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    Rhododendron

    Rhododendron ferrugineum

    Scientific classification

    Kingdom: Plantae(unranked): Angiosperms

    (unranked): Eudicots

    (unranked): Asterids

    Order: Ericales

    Family: Ericaceae

    Subfamily: Ericoideae

    Genus: Rhododendron

    L. Sp. Pl. i 392 (1753)[1]

    Type species

    Rhododendron ferrugineum

    L.

    Subgenera[2]

    Azaleastrum

    Choniastrum

    Hymenanthes

    Rhododendron

    Therorhodion

    - Former subgenera -

    Candidastrum

    Mumeazalea

    Pentanthera

    From Wikipedia, the free encyclopedia

    Rhododendron(from Ancient Greek rhdon"rose" and

    dndron"tree")[3][4]is a genus of 1,024 species of woody plants in theheath family (Ericaceae), either evergreen or deciduous, and foundmainly in Asia. Most species have showy flowers. Azaleas make up twosubgenera ofRhododendron. They are distinguished from "true"rhododendrons by having only five anthers per flower.

    1 Description

    2 Taxonomy

    2.1 Early history

    2.2 Modern era

    2.3 Phylogenetic analyses

    2.4 Subdivision

    2.4.1 Subgenera

    2.4.2 Sections and subsections

    2.4.3 Species

    3 Distribution and habitat

    4 Ecology

    4.1 Invasive species

    4.2 Insects

    4.3 Diseases

    5 Cultivation

    5.1 Commercial growing

    5.2 Horticultural divisions

    5.3 Planting and care

    5.4 Hybrids

    6 Uses

    6.1 Pharmacology

    6.2 Toxicology

    7 Culture

    7.1 Symbolism

    7.2 Literature

    7.3 Culinary

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    Tsutsusi7.4 Labrador tea

    8 See also

    9 References

    10 Bibliography

    10.1 Books and book chapters

    10.2 Articles

    10.3 Subdivisions

    10.3.1 Azaleas

    10.3.2 Tsutsusi

    10.3.3 Vireya

    10.3.4 Separate genera

    11 External links

    11.1 Databases

    11.2 Flora11.3 Rhododendron societies

    11.4 Botanical gardens

    Rhododendronis a genus characterised by shrubs and small to (rarely) large trees, the smallest species growing

    to 10100 cm (3.939.4 in) tall, and the largest,R. protistum var. giganteum, reported to 30 m (98 ft) tall.[5][6]

    The leaves are spirally arranged; leaf size can range from 12 cm (0.390.79 in) to over 50 cm (20 in),exceptionally 100 cm (39 in) inR. sinogrande. They may be either evergreen or deciduous. In some species, theundersides of the leaves are covered with scales (lepidote) or hairs (indumentum). Some of the best knownspecies are noted for their many clusters of large flowers. There are alpine species with small flowers and smallleaves, and tropical species such as section Vireyathat often grow as epiphytes. Species in this genus may be

    part of the heath complex in oak-heath forests in eastern North America.[7][8]They have frequently beendivided based on the presence or absence of scales on the abaxial (lower) leaf surface (lepidote or elepidote).These scales, unique to subgenusRhododendron, are modified hairs consisting of a polygonal scale attached by

    a stalk.[2]

    Rhododendronare characterised by having inflorescences with scarious (dry) perulae, a chromosome number ofx=13, fruit that has a septicidal capsule, an ovary that is superior (or nearly so), stamens that have no

    appendages, and agglutinate (clumped) pollen.[9]

    TheRhododendrongenus is the largest of the genera in the Ericaceae family, with 1,024 species,[10]thoughestimates vary from 850-1000 depending on the authority used, (Fayaz 2012) and is morphologically diverse.

    Consequently the taxonomy has been historically complex.[9]

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    Rhododendron wardiivar.puralbum

    Rhododendron in Japan

    A garden with tall Rhododendrons in

    Lynnwood, Washington

    Rhododendron forest in Nepal

    Early history

    Although Rhododendrons had been known since the description ofRhododendron hirsutumby Charles de l'cluse (Clusius) in the sixteenthcentury, and were known to classical writers (Magor 1990), and referredto as Chamaerhododendron(low-growing rose tree), the genus was first

    formally described by Linnaeus in his Species Plantarum in 1753. [1][11]

    He listed five species underRhododendron(Rhododendron ferrugineum(type species),R. dauricum,R. hirsutum,R. chamaecistus(nowRhodothamnus chamaecistus(L.) Rchb.) andR. maximum). At that time

    he considered the then known six species ofAzalea[12]that he haddescribed earlier in 1735 in his Systema Naturae as a separate genus.[13][14]

    Linnaeus' six species of Azalea wereAzalea indica,A. pontica,A. lutea,A. viscosa,A. lapponicaandA. procumbens(now Kalmia procumbens),which he distinguished fromRhododendronby having five stamens, asopposed to ten. As new species of what are now consideredRhododendronwere discovered, if they seemed to differ significantlyfrom the type species they were assigned to separate genera. ForinstanceRhodoraforRhododendron canadense(Linnaeus 1763), Vireya

    (Blume 1826)[15]andHymenanthesforRhododendron metternichhii,now R. degronianum (1826). Meanwhile other botanists such as

    Salisbury (1796)[16]and Tate (1831)[17]began to question the distinctionbetweenAzaleaandRhododendron, and finally in 1836,Azaleawas

    incorporated intoRhododendron(Don 1834)[18]and the genus dividedinto eight sections. Of these Tsutsutsi(Tsutsusi), Pentanthera,

    Pogonanthum, PonticumandRhodoraare still used, the other sectionsbeingLepipherum,Booram, and Chamaecistus. This structure largelysurvived till recently (2004), following which the development ofmolecular phylogeny led to major re-examinations of traditional

    morphological classifications,[13][14]although other authors such as

    Candolle (1838), who described six sections,[19]used slightly differentnumeration.

    As more species became available in the nineteenth century a betterunderstanding of the characteristics necessary for the major divisions.Chief amongst these were Maximovicz'sRhododendreae Asiae Orientali

    (1870)[20]and Planchon. Maximovicz used flower bud position and its

    relationship with leaf buds to create eight Sections.[21]Bentham and

    Hooker (1876) used a similar scheme, but called the divisions Series. [22]

    It was not until 1893 that Koehne appreciated the significance of scalingand hence the separation of lepidote and elepidote species. The largenumber of species that were available by the early twentieth centuryprompted a new approach when Balfour introduced the concept of

    grouping species into series, in The Species of Rhododendron(1930), referred to as the Balfourian system.[23]

    That system continued up to modern times in Davidian's four volume The Rhododendron Species

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    Table 1.: Taxonomic changes within genusRhododendron

    (Goetsch 2005)

    Chamberlain (1996) Goetsch

    Genus Subgenus Section Species Subgenus Section

    Menziesa

    Azaleastrum

    SciadorhodionCandidastrum

    Tsutsusi

    TsutsusiMumeazaleum

    Pentanthera

    Viscidula

    PentantheraHymenanthes Pentanethra

    RhodoraR.canadense

    R.vaseyi Azaleastrum Sciadorhodion

    Azaleastrum Choniastrum Choniastrum

    Subsequent research has supported the revision by Goetsch, although has largely concentrated on furtherdefining the phylogeny within the subdivisions.(Craven 2008) In 2011 the two species ofDiplarchewere alsoadded toRhododendron, incertae sedis.(Craven 2011) Similar findings were reported independently the

    following year by Brown et al.[9]

    Subdivision

    This genus has been progressively subdivided into a hierarchy of subgenus, section, subsection, and species.

    Subgenera

    Terminology from the Sleumer (1949) system is frequently found in older literature, with five subgenera and isas follows;

    SubgenusLepidorrhodiumKoehne: Lepidotes. 3 sections

    SubgenusEurhododendronMaxim.: Elipidotes.

    Subgenus PseudanthodendronSleumer: Deciduous azaleas. 3 sections

    SubgenusAnthodendronRehder & Wilson: Evergreen azaleas. 3 sections

    SubgenusAzaleastrumPlanch.: 4 sections

    In the later traditional classification, attributed to Chamberlain (1996), and as used by horticulturalists and the

    American Rhododendron Society,[39]Rhododendronhas eight subgenera based on morphology, namely thepresence of scales (lepidote), deciduousness of leaves, and the floral and vegetative branching patterns, after

    Sleumer (1980).[9][2][27]These consist of four large and four small subgenera. The first two subgenera(RhododendronandHymenanthes) represent the species commonly considered as 'Rhododendrons'. The nexttwo smaller subgenera (Pentantheraand Tsutsusi) represent the 'Azaleas'. The remaining four subgenera contain

    very few species.[40]The largest of these is subgenusRhododendron, containing nearly half of all knownspecies and all of the lepidote species.

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    SubgenusRhododendronL.: Small leaf or lepidotes (scales on the underside of the leaves). 3 sections,

    462 species, type species:Rhododendron ferrugineum.

    SubgenusHymenanthes(Blume) K.Koch: Large leaf or elepidotes (without scales). 1 section, 224

    species, typeRhododendron degronianum.

    Subgenus Pentanthera(G. Don) Pojarkova: Deciduous azaleas. 4 sections, 23 species, type

    Rhododendron luteum.

    Subgenus Tsutsusi(Sweet) Pojarkova: Evergreen azaleas. 2 sections, 80 species, typeRhododendron

    indicum.

    SubgenusAzaleastrumPlanch.: 2 sections, 16 species, typeRhododendron ovatum.

    Subgenus CandidastrumFranch. : 1 species,Rhododendron albiflorum.

    SubgenusMumeazalea(Sleumer) W.R. Philipson & M.N. Philipson: 1 species,Rhododendron

    semibarbatum.

    Subgenus Therorhodion(Maxim.) A. Gray: 2 species (Rhododendron camtschaticum,Rhododendron

    redowskianun).

    For a comparison of the Sleumer and Chamberlain systems, see Goetsch et al. (2005) Table 1.[2]

    This division was based on a number of what were thought to be key morphological characteristics. Theseincluded the position of the inflorescence buds (terminal or lateral), whether lepidote or elepidote,deciduousness of leaves, and whether new foliage was derived from axils from previous year's shoots or thelowest scaly leaves (Table 2.).

    Table 2.: Morphological classification ofRhododendron

    (Chamberlain 1996)[35]

    Inflorescence buds Leaf scales Leaf shoots Leaves Subgenus Section

    Terminal

    Present Rhododendron

    Absent

    Previous year

    Evergreen Hymenanthes

    Deciduous Pentanthera

    Pentanthera

    Rhodora

    Viscidula

    Lowest leaves Pentanthera Sciadorhodion

    Tsutsusi

    Lateral

    Evergreen Azaleastrum

    Deciduous

    Candidastrum

    Mumeazalea

    Therorhodion

    Following the cladistic analysis of Goetsch et al.(2005)[2]this scheme was simplified, based on the discoveryof three major clades (A,B,C) as follows.

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    Clade A

    SubgenusRhododendronL.: Small leaf or lepidotes (scales on the underside of the leaves). 3 sections,

    about 400 species, type species:Rhododendron ferrugineum.

    Subgenus ChoniastrumFranch. : 11 species

    Clade B

    SubgenusHymenanthes(Blume) K.Koch: Large leaf or elepidotes (without scales), including deciduous

    azaleas. 2 sections, about 140-225 species, typeRhododendron degronianum.

    Clade C

    SubgenusAzaleastrumPlanch.: Evergreen azaleas. 3 sections, about 120 species, type Rhododendron

    ovatum.

    Sister taxon

    Subgenus Therorhodion(Maxim.) A. Gray: 2 species (Rhododendron camtschaticum,Rhododendron

    redowskianun).

    Sections and subsections

    The larger subgenera are further subdivided into sections and subsections[40]Some subgenera contain only asingle section, and some sections only a single subsection. Shown here is the traditional classification, withspecies number after Chamberlain (1996), but this scheme is undergoing constant revision. Revisions by

    Goetsch et al.(2005)[2]

    and by Craven et al. (2008)[41]

    shown in (parenthetical italics). Older ranks such asSeries (groups of species) are no longer used but may be found in the literature, but the American

    Rhododendron Society still uses a similar device, called Alliances[39]

    SubgenusRhododendronL. (3 sections, 462 species: increased to five sections in 2008)

    (Discovereya (Sleumer) Argent , raised from Vireya)

    PogonathumAitch. & Hemsl. (13 species; Himalaya and adjacent mountains)

    (Pseudovireya (C.B.Clarke) Argent, raised from Vireya)

    RhododendronL. (149 species in 25 subsections; temperate to subarctic Northern Hemisphere)

    Vireya(Blume) Copel.f. (300 species in 2 subsections; tropical southeast Asia, Australasia. At one

    time considered separate subgenus[42])

    SubgenusHymenanthes(Blume) K.Koch (1 section, 224 species) (Increased to two sections)

    PonticumG. Don (24 subsections)

    (Pentanthera (G. Don) Pojarkova (2 subsections)new section, moved from subgenus Pentanthera)

    Subgenus Pentanthera(G. Don) Pojarkova (4 sections, 23 species) (Discontinued)

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    Pentanthera(G. Don) Pojarkova (2 subsections) (Moved to subgenus Hymenanthes)

    Rhodora(L.) G. Don (2 species; Rhododendron canadense, Rhododendron vaseyi) (Discontinued,

    redistributed)

    SciadorhodionRehder & Wilson (4 species) (Moved to subgenus Azaleastrum)

    ViscidulaMatsum. & Nakai (1 species; Rhododendron nipponicum) (Discontinued, added to

    section Tsutsusi, subgenus Azaleastrum)

    Subgenus Tsutsusi(Sweet) Pojarkova (2 sections, 80 species) (Discontinued, reduced to section and

    moved to subgenus Azaleastrum)

    BrachycalyxSweet (3 alliances, 15 species)

    Tsutsusi(Sweet) Pojarkova (65 species)

    SubgenusAzaleastrumPlanch. (2 sections, 16 species) (Increased to three sections)

    AzaleastrumPlanch. (5 species)

    (ChoniastrumFranch. (11 species) (Raised to subgenus))

    (Sciadorhodion Rehder & Wilson (4 species) (Moved from subgenus Pentanthera))

    (Tsutsusi (Sweet) Pojarkova (reduced from subgenus))

    Subgenus CandidastrumFranch. (1 species:Rhododendron albiflorum) (Discontinued, moved to section

    Sciadorhodion, subgenus Azaleastrum)

    SubgenusMumeazalea(Sleumer) W.R. Philipson & M.N. Philipson (1 species:Rhododendron

    semibarbatum) (Discontinued, moved to section Tsutsusi, subgenus Azaleastrum)

    Subgenus TherorhodionA. Gray (2 species)

    (* Subgenus Choniastrum Franch. (11 species))

    Species

    Species of the genusRhododendronare widely distributed between latitudes 80N and 20S and are consideredAlpine native plants from North America to Europe, Russia, and Asia, and from Greenland to Queensland,

    Australia and the Solomon Islands.[9]The centres of diversification are in the Himalayas and Malaysia,[37]withthe greatest species diversity in the Sino-Himalayan region, Southwest China and northern Burma, fromUttarakhand, Nepal and Sikkim to northwestern Yunnan and western Sichuan and southeastern Tibet, and with

    other significant areas of diversity in the mountains of Korea, Japan and Taiwan. More than 90% ofRhododendronsensuChamberlain belong to the Asian subgeneraRhododendron,Hymenanthesand sectionTsutsusi. Of the first two of these, the species are predominantly found in the area of the Himalayas and

    Southwest China (Sino-Himalayan Region).[2]

    The 300 Tropical species within the Vereyasection of subgenusRhododendronoccupy the Malay archipelagofrom their presumed Southeast Asian origin to Northern Australia, with 55 known species in Borneo and 164 inNew Guinea. The species in New Guinea are native to subalpine moist grasslands at around 3,000 metres above

    sea level in the Central Highlands.[42]SubgeneraRhododendronandHymenanthes, together with sectionPentanetheraof subgenus Pentanetheraare also represented to a lesser degree in the Mountainous areas of

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    Rhododendron fallacinum

    photographed in situon

    Mount Kinabalu, Borneo

    Rhododendron-Park Bremen,Germany

    North America and Western Eurasia. Subgenus Tsutsusiis found in the maritimeregions of East Asia (Japan, Korea, Taiwan, East China), but not in North

    America or Eurasia.[2][21]

    Invasive species

    Some species (e.g.Rhododendron ponticumin Ireland [43]and the UnitedKingdom) are invasive as introduced plants, spreading in woodland areasreplacing the natural understory.R. ponticumis difficult to eradicate, as its rootscan make new shoots.

    Insects

    A number of insects either target rhododendrons or will opportunistically attackthem. Rhododendron borers and various weevils are major pests of

    rhododendrons, and many caterpillars will preferentially devour them.

    Rhododendronspecies are used as food plants by the larvae of some members of the order Lepidoptera(butterflies and moths) (See List of Lepidoptera that feed on rhododendrons).

    Diseases

    Major diseases include Phytophthoraroot rot, stem and twig fungal dieback; Ohio State University Extension

    provides information on maintaining health of rhododendrons.[44]Rhododendrons can easily be suffocated byother plants.

    Both species and hybrid rhododendrons (including azaleas) are usedextensively as ornamental plants in landscaping in many parts of theworld, including both temperate and subtemperate regions,(Craven2008) while many species and cultivars are grown commercially for thenursery trade. Rhododendrons are often valued in landscaping for their

    structure, size, flowers, and the fact that many of them are evergreen. [45]

    Azaleas are frequently used around foundations and occasionally as

    hedges, and many larger-leafed rhododendrons lend themselves well tomore informal plantings and woodland gardens, or as specimen plants.In some areas, larger rhododendrons can be pruned to encourage moretree-like form, with some species such asR. arboreumandR. falconerieventually growing to 1015 m or more

    tall.[45]

    Commercial growing

    Rhododendrons are grown commercially in many areas for sale, and are occasionally collected in the wild, apractice now rare in most areas. Larger commercial growers often ship long distances; in the United States,

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    most of them are located on the west coast (Oregon, Washington state and California). Large-scale commercialgrowing often selects for different characteristics than hobbyist growers might want, such as resistance to rootrot when overwatered, ability to be forced into budding early, ease of rooting or other propagation, and

    saleability.[46]In the Indian state of Himachal Pradesh, rhododendron flowers have been used for some time tomake popular fruit and flower wines. The industry is promoted by the state government with tax benefits,

    looking to promote this industry as a full-fledged subclass of its economy.[47]

    Horticultural divisions

    Horticulturally, rhododendrons may be divided into the following groups:-[48]

    Evergreen rhododendrons: the main default category

    Vireya (Malesian) rhododendrons: epiphytic tender shrubs[49]

    Azaleas (section of generally small-sized, small-leaved and small-flowered shrubs):

    Deciduous hybrid azaleas:[50]

    Ghent (Gandavense) hybrids - Belgian raised[51]

    Knap Hill-Exbury hybrids - English raised[52]

    Mollis hybrids - Dutch & Belgian raised[53]

    New Zealand Ilam hybrids - derived from Knap Hill/Exbury hybrids

    Occidentale hybrids - English raised

    Rustica hybrids - sweet-scented, double-flowered

    Evergreen hybrid azaleas:

    Gable hybrids - raised by Joseph B. Gable in Pennsylvania, USA [54]

    Glenn Dale hybrids - USA raised complex hybrids

    Indian (Indica) hybrids - mostly of Belgian origin

    Kaempferi hybrids - Dutch raised

    Kurume hybrids - Japanese raised

    Kyushu hybrids - very hardy Japanese azaleas (to -30 C)

    Oldhamii hybrids - dwarf hybrids raised at Exbury, England

    Satsuki hybrids - Japanese raised, originally for bonsai

    Shammarello hybrids - raised in Northern Ohio, USA[55]

    Vuyk (Vuykiana) hybrids - raised in the Netherlands

    [56]

    Azaleodendrons - semi-evergreen hybrids between deciduous azaleas and rhododendrons

    Planting and care

    Like other ericaceous plants, most rhododendrons prefer acid soils with a pH of roughly 4.5-5.5; some tropicalVireyas and a few other rhododendron species grow as epiphytes and require a planting mix similar to orchids.Rhododendrons have fibrous roots and prefer well-drained soils high in organic material. In areas with poorlydrained or alkaline soils, rhododendrons are often grown in raised beds using media such as composted pine

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    Nova Zembla Rhododendrons

    growing in a nursery in New Jersey.

    bark.[57]Mulching and careful watering are important, especially beforethe plant is established.

    A new calcium-tolerant stock of rhododendrons (trademarked as'Inkarho') has been exhibited at the RHS Chelsea Flower Show inLondon (2011). Individual hybrids of rhododendrons have been graftedon to a rootstock on a single rhododendron plant that was found growingin a chalk quarry. The rootstock is able to grow in calcium-rich soil up to

    a pH of 7.5.[58][59]

    Hybrids

    Rhododendrons are extensively hybridized in cultivation, and natural hybrids often occur in areas where speciesranges overlap. There are over 28,000 cultivars of Rhododendron in the International Rhododendron Registry(http://www.rhs.org.uk/Plants/RHS-Publications/Plant-registers/Rhododendron) held by the Royal HorticulturalSociety. Most have been bred for their flowers, but a few are of garden interest because of ornamental leaves

    and some for ornamental bark or stems. Some hybrids have fragrant flowers[60]such as the Loderi hybrids,

    created by crossingR. fortuneiandR. griffithianum.[61]

    Other examples include the PJM hybrids, formed froma cross betweenRhododendron carolinianumandRhododendron dauricum, and named after Peter J. Mezitt of

    Weston Nurseries, Massachusetts.[62]

    Pharmacology

    Rhododendronspecies have long been used in traditional medicine.[63]Animal studies and in vitroresearch hasidentified possible anti-inflammatory and hepatoprotective activities which may be due to the antioxidant

    effects of flavonoids or other phenolic compounds and saponins the plant contains. [64][65][66]Xiong et al.have

    found that the root of the plant is able to reduce the activity of NF-B in rats.[67]

    Toxicology

    Some species of rhododendron are poisonous to grazing animals because of a toxin called grayanotoxin in theirpollen and nectar. People have been known to become ill from eating honey made by bees feeding onrhododendron and azalea flowers. Xenophon described the odd behaviour of Greek soldiers after havingconsumed honey in a village surrounded byRhododendron ponticumduring the march of the Ten Thousand in401 BC. Pompey's soldiers reportedly suffered lethal casualties following the consumption of honey made fromRhododendrondeliberately left behind by Pontic forces in 67 BC during the Third Mithridatic War. Later, it was

    recognized that honey resulting from these plants has a slightly hallucinogenic and laxative effect. [68]Thesuspect rhododendrons areRhododendron ponticumandRhododendron luteum(formerlyAzalea pontica), bothfound in northern Asia Minor.A brief documented video of this occurring in the modern day involves a group ofmen in Nepal foraging for this affected honey can be found here: http://eupterrafoundation.com/hallucinogenic-

    honey. Eleven similar cases have been documented in Istanbul, Turkey during the 1980s.[69]Rhododendron isextremely toxic to horses, with some animals dying within a few hours of ingesting the plant, although mosthorses tend to avoid it if they have access to good forage. The effects of R. ponticumwas mentioned in the 2009

    film Sherlock Holmes as a proposed way to arrange a fake execution. [70]It was also mentioned in the third

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    episode of Season 2 of BBC's Sherlock (TV series), and has been speculated to have been a part of Sherlock'sfake death scheme.

    Symbolism

    Rhododendron arboreum(lali guransh) is the national flower of Nepal.R. ponticumis the state flower ofIndian-administered Kashmir and Pakistan-controlled Kashmir.Rhododendron niveumis the state tree ofSikkim in India. Rhododendron is also the state tree of the state of Uttarakhand, India. Pink Rhododendron(Rhododendron campanulatum) is the State Flower of Himachal Pradesh, India.

    Rhododendron maximum, the most widespread rhododendron of the Appalachian Mountains, is the state flowerof West Virginia, and is in the Flag of West Virginia. Rhododendron macrophyllum, a widespread rhododendronof the Pacific Northwest, is the state flower of Washington.

    Literature

    In Joyce's Ulysses, rhododendrons play an important role in Leopold and Molly's early courtship: Mollyremembers them in her soliloquy - "the sun shines for you he said the day we were lying among therhododendrons on Howth head in the grey tweed suit and his straw hat the day I got him to propose to me".Jasper Fforde a British author, also uses rhododendron as a motif throughout many of his published books. See

    Thursday Next series,[71]and Shades of Grey.[72]Amongst the Zomi tribes in India and Myanmar,"Rhododendrons" called "Ngeisok" is used in a poetic manner to signify a lady.

    Culinary

    The rhododendron is the national flower of Nepal, where the flower is considered edible and enjoyed for itssour taste. The pickled flower can last for months and the flower juice is also marketed. The flower, fresh ordried, is added to fish curry in the belief that it will soften the bones. The juice of rhododendron flower is usedto make a squash called burans (named after the flower) in the hilly regions of Uttarakhand. It is admired for itsdistinctive flavor and color.

    Labrador tea

    Labrador tea is an herbal tea (not a true tea) made from three closely related species:

    Rhododendron tomentosum(Northern Labrador tea, previouslyLedum palustre),

    Rhododendron groenlandicum, (Bog Labrador tea, previouslyLedum groenlandicumorLedum latifolium)and

    Rhododendron neoglandulosum, (Western Labrador tea, or trapper's tea, previouslyLedum glandulosum).

    List of Award of Garden Merit rhododendrons

    List of Rhododendron diseases

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    14/24

    List of Rhododendron species

    ^abLinnaeus, C. (1753).

    "Rhododendron"

    (http://www.biodiversitylibrary.org/item/13829#page/404/mode

    /1up). Species Plantarum.

    Stockholm: Laurentii Salvii. p. i

    392. Retrieved June 15, 2014.

    1.

    ^abcdefghijklGoetsch,

    Loretta A.; Eckert, Andrew J.;

    Hall, Benjamin D. (July

    September 2005). "The

    molecular systematics ofRhododendron(Ericaceae): a

    phylogeny based uponRPB2

    gene sequences"

    (http://www.ingentaconnect.com

    /content/aspt/sb/2005/00000030

    /00000003/art00012). Systematic

    Botany30(3): 616626.

    doi:10.1600/0363644054782170

    (http://dx.doi.org

    /10.1600%2F036364405478217

    0).

    2.

    ^Harper, Douglas.

    "rhododendron"

    (http://www.etymonline.com

    /index.php?term=rhododendron)

    . Online Etymology Dictionary.

    3.

    ^

    (http://www.perseus.tufts.edu

    /hopper/text?doc=Perseus:text:1999.04.

    0057:entry=r(o/don),

    (http://www.perseus.tufts.edu

    /hopper

    /text?doc=Perseus:text:1999.04.

    0057:entry=de/ndron). Liddell,

    Henry George; Scott, Robert;A

    GreekEnglish Lexiconat the

    Perseus Project

    4.

    ^[1]

    (http://www.globaltrees.org

    /tp_bigtree.htm), Big Tree

    Rhododendron

    5.

    ^[2]

    (http://www.rhododendron-

    azalee.fr/classification_gb/fiches

    /protistum_giganteum.shtml)

    Rhododendron protistum var.giganteum

    6.

    ^The Natural Communities of

    Virginia Classification of

    Ecological Community Groups

    (Version 2.5), Virginia

    Department of Conservation and

    Recreation, 2012

    (http://www.dcr.virginia.gov

    /natural_heritage

    /natural_communities

    /ncintro.shtml)

    7.

    ^Schafale, M. P. and A. S.

    Weakley. 1990. Classification of

    the natural communities ofNorth Carolina: third

    approximation. North Carolina

    Natural Heritage Program, North

    Carolina Division of Parks and

    Recreation.

    8.

    ^abcdefGillian K. Brown,

    Lyn A. Craven, Frank Udovicic

    and Pauline Y. Ladiges.

    Phylogenetic relationships ofRhododendron section Vireya

    (Ericaceae) inferred from the

    ITS nrDNA region. Australian

    Systematic Botany 19, 329342.

    August 2006. 10.1071/SB05019

    (http://www.researchgate.net

    /publication

    /235601539_Phylogenetic_relati

    onships_of_Rhododendron_secton_Vireya_(Ericaceae)_inferred

    _from_the_ITS_nrDNA_region

    /file/72e7e51e347034a1c1.pdf)

    9.

    ^Sorting out the Genus

    Rhododendron: A Taxonomic

    Overview of this Large Complex

    of Species. University of

    California Botanical Garden at

    Berkeley, March 27 2014.

    (http://events.berkeley.edu

    /index.php/calendar

    /sn/bot.html?event_ID=73328)

    10.

    ^Linnaeus Gen. Pl. ed. 5, 185.

    1754.

    11.

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    15/24

    ^Linnaeus, C. (1753). "Azalea"

    (http://www.biodiversitylibrary.o

    rg/item/13829#page/162/mode

    /1up). Species Plantarum.

    Stockholm: Laurentii Salvii. p. i

    150. Retrieved June 15, 2014.

    12.

    ^a

    b

    American RhododendronSociety (Azalea Chapter): What

    is an Azalea?

    (http://azaleachapter.com/plant-

    info/plants/what-is-an-azalea)

    13.

    ^abcHerbert A. Spady.

    GROPING FOR GROUPING:

    Rhododendron Taxonomy.

    Quarterly Bulletin of the

    (http://scholar.lib.vt.edu/ejournals/JARS/v35n4/v35n4-

    spady.htm) ARS Volume 35,

    Number 4, Fall 1981

    14.

    ^Chris Callard. The History of

    Vireya Rhododendron Culture.

    Vireya Rhododendrons. 1998

    onwards (http://www.vireya.net

    /history.htm)

    15.

    ^Salisbury RA. ProdromusStirpium in horto ad Chapel

    Allerton, 1796. p. 286

    (http://www.biodiversitylibrary.o

    rg/item/9522#page

    /293/mode/1up)

    16.

    ^James Charles Tate,

    Nurseryman at Sloane Square,

    founder of the Nursery and

    Botanic Garden and importer of

    Rhododendrons. British History

    Online: Farm Gardening and

    Market Gardening

    (http://www.british-history.ac.uk

    /report.aspx?compid=28707)

    17.

    ^Don G. General History of

    Dichlamydious Plants. 1834 iii

    843

    (http://www.biodiversitylibrary.o

    rg/item/9905#page

    /851/mode/1up)

    18.

    ^A. P. de Candolle et al. (1838)."Rhododendron"

    (http://biodiversitylibrary.org

    /item/7156#page/724/mode

    /1upurl=). Prodromus systemati

    naturalis regni vegetabilis sive

    enumeratio contracta ordinum,

    generum specierumque

    plantarum huc usque

    cognitarum, juxta methodinaturalis normas digesta. Paris:

    Treuttel et Wrtz. p. 719.

    19.

    ^Maximowicz, CJ.

    Rhododendreae Asiae Orientalis

    St Ptersbourg, 1870

    (http://books.google.ca/books

    /about

    /Rhododendreae_Asiae_orientali

    s.html?id=s7MaAAAAYAAJ&redir_esc=y)

    20.

    ^abJin Xiao-Feng, Ding

    Bing-Yang, Zhang Yue-Jiao, and

    Hong De-Yuan. A Taxonomic

    Revision OfRhododendron

    subg. Tsutsusisect.Brachycalyx

    (Ericaceae). Annals of the

    Missouri Botanical Garden

    97(2):163-190. 2010 doi:

    http://dx.doi.org/10.3417

    /2007139

    (http://www.bioone.org/doi/abs

    /10.3417

    /2007139?journalCode=mobt)

    21.

    ^Bentham, G.; J.D. Hooker;

    Hooker, JD (1876). Genera

    plantarum ad exemplaria

    imprimis in herbariis

    kewensibus servata definita. Vol.

    2 Part II

    (http://www.biodiversitylibrary.org/item/14687#page/74/mode

    /1up). London: L Reeve & Co.

    Retrieved 10 July 2014.

    22.

    ^Stevenson JB (ed) The Species

    of Rhododendron. The

    Rhododendron Society,

    Edinburgh 1930

    23.

    ^Davidian, H. H. The

    Rhododendron Species. In fourvolumes from 1982-1995.

    Timber Press. 1.Lepidotes, 2.

    Elepidotes. Arboreum-Lacteum,

    3.Elepidotes Continued,

    Neriiflorum-Thomsonii,

    Azaleastrum and

    Camtschaticum, 4.Azaleas

    (http://books.google.ca/books

    /about/The_Rhododendron_Species_A

    zaleas.html?id=CBQmAQAAM

    AAJ&redir_esc=y)

    24.

    ^Sleumer, H. 1949. Ein System

    der Gattung Rhododendron L.

    Bot. Jahrb. Syst. 74: 511553

    25.

    ^Sleumer H (1966) Ericaceae 1

    Rhododendron. In Flora

    Malesiana. (Ed. CGGJ van

    Steenis) pp. 469656. (Wolters-

    Noordhoff Publishing:

    Groningen, The Netherlands

    (http://archive.org/stream

    /floramalesiana64stee

    /floramalesiana64stee_djvu.txt)

    26.

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    16/24

    ^abSleumer H (1980) Past and

    present taxonomic systems of

    Rhododendronbased on

    macromorphological characters.

    In Contributions toward a

    classification of rhododendron:

    proceedings internationalrhododendron conference. (Eds

    JL Luteyn, ME OBrien) pp.

    1926. (The New York

    Botanical Garden: New York)

    27.

    ^J. Cullen. A revision of

    Rhododendron I. Subgenus

    Rhododendron sections

    Rhododendron and

    Pogonanthum, RBGE, Vol. 39,No. 1 (1980).

    28.

    ^Philipson, W. R. and M. N.

    Philipson. 1986. A revision of

    Rhododendron III. Subgenera

    Azaleastrum, Mumeazalea,

    Candidastrum and

    Therorhodion. Notes Roy. Bot.

    Gard. Edinburgh 44: 1-23.

    29.

    ^SPETHMANN, W (1987): Anew infrageneric classification

    and phylogenetic trends in the

    genusRhododendron

    (Ericaceae), Plant Systematics

    and Evolution 157: 9-31

    (http://link.springer.com/article

    /10.1007%2FBF00939178)

    30.

    ^Stevens PF. A classification of

    the Ericaceae: subfamilies and

    tribes. Botanical Journal of the

    Linnean Society.Volume 64,

    Issue 1, pages 153, January

    1971. DOI:

    10.1111/j.1095-8339.1971.tb02133.x(http://onlinelibrary.wiley.com

    /doi/10.1111

    /j.1095-8339.1971.tb02133.x

    /abstract)

    31.

    ^Chamberlain, D.F. & Rae, S.J.

    (1990). A revision of

    Rhododendron. IV. Subgenus

    Tustsusi. Edinburgh Journal of

    Botany. 47(2) 89-200

    32.

    ^Kron, K. A. & W. S. Judd.

    1990. Phylogenetic relationships

    within the Rhodoreae

    (Ericaceae) with specific

    comments on the placement of

    Ledum. Systematic Botany 15:

    57-68. (http://www.jstor.org

    /discover/10.2307

    /2419016?uid=3739432&uid=2129&uid=2&uid=70&

    uid=3737720&uid=4&

    sid=21103902140831)

    33.

    ^Judd, W. S. & K. A. Kron.

    1995. A revision of

    Rhododendron. VI. Subgenus

    Pentanthera(sections

    Sciadorhodion,Rhodora, and

    Viscidula). Edinburgh J. Bot.

    52(1): 1-54.

    34.

    ^abChamberlain DF, Hyam R,

    Argent G, Fairweather G, Walter

    KS (1996). The genus

    Rhododendron: its classification

    and synonymy. (Royal Botanic

    Garden Edinburgh: Edinburgh)

    ISBN 1 872291 66 X(http://www.rbge.org.uk/about-

    us/publications/publications-

    catalogue/botanical-publications

    /rhododendron-publications)

    35.

    ^James Cullen. Hardy

    Rhododendron Species: A Guide

    To Identification. Timber Press,

    2005. ISBN 0881927236

    (http://books.google.ca/books/about

    /Hardy_Rhododendron_Species

    html?id=jxY4dVOm1VUC&

    redir_esc=y)

    36.

    ^abGAO Lian-Ming, LI

    De-Zhu, ZHANG Chang-Qin,

    YANG Jun-Bo. Infrageneric and

    Sectional Relationships in the

    Genus Rhododendron(Ericaceae) Inferred from ITS

    Sequence Data. Am Botanica

    Sinica 2002, 44(11) 1351-1356

    (http://www.jipb.net/pubsoft

    /content/2/2316

    /X010407(PS2).pdf)

    37.

    ^Loretta Goetsch, Andrew

    Eckert and Benjamin Hall.

    Classification of genus

    Rhododendron. 2005 Annual

    ARS Convention

    (https://web.archive.org

    /web/20130227185832/http:

    //www.flounder.ca/FraserSouth

    /Goetsch-Eckert-Hall.asp)

    38.

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    17/24

    ^ab"Genus Rhododendron

    Taxonomic Tree"

    (http://www.rhododendron.org

    /taxonomictree.asp).American

    Rhododendron Society.

    Retrieved 18 June 2014.

    39.

    ^a

    b

    Irving, E.; R. Hebda(1993). "Concerning the Origin

    and Distribution of

    Rhododendrons"

    (http://www.rhododendron.org

    /v47n3p139.htm).Journal of the

    American Rhododendron

    Society. American

    Rhododendron Society.

    Retrieved 4 November 2012.

    40.

    ^Craven, L.A.; Goetsch, L.A.;

    Hall, B.D.; Brown, G.K. (2008).

    "Classification of the Vireya

    group ofRhododendron

    (Ericaceae)".Blumea -

    Biodiversity, Evolution and

    Biogeography of Plants53(2):

    435.

    doi:10.3767/000651908X608070 (http://dx.doi.org

    /10.3767%2F000651908X60807

    0).

    41.

    ^abArgent, G.Rhododendrons

    of subgenus Vireya. 2006. Royal

    Horticultural Society. ISBN

    1-902896-61-0

    42.

    ^Dramatic rescue of couple

    trapped in rhododendron forest,

    Irish Central, 18th of July 2014

    (http://www.irishcentral.com

    /news/Dramatic-rescue-

    of-couple-trapped-

    in-rhododendron-forest.html?utm_source=outbrai

    n&utm_medium=content&

    utm_campaign=paid)

    43.

    ^"AG.ohio-state.edu"

    (http://www.ag.ohio-state.edu

    /~ohioline/hyg-fact

    /3000/3043.html). AG.ohio-

    state.edu. Retrieved 2013-02-26.

    44.

    ^a

    b

    Huxley, A., ed. (1992).New RHS Dictionary of

    Gardening. Macmillan

    45.

    ^Peter A. Cox (1993). The

    Cultivation of Rhododendrons.

    B. T. Batsford, London ISBN

    0-7134-5630-2 (pp80-1)

    46.

    ^The-fuga-experience.com

    (http://the-fuqua-experience.com

    /2010/02/17/waterfall-rhododendron-wine-himalayas-

    india/)

    47.

    ^RHS A-Z encyclopedia of

    garden plants. United Kingdom:

    Dorling Kindersley. 2008.

    p. 1136. ISBN 1405332964.

    48.

    ^"Vireya Rhododendrons -

    Welcome"

    (http://www.vireya.net/).

    Vireya.net. Retrieved

    2013-02-26.

    49.

    ^"Deciduous Azaleas"

    (http://www.rhodyman.net

    /rhodyde.html). Rhodyman.net.

    Retrieved 2013-02-26.

    50.

    ^"JARS v38n3 - Rescuing the

    Ghent and Rustica Flore Pleno

    Azaleas"

    (http://scholar.lib.vt.edu

    /ejournals/JARS/v38n3/v38n3-

    skinner.htm). Scholar.lib.vt.edu.

    2012-04-03. Retrieved2013-02-26.

    51.

    ^"Don Hyatt's Knap Hill

    Azalea Page"

    (http://www.tjhsst.edu/~dhyatt

    /knaphill.html). Tjhsst.edu.

    Retrieved 2013-02-26.

    52.

    ^"Quarterly Bulletin of the

    ARS - Vol 14 No 1, Living,

    Mollis Azaleas"(http://scholar.lib.vt.edu

    /ejournals/JARS/v14n1/v14n1-

    living.htm). Scholar.lib.vt.edu.

    1960-01-01. Retrieved

    2013-02-26.

    53.

    ^"The Rhododendron Legacy

    of Joe Gable"

    (http://www.tjhsst.edu/~dhyatt

    /gable.html). Tjhsst.edu.Retrieved 2013-02-26.

    54.

    ^"Quarterly Bulletin of the

    ARS - Vol 9 No 4, Baldsiefen,

    Shammarello's Wonderland"

    (http://scholar.lib.vt.edu

    /ejournals/JARS/v9n4/v9n4-

    baldsiefen.htm).

    Scholar.lib.vt.edu. 1955-10-04.

    Retrieved 2013-02-26.

    55.

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    18/24

    ^"Quarterly Bulletin of the

    ARS - Vol 33 No 1, Nosal, The

    Vuykiana Azaleas"

    (http://scholar.lib.vt.edu

    /ejournals/JARS/v33n1/v33n1-

    nosal.htm). Scholar.lib.vt.edu.

    1979-01-01. Retrieved2013-02-26.

    56.

    ^"Soil information for planting

    rhododendrons"

    (http://www.rhododendron.org

    /soil.htm). Rhododendron.org.

    Retrieved 2013-02-26.

    57.

    ^"Buy Lime Tolerant Inkarho

    Rhododendrons Online"

    (http://www.rhododendrons.co.uk/Category/41/lime-tolerant-

    rhododendrons.aspx).

    Rhododendrons.co.uk. Retrieved

    2013-02-26.

    58.

    ^Gardening. "The new

    lime-tolerant rhododendrons"

    (http://www.telegraph.co.uk

    /gardening/6581138/The-new-

    lime-tolerant-rhododendrons.html). Telegraph.

    Retrieved 2013-02-26.

    59.

    ^http://plants.wildeel.com

    /rhodofrag.html

    60.

    ^Davidian, H.H.,

    "Rhododendron Species, Vol. II:

    Elepidotes, Part 1 Arboreum-

    Lacteum" Timber Press, Inc.,

    Portland, Oregon, plate 44 & 45

    61.

    ^University of Connecticut:

    Rhododendron 'PJM' Hybrids

    (http://www.hort.uconn.edu

    /plants/r/rhopjm/rhopjm1.html)

    62.

    ^Popescu R, Kopp B. The

    genus Rhododendron: an

    ethnopharmacological and

    toxicological review. J

    Ethnopharmacol. 2013 May

    2;147(1):42-62. doi:

    10.1016/j.jep.2013.02.022. Epub2013 Feb 27.

    (http://www.ncbi.nlm.nih.gov

    /pubmed/23454683)

    63.

    ^Erdemoglu, Nurgun; Akkol,

    EK; Yesilada, E; Cali, I (2008).

    "Bioassay-guided isolation of

    anti-inflammatory and

    antinociceptive principles from a

    folk remedy,RhododendronponticumL. leaves".Journal of

    Ethnopharmacology119(1, 2):

    1728.

    doi:10.1016/j.jep.2008.06.021

    (http://dx.doi.org

    /10.1016%2Fj.jep.2008.06.021).

    PMID 18638535

    (https://www.ncbi.nlm.nih.gov

    /pubmed/18638535).

    64.

    ^Prakash, T. et al. (2008).

    "Hepatoprotective activity of

    leaves ofRhododendron

    arboreumin CCl4induced

    hepatotoxicity in rats"

    (http://www.academicjournals.or

    g/jmpr/abstracts/abstracts

    /abstrats2008

    /Nov/Prakash%20et%20al.htm).

    Journal of Medicinal Plants

    Research2(11): 31520.

    65.

    ^Agarwal, S.S.; Sharma

    Kalpana (1988). "Anti-

    inflammatory activity of flowers

    ofRhododendron arboreum

    (SMITH) in rats hind paw

    oedema induced by various

    phlogistic agents"(http://www.ijp-online.com

    /article.asp?issn=0253-7613;

    year=1988;volume=20;issue=2;

    spage=86;epage=89;

    aulast=Agarwal;type=0).Indian

    Journal of Pharmacology20(2)

    869.

    66.

    ^Xiong, Jing; Zhu, Zhonghua;

    Liu, Jianshe; Wang, Yang(2009). "The effect of root of

    rhododendron on the activation

    of NF-B in a chronic

    glomerulonephritis rat model".

    Journal of Nanjing Medical

    University23: 73.

    doi:10.1016/S1007-4376(09)600

    31-9 (http://dx.doi.org

    /10.1016%2FS1007-4376%2809%2960031-9).

    67.

    ^U S Food & Drug

    Administration Center for Food

    Safety & Applied Nutrition

    Food borne Pathogenic

    Microorganisms and Natural

    Toxins 1992 (Bad Bug Book)

    68.

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    19/24

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    20/24

    0-917304-71-3, ISBN 0-88192-109-2, ISBN 0-88192-168-8, ISBN 0-88192-311-7.

    (http://books.google.ca/books/about

    /The_Rhododendron_Species_Azaleas.html?id=CBQmAQAAMAAJ&redir_esc=y)

    Cox, P. A. & Kenneth, N. E. The Encyclopedia of Rhododendron Species. 1997. Glendoick Publishing.

    ISBN 0-9530533-0-X. (http://books.google.ca/books/about

    /The_encyclopedia_of_rhododendron_species.html?id=yftGAAAAYAAJ&redir_esc=y)

    James Cullen. Hardy Rhododendron Species: A Guide To Identification. Timber Press, 2005. ISBN

    0881927236 (http://books.google.ca/books/about

    /Hardy_Rhododendron_Species.html?id=jxY4dVOm1VUC&redir_esc=y)

    Blazich, F.A. and D.B. Rowe. 2008. "Rhododendron L., rhododendron and azalea." p. 943-951. In: F.T.

    Bonner and R.P. Karrfalt (eds.). The woody plant seed manual. Agr. Hdbk 727. U.S. Dept. Agr. For. Serv.

    Washington, D.C. (http://www.rhododendron.dk/rhododendron.pdf)

    Articles

    Walter Magor. A History of Rhododendrons. (http://scholar.lib.vt.edu/ejournals/JARS/v44n4/v44n4-

    magor.htm) JARS Volume 44, Number 4, Fall 1990

    GAO Lian-Ming, LI De-Zhu, ZHANG Chang-Qin, YANG Jun-Bo. Infrageneric and Sectional

    Relationships in the Genus Rhododendron (Ericaceae) Inferred from ITS Sequence Data. Am Botanica

    Sinica 2002, 44(11) 1351-1356 (http://www.jipb.net/pubsoft/content/2/2316/X010407(PS2).pdf)

    Goetsch, Loretta; Eckert, Andrew J.; Hall, Benjamin D. (JulySeptember 2005). "The molecular

    systematics ofRhododendron(Ericaceae): a phylogeny based uponRPB2gene sequences"

    (http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00012). Systematic Botany30(3): 616626. doi:10.1600/0363644054782170 (http://dx.doi.org/10.1600%2F0363644054782170).

    Subdivisions

    Azaleas

    Wilson EH, Rehder A. A MONOGRAPH OF AZALEAS RHODODENDRON SUBGENUS

    ANTHODENDRON. THE UNIVERSITY PRESS, CAMBRIDGE April 15 1921. PUBLICATIONS OF

    THE ARNOLD ARBORETUM, No. 9 (http://www.biodiversitylibrary.org/bibliography/55595#/summary)

    John L. Creech. An Embryological Study in the Rhododendron Subgenus Anthodendron Endl. Botanical

    Gazette Vol. 116, No. 3 (Mar., 1955), pp. 234-243 (http://www.jstor.org/discover/10.2307

    /2473343?uid=3739432&uid=2129&uid=2&uid=70&uid=3737720&uid=4&sid=21103875679061)

    Tsutsusi

    odendron - Wikipedia, the free encyclopedia http://en.wikipedia.org/wiki/Rhod

    24 8/4/2014

  • 8/12/2019 Rhododendron - Wikipedia, The Free Encyclopedia

    21/24

    Chamberlain DF. A revision of Rhododendron. IV Subgenus Tsutsusi. Edinburgh Journal of Botany

    06/1990; 47(02):89 - 200. DOI: 10.1017/S096042860000319X (https://www.researchgate.net/publication

    /231958128_A_revision_of_Rhododendron._IV_Subgenus_Tsutsusi)

    Powell, E. Ann, Kron, Kathleen A. Molecular systematics of Rhododendron subgenus Tsutsusi

    (Rhodoreae, Ericoideae, Ericaceae). Botany 2004: 147 (http://2004.botanyconference.org/engine/search

    /index.php?func=detail&aid=147)

    A. Kron and E. A. Powell. MOLECULAR SYSTEMATICS OF RHODODENDRON SUBGENUS

    TSUTSUSI (RHODOREAE, ERICOIDEAE, ERICACEAE) Edinburgh Journal of Botany Volume 66

    Issue 1 March 2009, pp 81-95 (http://journals.cambridge.org/action/displayAbstract?fromPage=online&

    aid=4481360K.)

    Yue-Jiao ZHANG, Xiao-Feng JIN, Bing-Yang DING and Jing-Ping ZHU. Pollen morphology of

    Rhododendronsubgen. Tsutsusiand its systematic implications. Journal of Systematics and Evolution.

    Volume 47, Issue 2, pages 123138, March 2009 (http://onlinelibrary.wiley.com/doi/10.1111

    /j.1759-6831.2009.00011.x/abstract)

    Jin Xiao-Feng, Ding Bing-Yang, Zhang Yue-Jiao, and Hong De-Yuan. A Taxonomic Revision Of

    Rhododendronsubg. Tsutsusisect.Brachycalyx(Ericaceae). Annals of the Missouri Botanical Garden

    97(2):163-190. 2010 doi: http://dx.doi.org/10.3417/2007139 (http://www.bioone.org/doi/abs/10.3417

    /2007139?journalCode=mobt)

    Vireya

    Sleumer, Hermann Otto. An account of rhododendron in Malesia. P. Noordhoff, Groningen 1966

    David G. Leach. The Discovery of the Malaysian Rhododendrons. JARS Volume 32, Number 1 Winter1978 (http://scholar.lib.vt.edu/ejournals/JARS/v32n1/v32n1-leach.htm)

    Argent, G.Rhododendrons of subgenus Vireya. 2006. Royal Horticultural Society. ISBN 1-902896-61-0

    Gillian K. Brown, Lyn A. Craven, Frank Udovicic and Pauline Y. Ladiges. Phylogenetic relationships of

    Rhododendron section Vireya (Ericaceae) inferred from the ITS nrDNA region. Australian Systematic

    Botany 19, 329342. August 2006. 10.1071/SB05019 (http://www.researchgate.net/publication

    /235601539_Phylogenetic_relationships_of_Rhododendron_section_Vireya_(Ericaceae)_inferred_from_t

    he_ITS_nrDNA_region/file/72e7e51e347034a1c1.pdf)

    Hall, B.D., L.A. Craven & L.A. Goetsch. 2006. The taxonomy of subsection Pseudovireya twodistinctly different taxa within subsection Pseudovireya and their relation to the rooting of section Vireya

    within subgenus Rhododendron. Rhododendron Sp. 1: 72, 9197

    Craven, L.A.; Goetsch, L.A.; Hall, B.D.; Brown, G.K. (2008). "Classification of the Vireyagroup of

    Rhododendron(Ericaceae)".Blumea - Biodiversity, Evolution and Biogeography of Plants53(2): 435.

    doi:10.3767/000651908X608070 (http://dx.doi.org/10.3767%2F000651908X608070).

    Goetsch, L.A., Craven, L.A. and Hall B.D. 2011. Major speciation accompanied the dispersal of Vireya

    Rhododendrons (Ericaceae, Rhododendron sect. Schistanthe) through the Malayan archipelago: Evidence

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