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Life sciences

Welcome | 1

Life sciences at Hull | 4

Why study at Hull? | 6

The Hull Campus | 8

The ScarboroughCampus | 10

Choosing a degree andgetting a place | 12

Your questionsanswered | 13

Employability of ourgraduates | 14

Hull Campus courses | 16

Scarborough Campuscourses | 26

The four-year route | 29

Life sciencesmodules | 30

Staff and their researchinterests | 31

The Deep | 36

Admissions/enquiries

Hull CampusJon Harvey (Admissions Tutor):[email protected] Secretary:01482 465144 / [email protected]/biosciBiological SciencesUniversity of HullHull, HU6 7RX

Scarborough CampusSunil Shastri (AdmissionsTutor): 01723 357221Administrator: 01723 [email protected]/cemsCentre for Environmental and

Marine SciencesUniversity of HullScarborough CampusScarborough, YO11 3AZ

Hull CampusSingle Honours UCAS code

Aquatic Zoology C390 BSc/AqZ

Biology C100 BSc/B

Biology with Molecular Bioscience (3 years) C1CT BSc/BMBs

Biology with Molecular Bioscience (4 years) C1CB MBiol/BMB

Biomedical Science (3 years) BC99 BSc/BiSc3

Biomedical Science (4 years with placement) BC9X MBSci/BS

Human Biology C102 BSc/HB

Marine and Freshwater Biology C163 BSc/MFB

Zoology C300 BSc/Zoo

With a foundation year

Aquatic Zoology (4 years) C350 BSc/AqZ4

Biology (4 years) C101 BSc/B4

Biology with Molecular Bioscience (4 years) C1CA BSc/BMBs4

Biomedical Science (4 years)* BC9Y BSc/BioS4

Human Biology (4 years) C103 BSc/HB4

Marine and Freshwater Biology (4 years) C165 BSc/MFB4

Zoology (4 years) C301 BSc/Zoo4

Scarborough CampusSingle Honours UCAS code

Coastal Marine Biology* CD14 S BSc/CMB

Ecology* C180 S BSc/Ec

Environmental Science* F750 S BSc/Env

With a foundation year †

Coastal Marine Biology (4 years)* CD1K S BSc/CMB4

Ecology (4 years)* C181 S BSc/Ecol4

Environmental Science (4 years)* F754 S BSc/EnvS4

* Also offered for part-time study.† The foundation year is taught at the Hull Campus.

Typical offers• Single Honours, Hull Campus: 240–280 points including A level Biology grade C• Single Honours Biomedical Science: BBB–BCC• Scarborough Campus: 200+ tariff points• Four-year BSc programmes with foundation year: 140–180 tariff points

Key facts

Life scienceswww.hull.ac.uk 1

Hello and a very warm welcome to the Department of Biological Sciences andthe Centre for Environmental and Marine Sciences (CEMS). We invite you todiscover why we pride ourselves on having two of the country’s friendliestdepartments and why our undergraduates are highly satisfied with theirstudent experience. Whatever your area of interest, we have something foryou.

Biological SciencesThe Department of Biological Sciences, on the Hull Campus, is a modern anddynamic environment with internationally recognised research activities. It has along-standing tradition of teaching excellence and offers a wide range of Bachelorsdegree courses. The superb lab facilities, integral use of IT for teaching and broadvariety of expertise within the department mean that choice is rarely an issue.

We have a thriving campus community, with an award-winning nightclub and astudents’ union that is among the best in England.

CEMSThe Centre for Environmental and Marine Sciences is a specialist department basedat our Scarborough Campus, with particular strengths in marine, ecological andenvironmental teaching and research. The department’s small size means thatstudents learn in a variety of ways which are not really possible in a largerdepartment. Field and lab work constitute around 40% of our students’ experiencewhile they are here – whether that is sampling organisms on the beach, dissectingsquid in the lab or climbing trees in Dalby Forest.

Despite its rural location, state-of-the-art technology means that our wirelessScarborough Campus is superbly well connected, and the online resources areidentical to those on the Hull Campus.

A shared ethosOur world-class research, recognised in the recent national Research AssessmentExercise, has attracted a vibrant PhD student population that helps to inspire andinform our teaching. It also means that you may have the opportunity to getinvolved with staff research projects and develop your skills further.

Exotic field courses and training in skills such as diving mean that, in our opinion,our courses offer more added value than other comparable degrees in the UK. Asolid reputation and an emphasis on transferable skills are just two of the reasonswhy more than 95% of our students find employment within six months ofgraduation.

The Sunday Times called Hull ‘the best-kept secret in higher education’. Now thesecret is out.

Wishing you all the best in your studies,

Dr Graham Scott Dr Phil WheelerHead of Biological Sciences Head of CEMS

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Hull was the first university in the UKto offer dive training as a Year 1

undergraduate option. Successfulcompletion of that module enablesyou to participate in the final-year

field trip – studying coral reefs off theEgyptian coast – and gives you the

opportunity to train to EuropeanScientific Diver standard.

Since 2004 theUniversity hasinvested £3 million innew and improvedinfrastructure. Inparticular, ourteaching labs havebeen fully refurbishedto provide world-class learningfacilities.

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A tradition of excellence extending into the 21st centuryEstablished in 1928, the Department of Biological Sciences has an excellenttradition in both research and teaching. Our very first Head of Department, SirAlistair Hardy, FRS, is world-famous for his work in oceanography and research intothe implications of plankton distribution for fisheries. As Chief Zoologist on Scott’sfamous ship the Discovery (1926) he was one of the first marine biologists to exploretropical and Antarctic waters. The department also has a strong tradition inbiomedical science, with notable successes such as the discovery of the role ofprolactin in vertebrate osmoregulation and the creation of the IVF Unit at HullPrincess Royal Hospital.

But the department is not just about great traditions – it is a thriving, dynamic placethat provides a vibrant teaching and research environment. Since 2004 theUniversity has invested £3 million in new and improved infrastructure. Inparticular, our teaching labs have been fully refurbished to provide world-classlearning facilities at a cost of more than £700,000.

In terms of research, Biological Sciences is an active, focused department that is atthe forefront of many exciting fields of biology. Over the last decade it hasdeveloped an international reputation not only in biomedical science but also inevolutionary biology and functional ecology. Researchers are interested in projectsranging from the cellular basis of diseases such as cardiovascular disease andsleeping sickness to the ecology and genetics of invasive ladybirds and theevolution of sexual mode in tadpole shrimps. The marine sciences featureprominently and are a particular area of expertise in CEMS at the ScarboroughCampus.

Most importantly, we translate this cutting-edge knowledge into research-driventeaching that benefits students at all levels.

Teaching facilitiesPractical classes at the Hull Campus take place in two ultra-modern teaching labs,both of which have been completely refurbished in the last few years. In all thereare over 160 computer workstations and interactive screens, allowing for computer-assisted teaching. Demonstrations by lecturers can be projected on screen, and allinformation can be accessed directly by every student. Both labs are particularlywell suited to cater for disabled students, featuring special wheelchair benchingand facilities for visually and hearing impaired students.

Teaching facilities at Scarborough also take advantage of the latest technology, andthe campus is entirely wireless so that students and staff can be online whereverthey are working or learning. Web-assisted learning and assignments are a user-friendly teaching innovation used at both campuses, alongside more traditionaltechniques.

Life sciences at theUniversity of Hull

Once you havegained the basicskills required forbiological sciencesduring your first year,you can selectspecific modules toreflect your interestsand, in effect, buildyour own degree.

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In short: quality of teaching, friendliness of staff and students, value formoney, unrivalled flexibility and added value leading to employmentafterwards. This pamphlet explains why we – and our students – love thisplace, and describes the opportunities that it provides. We will try to help youchoose the degree (and campus) most appropriate for you.

One of the greatest advantages of studying at Hull is that you will become part of auniquely interactive department where the breadth of disciplines covered by staffallows for collaborations in both research and teaching. For example, molecularbiologists are collaborating with fisheries biologists to look at the genetic structureof fish stocks, and invertebrate physiologists are collaborating with biochemists tostudy the chemical basis of behaviour and its evolution.

Most importantly, this collaborative research will benefit your education. State-of-the-art knowledge acquired via research is built into the structure and delivery ofour degree courses. Once you have gained the basic skills required for biologicalsciences during your first year, you can select specific modules to reflect yourinterests and, in effect, build your own degree. For example, you might wish tostudy marine and freshwater biology but also specialise in molecular biology. AtHull you can do this, and even have the flexibility of choosing between twocampuses. Finally, all students are able to do their own research project and workclosely with research scientists in the lab or in the field.

In choosing your university you need to consider many factors, including

• teaching quality and the value of the experience• graduate employment prospects and the value of the skills taught• the quality of the facilities, including student accommodation• the content and structure of the course

All of these vary considerably from university to university and from department todepartment. We believe, however, that life sciences at the University of Hull hasmuch to offer in every important area. Consider following the facts and figures.

Student satisfactionFor the fifth year running, the University of Hull has emerged as one of the top 10mainstream English universities for overall student satisfaction. Our teaching,assessment, feedback and academic support were all highly rated. This is no flash-in-the-pan result – our students are truly satisfied with their experience at theUniversity of Hull.

Excellent employabilityThe University of Hull regularly achieves a high position in league tables recordinggraduate employability, with fewer than 8% of students still unemployed sixmonths after graduation. Biological Sciences usually matches, and sometimesimproves on, these figures.

Why study at theUniversity of Hull?


First-class facilities• Both campuses offer a student-friendly environment.• We are a long-established traditional university dedicated to excellence in both

teaching and research. But we are also highly innovative, with investment in newteaching labs, new degrees in Human Biology and Zoology, and new teachinginitiatives such as the Undergraduate Ambassadors Scheme.

• University accommodation is excellent, and both Hull and Scarborough benefitfrom a plentiful supply of inexpensive high-quality private-sectoraccommodation. See the University’s prospectus or visit www.hull.ac.uk/accom.

Dynamic degrees• Our degree courses will introduce you to the core subjects but also allow you to

develop and pursue your own particular interests. Your progress will bemonitored regularly, and you will have the opportunity to transfer between someof our specialist degree courses.

• Our teaching and research interests span the breadth of modern-day biology,from molecular biology to management of natural resources.

• You may attend field courses or undertake field-based projects both in the UK andin exotic locations such as Brazil, Tobago, Mallorca and Honduras.

• At the Hull Campus we run a third-year module called Biology in Educationwhich allows students to spend time in local schools and help teaching biology –an excellent experience for those considering a teaching career.

• We offer a well-structured and proven foundation year, taught at the HullCampus, for applicants who do not have appropriate entry qualifications.

Prime prospects• We have excellent links with local industry, hospitals and research units – almost

all our Biomedical Science students gain employment straight away.• You may continue your studies by working towards a research degree (MSc,

MRes, PhD) or one of several taught Masters degrees, all encouragingcollaboration with potential employers.

• Our courses give you ample opportunity to refine those ‘key skills’ so popularwith employers.

We are a long-establishedtraditional universitydedicated toexcellence in bothteaching andresearch. But we arealso highlyinnovative, withinvestment in newlabs, new degreesand new teachinginitiatives.

The department runsbotanic gardens andhas very good linkswith industrialbiotechnologycompanies, localhospitals and NHSresearch units.

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Biological Sciences occupies most of the Hardy Building and the adjacentWolfson Building. Within these are teaching and research laboratories, anElectron Microscope Suite, freshwater and marine aquaria, a state-of-the-artGenome Analysis Suite, lecture theatres and staff offices. The department alsoruns botanic gardens in Cottingham, near Hull, and has very good links withindustrial biotechnology companies, local hospitals and NHS research units,not least the world-renowned Magnetic Resonance Imaging Unit at Hull RoyalInfirmary.

Our degree portfolioWe offer three-year degree courses in Biology, Biomedical Science, Aquatic Zoology,Marine and Freshwater Biology, Human Biology, Zoology, and Biology withMolecular Bioscience. We also offer routes into these for applicants without thenormal qualifications (see page 29). We are constantly updating our portfolio, socontact us for the latest information or check our website.

Why choose to study on the Hull Campus?• A department offering the chance to study the most recent developments in a

wide range of current biological topics• A compact campus with all facilities close at hand• Top-quality teaching in a supportive and friendly environment• A degree that allows you to choose the modules that most interest you• The chance to travel overseas to conduct field-based work on coral reefs,

mangroves and exotic freshwater systems• A wide choice of free elective modules from other departments if you wish to

broaden your education• Great links with local industry, hospitals and NHS research units• The chance to learn to dive and move towards a professionally recognised

qualification• Close to the centre of one of the UK’s largest cities, with all the social and

recreational facilities you would expect

Undergraduate Ambassadors SchemeAre you thinking of teaching as a possible career? If so, Hull is the place for you. In apioneering venture, our department offers an undergraduate module called Biologyin Education. Developed under a national initiative called the UndergraduateAmbassadors Scheme, the module gives you the chance to visit a school, help inclasses and try teaching. In addition, you will devise and develop a biology-basedproject with your class.

This is a great way of gaining some teaching experience and finding out whether itis the right career for you. You gain a range of skills, improve your employabilityand become a role model inspiring more young people to choose science-basedcareers or education in the future. A highly rewarding experience!

The Hull Campus

Just a few hundredyards from the shoreof the Yorkshirecoast, and a fewmiles away from theNorth York MoorsNational Park, thecentre hasincomparable naturalfacilities for studyand research on itsdoorstep.

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The Centre for Environmental and Marine Sciences (CEMS) is situated in anideal position on the Scarborough Campus. Just a few hundred yards from theshore of the Yorkshire coast, and a few miles away from the North York MoorsNational Park, it has incomparable natural facilities for study and research onits doorstep.

Situated within the Worsley Building, the centre offers modern laboratory andaquarium facilities and a dedicated microscopy suite. The newly refurbished andextended library and IT facilities are easily accessed, and the halls of residence –providing excellent en-suite accommodation – are also located on campus.

Our degree portfolioAt Scarborough we offer three-year degrees in Coastal Marine Biology,Environmental Science and Ecology. We also offer routes into these for applicantswithout the normal qualifications (see page 29) and opportunities for part-timestudy.

Why choose to study on the Scarborough Campus?• Degree courses that emphasise the hands-on, applied aspects of biology thanks

to the easy access to a variety of marine and terrestrial ecosystems• Small, intimate campus that enables you to fulfil your potential as an individual,

rather than one among the crowd• Easily accessible staff who will know your name and provide support when you

need it• Links with the British Antarctic Survey and the Central Science Laboratory, who

provide expert input• Surfing opportunities when you need a break from work!

Dive trainingDive training for undergraduates is facilitated at both campuses, and there areacademics on both campuses with Scientific Diving expertise. Training to OpenWater level is offered as a free elective, and those who wish to do so can train toEuropean Scientific Diving standard. Students from CEMS have exclusive access totheir own training facility at Wykeham Lakes.

Although many students purchase their own equipment, CEMS has enough to trainas many students as wish to learn. A recent development has been the purchase offull-face masks and underwater communication technology that give student diversexperience of using commercial dive equipment.

Dive training not only lets you experience the wonderful underwater world at firsthand; it also encourages personal development (confidence, working in groups,planning, safety awareness, self-discipline), and it looks great on your CV –whatever career path you choose.

The ScarboroughCampus

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Our degrees are listed on the inside front cover.Details of these courses are given on subsequentpages. For up-to-date information call us or checkour websites.

There is no need to worry about choosing between ourcourses. Before you make your UCAS application, youcan call us (01482 465398 or 465198) for advice. We willhelp you choose between Hull and Scarborough, andgive advice about which degree may suit you best. Youmay also email either of the following people. Wepromise a speedy reply.

Dr Jon Harvey – Admissions Tutor, [email protected]

Sunil Shastri – Admissions Tutor, [email protected]

One other piece of advice: if you are thinking of applyingfor one of our degrees but are not certain whether youshould go for a three-year course or a four-year coursewith a foundation year, put either one on your UCASapplication and we will advise you during your open-dayvisit. If you have applied for the four-year route and dobetter than expected in your exams, we may be able totransfer you to the three-year degree if you wish.

Getting a placeIf we make you an offer we will invite you to one of ouropen days, at either the Hull or the Scarborough Campusas appropriate. Please note that all candidates for thefour-year Masters in Biomedical Science (BC9X) will beinterviewed. You will be given a choice of dates anddays, usually Wednesdays and Saturdays. Members ofyour family or partners will also be encouraged toattend. Late applicants or those who cannot attend opendays should contact the admissions staff identifiedabove, who will make arrangements for individual visits.

We welcome and encourage applications from studentswith disabilities.

Entrance requirements depend on the degree you wish totake. When we make our offer your suitability for thecourse applied for will be taken into account along withyour predicted examination performance. Candidatesoffering qualifications other than A levels (for example,BTEC, IB, GNVQ, Access courses, etc) will be givencareful and sympathetic consideration. Anyone withoutformal qualifications should contact us to see if theirapplication will be considered.

For the four-year BSc degrees with a foundation(including Biomedical Science)These degrees are designed for those who in some waydo not meet the usual entry requirements of the three-year courses. (You may, for example, be offering anunusual combination of A level subjects.) See page 29 orcontact our admissions officers for more details.

For Biomedical Science (three-year BSc and four-yearMasters with placement):We ask for good A level passes in Biology, Chemistry andone other, preferably science, subject. A typical offerwould be 240–280 points for the three-year BSc and 300or above for the four-year undergraduate Masters.

For all other coursesWe ask for A level passes (or equivalent qualifications) intwo science subjects, preferably including a biologicalsubject. In our experience general aptitude anddetermination to succeed are often more important thanA level subjects. If Biology is your only science, we willstill give your application serious consideration. We willeven consider applicants without science A levels as ourfour-year BSc courses can be used as ‘conversion’courses. Our typical offer is 240–280 points.

Changing your mindA major attraction of a University of Hull degree is thatyou have the flexibility to change your mind! The onlycourse with set target numbers is MBiomedSciBiomedical Science (four-year with placement). Becausethis is funded by a health authority (NHS) and provides abursary, the numbers must be fixed. Otherwise we willfill degree courses according to demand.

This means that if you apply for, let’s say, Biology butdecide later that your interests really lie in Marine andFreshwater Biology, it will be simple for us to transferyou to your new choice, even after you arrive in Hull.Similarly, if you apply to Hull but later decide that youwould be better suited to a Scarborough course, or viceversa, we will probably be able to change yourapplication. Because most degrees share a common firstyear (the skills year), a change of course is possible aftera year of your studies or even later. Also, if you apply fora three-year degree but have a disaster and do badly inyour A levels there is the built-in ‘safety net’ of an almostcertain place on one of our four-year BSc courses, givingyou a chance to get back on track in a supportiveenvironment.

Choosing a degree and gettinga place


Who can I turn to for help and advice?Every student is allocated a personal tutor, a member ofstaff who will look after you during your time at theUniversity of Hull. Both Biological Sciences in Hull andCEMS in Scarborough have an ‘open-door’ policy, whichmeans that you can call at most times on any member ofstaff and seek their help. Staff are busy, but you will findthat we are approachable and genuinely willing to help.If we can’t help we usually know someone who can.

Howmuch work do I have to do? Is there any time forsport or leisure activities?The Year 1 timetable has seven or eight one-hour lecturesand three practicals, each lasting three to four hours, perweek. Wednesday afternoons are free for all students.The compact nature of both campuses means that youwill not spend much time travelling around, and if youwere feeling energetic you could fit in a game of squashbetween lectures or go to the superb students’ union justnext to Biology at Hull (or even go surfing atScarborough!).

How do I get to know other people in the department?The induction week features a number of social events toensure that you meet staff and fellow students, andwhen teaching starts in a practical or tutorial you willwork in small groups and will soon get to know othermembers of your class.

Can I choose modules from outside the Department ofBiological Sciences?Yes, in most courses a wide range of free electivemodules are available from across the subjects offered atthe campus you are attending (see the inner back cover).The department itself offers a number of free electivemodules, including Dive Training, Animal Behaviourand Marine Field Studies.

How do I choose a research project?In your second year, and with guidance from advisers,you choose a research supervisor for your final year andchoose a project from the range available. This givesundergraduate students the opportunity to join aresearch group and gain research experience. We offer20- and 40-credit research projects, including lab-basedprojects, projects involving field studies abroad, andliterature-based projects; there is also the UndergraduateAmbassadors Scheme, where you work in local schools.This practical element of your studies can contributesignificantly to your final degree assessment. It is also ofvalue in showing prospective employers yourcommitment to specific fields of biology and your

potential as a research scientist, or it can be seen as astepping stone to a Masters or a PhD. The researchinterests of our academic staff are listed on pages 32–4.

Are there any field courses?Field courses form an integral part of the Marine andFreshwater Biology, Coastal Marine Biology, AquaticZoology, Environmental Science and Ecology degrees.Biology students may also attend (space permitting).Students may attend a residential field course: recentdestinations have included North Yorkshire, the Isle ofCumbrae, the Red Sea, Thailand, Brazil and LakeMalawi. The cost of the field course depends on thedestination you choose, but all are heavily subsidised bythe department and represent excellent value for money.

What will I have to buy?You’ll need a lab coat. You’ll need some books. Ourlibraries on both campuses do have multiple copies ofall-important textbooks, but it will be more convenient tohave your own. At the Hull Campus there is also a smallone-off charge to help offset the cost of the printedlaboratory manuals and guides. Those taking fieldcourses will need wellington boots and/or snorkel andfins, sunglasses and suntan lotion. Those choosing theDive Training module will be required to contribute tothe cost.

Can I transfer from another institution?One benefit of CATS (the Credit Accumulation andTransfer Scheme) is that it facilitates such transfers. Evenif your present institution isn’t yet running CATS, you arewelcome to contact our admissions officer (see page 12)to see if you can be considered.

Can I gain work experience during my degree?We allow students to suspend their studies for a yearbetween the second and third levels if they wish to gainwork experience in a relevant industrial or researchenvironment in the UK or abroad. In addition, a numberof institutions offer summer internships to our top-performing students. So you may be offered the chanceto work at The Deep, the Hull IVF Unit or the Institute ofEstuarine and Coastal Studies during your first summer.Many of our staff also look for students to undertakesummer projects and can arrange fellowships fromplaces such as the Nuffield Foundation – just approachstaff working on subjects you have a particular interestin and they will be more than happy to help.

Your questions answered

Hull is regularlyamong the top UKuniversities in leaguetables recordinggraduateemployability: morethan 86% of itsstudents areemployed or engagedin further studywithin six months ofgraduation.

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The University of Hull is nationally recognised for the success of its graduates.Regularly among the top UK universities in league tables recording graduateemployability, more than 86% of its students are employed or engaged infurther study within six months of graduation. Biological Sciences usuallymatches, and even improves on, these figures.

Key issues to considerWe cannot deal here with all the complexities of graduate employment – some ofwhich are discussed in the Careers Service’s entry in the University’s prospectus –but a few points deserve mention.

• Graduates looking for satisfying work always have been and will continue to be ata considerable advantage compared with non-graduates.

• Employers today look for ‘transferable skills’ and often provide specialisedtraining ‘in house’. We have therefore built such skills into our degree courses.This ensures that our students are suitably prepared for the work market andespecially for the 40% of opportunities that are completely ‘open’. For these thesubject of your degree is less important than how your practical, social andintellectual skills have developed under the stimulus of the University’s academicand social life.

• The University’s Careers Service has been notably successful in helping ourstudents find jobs that fulfil them.

• The Careers Service does not stop providing help when you graduate but will, ifyou wish, continue to help you throughout your working life.

Opportunities for biology graduatesMany graduates take up careers that draw on the various key skills developed andpractised during their time at the University, rather than on their biological training.Many employers recognise that biologists from a good university make excellentemployees. This university is at the forefront, for example, of environmentaltechnologies, a job market that has, according to the Natural Environment ResearchCouncil, an estimated skills gap of more than 1.5 million jobs in the next 10 years.

Our degree courses open up many such career possibilities in fields related tobiology: that is, in industries such as pharmaceuticals, chemicals, paramedicalwork, hospital services and food production, or in other important areas such aspollution control, exploitation of natural resources and environmentalconservation. Typical employers include the water industry, government agencies,industrial concerns, local authorities, nature conservation bodies, researchinstitutions and international agencies.

The improved management of fisheries is an important issue for many nations andis recognised as a major concern within the European Common Fisheries Policy.Careers will thus continue to be available for graduates in the fishing industry,aquaculture, national and international government agencies, and the fisheriesresearch institutions.

Those who graduate with an upper second or a first class degree may also considerbecoming specialist researchers within universities, industry or research institutessupported by the Government. We offer guaranteed places on our Masters courses(depending on performance) for all our graduates.

Employability of ourgraduates

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BSc Aquatic Zoology

The seas cover most of the earth’s surface and are ofgreat significance for reasons that range fromharvesting fish to the disposal of waste. Fresh wateris a similarly precious commodity that supports adiverse range of organisms, but the resource is underincreasing pressure from human use. This coursefocuses on the land/sea interface – as represented bymarine coastal areas, estuaries and associatedwetlands – and on freshwater environments.

In particular, the course explores the fascinatingdiversity, ecology and physiological adaptations oforganisms in the freshwater, estuarine and intertidalenvironments, emphasising how such scientificknowledge informs the management of theseenvironments. The commercial importance of suchresources is another key topic. The course is delivered byour successful research and teaching groups in aquaticecology and resource management, aquaticecophysiology, and evolutionary biology.

Key modules provide extensive coverage of evolutionarybiology, ecophysiology of aquatic organisms, fishecology, conservation biology, behavioural ecology,biodiversity, oceanography, and freshwater and coastalfisheries. State-of-the-art teaching in modules onmolecular ecology and the evolution of aquaticorganisms focuses on tropical fishes (cichlids andguppies) that we research in our impressive freshwateraquaria.

One or more residential field trips are available. In recentyears, students have had the choice of a UK-based trip orone to a more exotic foreign destination. All trips aresubsidised heavily, but students have to bear some of thecost. Foreign destinations include Tobago and Mallorca.Honduras may be a new destination for 2010.

Hull Campus courses

Focus on research

Impacts of hydropowerThe impacts of hydropower schemes on fishpopulations are well documented. Common examplesinclude changes in fish species composition orpopulation structure, loss of access to spawninggrounds, or even loss of spawning grounds themselvesbecause of changes in habitat associated with theengineering works.

Nevertheless, such schemes remain a favourableoption for power generation since they are ‘renewable’and do not pollute the environment to the extent thatmore traditional sources of power generation do. HIFI(the Hull International Fisheries Institute, one of ourspecialist units) was commissioned to survey the fishpopulations of a selection of upland streams in Wales,Scotland and Ireland that were under consideration forpower generation. The aim was to identify keyspawning and nursery areas for salmon and browntrout, so that such areas could be protected whenfuture hydropower schemes were designed.

Fisheries data were collected and analysed for agestructure, population density, and growth rates.Continuing research provides scientific analysis andresolution of the possible conflict between the need forrenewable energy and the conservation of importantaquatic species.

Chemical ecologyHow do animals communicate? This is a question asold as mankind, and it is of both academic andpractical interest. We want to understand how animalsignals have evolved but also whether they can bedisrupted by human influences – for instance, whetherpollutants such as fuel oil may alter odour-mediatedbehaviours. To this end our research has uncoveredmany aspects of the chemical nature of animal signals.For example, we have induced a male shore crab toattempt to mate with a stone by coating the stone withthe sex pheromone from a female crab. This is adramatic example of how important chemical signalsare in controlling behaviour, particularly in aquaticanimals. The research feeds into applied projects suchas studying the use of chemical signals in aquaculture,their potential use against invasive pest species(crayfish, zebra mussels, American lobster), and howto use signal molecules to monitor/manage otterpopulations.


BSc Biology

Our flagship degree course, Biology provides a broad education for those whochoose not to specialise. Depending on your choice of modules, you canincorporate molecular biology, human biology, ecology and biomedical-basedsubjects in your degree. You can thus keep your degree broadly based or givea particular emphasis to your studies by your choice of optional modules. Thelist of modules on page 30 shows that you have plenty to choose from.

Our modules are usually worth 20 credits, and you are required to take 120 creditseach year. You have the option of selecting 20 of these credits each year from a freeelective catalogue that lists a range of attractive modules offered by otherdepartments. In this way you could supplement your biological training, if youwished, with, for example, a language module or one in psychology.

Learning takes place in lectures, practical classes and tutorials. You will also havetutorials – small-group sessions involving other students and your personal tutor.In your second year you will meet with other staff and have the chance to explorecontemporary issues in biology at a scientific and an ethical level, and in your finalyear you will receive close personal tuition from your research project supervisor.

Assessment takes a number of forms. Lectures are normally assessed by writtenexamination at the end of the module, but multiple-choice tests and essays writtenduring the teaching period are also used. All laboratory-based work is continuouslyassessed by means of lab books, assignments or short tests. For most modules thesplit between continuous assessment and written exam is 50:50. If you fail a moduleyou will be reassessed at a later date. Our overall failure rate is low, and weencourage and expect all our students to perform at their highest possible standard.Normally more than half the class graduate with degrees classified at the top twolevels (first and upper second).

Depending on yourchoice of modules,you can incorporatemolecular biology,human biology,ecology andbiomedical-basedsubjects in yourdegree.

Focus on research

Cell biology of trypanosomesThe parasite Trypanosoma brucei causes African sleeping sickness, a disease thatclaims more than 50,000 lives per year in sub-Saharan Africa. Klaus Ersfeld’s labis working on aspects of the cell biology of T brucei using molecular cell biologytechniques to identify molecules involved in the pathogenesis of this organism,with the aim of exploiting them as potential drug targets. Specific topics ofresearch include mechanisms of cell division and chromosome segregation duringmitosis. This research integrates well with modules taught by Dr Ersfeld inmodular cell biology and in molecular and medical parasitology.

We have designed abroad-based coursewith a moderncurriculum where youwill be able to chooseoptions to match yourinterests across thebreadth of biology.

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BSc/MBiol Biology with Molecular Bioscience

These new courses fit closely with the existing and successful BSc Biologydegree but will especially appeal to students wanting to learn more moleculargenetic techniques and understand how to take molecular approaches to arange of modern biological questions.

Molecular genetic techniques are now widespread in all areas of biology, beingcommonly used by researchers in disciplines such as parasitology, microbiology,ecology, evolutionary biology, fisheries, conservation biology and biomedicalscience. Yet many universities have molecular biology degrees that are restricted to‘classical’ molecular biology systems. By contrast we have designed a broad-basedcourse with a modern curriculum where you will be able to choose options to matchyour interests across the breadth of biology.

The MBiol degree allows you to progress to an advanced fourth year in which youcan choose to work in the research laboratories of members of staff, and also takeboth practical and lecture courses on molecular techniques. One example might be‘Practical DNA Sequencing and Bioinformatics’, in which students will sequence asmall genomic library and interpret the data by working in the department’sGenome Analysis Suite and Bioinformatics Laboratory.

Focus on research

Molecular genetic approaches to biodiversityApproximately half of the Earth’s surface is deep ocean. In these depths 50–90%of all animals are nematode worms. Although they dominate the abyssalcommunity, little has been know about their biology or diversity. Are these speciesunique to the ocean depths or are they much the same as those on the continentalshelf? By DNA sequencing and bioinformatics Dr David Lunt, working with theNatural History Museum and other institutions, has been able to show extensivenew diversity in the ocean abyss.

Molecular genetic approaches like these are becoming more and more importantin biodiversity research, and students in the department have worked on projectsinvestigating many animals including owls, sea cucumbers and iguanas.

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BSc/MBiomedSci BiomedicalScience

The biomedical sciences are among the fastest-growing areas of knowledge, and all aspects ofbiomedical science and research are increasinglysophisticated. Graduates are needed to cope withthis complexity and to develop and implement newfundamental knowledge in the clinical environment.Many of our graduates are employed as biomedicalscientists or researchers in the NHS and in researchlaboratories, universities or the private sector.

Our degrees are designed to equip you with the skills tofollow a career in the health services, includingbiomedical scientist roles, research and development,clinical trials, education, and sales and marketing.

These courses have been designed in partnership withthe Hull and East Yorkshire Hospitals NHS Trust andembody a multidisciplinary laboratory-based approach,encouraging flexibility in professional outlook while stillallowing specialisation. Both courses are accredited bythe IBMS (Institute of Biomedical Science).

Students following the four-year MBiomedSci course,sponsored by NHS Yorkshire and the Humber, spend ayear on placement in one of our partner hospitallaboratories. The degree offers a high level ofexperiential training, and is validated by the IBMS andapproved by the Health Professions Council (HPC).Details on funding can be obtained from the University.

Year 1 and Year 2 are identical for the BSc andMBiomedSci courses. The first year consists of coremodules introducing the fundamentals of biochemistry,microbiology, physiology and cell biology. In Year 2 youstudy the biological basis of disease and fundamentalconcepts underlying the clinical disciplines: clinicalbiochemistry, haematology and transfusion science,cellular pathology, medical microbiology andimmunology. These modules include contributions bysenior staff from hospital laboratories in Yorkshire.

At the end of Year 2, the four-year course is extended bya short placement in a hospital laboratory.

In your third year you follow in-depth modules in twoclinical specialties as well as the importance of qualityassurance, gaining a broader perspective.

On the three-year BSc you gain experience in researchthrough a laboratory-based project in the final year andby choosing optional modules designed to develop yourbreadth of knowledge. Graduates from this degree are

eligible to apply for trainee biomedical scientist posts inhospital laboratories.

On the four-year degree,* you spend a year workingwithin one of the clinical disciplines in an approvedhospital training laboratory in the Yorkshire region.

Hospital staff act as practice tutors, training andmentoring you throughout the year. One of the aims hereis to provide training and experience appropriate to therequirements of the HPC, including fitness to practice,alongside University-based assignments and the IBMSPortfolio for the Certificate of Competence.

After the training year you return to the University toundertake a laboratory-based research project andmodules on cardiac and skeletal muscle disorders,control of infection and management of health services.

* The Department of Health is currently reviewing thetraining for health care scientists nationally, which mayhave implications for the four-year programme from 2011.Contact us for an update on this provision.

Meeting professional standardsThe MBiomedSci (BC9X)* has been approved by theHPC: on graduation, students can apply to the HPC forRegistered Practitioner status as biomedical scientists.Satisfactory assessment of the Registration Portfolioduring the placement year ensures that our graduatesmeet the professional standards required. Incollaboration with the University of the West ofEngland, we use ‘Profiles’ – an electronic version of theportfolio.

Widening access to biomedical scienceIf you are interested in studying Biomedical Sciencebut do not have conventional qualifications, our four-year course including a foundation year is designed formature applicants or those with qualificationsinappropriate for direct entry to BC99. All four yearsare taught at the University, where you have access toall facilities. Year 1 provides a firm foundation inchemistry, mathematics and biochemistry.

What happens after graduation?Our first cohort of students graduated in 1999. Sincethen the number of graduates has increased steadily,and 65% of graduates from 1999–2009 have taken upposts as biomedical scientists. Graduates have gainedemployment, or continued their education with a PhD,MSc, PGCE or medical degree.


BSc Human Biology

This degree course has been developed to meet the growing need forscientists who are able to understand and solve problems at the interface ofbioscience and medicine. Its content reflects the interdisciplinary nature ofbiomedical sciences and offers excellent job prospects in academic research,industry, medical institutions and education.

The degree has a flexible modular structure which allows students to define theirown emphasis on particular areas of human biology. Some of the modules areshared with the department’s highly successful Biomedical Science degrees.

During the first year you will study core disciplines of biology, such as physiology,molecular biology and genetics, and biochemistry.

In the second year these foundations are expanded in modules specific to humanbiology, covering topics such as molecular cell biology, pathology, immunology, thebiological basis of disease and human evolution. You will also be able to do aliterature study on a topic of your choice.

The third year offers opportunities to specialise in areas that particularly interestyou. Examples are infectious diseases and molecular parasitology, cellularpathology, applied molecular biology, sports physiology and drug discovery. Youwill have the opportunity to undertake a two-semester research project in an arearelevant to human biology. Students’ research projects are closely linked withongoing research in our department. Also, for students interested in getting a job inteaching and education, we offer a secondary school placement module that willallow you to develop teaching skills.

Year 2 containsmodules specific tohuman biology,covering topics suchas molecular cellbiology, pathology,immunology, thebiological basis ofdisease and humanevolution.

Focus on research

Human biologyHeart and vascular diseases are one of the major causes of death in the Westernworld.The teams around Dr Anne-Marie Seymour and Dr Sandra Jones investigatechanges in heart tissue metabolism underlying the development of heart failure.Approaches involve the use of molecular and cellular imaging together withmagnetic resonance techniques.The groups have close collaborative links with theAcademic Department of Cardiology and the Department of Nephrology at HullRoyal Infirmary.

Infectious diseases are one of the most important causes of mortality and disabilityworldwide, particularly in developing countries. The groups led by Dr Klaus Ersfeldand Dr Frank Voncken are working on the parasite Trypanosoma brucei, thecausative agent of sleeping sickness in sub-Saharan Africa. Using molecular cellbiology and imaging techniques, they are investigating how the parasite mutipliesand persists inside the human host.

Dr Camille Ettelaie and her research group are investigating the role of cell surfacereceptors and associated signal transduction processes in cardiovasculardisease.This research is relevant to the mechanisms of atherosclerosis andthrombosis.

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BSc Marine and Freshwater Biology

This course covers many aspects of marine andfreshwater animals and plants and their relation tothe environment, giving particular emphasis to theeffects of human activities on nearshore andestuarine systems. In order to prepare students forcareers in the field, due weight is given toenvironmental management. The course focuses onmarine and freshwater zoology, and to that extentshares many modules with Aquatic Zoology, but alsocovers microscopic and macroscopic plants andother microbial organisms.

We start by giving you a general biological background,but from the outset we take you into the field to allowyou to experience a range of habitats and techniques.Many of our students now broaden their experience andskills by taking our free elective module in Dive Training.Such skills can then be employed in the field coursesthat come later.

The final year includes a research project based in thefield or laboratory, and students are given the chance towork with the research teams in our Institute ofEstuarine and Coastal Studies (IECS) and InternationalFisheries Institute (HIFI) as well as others in thedepartment. Topics range from fundamentalphysiological studies to impacts of pollution, speciesconservation and behavioural studies. Some studentstake the opportunity to mix the marine biology withmolecular genetics and issues relating to naturalselection and evolution.

The course builds on a long tradition of marine andfreshwater biology at Hull. The department and itsinstitutes (IECS and HIFI) have many members ofacademic staff as well as researchers who specialise inmarine, freshwater and estuarine biology.

The course also prepares students to enter taught MSccourses at Hull and elsewhere. Those who take this routegain a further qualification which increases theiremployability and breadth of expertise.

Focus on field workOur Marine and Freshwater Biology students areexposed to field work through a residential course atWhitby, North Yorkshire, in their first semester andthen through courses in such locations as Portugal,Mallorca, Tobago and Scotland. During theseinternational field courses, which usually occur duringthe vacation between the second and third years, ourstudents acquire many skills that are highly relevant totheir future careers as well as seeing other habitats andspecies. They survey marine and freshwater systemsand do small projects which allow them to explore andunderstand these ecosystems.

Focus on research

Physical ecologyAll animals and plants live in a world defined byphysical laws. Through natural selection, organismsare constantly bumping up against the physical limitsof their environment, but they can also take advantageof these limits in interesting ways. For instance, manyaquatic organisms take advantage of the way in whichwater flows over their bodies to enhance theirrespiration, photosynthetic or feeding rates.

By using simple ideas from engineering and physics,we can increase our understanding of the biology andecology of living systems. Our research spans a widerange of biology, from the dynamics of particle captureby filter feeders, through swimming by microscopicorganisms, to using knowledge of physical laws in thepresent to predict the behaviour of extinct animals. Weapply these concepts to organisms as diverse as fishand plankton and to environments ranging from coralreefs to the open ocean. While our biological workdraws on engineering and physics, it also has thepotential to provide insights from biology that can leadto solutions to engineering problems.

BSc Zoology

This course combines elements of traditional zoology with contemporarybiology to create a zoology programme for 21st-century scientists. It includesanalyses of animal biology from cellular to population level, and coversmolecular, neuronal, morphological and behavioural approaches.

During the first year of study you will cover the basic building blocks of zoology –organism diversity, evolution, ecology, genetics and biochemistry – and will havethe option to widen your experience through the study of physiology or chemistry orby taking part in a field trip to learn about intertidal organisms. Some students willeven learn to dive by following one of our credit-bearing free elective modules.

For second- and third-year students, options broaden to include animal behaviour,aspects of vertebrate biology and ecology, molecular genetics, evolutionary biology,ecophysiology, human/animal interactions, conservation biology andenvironmental management. The highlights of your degree may be our final-yearpractical modules, which offer you opportunities to work as a member of one of theresearch groups in the department on an original research project. If your heart isset on teaching as a career but your head tells you to try it first, we can even arrangefor you to take a credit-bearing module, Biology in Education, that will allow you todevelop your subject knowledge in the school classroom (see pages 8 and 30).

Learning and assessment styles are varied in all courses offered by the department,and BSc Zoology is no different. You will be exposed to classroom-based learning insmall seminar groups or traditional lecture situations. You will take part inlaboratory practicals and have the opportunity to experience field work; you willlearn independently and as a member of a group; and you will benefit fromexcellent IT and literature resources in a friendly and supportive setting.


During the first yearof study you willcover the basicbuilding blocks ofzoology – organismdiversity, evolution,ecology, genetics andbiochemistry.

Focus on research

Evolutionary causes and consequences of dispersal and colonisationDispersal and colonisation of new habitats are among the most important events inan organism’s life. Dr Lori Lawson Handley is interested in the evolutionary causesand consequences of dispersal: for example, how does geography influencedispersal, and what makes certain species so successful at colonising new areas?Recent work has focused on global colonisation and local dispersal in humans andthe harlequin ladybird – species that have more in common than you might think!

Dr Lawson Handley and colleagues’ recent research has shown that geneticdiversity declines smoothly with distance from East Africa, the cradle of modernhumans. These simple patterns tell us that anatomically modern humans evolved inAfrica and that from there they colonised the world, beginning approximately60,000 years ago. We are now trying to improve our understanding of the role ofnatural selection in shaping human populations: for example, we have establishedthat adaptation to colder climates has left a footprint on our genomes.

Another very successful global coloniser is the harlequin ladybird, Harmoniaaxyridis. Native to Asia, it is currently invading the British Isles, continental Europe,the USA and several other parts of the world. We are investigating the spread of thisinvasive species using molecular genetic techniques, and trying to understand whyit is so invasive. Members of the public have made a fantastic contribution to ourwork by reporting sightings and sending samples to the Harlequin Ladybird Survey.

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BSc Coastal Marine Biology

Students spend 40% of their time in the lab or in thefield. Field work is easy when the beach is 10minutes’ walk from the campus. Our emphasis ondeveloping real skills and learning through actionrather than by rote makes our students stand outfrom the crowd.

It is estimated that almost half the world’s population(more than 3 billion people) live or work within 200 kmof the coast. Coastal waters and the estuarine systemswithin them form an important link between terrestrialand marine environments, exhibiting complex interfacesin terms of energy and material movements. In addition,they represent an environment that is essential to theglobal economy and significantly disturbed by humanactivities. Organisms that live in the intertidal zone needto be adapted to deal with a narrow corridor of availablehabitat that constantly changes character over distancesof centimetres and minute by minute.

The Coastal Marine Biology degree has a core of field-based modules. You are required to undertake field workin all three years of the course, some of it residential. Inyour first year, you carry out a range of field-basedactivities such as sampling organisms from intertidalsediments, beach profiling, and identifying a range oforganisms found on the beach and in scientific trawls.Many Coastal Marine Biology students opt for the DiveTraining module and spend a significant amount of timein the pool and our freshwater training facility atWykeham Lakes. During Year 3, you can participate inthe optional Field Studies module and take the skills thatyou have learned in the UK to a more exotic locationsuch as Egypt, Mallorca or Brazil.

Students on the course take six modules per year.Typically these involve 10–15 hours of lectures supportedby about 30 hours of lab study, field work, tutorials andseminars. Depending on your choice of optionalmodules over the three years, modules are assessed by avariety of means (examination, coursework, practicalexams, oral presentations, web pages, posters orreports).

Scarborough Campus courses

Focus on research

Life in the deep freezeTwo staff members and two honorary fellows at CEMShave particular interests in the biology and ecology ofthe Antarctic. Dr Magnus Johnson studies the ecologyand behaviour of krill (whale food), while Dr CathWaller is interested in the ecology and physiology oforganisms throughout the Antarctic intertidal. Bothbring their research into the classroom and use dataand anecdotes from their antipodean adventures toenthuse students.

Krill (Euphausia superba) are regarded as the Duracellbatteries of the Antarctic ocean, and there are enoughof them to cover Scotland in a one-krill-deep blanket.They are the main food source for many of the‘charismatic megafauna’ (whales, dolphins andpenguins) that are found in the region. Like mostplankton, they migrate to the surface to feed and sinkto depth to escape predation. Until recently, it wasconsidered that they fed at night and hid during theday – making only a single migration every 24 hours.Working with Dr Geraint Tarling from the BritishAntarctic Survey, Dr Johnson found that they mayactually migrate 3–4 times every 24-hour period as theytend to snooze after eating, drift down to depth,defecate and then return to the surface to feed. Thishas interesting implications for the global carbon cycleas each time they sink they take captured carbon todepth so that it can become locked up in deep-seasediments.

The Antarctic intertidal zone has always been thoughtof as an environment that is too extreme for anythingto live in permanently. Dr Waller’s research has shownthat this is not always the case and that – at many siteson the Antarctic Peninsula – a wide range ofinvertebrates can survive even in the harsh winterconditions, where temperatures can fall to −25o C. DrWaller studies how these animals are able to survivethere and how they respond to environmental changes.This is especially important as the surface watersaround the Antarctic Peninsula are one of the fastest-warming areas anywhere in the world, and manyAntarctic invertebrates may take a long time to adapt tothese changing conditions.

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BSc Ecology

Ecologists are by nature ‘hard-core’ field biologists, exploring the physicaland biological links between organisms and environments using datacollected in the field. Scarborough’s unique position between the sea and themoors provides a wide range of habitats and an excellent field environmentfor the study of ecology.

The degree offers the excitement of 40% field work within the robust framework of arigorous academic course. From day one, students are immersed in an academicenvironment designed to provide them with the field-based skills and knowledge topursue their chosen careers. There are residential and local field-trip opportunitiesin all three years of the degree course. Local field work might involve ground-truthing maps for a geographic information systems (GIS) module or climbing treesas part of the Canopy Science module. Destinations for the final-year residentialfield trips currently include Egypt, Brazil, Mallorca and the west coast of Scotland.

Many of the other modules – such as Conservation Biology, Upland Ecology,Ecological Monitoring and Environmental Ecology – have interesting day tripsassociated with them. In second- and third-year modules, students use the trips togather data for assessed projects of their own design. In recent years, students haveoccasionally assisted the centre’s research postgraduates in studying the ecologiesof hares, adders or intertidal fish.

We aim to produce graduates who hit the ground running and are equipped to carryout high-quality ecological investigations, either at postgraduate level or within theprivate or public sector. As with all of the courses at the Centre for Environmentaland Marine Sciences, some Ecology students choose to remain at university for afurther year to study for an MSc by research, under the guidance of tutors that theyhave known during their undergraduate years.

From day one,students areimmersed in anacademicenvironmentdesigned to providethem with the field-based skills andknowledge to pursuetheir chosen careers.

Focus on research

Ecological studiesStaff and students at all levels get involved in research at Scarborough, much of itcoming from strong links with local and national organisations. Researchers atScarborough work in a range of environments, exploring how organisms andecosystems function and putting this knowledge to practical use. Staff arecurrently investigating a wide variety of topics from coastal fisheries to seaweedcommunities, coral reefs and upland mammals.

The anoa is a dwarf buffalo found only in rainforest on the islands of Indonesia. Itis one of South-East Asia’s least known large mammals and is declining rapidlyowing to hunting and habitat loss. Phil Wheeler has been spending his summerson Buton living in makeshift camps, sleeping in a hammock and walking linetransects through mountainous rainforest to survey the distribution andabundance of this rare species. His field work aims to improve our knowledge ofthis rare and charismatic animal, the threats it is currently facing and its generalecology. During his field work Phil often encountered hunters carrying dead anoasand, through interviewing them, found that large adults of this species – thepopulation of which is estimated at around 3,000 – will sell for no more than £30.

Research in ecology at Scarborough is expanding our knowledge of species andtheir environments in a way that will help safeguard their future.

‘I’m in Year 3 and havethoroughly enjoyed my time atScarborough. The size of theplace makes it easy to get toknow like-minded people, andthe course provides lots ofinteresting opportunities. Theteaching is excellent, and thelecturers really do follow the‘open-door policy’. The week-longfield trip to Scotland in Year 1allowed us all to get to know eachother really well, then at the endof Year 2 we travelled toIndonesia. Here we investigatedrainforest and coral reef ecology– a highlight of the course and anexperience I’ll never forget. Lastyear a group of students,including myself, set up SHrUBS(the Scarborough and HullUniversity Biological Society),and this year I took over asChairman. The aim was toenhance our field-work skills andour CVs, allowing everyone’sinterests to be catered for. Whaleand dolphin watches, marinemammal rescue courses and birdsurveys were a few of theactivities organised this year. I’vealso taken up scuba diving andam currently working towards myPADI Divemaster qualification.Being able to explore theunderwater world is fantastic,and this is a skill that I’ll continueto use in the future.’

Peter RobinsonBSc Ecology


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BSc Environmental Science

This degree explores the wide range ofenvironmental issues and processes that underpinthe link between humankind and the naturalenvironment. Our emphasis on developing practicalskills and learning through field-based teaching willsee students spend 40% of their time in the lab or inthe field and leave them well equipped for a flyingstart in the job market.

Environmental Science is a broad-based, holistic coursewhich not only draws on physical sciences such asbiology, geography, geology, chemistry, ecology andanthropology but also deals with topics relating torecreational management and to legal, economic andethical issues. The teaching team reflects this diversity oftopics and consists of a terrestrial ecologist, aneconomist, environmental chemists, a geomorphologistand marine biologists.

The degree has a core of field-based modules. You willundertake field work in all three years of the course, someof it residential. Proximity to the North York MoorsNational Park, the Yorkshire Wolds and Britain’s‘dinosaur coast’ provides ample opportunities for field-based study. In Year 1 this encompasses ecologicalsampling, water quality analysis, stratigraphy, andcoastal and fluvial geomorphology monitoringtechniques. For the core final-year residential field trip,many of our students opt for Mallorca – where they applywhat they have learned in north-east England to the widerange of habitats and environments available there.

Students on the course take six modules per year.Typically these involve 10–15 hours of lectures supportedby about 30 hours of lab study, field work, tutorials andseminars. Depending on your choice of optionalmodules over the three years, modules are assessed by avariety of means (examination, coursework, practicalexams, oral presentations, web pages, posters orreports). The course has a strong applied component,and associations with environmental practitioners andmanagers (e.g. North York Moors National Park, ForestryCommission, environmental consultants) provide ourstudents with exceptional resources and exposure tofuture career opportunities.

Our students have gone on to study for PhDs or Mastersdegrees in specialist areas such as archaeology,environmental management and environmental law, andhave found employment as environmental planners,teachers, environmental consultants and field scientists,among other roles.

Focus on research

Post-industrial pollution: extent and treatmentStaff and students at all levels get involved in researchat Scarborough, much of it coming from strong linkswith local and national organisations. Researchers inthe Centre for Environmental and Marine Sciences arecurrently investigating a wide variety of topics. Theserange through coastal fisheries, the impacts of modernagriculture on mammal populations, issuessurrounding the effects of environmental pollution andthe development of green technologies to mitigateenvironmental impacts from industry.

Pollution arising from abandoned industrial sites cancause long-lasting and severe impacts on river systems.Mine sites and steelworks, for example, can generateeffluents with extreme chemistry, often rich in ecotoxicmetals that can persist for decades. One problematicissue surrounding such pollution is that often there isno single body with legal liability for clean-up costs.Therefore low-cost, environmentally sensitivetreatment technologies such as treatment wetlandshave been developed for site clean-up where suitable.Recent research has also shown that the extent ofpollution arising from our abandoned metal mines islikely to have been severely underestimated previously.The pollution burden for some toxic metals arisingfrom these is of an equivalent size to that arising fromall licensed industrial premises (e.g. factories, sewagetreatment works) nationally.

Highlighting the enduring effect that industrial activitycan have on the movement of pollutants in ourenvironment, this research is being used byenvironmental regulators and managers to determinewhich sites should have priority when deciding how tospend limited remediation funds.


The four-year BSc degree courses are designed for students who do not meetthe normal entry requirements of our three-year degrees. The first yearprovides a firm foundation and is taught at the Hull Campus by staff from theBiological Sciences Department and the University’s Faculty of Education.From the very start you will be a full member of the University, with access toall our facilities.

The areas covered by the foundation year include mathematics, chemistry, IT andstudy skills. You may then stay at the Hull Campus or transfer to Scarborough(according to which course you have chosen) to begin your degree studies. For a listof available courses, see the inner front cover.

The first students taking the four-year route graduated in 1997, and we are confidentthat over the following years we have improved and refined its structure andcontent until it perfectly matches the requirements of applicants unsuited for directentry to the three-year degrees. This route is ideal for

• overseas students who may require time to adjust to study in the UK (in somecircumstances modules in English language can be included in the course)

• mature students without recent formal qualifications (at interview we will belooking for evidence of a continuing interest in learning)

• applicants who may have made a choice of A levels inappropriate for the three-year route

• applicants with predicted performance just below that required for direct entry tothe three-year degree

We have improvedand refined the fouryear route’s structureand content until itperfectly matches therequirements ofapplicants unsuitedfor direct entry to thethree-year degrees.

The four-year route

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Hull Campus

Details of which modules make up which courses areavailable on our undergraduate pages atwww.hull.ac.uk/biosci. We cannot guarantee that aparticular module will be available, but you can restassured that our courses always provide diversityand stimulation and are always current. Assessmentmethods vary, but will usually comprise 50%coursework and 50% written exam.

* free elective module** Biomedical Science onlyE module taught wholly or in part on the web1, 2, 3 level (year) of study

ModulesChemistry for Biologists 1Diversity of Aquatic Organisms 1Dive Training* 1Ecology and Evolutionary Biology 1Ecology and Evolutionary Biology with

Marine Field Work 1Field Studies in Marine Biology* 1Genetics and Molecular Biology 1Human Physiology* 1Human Physiology and Metabolism A 1Human Physiology and Metabolism B 1Introductory Biochemistry and Microbiology E 1Laboratory Based Skills 1 1Laboratory Based Skills 2 1Plant and Animal Diversity 1Skills for Biologists 1Animal Behaviour* 2Biological Basis of Disease 2Clinical Disciplines I** 2Clinical Disciplines II** 2Conservation Biology 2Contemporary Issues in Biology 2Ecophysiology of Aquatic Plants and Animals 2Evolutionary Biology 2Fish Ecology 2Human Pathology 2Immunology and Proteomics 2Laboratory Based Skills 3 2Molecular Genetics 2Molecular Cell Biology 2People, Biology and Their Environment 2Physical, Chemical and Biological Oceanography 2Vertebrate Zoology 2Applied Molecular Biology and Regulation of Gene

Expression 3

Behavioural Ecology and Physiology ofAquatic Animals 3

Biology in Education 3Biomedical Science Research Project** 3Cellular Pathology 3Clinical Chemistry 3Coastal Fisheries Management 3Current Ornithology 3Field Studies 3Fisheries Resource Management 3Freshwater Fisheries and Conservation 3Haematology 3Human Evolution and Genomics 3Infection Control 3Introduction to Drug Discovery 3Laboratory Based Skills 4 3Marine Eco-Mechanics 3Medical Microbiology 3Molecular and Medical Parasitology 3Placement 1** 3Placement 2** 3Pre-Placement Studies** 3Professional Studies** 3Research Project 3Reviews in Biology and Biomedical Science 3Sociobiology 3Threats and Remedies in Aquatic Environments 3Whales, Dolphins and Sharks 3

Life sciences modules

Focus on teaching innovationWe pride ourselves on our ground-breaking provisionof new and exciting initiatives for our students.

We were the first biological sciences department in theUK to offer Dive Training as a Year 1 free electivemodule. It is taught in collaboration with Above andBelow, a 5* PADI Dive Training Facility, and studentsare trained up to PADI Advanced Open Water. While afee is payable to attend the module, this is heavilysubsidised by the department. Training includeslectures, self-study periods, pool training and your firstopen-water dives. This module has proved hugelypopular.

We are also justifiably proud of being the firstbiological sciences department in the country topioneer the Undergraduate Ambassadors Scheme.Under this scheme (a national initiative) we offer anundergraduate module called Biology in Education.This gives participating students the chance to visit aschool, help in classes and try some teaching, inaddition to devising and developing a biology-basedproject with their class.


Scarborough Campus

Details of which modules make up which courses areavailable on our website (www.hull.ac.uk/cems).Please note that the list below is subject to change,depending on staff expertise and student interest. Ifyou have a question about a programme or module,please email [email protected] and the relevant tutorwill do their best to respond: our open-door policystarts right here.

* free elective module1, 2, 3 level (year of study)

ModulesDiversity of Life 1Dive Training 1Earth and Environmental Systems 1Global Environmental Issues* 1Introductory Research 1Physical Sciences 1Practical Ecology 1Aquatic Zoology 2Canopy Science 2Conservation Biology* 2Environmental Ecology 2Environmental Pollution and Toxicology 2Environmental Resource Management 2Integrated Coastal Management 2Intertidal Ecology 2Oceanography 2Remote Sensing and GIS 2River Basin Management 2Animal Behaviour 3Ecological Monitoring 3Environmental Impact Assessment 3Environmental Law 3Environmental Monitoring and Analysis 3Evolutionary Ecology 3Field Studies 3Fish and Fisheries 3Independent Research Project / Independent Library

Project 3Tropical Marine Systems 3Upland Ecology 3

Teaching innovations at ScarboroughAs at the Hull Campus, we pride ourselves oninnovative approaches to teaching and are alwayslooking for new ways of getting students enthusedabout their chosen topic. One of the latest is acollaboration between the Centre for Environmentaland Marine Sciences and the Department of ComputerScience. Eco-Sub allows students to experience avirtual marine world where they can explore theecology of deep-sea organisms by piloting a scientificsubmarine. Doing this exercise gives students arealistic feel for the difficulties involved in exploringthe ocean depths.

For many students the virtual marine world is notenough. Fortunately, dive training is also available atScarborough. For some students, this is actuallyessential preparation for the final-year field trip toEgypt. During the first year you can train to Open Waterstandard, and if you wish you can go on to achieve aprofessional standard (Divemaster or Scientific Diver).

If the deep sea doesn’t offer enough of a challenge,then you can always try your hand at teaching. As inHull, there are opportunities to get practically involvedwith local schools through the UndergraduateAmbassadors Scheme, and even have a go at teachinga class.

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All staff are active in research areas which complement our degree courses.Postgraduate training is also given, leading to the degrees of MSc and PhD bythesis; and we offer one-year MSc taught courses. There is considerableoverlap and interaction between the various research groups, but theresearch interests of the individual members of staff will give you an idea ofthe range of projects available in the final year.

Biological Sciences Department, Hull CampusAlistair Anderson, Lecturer Production of biodegradable plastics usingmicroorganisms.

Aziz Asghar, Lecturer Use of electrophysiological and immunocytochemicalmethods to investigate spinal cord function; spinal neuronal networks in chronicpain; comparison of dorsal horn neurotransmission in normal and arthritic states.

Thomas Breithaupt, Lecturer Sensory ecology of aquatic animals (crustaceans andfish) – mechanism of chemical signalling and odour orientation; behaviouralstrategies in dominance and mate choice; the nature of chemical signals.

Ian Cowx, Professor and Director of HIFI, Director of Research Impact of man’sactivities on freshwater fisheries, particularly with reference to river regulation, landdrainage engineering works and stock manipulation; freshwater fisheries stockassessment; tropical fisheries.

Mike Elliott, Professor, Director of Institute of Estuarine and Coastal StudiesBiology of estuarine and coastal fish and benthic organisms; ecotoxicology at theindividual-organism, population and community level.

Klaus Ersfeld, Senior Lecturer Molecular cell biology of the human parasiteTrypanosoma brucei; mitotic mechanisms and their exploitation as drug targets; celldivision and the cytoskeleton.

Camille Ettelaie, Lecturer Blood coagulation, thrombosis and haemostasis, mainlyconcerning cardiovascular disease.

Africa Gómez, NERC Advanced Research Fellow Ecological genetics of rotifers;phylogeography; cryptic species; evolutionary genetics of asexual reproduction.

Ray Goulder, Senior Lecturer, Deputy Head of Department Microbial ecology ofnatural waters; bacteria and pollution; relations between epiphytic bacteria andaquatic plants; microbiology of fish farming and its effluents.

John Greenman, Professor Tumour immunology; cancer genetics; prognostic anddiagnostic factors.

Rob Hammond, Lecturer Evolutionary biology; the evolution of social behaviour;conflict and cooperation in animal societies; behavioural genetics.

Bernd Hänfling, Research Fellow Ecology and evolution of aquatic organisms.Specific areas of interest: phylogeography; gene flow in riverine ecosystems; use ofmolecular markers and adaptive traits in conservation genetics; evolutionaryconsequences of biological invasions.

Jörg Hardege, Reader Reproductive biology and chemical ecology of marineinvertebrates; ecology of invasive species, animal behaviour and evolution.

Staff and theirresearch interests


Jon Harvey, Lecturer, Admissions Tutor Electric fishing and fish stock assessment;impact of fish-eating birds; recruitment in fish populations; conservation offreshwater species; impacts of hydropower schemes; impacts of flow regulation onfisheries.

Rebecca Hill, Lecturer Connective tissue changes in diabetic nerves; barrierfunction within the peripheral nerve; the effect of growth hormone on skeletalmuscle mitochondrial function in growth hormone deficient adults.

Stuart Humphries, NERC Fellow and Lecturer Biological fluid dynamics; thebiology of suspension feeders; coral reef biology; comparative biomechanics ofmarine organisms.

Bill Hutchinson, Lecturer Genetic analysis of ecological and behaviouralprocesses; environmental/anthropogenic influences on population dynamics andgenetic structure; genetic approaches to fishery management; non-invasive geneticapplications in conservation.

Sandra Jones, Lecturer Changes in ion channel expression, the regulation of cellsignalling and the intervention of exercise in the ageing heart.

Domino Joyce, Nuffield Research Fellow Sexual selection and behaviour ofAfrican cichlid fish; speciation; biodiversity.

Lori Lawson Handley, Lecturer Evolutionary biology of invasive species(particularly the harlequin ladybird); human migration and settlement history;evolutionary causes and consequences of dispersal; sex chromosome evolution.

David Lunt, Senior Lecturer Evolutionary genetics; phylogeography; molecularsystematics of invertebrates.

Leandro R Monteiro, Lecturer Morphometrics; morphological and functionalevolution of vertebrates.

Chris Murphy, Senior Teaching Fellow

Graham Scott, Senior Lecturer and Head of Department Animal behaviour;intertidal ecology; biological education.

Heather Sealy-Lewis, Lecturer Regulation of gene expression in Aspergillusnidulans with special interest in the ethanol utilisation regulon, and carboncatabolite repression.

Anne-Marie Seymour, Reader and Deputy Head of Department Biochemicaladaptation in heart failure; the relationship between heart metabolism andabnormal physiological function and dietary therapeutic treatments; biochemicalchanges in the ageing heart.

Cock van Oosterhout, Lecturer Conservation biology and genetics using guppiesand cichlids as model organisms. Leading field experiments in Trinidad and atropical marine field course for third-year students in Tobago.

Frank Voncken, Lecturer Host–pathogen interactions in trypanosomes.

All staff are active inresearch areas whichcomplement ourdegree courses.

Life sciences34

University of Hull International Fisheries InstituteHIFI, as it is known, was incorporated into the department in 2001. It is a specialistunit which undertakes a wide range of education, training, consultancy andresearch work both within the UK and internationally. HIFI staff are widelyexperienced in fisheries planning and management, fish marketing, fisherieseconomics, post-harvest operations, fishing-systems technology, fish biology andaquaculture.

Institute for Estuarine and Coastal StudiesThe IECS is a multidisciplinary research and consultancy organisation employingthe facilities and expertise in coastal science and management within theUniversity. The IECS covers a wide range of specialisms within the coastalenvironment, ranging from the biological and physical environments (topography,vegetation, ornithology, benthic and pelagic fauna) to coastal planning, marine lawand environmental impact assessment. Its Director, Professor Mike Elliott, is basedat the Hull Campus.

Centre for Environmental and Marine Sciences, Scarborough CampusRalph Bublitz, Lecturer Chemical ecology; crustacean biology.

Sue Hull, Lecturer The evolutionary ecology and population genetics of intertidalinvertebrates (especially ostracods and littorinid snails) and fucoid algae; thecomposition and dynamics of intertidal communities.

Alex Ibhadon, Senior Lecturer Environmental chemistry; heavy metal analysis;speciation of metals; effluent and waste treatment; advanced materials and polymerprocessing and application; process engineering and environmental applications.

Magnus Johnson, Lecturer The ecophysiology of crustaceans; fisheriesbioanthropology; mimicry; diving.

Will Mayes, Lecturer Monitoring and remediation of aquatic pollutants; hydrology;land-use change and flooding.

Gilbert Morrey, Lecturer Statistics; environmental issues.

Pete Rawson, Lecturer Biodiversity and biogeography of ammonites.

Sunil Shastri, Lecturer Ocean governance; marine and environmental law andpolicy.

Cath Waller, Lecturer Antarctic eclogy; intertidal ecological processes;nudibranchs; ecophysiology.

Phil Wheeler, Head of Centre Ecology and conservation of mammals and birds inBritish uplands and tropical rainforests.

As a visitor centre,describing itself asthe world’s onlysubmarium, TheDeep offers a uniqueinsight into theoceans – theirstructure as well astheir inhabitants.

Life sciences36

Strong linksThe Department of Biological Sciences has very close links with this world-classfacility. As a visitor centre, describing itself as the world’s only submarium, TheDeep offers a unique insight into the oceans – their structure as well as theirinhabitants. This £45 million investment has provided Hull with a major touristattraction (there have been more than two million visitors since its opening in 2002)and the department with a valuable teaching resource.

Research project opportunitiesSeveral undergraduate students are currently completing their final-year researchprojects at The Deep. These studies range from attempts to propagate and grow softand hard coral, for potential display and for coral reef restoration projects, toobservational analyses of the behaviour of sand tiger sharks, particularly their inter-and intra-specific interactions.

Shark conservation researchThe Deep is strongly supportive of research which assists in the conservation ofendangered marine species. This has led to the participation of student volunteersand final-year project students in shark health and behavioural studies. Dr ThomasBreithaupt’s students investigate whether sharks use acoustic communication byexposing sharks to noise and using hydrophones to detect their responses.

The Deep

Focus on research

CichlidsThe cichlid fishes of the African Rift Lakes are of great interest to biologists. Eachlake contains hundreds of species of fish which are found nowhere else in theworld. Dr Domino Joyce and her co-workers have been studying the fascinatingbehaviour and ecology of cichlid fishes from Lake Malawi, in an effort tounderstand how so many species evolved. In the clear shallow waters of the lake, itis possible to use scuba equipment to observe these fishes in their naturalenvironment; we have also brought some back to the purpose-built tropicalaquarium facility at the University for further study.

We use DNA sequence analysis to try to reconstruct the evolutionary history ofspecies and their ancestors. In one particular group of cichlid species that westudy, the males build large sand structures (bowers) under water, which they useto attract females. Different species build different-shaped bowers, and it may bethat female preference for different bowers has been the driving force behindincreasing variation in the population, eventually leading to the evolution of newspecies. It is possible to test this theory in large tanks and pools, by giving malesartificial bowers of various shapes to defend and letting the females choose whichmale and bower they prefer. We then use DNA paternity testing in the GenomeAnalysis Suite to identify which male has fathered a female’s offspring.Understanding mate choice and female preference in this way aids theidentification of species, which may look very similar in all other aspects butbower shape.

Understanding how species evolve will enable us to conserve the processes whichlead to the generation of biodiversity and to help design strategies for themanagement of the lakes in Africa, where the needs of an impoverished andrapidly expanding human population may be in conflict with the conservation ofthe unique fish species.

Studying for a degree at the University of Hull is a unique experience. We aimto provide you with an education that offers both depth and breadth ofknowledge. To meet these ends the University has developed an optional FreeElective Scheme. This scheme enables the majority of undergraduate studentsto take one module a year from outside their main course of study.

So, how does it work?Each year you take 120 credits’ worth of modules.

What sort of subjects can I take?You can take almost any free elective module from outside your main course ofstudy, usually at your home campus. You can even take a module from anotherfaculty. The catalogue of free electives might include

Hull• Human Physiology• Field Studies in Marine Biology• Chemical Methods and Ideas• Animal Behaviour

Scarborough• Outdoor Recreation and Education Management• Passport languages• Introduction to Poetry• Event Management

Or you might choose to take a module in one of a wide range of other subjects fromacross the University. Guidance about your choice of free electives will be availablein your department.

What are the main reasons for participating?• The scheme gives you the opportunity to study a subject without having to

commit yourself to taking further modules in that subject area.• By taking a free elective you are able to follow up your interests as part of your

degree.• With a broader education you may acquire extra skills that will help you when

you enter the employment market.

Admissions policyAdmissions information provided

in this pamphlet is intended as a

general guide and cannot cover all

possibilities. Entry requirements

are generally stated in terms of A

level grades and/or UCAS points,

but we encourage applications

from people with a wide range of

other qualifications and/or

experience. Some further details of

the various entry routes are

included in our general prospectus.

Please contact the Admissions

Service (see below) with any

specific queries about admissions.

DisclaimerThis publication is intended

principally as a guide for

prospective students. The matters

covered by it – academic and

otherwise – are subject to change

from time to time, both before and

after students are admitted, and

the information contained in it

does not form part of any contract.

While every reasonable precaution

was taken in the production of this

brochure, the University does not

accept liability for any

inaccuracies.

AddressFor general enquiries, please

write to

Admissions Service

University of Hull

Hull, HU6 7RX

T 01482 466100

F 01482 442290

E [email protected]

Free Elective Scheme

SEMESTER 1 SEMESTER 2

20 credits 20 credits

20 credits 20 credits

20 credits

20 credits

Here you take modules from yourmain course of study.

Here you have the option to take afree elective or another module fromyour main course of study.

Since arriving on British shores in2004, the harlequin ladybird has

become the fastest-spreading‘alien’ species on record.

Our researchers are investigatingthis invasion using the latest

molecular genetic techniques.

Change the way you think.

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w.hull.ac.uk

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