia ..................... 69Millidium .......................... 19Minthea ........................... 56Miscodera ......................... 9Mniophila ........................ 84Mniusa ............................ 30Mocyta, subg. .................. 33Molorchus ....................... 77Monochamus ................... 78Mononychus .................... 95Monotoma ....................... 62Mordella .......................... 71Mordellistena .................. 71Mordellistenula ............... 71Mordellochroa ................. 71Morychus ........................ 50Mosotalesus ..................... 52Mycetaea ......................... 66Mycetina ......................... 66Mycetochara ................... 73Mycetophagus ................. 69Mycetoporus ................... 27Mycetota, subg. ............... 33Mycterus ......................... 74Myllaena ......................... 38Myrmechixenus ............... 73Myrmecocephalus ........... 36Myrmecopora .................. 35Myrmetes ........................ 17Myrrha ............................ 67Myzia ............................... 67Nacerdes ......................... 74


Nalassus .......................... 72Nanodes .......................... 87Nanophyes ...................... 87Nanoptilium ...................... 19Nargus ............................. 22Nartus ................................ 7Nathrius .......................... 77Nausibius ........................ 62Neatus ............................. 72Nebria ............................... 8Nebrioporus ...................... 7Necrobia .......................... 59Necrodes ......................... 23Necydalis ......................... 77Nedyus ............................ 97Negastrius ....................... 53Nehemitropia .................. 35Neliocarus ........................ 89Nemadus ......................... 22Nemozoma ...................... 58Neobisnius ....................... 44Neocoenorrhinus ............. 86Neocrepidodera ............... 84Neohilara, subg. .............. 33Neohypdonus .................. 53Neomida .......................... 73Neomysia ......................... 67Neophytobius .................. 95Neosirocalus .................... 96Nepachys ......................... 59Nephanes ........................ 19Nephus ............................ 66Nevraphes ....................... 23Nicrophorus .................... 23Niptus .............................. 57Nitidula ........................... 61Nivellia ............................ 77Normandia ...................... 50Nosodendron ................... 55Nossidium ....................... 19Notaris ............................ 91Noterus ............................. 5Nothorhina ...................... 76Notiophilus ........................ 8

Notolaemus ..................... 63Notothecta, subg. ............ 33Notoxus ........................... 75Nudobius ......................... 42Oberea ............................. 79Obrium ............................ 77Ocalea ............................. 31Ochina ............................. 57Ochrosis .......................... 84Ochthebius ...................... 18Ochthephilum .................. 42Ochthephilus ................... 38Octotemnus ..................... 70Ocypus ............................ 44Ocys ................................ 10Ocyusa ............................ 30Odacantha ...................... 14Odonteus ......................... 46Oedemera ........................ 74Oedostethus .................... 53Oenopia ........................... 67Oiceoptoma ..................... 23Olibrus ............................ 63Oligella ............................ 19Oligomerus ...................... 57Oligota ............................ 37Olisthaerus ...................... 27Olisthopus ....................... 12Olophrum ........................ 25Omalium ......................... 24Omaloplia ....................... 47Omiamima ...................... 88Omias .............................. 88Omonadus ....................... 75Omophlus ........................ 73Omophron ......................... 8Omosita ........................... 61Oniticellus ....................... 47Ontholestes ..................... 44Onthophagus ................... 47Onthophilus ..................... 18Oodes .............................. 13Oodescelis ....................... 72Oomorphus ..................... 80

Ootypus ........................... 65Opatrum .......................... 72Opetiopalpus ................... 59Ophonus .......................... 13Opilo ............................... 58Oplocephala ..................... 73Oplosia ............................ 78Orchesia .......................... 70Orchestes ......................... 92Orectochilus ...................... 5Oreina ............................. 81Oreodytes .......................... 6Orithales ......................... 52Orobitis ........................... 97Orochares ........................ 24Orsodacne ....................... 79Orthocerus ...................... 72Orthochaetes .................... 91Orthocis .......................... 70Orthoperus ...................... 68Orthotomicus .................. 98Oryctes ............................ 48Oryzaephilus ................... 62Osmoderma ..................... 48Osphya ............................ 71Ostoma ............................ 58Othius .............................. 43Otho ................................ 51Otiorhynchus ................... 88Oulema ............................ 80Oulimnius ......................... 50Ousipalia .......................... 32Ovalisia ............................ 49Oxylaemus ....................... 65Oxymirus .......................... 76Oxyomus .......................... 47Oxypoda ........................... 29Oxyporus .......................... 40Oxypselaphus ................... 12Oxypteris .......................... 49Oxytelus ........................... 39Oxythyrea ......................... 48Pachnephorus .................. 81Pachnida .......................... 35


Pachyatheta ..................... 35Pachybrachis ................... 80Pachygluta ......................... 36Pachyta ............................ 76Paederidus ....................... 41Paederus .......................... 41Palorus ............................. 72Panagaeus ....................... 13Paracardiophorus ............ 54Paracymus ....................... 16Paradromius .................... 15Paragabrius ....................... 43Paralister ........................... 18Paramecosoma ................ 63Parameotica, subg. .......... 33Paranchus ........................ 12Paranomus ........................ 53Paranopleta ..................... 32Paratachys ......................... 11Paratinus ......................... 60Parocyusa ........................ 30Paromalus ........................ 18Patrobus ............................. 9Pediacus ........................... 62Pedinus ............................ 72Pedius .............................. 11Pedostrangalia ................... 77Pelecotoma ...................... 71Pelenomus ........................ 95Pelophila ............................ 8Peltis ................................ 58Peltodytes .......................... 5Pentanota ........................ 31Pentaphyllus .................... 73Pentarthrum .................... 93Penthelispa ........................ 72Perigona .......................... 14Perileptus ......................... 10Peritelus ............................ 88Phacophallus ................... 42Phaedon ........................... 81Phaenops ......................... 49Phalacrolinus ..................... 42Phalacrus ......................... 63

Phaleria ........................... 73Pheletes ............................. 52Philhygra ......................... 32Philonthus ........................ 43Philopedon ....................... 89Philorhizus ....................... 15Philorinum ....................... 24Phloeobium ....................... 25Phloeocharis .................... 27Phloeodroma ..................... 31Phloeonomus ................... 24Phloeophagus .................. 94Phloeopora ...................... 31Phloeostiba ...................... 24Phloeostichus ................... 62Phloeotribus ..................... 97Phloiophilus ..................... 58Phloiotrya ........................ 70Phosphaenus .................... 54Phosphuga ....................... 23Phratora .......................... 82Phryganophilus ................ 71Phylan .............................. 72Phyllobius ........................ 88Phyllobrotica .................... 82Phyllodecta ........................ 82Phyllodrepa ...................... 24Phyllodrepoidea ............... 25Phyllopertha .................... 48Phyllotreta ....................... 82Phymatodes ..................... 78Phymatura ....................... 36Physoronia ....................... 61Phytobaenus .................... 75Phytobius ......................... 95Phytobius .......................... 95Phytodecta ........................ 82Phytoecia ......................... 79Phytonomus ....................... 90Phytosus ........................... 37Pidonia ............................. 76Pilemostoma .................... 85Pissodes ........................... 94Pityogenes ........................ 98

Pityokteines ..................... 98Pityophagus ..................... 62Pityophthorus .................. 99Pityphilus ........................... 78Placonotus ....................... 63Placusa ............................ 37Plagiodera ....................... 81Plagionotus ...................... 78Plagiosterna ..................... 81Planeustomus ................... 38Platambus .......................... 7Plataraea, subg. ............... 34Plateumaris ...................... 79Platycerus ........................ 46Platycis ............................ 54Platydema ........................ 73Platyderus ........................ 12Platydracus ...................... 44Platylomalus .................... 17Platynaspis ....................... 67Platynus ........................... 12Platypsyllus ...................... 22Platypus ........................... 99Platyrhinus ....................... 86Platysoma ........................ 18Platystethus ..................... 40Platystomos ..................... 86Plectophloeus ................... 26Plegaderus ....................... 17Pleurophorus .................... 47Pocadiodes ....................... 61Pocadius .......................... 61Podabrus .......................... 54Podagrica ........................ 84Poecilium ......................... 78Poecilonota ...................... 49Poecilus ............................ 11Pogonocherus .................. 78Pogonus ........................... 10Polydrusus ....................... 88Polygraphus ..................... 97Polyphylla ........................ 47Poophagus ....................... 95Porcinolus ........................ 50


Porhydrus .......................... 6Poromniusa ...................... 31Porotachys ....................... 11Porrhodites ...................... 24Potamonectes ...................... 7Potosia ............................. 48Pragensiella ....................... 37Prasocuris ........................ 81Pria .................................. 61Priobium .......................... 57Prionocyphon ................... 49Prionus ............................. 76Prionychus ....................... 73Pristonychus ...................... 12Procraerus ....................... 53Propylea ........................... 67Prosternon ....................... 52Prostomis ......................... 74Proteinus .......................... 25Psammodius ..................... 47Psammoecus .................... 62Pselactus .......................... 94Pselaphaulax ................... 27Pselaphorhynchites ............ 86Pselaphus ......................... 27Psephidonus ...................... 25Pseudanostirus ................. 52Pseudaphana ..................... 30Pseudathous ...................... 52Pseudeuglenes .................. 75Pseudeurostus .................. 57Pseudocistela ................... 73Pseudocleonus ................. 90Pseudomedon ................... 41Pseudomicrodota ............. 37Pseudoophonus ................. 13Pseudophloeophagus ....... 94Pseudoplectus .................. 26Pseudoptilinus .................. 57Pseudostyphlus ................ 91Psilothrix ......................... 59Psylliodes ......................... 84Psyllobora ........................ 67Pteleobius ........................ 97

Ptenidium ......................... 19Pteroloma ........................ 20Pteronius ........................... 25Pterostichus ..................... 11Pteryngium ...................... 63Pteryx .............................. 19Ptilinus ............................. 57Ptiliola ............................. 19Ptiliolum .......................... 19Ptilium ............................. 19Ptinella ............................. 19Ptinomorphus .................... 56Ptinus ............................... 57Ptomaphagus ................... 22Pullus ................................ 66Pycnoglypta ..................... 24Pycnomerus ..................... 72Pycnota ............................ 35Pyrochroa ......................... 74Pyroglossa ........................ 30Pyropterus ........................ 54Pyrrhalta .......................... 82Pyrrhidium ....................... 78Pytho ................................ 74Quedius ............................ 45Rabigus ............................ 43Rabocerus ........................ 75Reesa ................................ 56Reichenbachia .................. 27Reitterelater ....................... 53Remus ............................... 44Rhacopus .......................... 51Rhagium ........................... 76Rhagocneme, subg. ........... 33Rhagonycha ..................... 55Rhamnusium .................... 76Rhamphus ........................ 93Rhantus .............................. 7Rhaphitropis ..................... 85Rhinocyllus ....................... 90Rhinomacer ....................... 85Rhinoncus ......................... 95Rhinosimus ........................ 75Rhinusa ............................ 93

Rhizophagus ..................... 62Rhopalocerina .................. 37Rhopalomesites ................ 94Rhynchaenus .................... 92Rhynchites ........................ 86Rhyncolus ......................... 94Rhysodes .......................... 10Rhyssemus ........................ 47Rhyzobius ......................... 66Rhyzopertha ..................... 56Riolus ............................... 50Ripidius ............................ 71Ropalodontus ................... 70Ropalopus ........................ 77Rosalia ............................. 77Rossioptera ....................... 54Rugilus ............................. 41Rushia .............................. 70Rusticoclytus ..................... 78Rutidosoma ...................... 95Rybaxis ............................. 27Rypobius ........................... 68Sacium .............................. 68Salpingus ........................... 75Salpingus .......................... 75Saperda ............................ 79Saprinus ........................... 17Saulcyella ......................... 26Scaphidema ...................... 73Scaphidium ...................... 38Scaphisoma ...................... 38Scarodytes .......................... 7Schistoglossa .................... 32Schizotus .......................... 74Sciaphilus ......................... 89Sciaphobus ....................... 89Scintillatrix ......................... 49Sciodrepa .......................... 22Sciodrepoides ................... 22Scirtes .............................. 49Scleropterus ..................... 95Scolytus ............................ 97Scopaeus .......................... 42Scotodes ........................... 74


Scraptia ............................ 75Scydmaenus ..................... 23Scydmoraphes .................. 23Scymnus ........................... 66Selatosomus ..................... 52Semanotus ........................ 78Semiadalia ......................... 67Semiophonus ..................... 13Sepedophilus .................... 28Serica ............................... 47Sericoda ........................... 12Sericoderus ...................... 68Sericus .............................. 53Sermylassa ....................... 82Serropalpus ...................... 70Sibinia .............................. 92Silis ................................... 55Silpha ............................... 23Silusa ................................ 37Silvanoprus ...................... 62Silvanus ............................ 62Simo ................................. 88Simplocaria ...................... 50Sinodendron ..................... 46Sipalia ............................... 32Sipalia ............................... 36Sirocalodes ....................... 96Sitodrepa .......................... 57Sitona ............................... 89Sitophilus ......................... 97Smaragdina ...................... 80Smicronyx ........................ 91Smicrus ............................ 20Sogda ............................... 20Soronia ............................. 61Sospita ............................. 67Spatulonthus ...................... 43Spavius ............................. 64Spercheus ......................... 15Spermophagus ................. 85Sphaericus ........................ 57Sphaeridium ..................... 17Sphaeriestes ..................... 75Sphaerites ........................ 17

Sphaerius ........................... 5Sphaeroderma .................. 84Sphaerosoma .................... 66Sphindus ........................... 60Sphodrus .......................... 12Spondylis .......................... 76Stagetus ............................ 58Staphylinus ....................... 44Stegobium ........................ 57Stenagostus ...................... 52Stenagria ........................... 36Stenelmis .......................... 50Stenichnus ........................ 23Stenister ............................ 18Stenocarus ........................ 97Stenocorus ........................ 76Stenolophus ...................... 14Stenomax ......................... 72Stenostola ......................... 79Stenotrachelus .................. 74Stenus ............................... 40Stephanocleonus .............. 90Stephanopachys ............... 56Stephostethus ................... 68Stereocorynes ................... 94Stereonychus .................... 91Stethorus .......................... 66Stichoglossa ..................... 30Stictotarsus ........................ 7Stilbus ............................... 63Stilicus ............................... 41Stomis ............................... 11Strangalia ......................... 77Stricticomus ..................... 75Strophosoma .................... 89Subcoccinella ................... 68Sulcacis ............................ 70Sunius ............................... 41Suphrodytes ........................ 6Symbiotes ......................... 66Synaptus ........................... 54Syncalypta ......................... 50Synchita ........................... 72Syneta .............................. 81

Synharmonia ...................... 67Syntomium ....................... 38Syntomus .......................... 15Synuchus .......................... 12Tachinus ........................... 28Tachyerges ....................... 92Tachyporus ....................... 28Tachys .............................. 11Tachyta ............................ 11Tachyura .......................... 11Tachyusa ........................... 31Tachyusida ....................... 37Tanymecus ....................... 89Tanysphyrus ..................... 91Tapeinotus ........................ 95Taphrorychus ................... 98Tarsostenus ...................... 59Tasgius ............................. 45Telmatophilus ................... 63Temnocerus ...................... 86Temnorhinus .................... 90Temnoscheila ................... 58Tenebrio ........................... 72Tenebroides ...................... 58Teredus ............................. 65Teretrius ........................... 17Tetartopeus ....................... 42Tetraplatypus ..................... 14Tetratoma ........................ 70Tetropium ......................... 76Tetrops ............................. 79Thalycra ........................... 61Thamiaraea ...................... 35Thamiocolus ..................... 96Thanasimus ...................... 58Thanatophilus .................. 23Thea .................................. 67Theca ................................ 58Thecturota ........................ 37Theetura ............................ 37Thes .................................. 68Thiasophila ...................... 30Thinobaena, subg. ............ 35Thinobius ......................... 38


Thinodromus .................... 38Thorictodes ...................... 55Throscus ........................... 51Thryogenes ....................... 90Thyasophila ....................... 30Thylodrias ........................ 56Thymalus .......................... 58Tillus ................................. 58Timarcha .......................... 82Tinotus ............................. 29Tipnus ............................... 57Tomicus ............................ 97Tomoglossa ...................... 32Tomoxia ........................... 71Trachodes ......................... 94Trachypachus ..................... 5Trachyphloeus .................. 88Trachys ............................. 50Tragosoma ....................... 76Traumoecia, subg. ............ 34Trechicus ........................... 14Trechoblemus ................... 10Trechus ............................. 10Triarthron ......................... 20Tribolium .......................... 72Trichius ............................ 48Trichiusa .......................... 35Trichoceble ....................... 59Trichocellus ....................... 14Trichodes .......................... 58Trichohydnobius ................ 20Trichomicra ...................... 31Trichonyx ......................... 26Trichophya ....................... 29Trichosirocalus ................. 97Trigonogenius .................. 56Trimium............................ 26Trinodes ........................... 55Triphyllus ......................... 69Triplax .............................. 65Trissemus ......................... 27Tritoma ............................ 65Trixagus ........................... 51Trochanterella, subg. ........ 33

Trogoderma ...................... 56Trogophloeus ..................... 38Trogossita .......................... 58Trogoxylon ....................... 56Tropideres ......................... 85Tropiderinus ...................... 85Tropinota .......................... 48Tropiphorus ...................... 89Trox .................................. 46Trypodendron ................... 98Trypophloeus .................... 98Tychius ............................. 91Tychus .............................. 26Typhaea ............................ 69Typhaeus .......................... 46Tyrus ................................ 27Tytthaspis ......................... 67Uleiota ............................. 62Uloma .............................. 72Upis .................................. 74Urodon ............................. 86Wagaicis ........................... 70Valgus ............................... 48Vanonus ............................ 75Variimorda ....................... 71Velleius ............................. 45Veraphis ........................... 22Vibidia .............................. 67Vincenzellus ...................... 75Xanthochroa ...................... 74Xanthogaleruca ............... 82Xantholinus ...................... 42Xanthosphaera ................. 21Xenota, subg. .................... 33Xestobium ........................ 57Xyleborinus ...................... 98Xyleborus ......................... 98Xylechinus ........................ 97Xyletinus .......................... 57Xylita ................................ 70Xylobanellus ..................... 54Xylobius ............................ 51Xylocleptes ....................... 98Xylodrepa ......................... 23

Xylodromus ...................... 24Xylographus ..................... 70Xylolaemus ....................... 72Xylophilus ........................ 51Xylostiba .......................... 24Xyloterus ........................... 98Xylotrechus ...................... 78Zabrus .............................. 13Zacladus ........................... 96Zavaljus ........................... 65Zeugophora ...................... 79Zilora ............................... 70Zorochros ......................... 53Zyras ................................ 36

Enumeratio nova Coleopterorum Fennoscandiae, Daniae et ... · Enumeratio nova Coleopterorum...

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Transcript of Enumeratio nova Coleopterorum Fennoscandiae, Daniae et ... · Enumeratio nova Coleopterorum...