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Special PublicationsMuseum of Texas Tech Un iversi t yNumber 55 20 June 2008title
Authors.0
Special PublicationsMuseum of Texas Tech Un iversi t yNumber 56 18 September 2009Phyllostomid Bats of Colombia: Annotated
Checklist, Distribution, and Biogeography
Hugo Mantilla-Meluk, Alex Mauricio Jimnez-Ortega, and Robert J. Baker
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Front cover: Map of Colombian phyllostomid species richness obtained by the superimposition of models of potential
dro for h 118 c cormd for Coloma. Warm color rr hgh lvl of c rch; cool
color rr low lvl of c rch. A how o h ma, 80% of h Coloma rrory hold al
vrom for hylloomd whch ar al o oally rcr ovr 30 c a gl locao. Of arclar
r h orhr oro of h Coloma Amazo, whch rr o-forh of h ara of h cory
ad characrzd y a hgh dvry of hylloomd a. Or modl of hylloomd rch dca ha h
orhr oro of h Coloma Amazo hold ara whr o 57 hylloomd c oally ca fod
ymary. Covrly, h hghlad of h Coloma Ada ym, h avaa of h Coloma Oroqa,
o h ar oro of h cory, ad h ard vrom of La Gajra la, h orhrmo oro
of h Coloma Cara rgo, oally clo h l al vrom for hylloomd c ad
rr aral arrr for hr dro.
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Texas Tech University
and
Universidad Tecnolgica del Choc Diego Luis Cordoba
Hugo Mantilla-Meluk, alexMauricio JiMnez-ortega, and robertJ. baker
SpeCiAL PubLiCAtiOns
Museum of Texas Tech UniversityNumber 56
Phyllostomid Batsof ColomBia: annotatedCheCklist, distriBution, and BiogeograPhy
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Layo ad Dg: La bradly
Covr Dg: Hgo Malla-Mlk
Coyrgh 2009, Mm of txa tch uvry
All rgh rrvd. no oro of h ook may rrodcd ay form or y ay ma,
cldg lcroc orag ad rrval ym, xc y xlc, ror wr rmo
of h lhr.
th ook wa tm nw Roma ad rd o acd-fr ar ha m h gdl
for rmac ad draly of h Comm o prodco Gdl for book Logvy of
h Cocl o Lrary Rorc.
prd: 18 smr 2009
Lrary of Cogr Caalogg--plcao Daa
scal plcao of h Mm of txa tch uvry, nmr 56
sr edor: Ror J. bakr
phylloomd ba of Coloma: Aoad Chckl, Dro, ad bogograhy
Hgo Malla-Mlk, Alx Marco Jmz-Orga, ad Ror J. bakr
issn 0169-0237
isbn 1-929330-18-9
isbn13 978-1-929330-18-8
Mm of txa tch uvry
Lock, tX 79409-3191 usA
(806)742-2442
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1
Phyllostomid Batsof ColomBia: annotated CheCklist, distriBution, and
BiogeograPhy
Hugo Mantilla-Meluk, alexMauricio JiMnez-ortega, and robertJ. baker
aBstraCt
W ror 118 cormd hylloomd c for Coloma, ad 14 c o-
ally r h cory, for a oal of 132 c rrg 40 gra ad10
faml. All kow hylloomd faml ar rrd Coloma, wh
h xco of h famly Macroa (o r soh Amrca). A 118
kow c, Coloma ha h gra mr of hylloomd a of ay cory.
icldd h l ar 27 rcly rcogzd c (19 wly dcrd c ad
gh wly lvad c), 19 of whch ar cormd for Coloma, whl gh ar
oally r. th chckl accomad y 18 axoomc comm xla-g rc chag h axoomy of h gro. i addo, Gograhc iformao
sym (Gis)-ad modl of oal dro wr crad for oh cormd
ad oally r hylloomd c, ad c rch ar wr aa-
lyzd. Fally, comm o h ogograhy of h gro ar cldd.
Ky word: a, ogograhy, chckl, Chrora, Coloma, dro
modl, phylloomda, axoomy
resumen
Roramo 118 c d mrclago lomdo ara Coloma y 14 -
c ocalm r l a, ara oal d 132 c q rra
40 gro y 10 famla. toda la famla d mrclago lomdo
r Coloma co xcc d d la famla Macroa (a sr
Amrca). Co 118 c rcoocda, Coloma l a q o l mayor
mro d mrclago lomdo. nro lado cly 27 c rcoocda
rcm (19 rcm dcra y ocho rcm lvada), 19 d la
cal cormada ara Coloma mra ocho corrod a c o-
calm r l a. nra la acomaada or 18 comaro
q xlca lo camo rc la axooma dl gro. Adcoalm, mod-
lo d drc ocal aado sma d iformac Gogrca (siG)
fro grado ao ara la c d mrclago lomdo r como
ocal l a, al mo q aro d dvrdad fro aalzado ara
lo mrclago lomdo d Coloma. Falm lo aro gral d la
ogografa dl gro o comado.
palara clav: ogografa, Coloma, la aoada, modlo d drc,
mrclago, phylloomda, Qrro, axooma
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2 sPeCial PuBliCations, museumof texas teCh university
introduCtion
i rc yar, h avalaly of gc daa for
drmg hylogc rlaoh, a wll a hdcro of w c ad lvao of c
o c a, ha rld aal rvo a
may axoomc lvl of norocal mammal. phyl-
loomd a ar o of h mo vly dd
gro of norocal mammal ad a major orc of
w c dcro ad hghr axoomc chag
o accommoda moohyly. Hr, rvo work,
such asMamferos (Synapsida: Theria) de Colombia
(Alrco al. 2000),Mammal Species of the World
(Wlo ad Rdr 2005),Mammals of South America
(Gardr 2008), ad h Global Mammal Assessment
(iuCn 2008), wr d o gra a chckl of hyl-loomd a from Coloma. th chckl cld
rcly dcrd axa ad wly lvad c, a
wll a ho c ha mgh xcd o occr
Coloma. th ymac clacao of bakr
al. (2003) followd. th ymac arragm
is based on statistically supported monophyly which
wa o achvd rvo clacao of h
comlx of a (bakr al. 1989; Kooma 1993;
Wrr al. 2000). to facla comaro of or
chckl o ha of Alrco al. (2000) (h mocomrhv chckl of Coloma mammal
avalal o da), w follow hr forma. Dffrc
w Alrco al. (2000) ad h crr chck-
l ar hghlghd wh h l y ymol, whch
ar xlad mor dal h x. th chckl
alo cld rfrc. Modl of oal dr-
tion for each Colombian phyllostomid species also
ar rovdd; h ma wr grad ad o
margal local drvd from collcg local
of mm vochr cm from major mammal
collco Coloma ad h ud sa. th d-
ro ma wr h d o vga af c comoo amog Coloma corgo
a old y Hrdz-Camacho al. (1992). i
addo, dro modl wr d o drm
phyllostomid richness distribution in Colombia and
h rlaoh w vromal varal ad
ar of c rch. Fally, w dc h
org of h gograhc arog orvd amog
Coloma hylloomd a.
methods
Data gathering.A v arch for Co-
lombian phyllostomid bat records was conducted us-
g hr rmary orc: dgal mm daaa,
rcord rord h cc lrar, ad drc
co of mm vochr cm Colom-
a ad Amrca o. th arch rld
19,952 rcord oad from mm daaa ad
lrar rcord, a follow: Amrca Mm of
naral Hory (AMnH) (n = 950); brh Mm
of naral Hory (bM) (hr Coloma rcord
rord h lrar); io Alxadr vo
Hmold (iAvH) (n = 91); io d Cca na-ral, uvrdad nacoal d Coloma (iCn) (n =
9,477); Fld Mm of naral Hory (FMnH) (n
= 1,556); Mo d Hora naral uvrdad dl
Caca (MnHuC) (n = 543); Mo d Hora na-
ral uvrdad d lo Ad (MnHu) (x lcd
rcord rord h lrar); Mo d Hora
naral uvrdad Dral d bogo Fracco
Jo d Calda (MuD) (n = 330); Mo d Ho-
ra naral d la uvrdad tcolgca dl Choc
(MZCH) (n = 600); naoal Mm smhoa i-
o (usnM) (n = 5,980); Royal Oaro Mm
(ROM) (52); txa tch uvry Mm (ttu)
(n = 259); uvrdad dl Vall (uV) (n = 105). i
addo, a lcd of hylloomd rcord
wa axoomcally xamd y drc co of
mm vochr cm a follow: iCn (n = 788);
FMnH (n = 220); ttu (n = 75); usnM (n = 193).
th coml l of cm h lmary
maral co avalal ol a h wag of
h naral scc Rarch Laoraory, Mm oftxa tch uvry (www.rl..d).
Distribution models.Models of potential distri-
bution were created for each phyllostomid bat species
r Coloma. th modl wr grad
g a daa o h margal local Gardr
(2008) a wll a 17,778 rcord of Coloma hyl-
loomd c w wr al o gorfrc ad
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mantilla-meluketal.Phyllostomid Batsof ColomBia 3
o lcroc gazr avalal from Fallg Ra
Gomc, ic. (www.fallgra.com) ad h naoal
Goaal illgc Agcy (h://goam.ga.
ml/ggmavwr/MaFrams.a).
Gorfrcd Coloma hylloomd am-
lg local wr covrd o o olygo
layr ArcGis 9.3. each layr rrg c
droal o wa h lacd ovr a olygo
layr of norocal corgo oad from h w-
of h World Wldlf Fd (h//:www.wwf.org).
ecorgo rcg c droal rcord
wr lcd g h Select by location option of
ArcGis 9.3 ad xord a vcor l. th xord
l rrg lcd corgo wr a mak
h saal Aaly xo of ArcGis 9.3. Rarlayr rrg mmm ad maxmm c
lvaoal lm wr h crad for ach c
ad o lvaoal rag rord h lrar.
Dgal formao o lvao for h noroc wa
drvd from a Dgal elvao Modl (DeM) aval-
al a h w WorldClm (h://www.worldclm.
org/oclm.hm) (cr cororad o h saal
Aaly xo of ArcGis 9.3: [_am_lva-
oal_rag] = ([DeM] >= mmm lvaoal val
AnD [DeM]
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4 sPeCial PuBliCations, museumof texas teCh university
Table 1. List of phyllostomid bats from Colombia. Symbols are as follows: additions and changes to Alberico et al.
(2000) (+); endemic (); taxonomic comments and consideration (). Elevational ranges in meters. Abbreviations
of Colombian natural regions: Amazon (amz); Andean (and); Caribbean (car); Orinoquia (ori); and Pacic (pac).
Abbreviations of Colombian departments: Amazonas (ama); Antioquia (ant); Arauca (ara); Atlntico (atl); Bolivar
(bl) (2); Boyac (by); Caldas (cl); Caquet (caq); Casanare (cas); Cauca (cau); Cesar (ce); Choc (cho); Crdoba
(co); Cundinamarca (cun); Guaina (gn); Guaviare (gv); Huila (hu); La Guajira (gua); Magdalena (ma); Meta (met);
Nario (na); Norte de Santander (nsn); Putumayo (pu); Quindo (qui); Risaralda (ri); San Andrs (sand); Santander
(snt); Sucre (su); Tolima (to); Valle del Cauca (vc); Vaups (va); Vichada (vi). References of the scientic literature
used as sources for records of Colombian phyllostomid bats are included in Appendix III.
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference Collections
PHYLLOSTOMIDAE
Subfamily
Micronycterinae
LampronycterisSanborn
1949Lampronycteris brachyotisDobson 1879
amz andcar ant gua va bl to 0-700
Marinkelle &Cadena 1972
IAvH ICN 16955MHNU USNM
MicronycterisGray 1866
Micronycteris hirsuta
(Peters 1869)
amz and
car pac
caq cau cl cho
ma vc 20-1100 Sanborn 1932
AMNH ICN 17236
MHNUC UV
Micronycteris megalotis
(Gray 1842) co
ant caq cau cun
cl cho ri vc
bo cas ce co
hu gua ma me
qui sand snt vi 25-2400 Dobson 1878
AMNH FMNH IAvH
ICN 1597 MHNUC
USNH UV
Micronycteris microtisMiller, 1898
amz andcar ori ant caq cun 15-2100 Simmons 1996 AMNH 99344
Micronycteris minuta
(Gervais 1856) co
ant cl cho met
ma na vc
ama caq cas
co cun gua
nsn pu snt 5-1130 Koopman 1982
AMNH FMNH IAvH
ICN MZCH TTU
USNM UV
Micronycteris schmidtorum
(Sanborn 1935)
amz and
car pac ant cho snt gua ma 10-160 Koopman 1982 MZCH ICN 14751
SubfamilyDesmodontinae
Tribe Desmodontini
DesmodusWied-Neuwied
1826
Desmodus rotundus (E.
Geoffroyi St. Hilaire 1810) co
ant caq cau cl
cho cun gn na ri
snt to vc
ama bl bo cas
ce hu gua ma
me na qui vi 0-2600 J.A. Allen 1900
AMNH FMNH IAvHICN 466 MHNUC
MUD MZCH ROM
TTU USNM UV
Diaemus Miller 1906
Diaemus youngi (Jentick
1893) co
ant caq cau cho
pu vc cas hu me 0-500
de la Torre
1956 IAvH ICN 7580 UV
Tribe Diphyllini
Diphylla Spix 1823
Diphylla ecaudata Spix
1823 co ant caq cau ma gv me va 0-500 J.A. Allen 1900
AMNH FMNH IAvH
ICN 14568 MHNUC
USNMSubfamily
Lonchorhininae
Lonchorhina Tomes 1863
Lonchorrhina aurita
Tomes 1863 co
ant caq cau cho
met na vc cl co nst ri 25-1550
Nicforo Mara
in Sanborn
1949
AMNH FMNH IAvH
ICN 14716 MHNUC
MZCH USNM UV
Lonchorrhina marinkellei
Hernndez-Camacho andCadena 1978 amz caq va 0-500
Hernndez
Camacho &Cadena 1978 ICN (holotype)
Lonchorrhina orinocensis
Linares and Ojasti 1971 amz ori caq met ama va vi 75-620
Linares &
Ojasti 1971 IAvH ICN 14671
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mantilla-meluketal.Phyllostomid Batsof ColomBia 5
Table 1. (cont.)
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference Collections
Subfamily
Phyllostominae
Tribe Macrophyllini
Macrophyllum Gray 1838
Macrophyllum
macrophyllum (Schinz1821) co ant cho ma vc
bl gua snt tova 10-1070 J.A. Allen 1900
IAvH ICN 18773MUD UV
TrachopsGray1847
Trachops cirrhosus (Spix
1823) co
ant caq cau ma
vc
ama ara bl bo
ce gua met punsn snt to va
vi 10-1120 Dobson 1878
AMNH FMNH IAvH
ICN 544 MHNUCMZCH TTU USNM
UV
Tribe Phyllostomini
Lophostoma d'Orbygni1836 + (1)
Lophostoma brasiliensePeters 1866 +
amz carori pac
ant cho gn metvc
caq cun hugua met pu va 10-1100
Lemke et al.1982
FMNH ICN 17932USNM UV
Lophostoma carrikeri (J.A. Allen 1910) + amz ori caq met 45-700
McCarty et al.1983 ICN 5140 FMNH
Lophostoma silvicolum
d'Orbygni 1836 + co
ant caq cau cl
cho gn ma vc
gua me pu to
va 10-850 J.A. Allen 1900
AMNH IAvH ICN
17877 MZCH USNM
UV
Mimon Gray 1847
Mimon bennettii (Gray
1838) (2) amz ori cq gn me 0-500
Montenegro
and Romero-
Ruiz 2000
FMNH 113425 ICN
USNM
Mimon cozumelae
Goldman 1914 (2) + car pac ant cho co 0-500
Marinkelle &
Cadena 1972
FMNH 69427 USNM
43175
Mimon crenulatum (E.
Geoffroyi St. Hilaire 1803) co
ant caq cau cl
cho vc
co gn gua ma
me va vi 5-830 Handley 1960
IAvH ICN 10231MHNUC MZCH TTU
UV
Phylloderma Peters 1865Phylloderma stenops Peters
1865 co
ama ant caq cho
gn met vc gn pu 0-1100
Marinkelle &
Cadena 1972
IAvH ICN 4465 TTU
UV
PhyllostomusLacpde
1799
Phyllostomus discolor
(Wagner 1843) co
ant caq cau clcho cun gn met
ri to vc
bl bo cas hu
ma nsn su 10-1650
Valdivieso and
Tamsitt
AMNH FMNH IAvHICN 458 MZCH
USNH UV
Phyllostomus elogatus (E.Geoffroyi St. Hilaire 1810) amz caq cau gn pu cas co met va 10-850 Furman 1966
FMNH IAvH ICN
8320 MUD TTUUSNM
Phyllostomus hastatus
(Pallas 1767) co
ant bo caq caucl cho cun gn
na to vc
ama bl cas cehu gua met snt
su vi 0-1295 J.A. Allen 1900
AMNH FMNH IAvH
ICN 639 MHNUCMUD MZCH ROM
TTU USNM UV
Phyllostomus latifolius
(Thomas 1901) amz pac caq cau va vc 0-500
Marinkelle &
Cadena 1972
ICN 14603 MHNUC
MNHU MZCH UVTonatiaGray 1827
Tonatia saurophila
Koopman and Williams
1951
amz and
ori pac ant caq cau met hu ga pu va 10-140
Sanchez-
Palomino et. al
1993 FMNH ICN 10254 UV
Tribe Vampirini
ChrotopterusPeters 1865
Chrotopterus auritus
(Peters 1865) co
ant caq cau cho
me ma gn gua 0-850 J.A. Allen 1900
AMNH IAvH ICN
14566
Vampyrum Refinesque
1815
Vampyrum spectrum
(Linnaeus 1758) co
ant caq cau cl
cho na vc ama cu met vi 10-1065
Hall & Kelson
1959 IAvH ICN 176 USNM
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6 sPeCial PuBliCations, museumof texas teCh university
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference CollectionsSubfamily
Glossophaginae
Tribe Choeronycterini
AnouraGray 1838
Anoura aequatoris
(Lnnberg 1921) + and ant cau cl na ri hu to vc 1000-3000
Mantilla-Meluk
and Baker 2006
FMNH ICN 7615
MHNUC
Anoura cadenai Mantilla-Meluk and Baker2006 and pac cau vc 1000-1500
Mantilla-Melukand Baker 2006 ICN holotype NMNH
Anoura caudifer(E.
Geoffroyi St. Hilaire 1818)
amz and
car ori
ant cau caq cecl cho ma na ri
snt to va vc
cas cun hu guamet pu to va
vi 500-2800
Mantilla-Meluk
and Baker 2006
AMNH FMNH IAvH
ICN 13833 MHNUCMUD ROM TTU
USNM UV
Anoura cultrata Handley
1960
and car
pac
ant caq cau cl
cho nsn snt vc
cun met na
nsn 0-1800
Lemke and
Tamsitt 1979
FMNH IAvH ICN
7616 MHNUC UV
Anoura fistulata Muchhala,
Mena, and Albuja 2005 + and na cau ri 1000-1800
Mantilla-Meluk
and Baker 2008 FMNH 113512 ICN
Anoura geoffroyi Gray
1838 co
ant caq cau clcho cun ri snt
vc
cun gn met na
to va 500-3600 J.A. Allen 1916
AMNH FMNH IAvH
ICN 843 MHNUCMUD ROM USNM
UV
Anoura latidens Handley1984 and cau 1000-1500 Handley 1984 MHNUC 1552
Anoura luismanueli
Molinari 1994 + and cun snt 1000-1500
Mantilla-Meluk
and Baker 2006 ICN 6602 MUD
ChoeroniscusThomas1928
Choeroniscus godmani
(Thomas 1903) and car
ant cau cl na
met vc cun 0-1600
Tamsitt et al.
1965
ICN 8064 MHNUC
UV
Choeroniscus minor(Peters 1868) (3) amz and
cau caq cho cungn to pu va 0-1032
Valdivieso1964
ICN 9775 MHNUCTTU UV
Choeroniscus periosus
Handley 1966 pac cau cho vc na 0-500 Handley 1966
AMNH ICN MZCH
NMNS (holotype)UVLichonycterisThomas
1895
Lichonycteris degener
Miller 1931 + amz caq gv 0-500
Montenegro
and Romero-
Ruiz 2000 ICN 14571
Lichonycteris obscura
Thomas 1895 (4) amz pac
ant caq cl cho
vc co ma na to 0-500
Marinkelle &
Cadena 1972
FMNH IAvH ICN
MZCH USNM UV
Scleronycteris Thomas1912
Scleronycteris ega Thomas
1912 (5) amz va 0-500
Alberico et al.
2000 IAvH ?
Tribe Glossophagini
Glossophaga E. GeoffroyiSt. Hilaire 1818
Glossophaga commissarisi
Gardner 1962 amz and ama ant cho
bl caq ce gn
ma met su vi 0-1000
Webster and
Jones 1987
IAvH ICN MUD
MZCH TTU 9093 UV
Glossophaga longirostris
Miller 1898 and car ant cl cun ma
atl bl bo ce
gua su vi 5-1050 Miller 1898
AMNH FMNH IAvH
ICN 5361 MUD
USNM UV
Glossophaga soricina
(Pallas 1766) co
ant cau caq cl
cho ma met gn
to vc
ara caq cas
cun gn vi 0-1560 J.A. Allen 1900
AMNH FMNH IAvH
ICN 1749 MHNUC
MZCH ROM TTU UV
Leptonycteris Lydekker1891
Leptonycteris curasoae
Miller 1900 and car bl snt bo gua ma 0-900
Marinkelle and
Grose 1966
FMNH MHNU ICN
2521 USNM
Table 1. (cont.)
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mantilla-meluketal.Phyllostomid Batsof ColomBia 7
Table 1. (cont.)
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference CollectionsSubfamily
Lonchophyllinae
LionycterisThomas 1913
Lionycteris spurrelli
Thomas 1913
amz and
car pac
ant caq cau cho
cun na me vc
gn cas gn va 90-1400 Thomas 1913
BM holotype IAvH
ICN MZCH USNM
UV
LonchophyllaThomas
1903
Lonchophylla cadenai
Woodman and Timm 2006+ and pac ri vc 0-1500
Woodman andTimm 2006 ICN USNM holotype
Lonchophylla chocoana
Dvalos 2004 + pac vc 500-1000 Dvalos 2004
ICN ROM holotype
MZCH USNM
Lonchophylla concavaGoldman 1914 + (6)
and carpac
cau cho na quisnt vc 0-1000
Albuja andGardner 2005
FMNH ICN 7009MHNUC USNM
Lonchophylla fornicata(Woodman 2007) pac cau 500 - 1560 Woodman 2007 ICN 13647 MHNUC
Lonchophylla handleyi Hill1980 + (7) and
ant cau cho hunar vc 500-1000
Alberico andOrejuela 1983 MHNUC 718 UV
Lonchophylla
orienticollina Dvalos andCohortals 2009 + (8) and ori met 0-1500
Dvalos &Corthals 2009 ICN holotype
Lonchophylla robustaMiller 1912
and carpac
ant cl cho cunna snt vc
bo ce gua mansn to 0-1900 Sanborn 1949
AMNH FMNH IAvH
ICN 4397 MHNUCMZCH USNM UV
Lonchophylla thomasi J. A.
Allen 1904 co ant caq vc
ama bo gua
met pu ri va vi 0-1000 Koopman 1982
FMNH IAvH ICN
9479 MZCH TTU
USNM UV
Subfamily Carollinae
CarolliaGray 1838 (9)
Carollia brevicauda
(Schinz 1821) co
ant cau caq clcho gn met na ri
snt to vc
ama ara bo cas
ce cun gn guahu ma me nsn
qui to va vi 500-2000 Bangs 1900
AMNH FMNH IAvH
ICN 5228 MHNUCMUD MZCH ROM
TTU USNM UV
Carollia castanea H. Allen
1890 co
ant caq cau cl
cho gn nsn ri vc
ama bo cas cocu ma me pu
snt to 0-1500
Hershkovitz
1949
AMNH FMNH IAvH
ICN 7049 MHNUCMUD MZCH TTU
USNM UV
Carollia monohernandeziCuartas and Gonzlez 2004
+
and car
pac ant bl caq cho 30-2660
Muoz, Cuartasand Gonzlez
2004 MUA holotype MZCH
Carollia perspicillata
(Linnaeus 1758) co
ant caq cau cl
cho cun gn ma
met ri snt to vc
ama ara atl bl
bo cas ce cogn gua gv na
nsn pu su va
vi 0-2000 Dobson 1878
AMNH FMNH IAvHICN 555 MUD MZCH
ROM TTU USNMH
UV
Subfamily
Rhonophyllinae
RhinophyllaPeters 1865
Rhinophylla alethina
Handley 1966 pac ant cau vc cho 0-1000 Handley 1966
AMNH FMNH ICNMHNUC MZCH
NMNM (holotype) UV
Rhinophylla fischerae D.C. Carter 1966 amz ama caq cau gn met pu 0-500
Marinkelle &Cadena 1972
FMNH IAvH ICN5161TTU
Rhinophylla pumilio Peters
1865 amz ori
ama caq cau gn
met pu vi gn va 0-500
Barriga-Bonilla
1965
MHNU IAvH ICN 671
TTU UV
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8 sPeCial PuBliCations, museumof texas teCh university
Table 1. (cont.)
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference Collections
Subfamily
Glyphonycterinae
GlyphonycterisThomas
1896 (10)
Glyphonycteris sylvestris(Thomas 1896) amz car ant caq co 15-1100
Montengro and
Romero-Ruiz2000 ICN 172392
NeonycterisSanborn 1949
Neonycteris pusilla
(Sanborn 1949) + (11) amz va 0-500 Sanborn 1949
AMNH 78830
Brazil-Colombia
TrinycterisSanborn 1949
Trinycteris nicefori
Sanborn 1949
amz and
pac
caq cho nsn to
vc ama gn met 15-150 Sanborn 1949
AMNH holotype ICN
MZCH UV
Subfamily
Stenodermatinae
Tribe Sturnirini
SturniraGray 1842
Sturnira aratathomasi
Peterson and Tamsitt 1968 and ant cau cl ri vc 1600-2800
Peterson &
Tamsitt 1968
IAvh UV ROM
(holotype) USNM
Sturnira bidens (Thomas
1915) and
ant cau cl cho
na ri to
bo hu met qui
vc 1800-3100
Marinkelle &
Cadena 1972
IAvH ICN 1700 ROM
MHNU MHNUC UV
Sturnira bogotensis Shamel1927 and ant na to cun met ris 1500-3100 Shamel 1927
IAvH ICN MUDUSNM holotype
Sturnira erythromus
Tschudi 1844 and
ant cau cl nar ri
snt to
bo cas ce cun
ma met nsn
qui to vc 1800-3500
Lemke et al.
1982
FMNH IAvH ICN
5365 MHNUC TTU
USNM UV
Sturnira koopmanhilli
McCarthy and Albuja 2006
+ (12) and pac ant cau cho na 1000-1500
McCarty,
Albuja, and
Alberico 2006
MHNUC MZCH UV
4442
Sturnira lilium (E.Geoffroyi St. Hilaire 1810) co
ant cau cho cun
gn ma na ri sntto vc
ama bl bo clcas gn gua hu
met nsn pu suva 0-1900
Hershkovitz1949
IAvH ICN 683
MHNUC MUDMZCH TTU UV
Sturnira luisi Davis 1980 and pac
ant cau cho to
vc atl ce met snt 0-500
Alberico &
Negret 1992
FMNH ICN MHNUC
MZCH USNM
499420UV
Sturnira magna de la Torre1966 amz
ama cau guamet pu gv 0-500
Marinkelle &Cadena 1972
FMNH ICN 6880MHNU UV
Sturnira mistratensis
Contreras-Vega and Caden,2000 + and ri 980
Contreras andCadena 2000 ICN holotype
Sturnira oporaphilum
Tschudi 1844 (13) and pac
ant cau cl cun
na ri snt to vc
bo ce hu met
nsn pu vc
Valdivieso
1964
AMNH FMNH ICN
MHNUC MUD TTU
USNM 483510 UV
Sturnira tildae de la Torre
1959
amz and
ori
ama ant gn met
pu to
bo caq cun gn
ri va vi 0-500
Marinkelle &
Cadena 1972
IAvH ICN 5206 MUD
TTU UV
Tribe Stenodermatini
Subtribe VampyrissinaChirodermaPeters 1860
Chiroderma salvini
Dobson 1878 and pac
ant caq cau cl
cho na snt to ce met pu ri vc 0-2000 Dobson 1880
FMNH IAvH ICN
6193 MHNUC MZCH
USNM UV
Chiroderma trinitatum
Goodwin 1958
amz car
pac
ant caq cau cho
gn
ama bo ce ma
snt va vc vi 0-500
Barriga-Bonilla
1965
FMNH IAvH ICN
18852 MZCH TTU
USNM UV
Chiroderma villosumPeters 1860
amz carpac
ant caq cau choma na vc
ama ara metpu va 0-500 Allen 1900
AMNH FMNH IAvHICN 2188 USNM UV
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mantilla-meluketal.Phyllostomid Batsof ColomBia 9
Table 1. (cont.)
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference Collections
PlatyrrhinusSaussure
1860
Platyrrhinus albericoi
Velazco 2005+ and ori met
ant bo cun ma
nsn qui ri snt
vc 1000 - 1800 Velazco 2005
IAvH ICN 8151 MUA
USNM
Platyrrhinus aquilus
(Handley and Ferris 1972)
+ car 0-500
Velazco in
press
Platyrrhinus
brachycephalus (Rouk and
Carter 1972) amz ama ara caq met
bo cas cl cocun hu pu ri
va 0-500
Rouk and
Carter 1972
FMNH IAvH ICN121261 MZCH TTU
USNM UV
Platyrrhinus chocoensis
Alberico and Velasco 1991 pac cau cho na vc 0-1000
Alberico &
Velasco 1991
FMNH ICN IAvH
MHNUC MZCHUSNM UV
holotype
Platyrrhinus dorsalis
(Thomas 1900) co
ant caq cau clcho cu na ri to
vc
bo cau cun hu
met qui 1000-1300 Dobson 1878
FMNH IAvH ICN 656KU MHNUC MUA
MZCH TTU UV
Platyrrhinus helleri (Peters
1866) co
ant caq cau cho
cun gn met to
bo cl co hu ma
na nsn pu qui
ri snt vc va vi 0-1500 Sanborn 1955
FMNH IAvH ICN
13027 MZCH
MHNUC MUA TTU
USNM UV
Platyrrhinus incarum(Thomas 1912) + amz ama 0 -500
Velazco (inpress) USNM 483642
Platyrrhinus infuscus(Peters 1880) amz caq cau met pu ama pu 0-1000
Marinkelle1970
FMNH IAvH ICN
MHNU TTU USNMUV
Platyrrhinus ismaeli
Velazco 2005 + and hu
ant bo caq cun
met nsn pu qui
ri vc 1000-1500 Velazco 2005
FMNH 58733 IAvH
ICN MUA USNM UV
Platyrrhinus nigellus(Gardner and Carter 1972)+ and car ce na
bo caq cau
cun hu ma metna nsn pu quiri snt vc 620-2757 Velazco 2005
FMNH 6948 IAvHICN USNM
Platyrrhinus umbratus
(Lyon 1902) + and car ant ma to
bo ce cho cun
hu ma ri snt vc 400-2550
Velazco (in
press) MUD holotype
Platyrrhinus vittatus
(Peters 1859) co
ant cau cl cun
ma na ri to ma met na vc 1000-3000 Allen 1900
AMNH 1500 IAvHICN MHNUC MUD
TTU UV
Platyrrhinus sp. nov.
Velazco and Gardner (inpress) pac cho, vc
Velazco and
Gardner (inpress)
UrodermaPeters 1866
Uroderma bilobatum
Peters 1866 co
ant caq cl cho
cun gn ma met
ama ara bl bo
cas ce co cunhu gua ma met
pu snt su to va
vc 0-1500 Allen 1900
AMNH FMNH IAvHICN 3993 MUD
MZCH TTU USNM
UV
Uroderma magnirostrum
Davis 1968 co ant caq cho met
ama bl caq cas
cun gua me
nsn to vc 0-500 Davis 1968
AMNH IAvH ICN
12661 MUD TTU
USNM UV
Vampyressa Thomas 1900
(14)+
Vampyressa melissa
Thomas 1926 (15) + amz and caq hu na snt vc
ama cau met
na pu ri snt vc 0-2000
Lemke et al.
1982 Ospina
and Gmez
1999
FMNH IAvH 2282
ICN TTU
Vampyressa thyone
Thomas 1909 co
ant caq cau cho
gn ma na ri to
vc
ama bo cl cun
met pu snt va 0-1900 Thomas 1909
AMNH BMNHFMNH ICN
MHNROM UC
USNM 483735
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10 sPeCial PuBliCations, museumof texas teCh university
Table 1. (cont.)
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference Collections
VampiriscusThomas
1900 (14)
Vampyriscus bidens
(Dobson 1878) + amz and caq cau va met pu 0-1000
Marinkelle &
Cadena 1972
IAvH ICN 18646
ROM USNH UV
Vampyriscus brocki
(Peterson 1968) + amz ama ce ma 0-500
Baker et al.
1972
CSJ ICN 14913 UV
IAvH ROM TTU
Vampyriscus nymphaea
(Thomas 1909) + pac ant cau cho ri vc 0-1900 Thomas 1909
AMNH FMNH ICN
16165 MHNUC
MZCH USNM UV
VampyrodesThomas 1900
Vampyrodes caraccioli
(Thomas 1889) co cau cho gn ant pu vc va 0-1000
Arata et al.
1968
FMNH IAvH ICN
11274 TTU USNM
UV
Tribe
Mesostenodermatini
Subtribe EnchisteninaEnchistenes Andersen
1906
Enchistenes hartii Thomas
1892 + co
ant caq cau cl
cho na ri vc
ama bo cas
cun met snt pu
to 0-2000
Arata et al.
1968
FMNH ICN 6771
MHNUC MZCH TTU
USNM
Subtribe Ectophyllina(16)
MesophyllaThomas 1901
Mesophylla macconnelli
Thomas 1901 co
ant caq cau cl
cho ri vc gn
ama gn met na
pu ri va vc vi 0-1500 Laurie 1955
FMNH IAvH ICNMZCH TTU USNM
499483 UV
Subtribe Artibeina
Artibeus Leach 1821
Artibeus amplus Handley1987 and car ant caq cau bo ce ma met 0-1300 Handley 1987 IAvH ICN 13344 UV
Artibeus concolorPeters1865 (17) + ama and snt gn
ama caq gn vavi 0-500 Dobson 1880 IAvH ICN 15014 UV
Artibeus jamaicensis Leach
1821
car ori
pac
ant cau cl cho
cun ma ri to vc
bl bo cl cas ce
co cun ma met
nsn sand pro 0-2100 J.A. Allen 1890
AMNH FMNH IAvHICN MHNUC MUD
MZCH ROM TTU
USNM 499488 UV
Artibeus lituratus (Olfers
1818) co
ant cau cl chocun ma met snt
to
ama ara bo cas
ce co gn gua
hu na nsn puqui ri snt to vc
va vi 0-2600 Dobson 1878
AMNH FMNH IAvH
ICN 447 MHNUCMUD MZCH TTU
USNM UV
Artibeus obscurus (Schinz
1821) amz and ant cau met gn
ama ara cl caq
gn met ri va vi 0-1000 Muoz 1986
FMNH 113407 IAvH
ICN TTU USNM UV
Artibeus planirostris (Spix
1823)
amz and
ori caq
ama cun hu
met nsn pu snt
vi 0-1300 Koopman 1982
IAvH ICN 17307 TTU
UV
Dermanura Gervais 1856+ (18)
Dermanura anderseni(Osgood1916) + amz ama
ama ant bl caq
ce cho co cunmet nsn 0-500
Cuervo et al1986
ICN 17180 ROM63061
Dermanura bogotensis(Andersen 1906) + and cun ce ri 2600
Acta
chiropterologica ICN holotype
Dermanura glauca
(Thomas 1893) + co
caq cau cl cho
cun ma met snt ant 0-2100 Dobson 1880
IAvH ICN MHNUC
UV
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mantilla-meluketal.Phyllostomid Batsof ColomBia 11
Taxa Region
Departments
(records from
reviewed
literature)
Departments
(records from
museums)
Elevational
range Reference CollectionsDermanura gnoma
(Handley 1987) + amz caq ant 0-600
Muoz-Saba
1999
IAvH ICN UV MHN
ROM UC 545
Dermanura phaeotis Miller
1902 +
and car
pac
ant cau cl cho
na ri vc ce co ma qui 0-1700 J.A. Allen 1916
FMNH 113998IAvH
ICN MHNUC MZCH
TTU UV
Dermanura rava Miller1902 + and pac cho 0 -1000
Solari et al.2008 ICN 8738
Dermanura tolteca
(Saussure 1860) + and pac ant cau snt ri 1500-2500
Cuervo et al
1986 ICN 11516 MHNUC
Dermanura watsoni
(Thomas 1901) + and pac cau cho met ant na vc 0 - 1000
Solari et al.
2008
AMNH BM ICNMZCH MHNUC 650
ROM TTU
Subtribe
Stenodermantina
Ametrida Gray 1847
Ametrida centurio Gray1847 amz vi 0-500
Alberico et al.2000 ICN 13983
SphaeronycterisPeters
1882
Sphaeronycteris
toxophyllum Peters 1882
amz and
car
ama cun met
snt caq ma va vi 0-2600 Sanborn 1941
IAvH ICN 499 ICN
499 TTU UV
Table 1. (cont.)
Addoal gc daa wll dodly rovd
hghr rolo ad oly dca ohr axoomc
chag, a h o m h molclar-ad
r ar h ord clacao avalal ad
ar aroraly d a a arg o o drad
h dvry, volo of morhology ad karyoy,cologcal aocao, ad ohr ac of h ology
of nw World laf-od a.
List summary.W ror 118 cormd hyl-
loomd c for Coloma (tal 1) ad 14 c
oally r h cory (tal 2) for a oal of
132 c rrg 40 gra ad10 faml.
All kow hylloomd faml ar rrd
Coloma, wh h xco of h famly Mac-
roa (o r soh Amrca). A 118 kow
c, Coloma ha h gra mr of a -
c of hylloomd a of ay cory, ragcor of largr gograhc ara ad grar ovrall
mammala dvry ch a brazl, Mxco, ad pr
(tal 3). th mr of hylloomd c rord
for Coloma xcd o cra h ar fr
a a rl of v ld work (.g., codcd y
Coloma mm ad c), a wll a rv-
oary ymac of hylloomd a.
Taxonomic comments.egh axoomc
comm (dgad y tal 1) ar cldd:
(1) W follow L al. (2002) ad dfy a mmr
of h gLophostoma:L. brasiliense,L. carrikeri,
and L. silvicolum, whch wr cldd a mmr
of h g Tonatiay Alrco al. (2000). (2) Wfollow smmo ad Vo (1998) ad rcogzMimon
cozumelae Goldma 1914 a a dffr axo from
M. bennettii. Alrco al. (2000) rcogzdMimon
cm from Chgorod, Aoqa, orgally
reported asM. cozumelae y Markll ad Cada
(1972), aM. bennettii.Mimon cm from Chg-
orod rord Markll ad Cada (1972) wr
dod a h usnM collco ad h Mm
of naral Hory of h uvrdad d lo Ad
bogo (MHnu), acroym d Markll ad
Cada (1972). Mimon bennettii cm() from
h darm of Ma, o h ar vra of h
Coloma Ad, rord y Alrco al. (2000)
should correspond to the material supposedly deposited
a h AMnH y Markll ad Cada (1972). How-
vr, afr mg h collco daaa of h AMnH
w wr o al o d Coloma cm ofM.
cozumelae orM. bennettii. (3) W follow smmo ad
Vo (1998) ad ra Choeroniscus intermedius J.A.
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12 sPeCial PuBliCations, museumof texas teCh university
Table 2. Species potentially present in Colombia. Abbreviations for Colombian regions are the same as those used
in Table 1. Abbreviations of countries: Ecuador (ecu); Peru (per); Venezuela (ven). Elevations in meters.
Table 3. Number of phyllostomid species by country, as reported in the most
recently updated literature.
Country Number of species Reference
Colombia 118 This work
Brazil 90 dos Reis et al. (2007)
Venezuela 88 Linares (1997)
Peru 88 UICN (2009)
Ecuador 86 Tirira (2007)
Bolivia 72 Aguirre (2008)
Costa Rica 63 LaVal and Rodrguez (2002)
Mexico 55 Ceballos and Oliva (2005)
Paraguay 20 Lpez-Gonzlez (2005)
Argentina 17 Barquez et al. (1999)
Taxa Region
Elevational
range Reference CountrySubfamily Micronycterinae
Micronycteris brosseti Simmons and Voss 1998 amz 0-500 Simmons and Voss 1998 per
Micronycteris giovanniae Baker and Fonseca 2007 pac 0-500 Fonseca et al. 2007 ecu
Micronycteris homezi Pirlot 1967 and ori 0-1500 Pirlot 1967 ven
Micronycteris matses Simmons et al. 2002 amz 0-500 Simmons et al. 2002 per
Subfamily Lonchorhininae
Lonchorhina fernandezi Ochoa and Ibaez 1982 amz ori 0-500 Ochoa and Ibaez 1982 ven
Lonchorhina inusitata Handley and Ochoa 1997 amz ori 0-500 Handley and Ochoa 1997 ven
Subfamily Phyllostominae
Lophostoma aequatorialis Baker et al. 2004 pac 0.800 Baker et al 2004 ecu
Lophostoma yasuni (Fonseca and Pinto 2004) amz 0-500 Fonseca and Pinto 2004 ecu
Subfamily Lonchophyllinae
Lonchophylla orcesi Albuja and Gardner 2005 pac 0-1000 Albuja and Gardner 2005 ecu
Lonchophylla pattoni Woodman and Timm 2006 amz 0-500 Woodman and Timm 2006 perSubfamily Glyphonycterinae
Glyphonycteris daviesi Hill 1965 amz ori 0-1000 Gardner 2008 bra per ven
Subphamily Stenodermatinae
Platyrrhinus aurarius Handley and Ferris 1972 amz ori 700-2200 Velazco 2005 ven
Platyrrhinus matapalensis Velazco 2005 pac 200-1500 Velazco 2005 ecu
Dermanura rosenbergi Thomas 1897 pac 0-1000 Hoofer et al. 2008 ecu
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mantilla-meluketal.Phyllostomid Batsof ColomBia 13
All ad Chama 1893 a a jor yoym of C.
minor. (4) Alrco al. (2000) mad rfrc o h
holotype ofLichonycteris obscura (type locality Ma-
aga, ncaraga) wh a Coloma org. th may a doral mak ad h ahor may hav
rfrrg o h holoy ofLionycteris spurrelliwhich
adjac oL. obscura hr l. th holoy of
L. spurrelli dod h bM (bM-13.8.10.1) ad
wa collcd Codoo, Choc, Coloma (Carr
ad Dola 1978). (5) W follow Alrco al. (2000)
and include Scleronycteris ega thoma 1912 a r
Coloma. Howvr, w fald o d h mm
maral odly dod a h iAvH or-
g h hyoh y Alrco al. (2000). Frhr
vgao cary o corm h rc
of h axo Coloma. (6) Alrco al. (2000)followd h crra of Hadly (1966) ad rord
L. mordax for Coloma; Lonchophylla mordax was
rrcd o ohar soh Amrca y Alja ad
Gardr (2005). W follow Alja ad Gardr (2005)
and treat Colombian populations ofLonchophylla
rvoly dd aL. mordax asL. concava. (7)
Grfh ad Gardr (2008) rrcd h dr-
tion ofL. handleyi o pr ad ragdL. handleyi
rcord rord y Alrco ad Orjla (1992) a
L. chocoana. W ror a rcord ofL. handleyi from
Hla dod a h collco of h MHnuC.(8)
Dvalo ad Corhal (2009) xdd h dro
ofL. orienticollina on the western side of the Eastern
Cordllra of h Coloma Ad. th drc aaly
of h morhology of cm iCn 16238 from h
wr vra of h ear Cordllra, cldd
the type series ofL. orienticollina, rvald ha h
specimen does not differ from typicalL. robusta. Fr-
ther research is necessary to determine the actual limits
of h wo axa Coloma. (9) W follow McLlla
ad Kooma (2008) ad ra Carollia colombiana
Cara al. 2001,wh y localy La Cja road,
baroa, Aoqa, Coloma, a jor yoym of
C. castanea; (10) Alhogh Glyophonycteris daviesi
(Hll 1965) wa cldd Alrco al. (2000), h
ahor dd o rovd formao o mm ma-
ral org rc Coloma. Howvr, G.
daviesi ha docmd for brazl, ecador, pr,
ad Vzla from local adjac o Coloma
ad holdg vrom mlar o ho ycal of
h Coloma Amazo ad Oroqa. (11) Alrco
al. (2000) dd o rovd a caalog mr for
Neonycteris pusilla (saor 1949) cm do-
d a h AMnH. Alrco al. (2000) may rfr o
cm AMnH 78830-31 h y r of h
c, from tahaa, Ro Va, a h Colom-
a ordr, Amazoa brazl, collcd y h Olallarohr o 14 Jly 1929 (saor 1949). (12) sc-
m rord y Alrco al. (2000) a Sturnira
mordax are treated as S. koopmanhilli followg h
crra rood y McCarhy al. (2006). (13) W
follow Gardr (2008) codrg Sturnira ludovici
as a subspecies of S. oporaphilum (S. o. ludovici).
Sturnira ludovici wa rad a a c y Alrco
al. (2000). (14) W follow bakr al. (2003) ad
rcogz a mmr of h g Vampyriscus: V.
bidens, V. brocki, ad V. nymphaea, rvoly cldd
h g Vampyressay Alrco al. (2000). (15)
W follow Lm al. (2003) ad codr Colomacm rvoly dd a V. pusilla as V.
thyone. Gardr (2008) cod h dro ofV.
pusilla o ohr brazl ad paragay.(16)Alrco
al. (2000) rcogzdEctophylla alba as present in
Coloma ad o maral dod a h uV wh
o caalog mr rovdd hr lcao. W
cold o d cm ofE. alba supposedly depos-
d a h uV; h work w follow Gardr (2008)
who restrictedE. alba o Cral Amrca. i lkly
that ColombianE. alba cm rord y Alrco
al. (2000) corrod oMesophylla macconnellii.
(17) W follow Hoofr al. (2008) ad rcogz
Koopania concolorasArtibeus concolor. (18)W fol-
low Hoofr al. (2008) codrgDermanura as
a dc g fromArtibeus; Alrco al. (2000)
consideredDermanura a a g whArtibeus.
A larg mr ofDermanura specimens deposited in
Coloma collco (. . 236 of h cm or
daaa of h iCn collco) hav mdd
asD. cinerea, an epithet which enclosedD. anderseni,
D. cinerea, D. glauca,D. gnoma, adD. watsoni, a
mod y Gardr (2008).
Newly described and elevated species.W
cldd or l 27 wly rcogzd c (19
wly dcrd c ad gh wly lvad -
c), 19 of whch ar cormd for Coloma, whl
gh ar oally r (*) h cory. th
w c ar a follow: Micronycteris matses*
smmo al. 2002 wh y localy nvo sa
Ja, Loro, pr; M. brosseti* smmo ad Vo
1998 wh y localy paraco, Frch Gaa; M.
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14 sPeCial PuBliCations, museumof texas teCh university
giovanniae*bakr ad Foca 2007 wh y localy
emralda sa Lorzo (oward La), Fca sa
Jo, ecador; Lophostoma aequatorialis*bakr
al. 2004 wh y localy eac exrmalLa Chqa, ar sa Lorzo, emralda, ecador;
L. yasuni*(Foca ad po 2004) wh y local-
y h vcy of Ya Rarch sao, Ya
naoal park ad bohr Rrv, Orllaa, eca-
dor;Anoura aequatoris with type locality in Illambo
Gala, ecador (Lrg 1921) a formr c
ofA. caudiferlvad y Malla-Mlk ad bakr
(2006) ad cormd Coloma y h am a-
hor;A. cadenai Malla-Mlk ad bakr 2006 wh
y localy Calma, Vall dl Caca; A. stulata
Mchhala al. 2005 dcrd from Codor Mrador,
ecador, rord for Coloma y Malla-Mlkad bakr (2008) ad Malla-Mlk al. (2009);A.
luismanueli Molar 1994 wh y localy Mrda,
Vzla, cormd Coloma y Malla-Mlk
ad bakr (2006). Alrco al. (2000) mod
Perez? (kow da) a lograhc rfrcfor
A. luismanueli Coloma; h work corrod o
a draf vr lhd ad o rlmary dca-
tions (some of which were made by the senior author
a a d a h iCn). Malla-Mlk al. (2009)
rovd addoal formao o h axoomc df-
ferentiation betweenA. aequatoris andA. luismanueli
Coloma;Lichonycteris degenerMllr 1931 wh
y localy par, blm, brazl, rcogzd a a
dffr axo fromL. obscuray Gardr (2008). th
only published reference of a Colombian specimen of
L. degener fod Mogro ad Romro-Rz
(2000); Lonchophylla cadenai Woodma ad tmm
2006 wh y localy bavra, Vall dl
Caca;L. chocoana Dvalo 2004 wh y localy
h vcy of Alo tamo, ecador;L. orcesi* Alja
ad Gardr 2005 wh y localy La paml,
Ro pdra, Cordllra d to, emralda, eca-
dor; L. orienticollina Dvalo ad Corhal 2009 wh
y localy a h rco of Cao Gamalo ad
Cao La Cra, Ma, Coloma;L. pattoni* Woodma
ad tmm 2006 wh y localy Rrva Cco
Amazco, orh ak Ro Madr d Do, 14 km
a pro Maldoado, tamoaa, pr. th cl-
sion ofL. pattoni within the list of species potentially
present in the Colombia was based on specimens that
partially matchedL. pattoni dcro from Lca,
Amazoa dod a h iCn; howvr, frhr r-
vw cary o corm rc h cory.
i addo, h dro ofL. pattoni was recently
xdd o ecadora rovc of paaza (Malla-
Mlk al. 2009) h ecadora Amazo ha
har h cologcal characrc of h Colomaohr Amazo; Carollia monohernandezi Cuartas
ad Gozlz 2004 wh y localy Vllaraz,
Florca, Caq, Coloma; Sturnira koopmanhilli
McCarhy al. 2006 wh y localy La paml,
Rrva ecolgca Coacach-Cayaa, emralda,
ecador; S. mistratensis Corra ad Cada 2000
y localy pro d Oro, Raralda, Coloma;
Platyrrhinus albericoi Vlazco 2005 wh y localy
sa pdro, pacaramo-plcoaa road, Darm
of Czco cormd Coloma y Vlazco al. (
r);P. aquilus (Hadly ad Frr 1972) wh y
localy h had of Ro pcro, 4,100 f Crro Mal,Dar, paama lvad ad cormd Coloma
y Vlazco ad Gardr ( r);P. incarum, rv-
ously considered a subspecies ofP. helleri (Vampyrops
zarhinus incarum) (thoma 1912) wh y localy
pozo, paco, pr, lvad o c ad co-
rmd Coloma y Vlazco al. ( r) ad
o maral from Lca, Amazoa; P. ismaeli V-
lazco 2005 wh y localy bala, Chachaoya,
Amazoa pr;P. matapalensis*Vlazco 2005 wh
y localy Maaalo, Zarmlla, tm, pr;P.
nigellus, dcrd a Vampyrops nigellusy Gardr
ad Carr (1972) wh y localy Hahachayo,
Ayaccho, pr, lvad ad cormd Coloma
y Vlazco (2005); P. umbratus (Lyo 1902), rcog-
zd y smmo (2005) wh y localy sa
Mgl, Macoama Rvr, Magdala Coloma; P.
sp. nov. (Vlazco ad Gardr r) dcrd
from h bogograhc Choc rgo of Coloma
and Ecuador and closely related toP. vittatus;Derma-
nura bogotensis (Adr 1906) wh y localy
Crch, ar bogo, Cdamarca, Coloma,
lvad y Lm al. 2008; D.rosenbergi* thoma
1897 oally r Coloma, wh y localy
Cachav, emralda, ecador, lvad y solar
al. ( r); h axo wa rvoly codrd a
jor yoym ofD. glaucay smmo (2005) ad
cldd dr h gArtibeus (asA. glaucus) y
Alrco al. (2000).
Models of potential distribution.Models of
potential distribution of phyllostomid species present
ad oally r Coloma ar how A-
dx i ad ii, rcvly. th modl of oal
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mantilla-meluketal.Phyllostomid Batsof ColomBia 15
distribution of species of dubious records in Colombia
such as N. pusilla and S. ega were included within
Adx ii. Or modl of oal dro r-
r h r am o corora h avalalcomaoal chology, arclarly Gograhc
iformao sym (Gis), o rodc mor olog-
cally magfl rdcd dro for a larg
of gograhcally rlad hylloomd c, g
a homogo aalycal rocdr whch allow
raaly ad mrovm. th rocdr allowd
o mor rgoroly dla oal droal
lm comaro wh rvo ffor g oly
margal rcord, ch a ho y paro al.
(2007) ad Gardr (2008). Alhogh for om c
h modl cold ovrma h acal dro,
h mhodology may rov fl dlmg dr-oal rag of Coloma hylloomd. Modl of
potential distribution of Colombian phyllostomid bats
rd h work ar avalal varo forma
(grd, ac, df, ad jg) a h w of h naral
scc Rarch Laoraory, Mm of txa tch
uvry (www.rl..d).
General phyllostomid distributional patterns.
th Coloma rrory coymcally dvr d
o a comao of comlx gologc, cologcal, ad
ogograhc roc ha rl a vary of ch
oor for hylloomd a. O of h mormarkal ac of h Coloma gograhy
locao h orhwr corr of soh Amrca,
adjac o ad commcag wh h paamaa
ihm. norhwr soh Amrca ha d-
d a h cr of h gra drcoal xchag
of faa w norh ad soh Amrca drg
h La trary (Hrhkovz 1972). i addo, h
roxmy of h Coloma rrory o av cr
of cao soh Amrca, Cral Amrca, ad
h Cara (Hrhkovz 1972) ha cord o
oly o h hgh mr of c rord for Co-
loma, alo o h comlxy of rgoal c
arragm fod or aaly. th gograhc
dvry of h Coloma rrory ca dvdd
o v aral rgo a follow: Amazo, Ada,
Cara, Oroqa, ad pacc (Choc), ach o
characrzd y q arragm of colog-
cal varal (Hrdz Camacho al. 1992). Or
aaly aocad h gra mr of c
wh h Ada (85 c) ad Amazo rgo (74
c), followd y h pacc wh 64 c, h
Cara wh 62 c, ad h Oroqa wh 41
c (tal 4). Alhogh a larg mr of c
wr hard amog rgo (tal 4), om c
wr docmd from oly o rgo (AdxiV) a follow: Amazo rgo, 14 q c,
cldg h dadN. pusilla and S. ega;Ada,
12; ad pacc, 7.
Multiple correlation analyses between phyllosto-
md rch ad vromal varal howd ha
hylloomd rch Coloma had a gca,
ov corrlao wh mrar ad rcao
ad a gca ad gav corrlao wh lvao
(tal 5). th hghr mr of c wr fod
ara wh avrag rcao grar ha 2,000
mm, avrag mrar grar ha 25C, ad lva-o lowr ha 1,000 m (Fg. 1a, ad Covr Fgr)
(tal 5). th aov mod codo ar commo
across the country and dominate the lowland forested
ara of h dmo of h Ada ym ad h
Amazo rgo.
th comaro w rl from or
rgoal c cog ad rl o c ac-
cmlao or rch modl gg a grar
gograhc arog amog c drd
alog h Ada rgo wh rc o h Amazo
rgo. Alhogh h ovrall mr of c o-ally r h Ada rgo wa grar ha
ha h Amazo rgo, h oal mr of
sympatric species per unit of area in our richness model
wa grar h Amazo. th ar orvd
or rch modl ca rlad o h vrom-
al hrogy ycally aocad wh lvaoal
rag, a wll a h lowr ara occd y oally
al vrom. Covrly, rad for
rval h lowlad of Coloma ar wdly
drd acro h Amazo offrg mll ch
oor for hylloomd a, arclarly for
amalvoro ad frgvoro form (Mdll al.
2000; Jmz-Orga ad Malla-Mlk 2008). th
gra cocrao of c or c rch
modl occrrd o h Gaa-Amazo corrdor (Covr
Fgr). th corrdor characrzd y rrac (200
o 500 m) of Camrc org aocad wh h Gaa
shld. W hyohz ha h rrac cod
mrg lad ma ha rcrd a c drg
h rad vao of war ma o h lowlad
of orhwr soh Amrca drg h Moc.
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16 sPeCial PuBliCations, museumof texas teCh university
Table 4. Comparisons of phyllostomid species composition among Colombian natural regions.
Fgr 1. scar lo howg h ov corrlao w hylloomd c rch Coloma, xrd
mr of c, ad ma aal mrar of h cold moh of h yar, xrd Cl dgr
(lf); ad h gav corrlao w hylloomd c rch Coloma, xrd mr of
c, ad lvao, xrd mr aov a lvl (rgh).
Species Unique Species Andean Caribbean Orinoquia Pacific
Amazon 74 14 50 46 40 41
Andean 85 12 - 56 36 51
Caribbean 62 0 - - 36 46
Orinoquia 41 0 - - - 35
Pacific 64 7 - - - -
Richness Elevation MAP TCMY
Richness 0
Elevation -0.38978 ** 0 1.4565E-29
Precipitation 0.37947 ** -0.10209 0 9.4415E-56
Temperature 0.48327 ** -0.76253 0.14187 0
Table 5. Results of Spearman correlation between richness and environmental variables (** signicant at
p>0.01). Correlations were based on values extracted from raster layers (N = 14,560). MAP = mean annual
precipitation; TCMY = mean annual temperature of the coldest month of the year.
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mantilla-meluketal.Phyllostomid Batsof ColomBia 17
For a gca rod of h arly o md Moc
(23-17 Mya), h Coloma Amazo wa dr h
c of h lak formao of pa (Wlgh
ad salo 2006). poror dao wr aarlyfrq drg h md ad la Moc (Hovkok
al. 2007, ad rfrc hr) ad lvao
grar ha 200 m cold lay a mora rol
h alhm of ford vrom ycally
rfrrd y hylloomd.
Amog Coloma rgo, h Ada, h Ca-
ra, ad h pacc wr mo mlar rm of
c comoo (Fg. 2a) (tal 4). Hgh lvl of
mlary amog h Ada, Cara, ad pacc r-
go ar h coqc of h covrgc of c
dro h orhwr corr of h cory.th oro of h cory akd y h pacc
Oca ad h Cara sa, ad olad from
h ar oro of cory y h hghlad of h
Ad. Hradz-Camacho al. (1992) dd h
southwestern portion of the Colombian Caribbean as an
dd cologcal . Hrdz-Camacho al.
(1992) dvdd h Cara o hr zo: 1) h
Fgr 2. a) Coloma aral rgo a dd y Hrdz-Camacho al. (1992) ad hogram of or hr-
archcal clr aaly howg mlar c comoo amog rgo. ) Ma of orhwr soh
Amrca howg h oal ogograhc for Coloma hylloomd a; dahd l how h hyo-
hcal dvo w h orhr Coloma Amazo (Gaa Amazo) ad h ohr Coloma Amazo.
ecrcld ara rr h ohrmo lm of h hyr hmd codo of h bogograhc Choc (lack),
whch ac a a lr of c o h wr d of h cory. th ara gry rr h mo hmd oro
h Cara rgo codrd a a xo of h orhr oro of h bogograhc Choc.
olad moao ym of h srra nvada d
saa Mara; 2) h Ard pr-Cara l, wh low
rcao, characrzd y h rc of ard ad
m ard vrom; ad 3) a rgo aocad whh oodd fa of h Caca ad Magdala Rvr.
th lar zo rood o cologcally cocd
wh h orhr oro of h bogograhc Choc
cog h Choc-Magdala bogograhc
zo. Malla-Mlk ad Jmz-Orga (2006)
dd a ladal a c rovr alog h
Coloma bogograhc Choc, ad rcogzd h
orhr oro of h Coloma pacc Coa a a
ogograhc for chrora. i or modl,
phyllostomid lowland species distributed across the
pacc ad h Cara rgo Coloma y-
cally xdd hr dro o adjac CralAmrca (.., Dermanura phaeotis, D. rava, ad D.
watsoni) (solar al. r). th olao of h
wr oro of Coloma cad y h hghlad
of h Ada ym alo rol for h d-
roal ar orvd amog mmr of h
famly Lochohylla. ncar fdr h g
Lonchophylla ar ycal rrav of h Colom-
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18 sPeCial PuBliCations, museumof texas teCh university
hghlad cal ar rrav of h r
Aora,g Anoura, a wll a rrav
of h gra Platyrrhinus and Sturnira. All c
h g Anoura are represented in Colombia(Malla-Mlk ad bakr 2006, 2008) ad hav a
aar gograhc dvo hogh o aoc-
ad wh h rg ar of h hr ma
of h Coloma Ad. Rag of hghlad hyl-
lostomid specialists such as Anoura, Platyrrhinus,
and Sturnira gg ha h Coloma Cral ad
Wr Cordllra ac a a gl arad from
h ear Coordllra y h ir-Ada Vally of
h Magdala Rvr.
th gra dffrc hylloomd c
comoo amog Coloma rgo wa fod -w h pacc ad h Amazo-Oroqa rgo,
whch ar arad y h Ad (tal 4). Alhogh
vral av hylloomd c hav a ra-
Ada dro, molclar aaly codcd
o cocc olao from oh vra of h
Ad hav rld h dcao of hghr lv-
l of gc dffrao ha wold xcd
for cocc axa (Hoffma ad bakr 2003; bakr
al. 2004; bakr ad bradly 2006; Vlazco ad pa-
ro 2008). th aov mod d cocldd
ha h Ada ym a ffcv arrr o g
ow, h romog cao.
th Coloma Oroqa ha h fw mr
of hylloomd c. th Llao Oral of h
Coloma Oroqa co a larg low-lyg la,
ad amog hylloomd a hr ar vry fw -
c calzd for of o ac. i aar ha
h o avaa of h Oroqa do o offr may
ch oor for hylloomd for cal.
th Coloma Oroqa characrzd y dramac
cao of cologcal varal ad h rog -
vromal grad ha rval alog gograhy
ha a dahc, rahr ha a clmac, a: xv
ado ara h a of Coloma rovd a fr-
drag ra, ad hc a war-rd vro-
m for h vgao ha rld o rocal dry
for ad om (Marcha al 2006).
edahc codo of h Oroqa rv h
dvlom of rad for h rgo. Howvr,
gallry for ar commo o h marg of h war
a Chocoa faa ad vral c xd hr
dro o orhr ecador (..,Lonchophylla
cadenai,L. chocoana,L. concava,L. fornicata andL.
orcesi*) (Malla-Mlk 2007). th comao ofcommo rgoal droal ar ad hylog-
c af llra h c of Coloma
gograhy o h voloary hory of h aov
mod gro. th formao of h Chocoa ad
Cara rgo ar rlavly rc v aoc-
ad wh h al rod of h Ada lfg (5 o
3 Mya) ad h crao of h ir-Ada vally ad
hr aocad oodd fa ha lld h lowlad of
rmal orhwr soh Amrca (Marz 1997;
erclla al. 2002). pror o h al lfg of h
Ad, xrohyc vrom aocad wh h
ffc of h Alo wd xdd hr dr-o alog h orhr coa of soh Amrca. th
comlo of h ihm of paama ad h dva-
tion of the Humboldt marine current resulted in the
rdoma hyr-hmd vrom of Cral
Choc (Fg. 2). th rc of h Chocoa hyr-
hmd for rr a cologcal ha w
h Cral Amrca-Cara dry corrdor ad h
xrohyc for cral ad ohr ecador. O
armo oro, xrohyc vrom
h lowlad of h Coloma Cara rgo ar
rrd y h hghlad of h srraa dl prj,
whch ola h Coloma ad h Vzla Ca-
ra coa. i lkly ha c wh a Cral
Amrca org colozd h cory followg ard
coym, whl lal ha c wh a
Gaa-Amazoa org colozd wr Coloma
g h lowlad a of h r-Ada ym.
th crr rdoma hyr-hmd codo of
cral Choc co a addoal aral arrr
olag hylloomd olao from h ohr
Coloma Choc ad ecador.
th Coloma Ad lay a dol rol a a
ffcv arrr for olag aral olao o h
ar ad wr vra of h ym a wll a
a source of innumerable niche opportunities for phyl-
loomd a, rcrg c from h ohr rgo.
Or aaly rvald ha rch dro amog
hylloomd dcrad wh lvao, wh a larg
number of species concentrated on the piedmonts of
h Ad ad fw gro adad o hghlad coy-
m (Covr Fgr ad Fg. 1). Amog hylloomd
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mantilla-meluketal.Phyllostomid Batsof ColomBia 19
cor. W hyohz ha gallry for hav a
lmd carryg caacy for rcrm ad alh-
m of c dvry ycal of mar for. i
addo, clma h Llao Oral ca cla-d a rocal dral wh dffrc mohly
mrar grally mall wh daly cao
ha may larg (20C), cally drg h dry
ao. th clma domad y h ir-trocal
Covrgc Zo (itCZ) ad mor drvd from
h Alac Oca. th wo ray ao, from March
o May ad from Ocor o Dcmr, ar arad
y dry ao from Jaary o Frary ad from J
o smr. th avrag rcao 950 mm
wh a avrag of 175 mm March; howvr, hr
codral varao (brro al. 2003). Crr
clmac codo orvd o h avaa of h
Coloma Oroqa hav domad h rgo for a
la 4000 yar, c h early ad Mddl Holoc
(Marcha al. 2006).
Status of the knowledge on Colombian phyllosto-
mid bats.A ad y rl oad h
work, h Coloma rrory hold a q ad rch
mammala faa, ad a aoal la for dy
rgly dd. A aaly of 17,778 hylloomd
a mm rcord (Fg. 3) rvald a amlg a
of hylloomd, rmarly a a rl of major Colom-
a acadmc cr g locad wh c h
Ad ad collcg local g rmarly adjac
o h c. May Coloma gograhc ara ad
Fgr 3. Ma of Coloma howg h dro of amlg local of phylloomd a
aocad wh h 17,778 rcord aalyzd h dy. ecrcld ara rr h amlg
a of hylloomd collcg local rmarly adjac o major c Coloma a fol-
low 1) saaf d bogo, 2) saago d Cal, ad 3) Mdll.
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20 sPeCial PuBliCations, museumof texas teCh university
axa rma drrrd, ad h arclarly
r for ara ad axa ha oally coa h hgh-
dvry of hylloomd c. th work rov
ha h xv collco of mammal dod Coloma o co a crdly val-
al rorc. Howvr, craoral work
rgly rqrd. W v h raoal cc
commy o valda h collco hrogh hr
a a cc rorc. symac a radly
dvlog ld soh Amrca (Vo 2009) ad
the participation of the international community is
crcal for h dvlom ad cooldao of a
roacv Coloma chool ymac ad vol-
o. iraoal o whch hav d
and enriched their own collections in the past with
Coloma cm, ad soh Amrca for hamar, hold cra h mcham o facla ac-
c o cm, arclarly holoy maral whch
graly mrov ymac d Coloma. W
clra ffor y o ch a h AMnH ad
ttu for ladg h ah dgzg cm mad
accl va r. W alo alad local ffor docmg h Coloma mammala dvry
mad y h varo acadmc o holdg col-
lco. i alo rmarkal h laor of h io
Alxadr vo Hmold hrogh h crao of h
r of chckl ofBiota Colombiana. i h work
w follow Alrco al. (2000) a wll a h r
of darmal chckl lhd Biota Colom-
biana as primary source of reference for phyllostomid
rcord. iav lk h r h grao of
h kowldg rqrd for h adqa corvao
ad maagm of Coloma aral rorc. i
hod h work wll mla addoal rarcho flll h ga of formao o Coloma hyl-
loomd a dd hr.
aCknowledgments
th work wold o ol who h
ady ld work ad craoral roc of aff a
h varo mm collco d a rorc. W
ar arclarly hakfl o h aff of h io
d Cca naral of h uvrdad nacoal
d Coloma ad cally Y. Moz-saa ad A.Cada for hr advc ad aac drg h var-
o ag of h work. W alo hak b. D. paro,
W. b. saly, L. R Hay, p. Vlazco, ad C. Dck
for aac a h collco of h FMnH, a wll
a h thoma J. D ad Karl p. schmd Fd of
Fld Mm scholarh Comm for acal
or o v hr collco; D. e. Wlo, A. L.
Gardr, ad L. Gordo for aac a h mam-
mal collco of h smhoa io; A.
Rodrgz-bolao, for acc ad aac a h
collco of h uD; A. Arlla ad L. palaco for
aac a h collco of h utCH; H. Garr for
aac a h nsRL. W arca h coorao
of J. Zak ad h Darm of bologcal scc of
ttu. W ar graly dd o J. Marz brro, p.
Gmz-sadoval, M. A. Hoyo, F. A. sarz, ad A.Mar for hr aac daa collco. W hak
Cl Cao, Lz sl, M. McDoogh, R. Marcha, J.
parlo, L. Dvalo, s. solar, J. p. Carrra, M. po, p.
Lar, p. Vlazco, H. e. Ramrz, ad M. e. Rodrgz
for hr ghfl comm o h axoomy ad
dro of Coloma hylloomd. Fally, w
hak L. bradly ad n. d la sacha for hr valal
doral aac, a wll a J. salazar-bravo ad s.
solar for xcll crcal rvw of h work.
literature Cited
Alrco, M., ad A. J. ngr. 1992. prmr aor or lo
mamfro dl Vall dl paa. novdad Colomaa,
Mo d Hora naral, uvrdad dl Caca, nva
eoca 5:66-71.
Alrco, M., ad e. Vlaco. 1991. Dcro of a w road-
od a from Coloma. bor zoologch brg
42:237-239.
Alrco, M. ad J. Orjla. 1983. Dvrdad cca d
do comdad d mrclago naro, Coloma.
Cda 11:31-40.
Alrco, M., A. Cada, J. Hrdz-Camacho, ad Y. Moz-
saa. 2000 Mamfro (syada: thra) d Coloma.
boa Colomaa 1(1):43-75.
8/23/2019 Filostomidos de Colombia.pdf
25/44
mantilla-meluketal.Phyllostomid Batsof ColomBia 21
Alja, V. L., ad A. L Gardr. 2005. A w c ofLon-
chophylla thoma (Chrora: phylloomda) from
ecador. procdg of h bologcal socy of Wah-
go 118(2):442-449.
All, J. A. 1890. no o a mall collco of W ida a,
wh dcro of a aarly w c. bll of
h Amrca Mm of naral Hory 3:203-18.
All, J. A. 1900. L of a collcd y Mr. H. H. smh
h saa Mara rgo of Coloma, wh dcro of
w c. bll of h Amrca Mm of naral
Hory 13:89-94.
All, J. A. 1916. L of mammal collcd Coloma y
h Amrca Mm of naral Hory xdo,
1910-1915. bll of h Amrca Mm of naral
Hory 35:191-238.
All, J. A., ad F. M. Chama. 1893. O a collco of mam-
mal from h lad of trdad, wh dcro of wc. bll of h Amrca Mm of naral
Hory 5:203234.
Adr, K. 1906. brf dago of a w g ad w
form of odrmao a. Aal ad Magaz of
naral Hory r 7,18:419-423.
Araa, A. A., J. b. Vagh, K. e. nwll, R. A. J. barh, ad M.
Graca. 1968. Salmonellaand Shigella infections in bats
lcd ara of Coloma. th Amrca Joral of
trocal Mdc ad Hyg 17:92-95.
bag, O. 1900. L of h mammal collcd h saa
Mara rgo y W. W. brow Jr. procdg of h nw
Zlad Zologcal Cl 1:87-102.
bakr, R. J., H. H. Goway, ad A. Cada. 1972. th hyl-
loomad a, Vampyressa brocki, Coloma. bll
of h sohr Calfora Acadmy of scc 71:4.
bakr, R. J., C. s. Hood, ad R. L. Hoyc. 1989. phylogc
rlaoh ad clacao of h hghr cagor
of h nw World a famly phylloomda. symac
Zoology 38:228-238.
bakr, R. J., s. R. Hoofr, C. A. porr, ad R. A. Va D
bch. 2003. Dvrcao amog nw World Laf-
od ba: a voloary hyoh ad clacao
frrd from dgomc cogrc of DnA qc.
Occaoal par, Mm of txa tch uvry
230:+1-32.
bakr, R. J., R. M. Foca, D. A. parh, C. J. phll, ad F.
G. Hoffma. 2004. nw a of h g Lophostoma
(phylloomda: phyllooma) from orhwr
ecador. Occaoal par, Mm of txa tch
uvry 232:+1-16.
bag, O. 1900. L of h mammal collcd h saa Mara
rgo of Coloma y W.W. brow, Jr. procdg of
h nw eglad Zologcal Cl 1:87-102.
barrga-bolla, e. 1965. edo maozoolgco coloma-
o, i. Chrora. Caldaa 9:241-271.
barqz, R. M., M. A. Mar, ad J. bra. 1999. th a of
Arga. scal plcao, Mm of txa tch
uvry 42:1-275.
brro, J. C., Arlaz M. V., Dvvoord J. F., Clf, adA. Hooghmra, H. 2003. poll rrao ad
ccoal vgao chag o h ado laa
of Araracara, Coloma Amazoa. Rvw of palao-
oay ad palyology. 126, 163181.
bradly, R. D., ad R. J. bakr. 2001. A of h Gc s-
c Coc: cyochrom- qc ad mammal.
Joral of Mammalogy 82(4):960-973.
Carr, D. C., ad p. G. Dola. 1978. Caalog of y c-
m of norocal a lcd eroa mm.
scal plcao, Mm of txa tch uvry
15:1-136.
Callo, G., ad G. Olva. 2007. Lo mamfro lvr d
Mxco. socdad ara l coocmo y o d la dvr-dad fodo d clra comca, Mxco. 986 .
Coa, A. G., ad J. A. Oado. 1996. th gologc volo of
h Cral Amrca ihm. i: evolo ad vro-
m of trocal Amrca. th uvry of Chcago
pr, Chcago. 425 .
Corra, M., ad A. Cada. 2000. ua va c dl
gro Sturnira (Chrora: phylloomda) d lo
Ad colomao. Rva d la Acadma d Cca
exaca, Fca y naral XXiV(91):285-287.
Cara, C. A., J. Moz, ad M. Gozlz. 2001. ua va
especie de Carollia Gray, 1838 (Chrora: phylloo-
mda) d Coloma. Acaldad bolgca 23(75):63-
73.
Crvo Daz, A., J. Hrdz Camacho, A. Cada G. 1986.
La acalzada d lo mamfro d Coloma: aoa-
co or drc. Caldaa 15:471-501.
Dvalo, L. 2004. A w chocoa c ofLonchophylla
(Chrora: phylloomda). Amrca Mm no-
va 3426:1-16.
Dvalo, L. ad A. Corhal. 2009. A w c ofLon-
chophylla (Chrora: phylloomda) from h ar
Ad of orhwr soh Amrca. Amrca M-
m nova 3635:1-16.
Dav, W. b. 1968. Rvw of h G Uroderma (Chrora).
Joral of Mammalogy 49:676-698.
d la torr, L. 1956. th dal formla of h a of h g-
nusDiaemus. procdg of h bologcal socy of
Wahgo 69:191-192.
Doo, G. e. 1878. Caalog of h Chrora h collc-
o of h brh Mm. brh Mm (naral
Hory). Lodo, xl + 567 + pl. i-XXX.
Doo, G. e. 1880. O om w or rar c of Chrora
h collco of h Gg Mm. procdg
of h Zoologcal socy of Lodo 1880:461-465, pl.
XLVi.
8/23/2019 Filostomidos de Colombia.pdf
26/44
22 sPeCial PuBliCations, museumof texas teCh university
do R, n. R., A. L. prach, W. A. pdro, ad i. p. d Lma
(d). 2007. Morcgo do bral. uvrdad eadal
d Lodra, Lodra 253 .
erclla, G. b. Aloo, F. erada, F. L. Chocc, J., baraza, adM. Farra. 2002. th Magdala rd ym (Ca-
ra sa): r-day morhology ad archcr
modl. Mar Gology 185, 303318.
Foca, R. M. ad C. M. po. 2004. A w Lophostoma
(Chrora: phylloomda: phyllooma) from h
Amazoa of ecador. Occaoal par Mm of
txa tch uvry 242:1-9.
Foca, R. M., al. 2007. Morhologcal ad molclar vara-
o wh ll g-ard a of h gMicronycteris
(phylloomda: Mcroycra) from sa Lorzo,
ecador. p. 721746 th qal aral:
hoorg h lf ad lgacy of Olvr p. paro (D.
A. Kl, e. p. La, J. salazar-bravo, ad J. L. pao,
d.). uvry of Calfora plcao Zoology
134:1981.
Frma, D. p. 1966. th rcd m of paaa (Acara:
srcda). p. 125-166 ecoara of paama
(R.L. Wzl ad V.J. to, d.). Fld Mm of
naral Hory, Chcago.
Gardr, A. L. 1977. Fdg Ha. p. 293-350 bology of
a of h nw World Famly phylloomda par ii (R.
J. bakr, J. K. Jo, Jr. ad D. C. Carr, ed.). scal
plcao Mm txa tch uvry13:1-364.
Gardr, A. L. 2008. Mammal of soh Amrca, Volm 1
Maral, Xarhra, hrw, ad ba. th uvr-
y of Chcago pr, Chcago ad Lodo. 669 .
Gardr, A. L. ad D. C. Carr. 1972. A w sodrmaa
a (phylloomda) from pr. Occaoal par,
Mm txa tch uvry 2:1-4.
Goldma, e. A. 1914. Dcro of v w mammal from
paama ad Mxco. smhoa Mcllao Col-
lco, 60(2): 1-18.
Grfh t., A. ad A. L, Gardr. 2008. sfamly Gloohag-
a boaar, 1845. p. 224-244 Mammal of soh
Amrca, Volm 1 Maral, Xarhra, hrw,
ad ba (A. L. Gardr ed.). th uvry of Chcago
pr, Chcago ad Lodo.
Hadly, C. O. 1960. Dcro of w a from paama.
procdg of h ud sa naoal Mm112:459-479.
Hall, e. R. ad K. R. Klo. 1959. th mammal of norh
Amrca. th Roald pr nw York. 546 .
Hadly, C. O. 1984. nw c of mammal from orhr
soh Amrca: A log og a g Anoura Gray.
procdg of h bologcal socy of Wahgo
97:513-521.
Hadly, C. O. 1987. nw c of mammal from norhr
soh Amrca: fr-ag a, gArtibeus Lach.
p. 163-172 sd norocal Mammalogy (b.D.
paro ad R. M. tmm, d.). eay Hoor of
phl Hrhkovz, Fldaa: Fld Mm of naral
Hory, Chcago.
Hadly, C. O. 1966. Dcro of w a (Choroc
ad Rhohylla) from Coloma. procdg of h
bologcal socy of Wahgo 79:83-88.
Hadly, C. O., ad G. J. Ochoa. 1997. nw c of mam-
mal from orhr soh Amrca: A word-od a
gLonchorhinatom (Chrora: phylloomda).
Mmora d la socdad d Cca naral La sall
57:71-82.
Hrdz-Camacho, J., ad A. Cada. 1978. noa ara la
rv dl Gro Lonchorhina (Chrora, phyllo-
omda). Caldaa 12:199-251.
Hrdz-Camacho, J., A. Hrado , R. Orz, ad t. Walch-
rgr. 1992. udad ogogrca d Coloma. p.105-151 La Dvrdad bolgca d iroamrca (i.
G. Halffr, d.). Aca Zoolgca Mxcaa, io d
ecologa, A.C., Mxco, Mxco.
Hrhkovz, p. 1949. Mammal of orhr Coloma. prlm-
ary ror no.5: ba (Chrora). procdg of h
ud sa naoal Mm 99:429-454.
Hrhkovz, p. 1972. th rc mammal of h norocal
Rgo: A zoogograhc ad cologcal rvw. p. 311-
431 evolo, Mammal, ad sohr Co
(A. Ka, F. C. erk ad b. Gla, d.), Alay sa
uvry pr, nw York.
Hll, J. e. 1965. no o a from brh Gaa, wh h
dcro of a w g ad c of phylloo-mda. Mammala 28:553-572. [Dad 1964; mr 4
lhd 1965.]
Hoffma, F. G., ad R. J. bakr. 2003. Comarav hylo-
gograhy of hor-ald a (Carollia: phylloomda).
Molclar ecology 12:3403-3414.
Hoofr, s. R., s. solar, p. A. Lar, R. D. bradly, ad R. J.
bakr. 2008. phylogc of h fr-ag a (phyl-
loomda: Ara) frrd from mochodral DnA
qc. Occaoal par, Mm of txa tch
uvry 277:1-15.
Hoofr, s. R., ad R. A. Va D bch. 2003. Molclar
hylogc of h chrora famly Vrlloda.
Aca Chrorologca 5, lm:1-63.
Hovkok, J., M. Grga, M. R, L. A. Raa, J. Gr-
rro, A. Raz, J. Mlo, L. Romro, F. Jam, ad s.
Lz. 2007. th ar of Moc Amazoa -
coal maym: Hgh frqcy hf of low
grad coal. Gologcal socy of Amrca bl-
l. 119:1506-1520.
iuCn, Corvao iraoal, Arzoa sa uvry,
txa A&M uvry, uvry of Rom, uvr-
y of Vrga, Zoologcal socy Lodo. 2008.
A Aaly of Mammal o h 2008 iuCn Rd L
8/23/2019 Filostomidos de Colombia.pdf
27/44
mantilla-meluketal.Phyllostomid Batsof ColomBia 23
. Dowloadd o 9
Ocor 2008.
Jmz-Orga A. M. ad H. Malla-Mlk. 2008. el al d
la ala lcva la corvac d oq rocaly al ldad d lo mrclago como odcador d
dro. Rva icoal uvrdad tcolgca
dl Choc Dgo L Crdoa 27(1):100-108.
Kooma, K. F. 1982. bogograhy of h a of soh
Amrca. p. 273-302 Mammala ology soh
Amrca (M. A. Mar ad H. H. Goway, d.). u-
vry of prg, prg, pylvaa.
Kooma, K. F. 1993. Ordr Chrora. p. 137-241 i Mam-
mal sc of h World: a axoomc ad gograhc
rfrc. scod do (D. e. Wlo ad D. M. Rdr,
d.). smhoa io pr, Wahgo.
LaVal, R. K. ad b. Rodrgz-H. Lo mrclago d Coa
Rca. 2002. io nacoal d bodvrdad, inbo320 .
Lar, e. M. O. 1955. no o om mammal from eca-
dor. Aal ad Magaz of naral Hory r 12
8:268-276.
L, t. e., Jr., s. R. Hoofr, ad R. A. Va D bch. 2002.
Molclar hylogc ad ymac rvo of h
g toaa (Chrora: phylloomda). Joral of
Mammalogy. 83:49-57.
Lmk, t. O. ad J. R. tam. 1979. Anoura cultrata (Chi-
rora: phylloomada) from Coloma. Mammala
43:579-581.
Lmk, t. O., A. Cada, R. H. p ad J. i. Hrdz-Cama-
cho. 1982. no o oom, a, ad rod w o
h faa of Coloma. Mammala 2(46):225-234.
Lm, b. K., W. A. pdro, ad F. C. pao. 2003. Dffrao
a c a of h norocal yllow-ard a
Vampyressa pusilla and V. thyone (phylloomda) wh
molclar hylogy ad rvw of h g. Aca
Chrorologca 5:15-29.
Lm, b. K., M. D. egrom, J. C. pao, ad J. W. bckham.
2008. symac rvw of mall fr-ag a (Arti-
beus) from h Gaa, ad a r-valao ofA. glaucus
bototensis. Aca Chrorologca 10(2):243-256.
Lar, O. J., ad J. Oja. 1971. ua va c d mr-
clago dl gro Lonchorhina (Chrora: phyllo-
omda) dl r d Vzla. novdad Cca,
sr Zoolgca 36:1-8.
Lar, O. J. 1997. Lo mamfro d Vzla. socdad d
Corvac Ado d Vzla. 688 .
Lrg, e. 1921. A cod coro o h mammalogy of
ecador, wh om rmark o Coenolestes. Arkv fr
Zoolog sockholm 14(4):1-104.
Lz-Gozlz, C. 2005. Mrclago dl paragay. bfra
lcaco dl com aol dl rograma Mab y
d la Rd iroMab d unesCO 300 .
Lyo, M. W., Jr.1902. Dcro of a w hylloom a from
h ihm of paama. procdg of h bologcal
socy of Lodo 15:151.
Malla-Mlk H. 2007. Lochohll, h Chocoa a.Rva icoal uvrdad tcolgca dl Choc
Dgo L Crdoa 26:49-57.
Malla-Mlk H. ad A. M. Jmz-Orga. 2006. eado d
corvac y alga codraco ogogrca
or la qrorofaa dl Choc bogogrco Co-
lomao. Rva icoal uvrdad tcolgca
dl Choc 25:10-17.
Malla-Mlk, H. ad R. J. bakr. 2006. symac of mall
Anoura (Chrora: phylloomda) from Coloma,
wh dcro of a w c. Occaoal par,
Mm of txa tch uvry 261:1-18.
Malla-Mlk, H. ad R. J. bakr. 2008. Mammala, Ch-
rora, phylloomda,Anoura stulata: Fr cormdrcord for Coloma wh hylogograhc o. Chck
L 4(4):427-430.
Malla-Mlk, H., A. A. M. Jmz-Orga, ad R. J.
bakr. 2009. Mammala, Chrora, phylloomda:
Lonchophylla pattoni: r rcord for ecador. Rva
icoal uvrdad tcolgca dl Choc Dgo
L Crdoa 28 (1):152-155.
Malla-Mlk, H., A. A. M. Jmz-Orga, ad R. J. bakr.
2009. Rag xo ofAnoura aequatoris and notes
on distributional limits of small Anoura Coloma.
Rva icoal uvrdad tcolgca dl Choc
Dgo L Crdoa 28 (1):107-112.
Malla-Mlk, H, H. Ramrz-Chav, C. Frdz-Rodrgz, ad R. J. bakr. (in press). no o gograh-
c dro; Mammala, Chrora, phylloomda,
Anoura stulata. Chck L.
Marcha, R., J. C. brro, H. bhlg, A. boom, ad H. Hoogh-
mra. 2006. Coloma dry mo for rao
h Llao Oral, a comaro of modl ad
oll-ad om rcorco. palaogograhy,
palaoclmaology, palaocology 234:2844.
Markll, C. J. 1970. Vampyrops intermedius . . from Co-
loma (Chrora, phylloomada). Rva bralra
d bologa 30:49-53.
Markll, C. J., ad A. Cada. 1972. no o a w o
h faa of Coloma. Mammala 36:49-58.
Markll, C. J., ad e. Gro. 1966. imoraca d lo mr-
clago ara la ald lca co cal rfrca a la
mco zooca. Aoqa Mdca 16:179-94.
Marz, J. O., 1997. Zoa mara, inGeOMinAs, Mmo-
ra xlcava dl Ala Golgco Dgal d Coloma
(Vr 1.0).
McLlla L. J., ad K. F. Kooma. 2008. sfamly Carol-
la. p. 208-218 Mammal of soh Amrca,
Volm 1 Maral, Xarhra, hrw, ad ba
8/23/2019 Filostomidos de Colombia.pdf
28/44
24 sPeCial PuBliCations, museumof texas teCh university
(A. L. Gardr ed.). th uvry of Chcago pr,
Chcago ad Lodo.
McCarhy, t. J., L. Alja, M. s. Alrco. 2006. A w c
of Chocoan Sturnira (Chrora: phylloomda) fromwr ecador ad Coloma. Aal of h Carg
Mm 75(2):97-110.
McCarhy, t. J., A. Cada G., ad t. O. Lmk. 1983. Com-
m o h r Tonatia carrikeri (Chrora: phyl-
loomada) from Coloma Lozaa. Aca Zoolgca
Colomaa 40:1-6.
Mdll, R. A., M. eqha, ad M. A. Am. 2000. ba dvr-
y ad adac a dcaor of drac noro-
cal rafor. Corvao bology. 14:1666-1675.
Mllr, G. s., Jr. 1931. two w soh Amrca a. Joral
of Mammalogy 12:411-412.
Mllr, G. s., Jr. 1898. Dcro of v w hylloom
a. procdg of Acadmy of naral scc ofphladlha 50:326-337.
Mogro, O. L. ad M. Romro-Rz. 2000. Mrclago
dl cor r d la rraa d Chrq, Caq, Co-
loma. Rva d la Acadma Colomaa d Cca
23(slmo ecal):641-49. [Dad 1999; lhd
Frary 2000].
Mchhala, n., p. Ma V., ad L. Alja V. 2005. A w c
ofAnoura (Chrora: phylloomda) from h eca-
dora Ad. Joral of Mammalogy 86:457-461.
Moz-saa, Y. 1999. Lo mrclago dl gro Artibeus
Lach, 1821 (Chrora: phylloomda: sordr-
maa) d Coloma. t d Mara, io d
Cca naral, uvrdad nacoal d Coloma,
saaf d bogo.
Moz, J. 1986. Mrclago dl parq naral el Rf-
go (Aoqa, Coloma). Acaldadd bolgca
15:66-76.
Moz, J., C. A. Cara, ad M. Gozlz. 2004. s dcr
a va c d mrclago dl gro Carollia
Gray, 1838 (Chrora: phylloomda) d Coloma.
Acaldad bolgca 26:80-90.
Ochoa G, J., ad C. iaz. 1984. nvo mrclago dl gro
Lonchorhina(Chrora: phylloomda). Mmora d
la socdad d Cca naral La sall 42:145-59.
[Dad 1982; lhd 1984.]
Oa-A, O., ad L. G. Gmz. 2000. Rqza, adaca
rlava y aro d acvdad moral d la comdad
d lo mrclago qrro d la Rrva naral La
plaada, naro, Coloma. Rva d la Acadma Co-
lomaa d Cca 23 (slmo ecal): 659-69
[Dad 1999; lhd Frary 2000.]
paro, b. D., G. Callo, W. schr, M. F. togll, t.
brook, L. La, p. Orga, i. salazar, ad b. e. Yog.
2007. Dgal dro ma of h mammal of h
wr hmhr, vro 3.0. narsrv, Arlgo,
Vrga.
pro, R. L. ad J. R. tam. 1968. A w c of a
of h G Sturnira (Famly phylloomada) from
orhwr soh Amrca. Lf scc Occaoal
par, Royal Oaro Mm 12:1-8.
Rok, C. s. ad D. C. Carr. 1972. A w c ofVampyrops
(Chrora: phylloomada) from soh Amrca.
Occaoal par, Mm of txa tch uvry
1:1-7.
saor, C. C. 1932. norocal a h Carg Mm.
Aal of h Carg Mm 21:171-183.
saor, C. C. 1941. Dcro ad rcord of norocal
a Fld Mm of naral Hory. Zoologcal sr
27:371-387.
saor, C. C. 1949. ba of h g Micronycteris and its
gra. Fldaa: Zoology, w r 31:215-233.
saor, C. C. 1955. Rmark o h a of h g Vampy-
rops. Fldaa: Zoology 37:403-413.
sachz-palomo, p., M. dl p. Rva-pava, ad A. Cada.
1996. Dvrdad olgca d a comdad d
qrro y rlac co la rcra dl ha d
oq d galra, srraa d la Macara, Coloma.
Caldaa 18:343-353.
shaml, H. H. 1927. A w a from Coloma. procdg of
h bologcal socy of Wahgo 40:129-130.
smmo, n. b. 2005. Ordr Chrora. p. 312-529 Mam-
mal sc of h World: a axoomc ad gograhc
rfrc, 3rd do (D. e. Wlo ad D. M. Rdr.
ed.) Joh Hok uvry pr, balmor, Mary-
lad.
smmo, n. b. 1996. A w c ofMicronycteris (Chi-
rora: phylloomda) from orhar brazl, wh
comm o hylogc rlaoh. Amrca
Mm nova 3158:1-35.
smmo, n. b., ad R. s. Vo. 1998. th mammal of paraco,
Frch Gaa: A norocal lowlad rafor faa,
par 1. ba. bll of h Amrca Mm of naral
Hory 237:1-219.
smmo, n. b., R. s. Vo, ad D. W. Flck. 2002. A w Ama-
zoa c ofMicronycteris (Chrora: phylloo-
mda) wh o o h roog havor of ymarc
cogr. Amrca Mm nova 3358:1-14.
solar, s., s. R. Hoofr, p. A. Lar, A. brow, R. J. bll, J. A.Grrro, J. Orga, J. p. Carrra, R. D. bradly, ad R.
J. bakr. ( r). Oraoal crra for gcally
dd c: aaly of dvrcao of h mall
fr-ag a,Dermanura (phylloomda: sodr-
maa). Aca Chrorologca.
tam, J. R., D. Valdvo, ad J. i. Hrdz-Camacho.
1965. Addoal rcord ofChoeroniscus Coloma.
Joral of Mammalogy 46:704.
tlg, e. C., M. s. srgr, O. Mad, p. ba, s. J. Obr,
ad W. J. Mrhy. 2005. A molclar hylogy for a
llma ogograhy ad h fol rcord. scc
307(5709):580-584.
8/23/2019 Filostomidos de Colombia.pdf
29/44
mantilla-meluketal.Phyllostomid Batsof ColomBia 25
trra, D. 2007. Ga d camo d lo mamfro dl ecador.
edco Mrclago blaco. plcac cal or
lo mamfro dl ecador 6. Qo. 576 .
thoma, O. 1909. no o om soh-Amrca mammal,wh dcro of w c. Aal ad Magaz
of naral Hory r 8(4):230-242.
thoma, O. 1912. nw a ad rod from soh Amrca.
Aal ad Magaz of naral Hory r 8 10:403-
11.
thoma, O. 1913. A w g of gloohag a from Co-
loma. Aal ad Magaz of naral Hory r
8, 12:270-71.
thoma, O. 1897 Dcro of w a ad rod from
Amrca. Aal ad Magaz of naral Hory r
6 20:544-53.
tchd, J. J. vo. 1844. Mammalm coc. Archv na-
rgch 10(1):244-55.
Valdvo, D. 1964. La faa qrra dl Daramo d
Cdamarca, Coloma. Rva d bologa trocal
12:19-45.
Valdvo, D., ad J. R. tam. 1962. Fr rcord of h al
ar-od a Coloma. Joral of Mammalogy
43:422-423.
Vlazco, p. M. 2005. Morhologcal hylogy of h a g
Platyrrhinus sar, 1860 (Chrora: phylloom-
da) wh h dcro of for w c. Fldaa:
Zoology 105:1-53.
Vlazco, p. M. ad b. D. paro. 2008. phylogc ad o-
gograhy of h road-od a, GPlatyrrhinus(Chrora: phylloomda). Molclar phylogc
ad evolo 49:749-759.
Vlazco, p. M., ad A. L. Gardr. ( r) A w c of
Platyrrhinus (Chrora: phylloomda) from wr
Coloma ad ecador, wh mdd dago ofP.
aquilus,P. dorsalis, adP. umbratus. procdg of h
bologcal socy of Wahgo.
Vlazco, p. M., A. L. Gardr, ad b. D. paro. ( r).
Systematics of thePlatyrrhinus helleri c comlx
(Chrora: phylloomda), wh dcro of wo
w c. Zoologcal Joral of h Zoologcal L-
a socy.
Vo, R. s. 2009. book Rvw. Mammal of soh Amrca,
Volm 1 Maral, Xarhra, hrw, ad a.
Joral of Mammalogy. 90(2):521-523.
Wr, W. D. ad J. K. Jo Jr. 1987. A w c of
Glossophaga commissarisi (Chrora: phylloomda)
from soh Amrca. Occaoal par, Mm of
txa tch uvry 109:1-6.
Wlgh, F. p., ad J. A. salo. 2006. A Moc rcv o
h volo of h Amazoa oa. scra Gologca
133:439-457.
Wrr A. L. M. V. Rockma, ad n. b. smmo. 2000.
phylogy of phylloomd a (Mammala: Chrora):daa from dvr morhologcal ym, x chromo-
om, ad rrcd . bll of h Amrca
Mm of naral Hory 248:1-200.
Wlo, D. e., ad D. M. Rdr (d.). 2005. Mammal sc
of h World: A taxoomc ad Gograhc Rfrc
(3rd d). Joh Hok uvry pr.
Woodma, n. 2007. nw c of car-fdg a, g
Lonchophylla, from Wr Coloma ad Wr
ecador (Mammala: Chrora: phylloomda).
procdg of h bologcal socy of Wahgo
120(3):340-358.
Woodma, n., ad R. M. tmm. 2006. Characr ad hy-
logc rlaoh of car-fdg a, whdescriptions of new Lonchophylla from western South
Amrca (Mammala: Chrora: phylloomda: Lo-
chohyll). procdg of h bologcal socy of
Wahgo 119:437476.
Addresses of authors:
hugo mantilla-meluk
Darm of bologcal scc
box 43191
txa tch uvry
Lock, tX 79409
alex mauriCio Jimnez-ortega
Gro d ivgac Majo d Faa slvr
Chocoana
uvrdad tcolgca dl Choc, D.L.C.
Qd, Coloma
roBert J. Baker
Darm of bologcal scc
box 43191
txa tch uvry
Lock, tX 79409
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26 sPeCial PuBliCations, museumof texas teCh university
aPPendix i
Modl of oal dro for cormd Coloma hylloomd c.
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mantilla-meluketal.Phyllostomid Batsof ColomBia 27
aPPendix i (Cont.)
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aPPendix i (Cont.)
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mantilla-meluketal.Phyllostomid Batsof ColomBia 29
aPPendix i (Cont.)
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aPPendix i (Cont.)
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aPPendix i (Cont.)
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aPPendix ii
Modl of oal dro for hylloomd c oally r Coloma.
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mantilla-meluketal.Phyllostomid Batsof ColomBia 33
aPPendix iii
L of lograhc rfrc of lhd mammala c chckl ha cld hylloomd darmal
rcord for Coloma. th ld rfrc wr d a orc corcg colm wo of tal 1.
Colombia in general.Alrco, M., A. Cada, J. Hrdz-Camacho, ad Y. Moz-saa. 2000 Ma-
mfro (syada: thra) d Coloma. boa Colomaa 1(1):43-75; All, J. A. 1900. L of a collcd
y Mr. H. H. smh h saa Mara rgo of Coloma, wh dcro of w c. bll of h
Amrca Mm of naral Hory 13:89-94; All, J. A. 1916. L of mammal collcd Coloma y
h Amrca Mm of naral Hory xdo, 1910-1915. bll of h Amrca Mm of naral
Hory 35:191-238; Araa, A. A., J. b. Vagh, K. e. nwll, R. A. J. barh, ad M. Graca. 1968. Salmonella
and Shigella fco a lcd ara of Coloma. th Amrca Joral of trocal Mdc ad
Hyg 17:92-95; bag, O. 1900. L of h mammal collcd h saa Mara rgo y W. W. brow Jr.
procdg of h nw Zlad Zologcal Cl 1:87-102; barrga-bolla, e. 1965. edo maozoolgco
colomao, i. Chrora. Caldaa 9:241-271; Crvo Daz, A., J. Hrdz Camacho, A. Cada G. 1986.
La acalzada d lo mamfro d Coloma: aoaco or drc. Caldaa 15:471-501; Gardr,
A. L. 2008. Mammal of soh Amrca, Volm 1 Maral, Xarhra, hrw, ad ba. th uvry ofChcago pr, Chcago ad Lodo. 669 ; Hrhkovz, p. 1949. Mammal of orhr Coloma. prlmary
ror no.5: ba (Chrora). procdg of h ud sa naoal Mm 99:429-454; iuCn, Corva-
o iraoal, Arzoa sa uvry, txa A&M uvry, uvry of Rom, uvry of Vrga,
Zoologcal socy Lodo. 2008. A Aaly of Mammal o h 2008 iuCn Rd L . Dowloadd o 9 Ocor 2008; Markll, C. J., ad A. Cada. 1972. no o a w o h
faa of Coloma. Mammala 36:49-58.
Regional and departmental records.Antioquia: Cara-Call, C. A., J. Moz-Arago. 2003. La d
lo mamfro (Mammala: thra) dl daramo d Aoqa, Coloma. boa Colomaa. 4(1):65-78;
Moz, J. A. 1986; Mrclago dl arq aral el Rfgo (Aoqa, Coloma). 1986. Acaldad
bolgca. 15(57):65-76; Moz-saa, Y., H. F. Lz-Arvalo, ad A. Cada. 1999. Aor al coocmo
d la cologa d lo mrclago d lo aoramo d mrmol y calza, cor d Ro Claro (Aoqa,Coloma). Rva d la Acdma d Cca naral Fca y exaca d Coloma. slmo cal:
651-658; Caldas: Caao, J. H., Y. Moz, J. e. boro, ad J. H. Vlz. Mamfro dl daramo dar-
amo d Calda-Coloma. 2003. boa Colomaa. 4(2):247-259; Caqueta: Mar-Vazqz, A., ad A. V.
Aglar-Gozlz. 2005. Mrclago (Chrora) dl daramo d Caq-Coloma. boa Colomaa.
6(2):211-218;Cauca: Alrco, M., ad A. J. ngr. 1992. prmr aor or lo mamfro dl Vall dl paa.
novdad Colomaa, Mo d Hora naral, uvrdad dl Caca, nva eoca 5:66-71; Rva-pava,
M. p., H. e. Ramrz-Chav, Z. i. Alvarz, b. L. no Valca. 2007. Calogo d mamfro r la
colcco d rfrca y xhc dl Mo d Hora naral d la uvrdad dl Caca. tallr doral
uvrdad dl Caca, poay. 96 ; Choc: Arlla-Aglar, A. A., A. M. Jmz-Orga, H. Malla-
Mlk. 2007. Aaly of h o-hmaohag a c card wh of radcao of Dmod rod
(e. Goffroy, 1810) h Coloma ogograhc Choc.Revista Institucional Universidad Tecnolgica Del
Choc. 26:42-48; Jmz-Orga, A. M., J. Moz-Arago, A. M. Gozal, O. Y. scada-Ma, n. Gl-pao.2002. edo d alga comdad d qrro oq lval rocal y oq my hmdo ro-
cal dl Choc (Coloma).Revista Institucional Universidad Tecnolgica Del Choc.15:20-25; Malla-Mlk
H. ad A. M. Jmz-Orga. 2006. eado d corvac y alga codraco o
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