New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff...

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- Novem b er 1997 New letter NSW Group Activities: 2.+-30 October: Soutl) Coast Fie ld Trip Meet 7 a.m. outside gates Gre vi llea Park t:lllle: Hnillc II I Pa ul & Ju -d HIlr11IW!U n 1 ' 1. . n 'I(' '\ ' Phone: ((12) lJ 44n 377) Subject: a u.1J.:n \ Jsi!. u. lng the key. I l)() ;I Ltil. l1 1 L:s. 2 p.m. Suhject: Pro pa ga tion b. C utting S.E. Old Gr ou p Meetings to 1997 All meetings commence at 9.30 am . /fl tc . notified. Further information contact " (07) 5546 3322 Sunday 30 Novemb er Venue: Home 01 Joh n ,\: I'll It I{ ._ 10 Smilhs Wights MllUnla i!1 4" _11 (07) 3289 J43 I Subject: A review of mulches 101 gI L' \ tilL! :11 ; planting techniqu e.' . Sunday 25 January 1998 Venue: Home of John & Irenl' SI Il! \ 17 Trafalgar Street. Toowoomba 4:)50 Phone: (07) 4636 4325 Subject: Maintaining a large Grcvilkaclllkl'll l) ll Small hacbar d nursery I Marcil 1 98 \ e nu e: \tlount An nan Botanic Garden Subject: E 'amine the new Grevillea planting. Finalise arrangements for the Giant Plant Sale Sat urday & Sund ay Apr il 18-19 Giant A utu mn Plant Sale & Art Sho w. Set lip Friday April 17, Pu ll Down Mon day April 20 Best Wishes for a Merry Chri st mas and a successfu l 1 998 for Growing Grevi ll eas INSIDE GROUP ACTIVITY - New South Wales Giant Autumn Plant Sa le >- TAXONOMY - New Species of (ircvilka - Grcvillca cefalO CONSERVATI01\" - Wee Jasper Grcvillea I )0> IN YOUR GARDEN Items of Interest from Ii ec llher Cllirkc Repo rt un Guided uf llunlu RC !:,- II )1i BOlanic Garden by H eoflrer Clork e >- PROPAGATION The Greal Grafting Debatc hI' Ric!wrd Tomkin Seed Bank by Jud\' , 'imiflr )0> FINANCIAL REPORT

Transcript of New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff...

Page 1: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

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November 1997

New letter NSW Group Activities

2+-30 October

Soutl) Coast Field Trip Meet 7 am outside gates Grevillea Park

tlllle Hnillc II I Pa ul amp Ju -d HIlr11IWU n 1 1 n I(

Phone ((12) lJ 44n 377)

Subject au1Jn Jsi u lng the idcllt i fic~lLion key I l)() ILtil l1 1Ls

2 pm

Suhject Propaga tion b C utting

SE Old Group Meetings to 1997 All meetings commence at 930 am fl tc

notified Further information contact (07) 5546 3322

Sunday 30 Novem ber

Venue Home 01 Joh n Ill It I _

10 Smilhs R()~hl Wights MllUnla i1 4 _11

I~hone (07) 3289 J43 I

Subject A review of mulches 101 gI L tilL 11

planting technique

Sunday 25 January 1998

Venue Home of John amp Irenl SI Il

17 Trafalgar Street Toowoomba 4)50

Phone (07) 4636 4325

Subject Maintaining a large Grcvilkaclllklll l)ll Small hacbard nursery

~unday I Marcil 1 98

enue tlo unt An nan Botanic Garden

Subject E amine the new Grevillea planting Finalise arrangements for the Giant Plant Sale

Saturday amp Sunday April 18-19

Giant Autumn Plant Sale amp Art Show Set lip Friday April 17

Pu ll Down Monday April 20

Best Wishes for a Merry Christmas

and a successful 1998 for Growing Grevilleas

INSIDE ~ GROUP ACTIVITY - New South Wales

Giant Autumn Plant Sa le

gt- TAXONOMY - New Species o f (ircvilka - Grcvillca cefalO

~ CONSERVATI01 - Wee Jasper Grcville a

I )0gt IN YOUR GARDEN Items of Interest from Iiecllher Cllirkc

Report un Guided T()ur~ uf llunlu RC - II )1i

BOlanic Garden by H eoflrer Clorke

gt- PROPAGATION The Greal Grafting Debatc hI Ricwrd Tomkin

Seed Bank by Jud imiflr

)0gt FINANCIAL REPORT

Prugress RCP Llft

h~ Giant

Gian A t m Plant Sa e 18-19 April 1998 - Mt Annan Botanic Garden

The organi cr will aL 0 have AhritJinul painting anu w r~~ by Pro Hart and Norman Linu ay l(lr~aIL Rcpr luuLl llln

u tUl11rl Plant ule 1 hI held al the Euucalion eL hing of the b ter featuring nuuL (1111111 ill 1 1 ~1 ) he e nt n 01 the Ml nnan Botanic Gardens un 1 -1~ April availahle for less tha n ~2()O While VilLI rnu rcasunahl y

1991- ha hoen confirmed rhL sale will not ht nstriltl~d qULly the r levance 01 thesl wnrk lail -sunJ the mduls tll re ilk a alth lugh it i h pet tha l ) wiue ran1C 111 wtre all Austra lian-born Ant III C11l11~ lhlluJ) (ilJp he on alt Al l genera ar welcomt The pTU ced~ will hen hl wi ll benefit from any pr (it q ak 1 1m lIs1 lryll1g It Ithe stud group whilth dl ~pcnd the money n c n crvaLion interest someone to hold an npLn air inl-las lmg In COIIshyprojects a~ weB as ho rticultural and tff anomie research junctjnn with lht cxhibl t 10 10 the genus Grcvlllea I halc managld to find an organiser

We hav two olher roum~ and kilhen ) uilabk 1lfC di~shyto run an art 1l1O in i ne l)f tht hUJrdlng~ u ing lhe (heme of nati e plant and lantiscaptls II YIJU arc a painter l)r play talk and auulcl-visurth v ill I~ f1 lu ku IILilli

lhe re be anyone willmg It) mlJunl ~1kn W SOl cnne who jy ami would hk LO exhihil work cUI-lll c Jl~plt )1

tm sail please contact me un (02) 9543 2242 Banksia and other proteaceueJ

The fo lio v in t pTllgramml i bullIf au IT1l11 11

Displays amp Activities Thl Gr vilka Study Group rh le)graphlc and cut-l1 wer di play I Grevll1ea

middot) lmiddot) ) Cut tl Wr uispla 01 Banksia and other proteaceae

Merv I1 odge Demnnstnltinn and Idlk Prupagali)n b graJung

d tarr iolt l Dernl nstraUon and Talk Propagation by ullin l 2 lIIustn t u lId ltIlk Tht Grcvil lea Gardlo

He ather Clarkt Grevilleas at llumcr Region Botanic G ardl o

Peter O lde (tU lk ) 1 The genus Grevl11ea 2 Rare amp Endangered G revilleas

erv Hodge ell Marri l)ll tiue dlif1I y f Grtvilleas

R Brown IIlawarra Grevillea Park i ) Banksia Audl -vi tJaJ

Mount Annan Stalt Laning lrom S(ratdl (dcmonitrat inn and talk I

Mount Annan taft Guided Walks through tht nursery ancl coll cct r)t1

Mount Ann an tafi GUIded walk lhmugh lh Botanic Gardens

The ve nue is superh and the eaLlhr Wlll Ol wllnderful I t i ju t tbe righl time to bu~ plunts and plant thelll out

Seed Bank by Judy Smith

The seed bmk h ilS gone inLcrnatioD II I stnt some s cd to Germany ia Ray Brown an d the internet - cry e iling

Any donations of seed belp La rnakt a bette r seed bank or everyone to beneshyfit T h nk you lu our nly U nors this year - Bill Lovett and Ian MiLcheLl

To ob tain free seed please send a selfshyaddressed envelop with n70cent stamp to Judy Smith 15 Cromddlt Lreel Mortdale 2223

FREE SIO

G Ql[Jleniifolia

bUllksii red (gT Y leaf shru b form ) tIdicherw lla jomsonii jWliperill(l (red upright ) lollglStya

macieayana pelrophiloid rivu mis Ihelena1fl i rcll a

triloba

CUTTING EXCHANGE

Cutting matcl ial ii avlriahk to iinanclal mem crs only from

Dav Ma~on Box l)1 Co aki 2471

Hundrellli of species available

Pkasc contac t me 1 n1JV lu vc thl plant au rcqulrt The cn~ t is

650 p yab k with orde r which CO I rs lht co 1 ( r p 1 I ing and

relurn VIa lf ss Pns t

Grevi llea Study Group NQ 4 2 l1vembcr ll)l)7

New Species of Grevi llea P ade October 7997

Sincc puhllcati on ofV ul I of the G rcvillea Boo k (Dcccm her 1l)l)4) Jescriptions of 4 new speci s (C celato W M Molyshyneux G n1aCClliCheol1ii G 1 K igherv amp RJ Cranfie ld G oxyollilw ssp oryo lliiza Makins(~n and G rhyolitica ssp rhvoirica Makinson IT -ranking of 1 ubspecies to species ( G p nlicoicics R H I) and descriptions of three new subshyP L c ICS ((J ()JIII If7orl ssp jecLllua G 0 J( IIIW 11]7 fea rishyIlata C rrolill ca ssp lcmivestilO) have heen published

T hi now brings the tn tal numhe r of accepted peci es to 3-1 8 The Flora Ll f Australia treatment of revillca by Bob llakin son wi ll be published probably in 1998 and this ill incluJ e lt1 num he r of additional new specie

Ithough it is not nrt cl in that all specis accepted b O ldc amp larrilltt will be acccr Lcd in th e n ora treatme nt I am u 1nl idcnt th at thl I-C wil l only be minm changes It is likely lh t the number uf acce pted spcli cs after the Flora trLatshyn1Lnl will excel lfl()

1~95 ( ircllicli cdu( (sec separatc article this newsct ter) 11)96 Grn illca I7ll1cclllcheof1ii

n ahb ni alcd d ~~cnptlo n of l71[ cculchcOllii was in shy1

cluded 111 Ncwsle tkr No 46

Commentary The re is no douht as to the distin ll1 (111 1 1

thi s species through its fidJIc-s hapcd ampleXicaul le aves wh ich are unlike any o ther in the genus My upi nion thilt this spccies belongs in our Group 14 ()ne III its closest relatives bcing G ripicola) has been lllnfirrmd hv Iiuits sent to me from a cultivated spccimen hy IIiabeth (l l1r [Cl

The fruits cxhihit an oh lique ly trullcal ed hI witl lt 1 I

ridging a major point separating this grlllip lrul1l our CirUllI 16 which incluJes c mllnglesiuitcs Apart iru l1l its kdll

G l71occll lcizeonii differs fwm C (l~csi( id(s 111 Its Il 111 ~ 1

pistils and the shape of its fruit s

A me ndme nts to the 0 amp M Kcv arc

Kev to Groups P 19 1 Insert 3ia Pistil IN mm long icdves nllt pltllldurifml1l (il (l u[1 I[i Iia Pistil 2() I1lm IOllg 1c1VlS PII1 J Lll ilp lTll (r l Illj I

K v to Species rruup 14 ( I =O)

iter I inse rt the f()lllWI1g bill

5a Leaves amplcxilaul ( CCIIc(ril ii Sa Leaves not amp lexiearl

(10 he cunl il1ucdj

co Wee Jasper Grevillea

The future of the end angered Grevillea iaJpicula which is restricted to th e Wee Jc sp r area about 40 km north wes t of Canberra IS looking much brighte r following the comb ined efforts of a small group of lanJholde rs and conshyse rva tion agenci s

The plant is the focus orth Wee Jasp rGrevill a R ecovery Team which invo lv s four landowners th local school teacher and representatives from the oc iety fo r G rowing Australian Plan t the G oobarra and ra-G oodradigbee Re shyserve Trust Burr inJuc Sta te Recreat ion Are a N W ashytiona I Parks amp Wildlife Se rvice th Biodiversity G roup of Environme nt Australia and the NSW Departmen t of Land and Water Conservation

There are less than 250 mature plan t in the wild which are subjec t to grazing by dom tic stock fe ral goats and blackbe rry and swe t briar weed infesta tion Many of the plants onl y survive in precipitous Inaccessible 10 ations

Rece ntly all signific t c llonics o f Grevillea iasp icula wen fenced off from graLing animals nd extensive control of weed infes tations has b en comple ted A nnual monitoring of the populationswi ll provide important information bout the effectiveness of recovery aclions and in particu lar the recruitment of eedlings

The cost to datt is $18000

~()urce J Briggs Dan thon ia 6( 1) June 1997 reprint fr 1m O n the Brink 9 May 1997

RVA ION

Grevilea iaspicula

ovembcr 1997 I G rev illc a StuJy G roup ltgt 4N

Grevillea ceata (PROTEACEAE) A NEW SPECIES FROM CENTRAL EASTERN GIPPSLAND VICTORIA

wM Molyneux PO Box 386 Yarra Glen Victoria 3775

ABSTRACT

~middot1olJ)eux v M Crelillea celata (Proteaceae) a new species from central eastern Gippsland Victoria Muelleria 8(3) 311-316 (1995)- Grcvilfea cclata is dcscribed and named and distinguished from G alpina Lindley and G chrysophaca F Mueller ex Meisner species with which it has been conshyfused by arious au thors Morphological varia tion affini tics ecology and conservation status are discussed

INTRODUCTION

Sevcral authors have noted the existence of a distinctive (jrcliflco in the Bruthen region assigning it to either G alpirw Lindley (eg Willis 197246) or to G cllIysophaca F 1ueller ex Meisner (eg cGillivray 1993 270-1 ) or to both taxa (Costcrmans 198 1 162 in circumscription and maps) The taxon has been informally known for some years as G sp nov Nowa Nowa from a nearby locality

The author has made observations of this taxon in the field since 197R observing especially its modes of regeneration and polymorphism in t10wer morphology and t10wer colour and in leaf shape and size These observations together with detailed measurements of live and herbarium material ol all three species arc detailed below and provide evidence for recognition of this taxon as a distinct species Some diagnostic features are not readily apparent from dried hershybarium specimens and their label data this difficulty probshyably underlies the uncertainty of diagnosis and assignmen t of this taxon in the past

Polymorphism within the taxon does exist with some apshyparently unstable character states it is possible that this species is of relatively recent hybrid origin The most typical form has red and yellow flowers and type material has been scleeted from a large population of this form as being the most representative

Definition of morphological characters and states follows that of McGillivray (1993)

TAXONOMY

Grevillea celata Molyneux sp nov

G alpillae Lindley et cluysophaea F Mueller ex Meisner affinis sed surculis radicibus cont1orescentiis magis ramosis forma amplitudine stato nectarii et pistilis longioribus saepe differt

TYPUS Victoria East Gi[lpsland Botanicil Rql( lll Colquhoun State Forest 1315 km cast north cast aillng Dclttd Horse Creek Road form the turnoff on the Bruthen-Buch111 Road c55 km south cast ()j BrullKn ~vLlp rc-I IldlllhL k R422 840306 13 Oct ISJ93 WM M(IIl(Ux (HU10Ir)l MEL ISOTYPI AD BRL CAN 13 K NSV IL](1I I)

Upright and open to low Clnd dense slzruh ()A-IS Il1 Lti l suckering from roots branchlets shortly tome 11 tuse II

mixed white and fawn hairs Juvenile le(Hs pink lH 111 1~ soon becoming green Adult leavcs mostly spread ing (11 rlti re 1 ascending shortly petiola te simple and en ti re (1hlung -L 1shyliptical to broadly ellipt ical or broadly linear often sUhCilIlVe upwards in eross seetiun ( 15- )2()-44( -51) mm long (4-) 7 -IS mm wide petiole (l5-15 mm long hase lttenualL 11L softly mucronate to apiculate margin~ irregularly wavy rLshycurved to loosely revolute seldom obscuring the lower surshyface upper surface tuberculate with a scattering or shurt hairs dull green or yellow-glecn lower surfacL tOI11LmiddotntlSL the hairs mainly white with an admixture of lightly krshyruginous ones hairs variably straight or twisted VL11lti(111 brochididromous obvious on upper surface less Sll helc11 due to indumentum Conflorescellces mainly terminal LliUshy

ally on short lateral branchlcts or occasionally axillar) (111

older wood floral rachis apically decurved or deflcxed llr rarely straight simple or up to 3-branched with no c()mmon peduncle (2-)4-8-t1owered forming a loose clustercentrifushygal (basipetal) peduncles (1- )2-3( -7) mm I()ng rachis 1-5 mm long both with an open indumentum of short mixLd hairs the hairs mainly white with some lightlv lcrruginuul

- - - shy

bracts triangular to broadly linear 11-25 mm long 12-14 mm wide tomentose outside with white hairs overlain hI

more erect lightly ferruginous hairs cm and ncar the mclrgins and mid-line glabrous inside except along the margins dl1Li

at the tip deciduous when buds are c 3 mm long fediccll 4-9 mm long with white and lightly ferruginous hairs (tlw proportion variab le ) pedicels ascending at c 35deg-45deg tll the rachis Torus oblique at 250-40deg more or less square in plan vicw (ie distinctly angled below the tepa I sutures) 17-22 mm across Perianth often tardily decidullus dilatcd at the base obliquclyoblong to ovate often with prunounced ribbing of the tepal mid-lines especially below the curve (3-)48-7(-8) mm across outer surface with an open short irregular indumentum of white and lightly ferruginous hairs inner surface glabrous in the basal 2-3 mm then densely bearded for c 2 mm towards the curve with retrorse white hairs distributed evenly on the dorsal and ventral tepals then with scattered white hairs above the the beard limb obliquely ovoid 2-3 mm long 22-4 mm wide tepals (nften only the dorsal pair) variably horned with a short villous appendage (sometimes scarcely apparent) dorsal tepals (13-)14-16(-18) mm long NectalY conspicuous at c 45deg to the pedicel arcuate-rectangular to oblong in plan view 13shy25 mm long 18-25 mm wide 02-07 mm thick margin entire or irregular sometimes concave and cupping the base of the ovary Pistil lR-20( -25)mm long uVCily sessife 2-25

Grevillca Study Circup Q 48 4 NllVemher 1ljlJ7

mm long villous ovules attached at about the midpoint hetween the basal and medial positions style dorsally villous in the lower half with mixed white and ferruginous hairs and with the distrihution and density of indumentum someshywhat variahle ventral side glabrous or with a few scattered hairs style-end lateral Pollen-presenter elliptical to almost round 28-4 mm long 2-32 mm wide face shallowly concave to slightly convex with the margins often thickened stigma slightly and distatly off-centre Fruits follicular obliquely Oblong-ellipsoid to ovoid-ellipsoid 12-14 mm long 5-8 mm wide tomentose surface beneath indumentum usually granulose and irregularly longitudinally ribbed style pershysistent pericarp c 025 mm thick suture opening to 4-6 mm wide opcned fruits often retained until following flowshycring season Seeds narrowly elliptical c 9 mm long 3 mm widc 19 mm thick outer surface convex irregularly rugose and granulate inner face more or less flat seed curved inwards to hoth ends each end with a prominent pulvinus 09-15 mm long elaiosome irregularly triangular c 2 mm long c 13 mm wide at base

SPECIMENS EXAMINED (6 CITED)

ETYMOLOGY

The specific epithet is from the Latin cclatus hidden or concealed within referring to the -previQus nfus ion of the taxon with both G alpina and G cllIysophaea

VARIATION

Flowercolour variation appears to be a feature of the species and is detailed under Flower Colour (below) all the colour variations there noted except the lemon-yellow occur in all populations surveyed The species also shows considshyerable variation ( especially as compared with its presumed closest relatives - sec under Affinities below) in flower size particularly in perianth width

FLOWERING PERIOD

Flowering has been observed from July to Decemher and in late January and in February

FLOWER COLOUR

Typically the perianth is red in the basal half shading to yellow from the eurve to the apex of the tepa Is and the style is green at the base shading to pink or cherry-red in the apical third with a green style-end Type material was selected as having flowers of this colour pattern as being the most representative of ( and apparently dominant in) all populations observed

Flower colour is however variable even within populations that appear to be wholly root-suckering and therefore probshyably clonal An example of the latter is the population (measshyuring c 40m 2) from which the Type material was selected within this population perianth colour may be red basally and yellow apically as above or (respectively) red and white pink and white or apricot and white A single plant with plain lemon-yellow perianths was found about 100m from

the Type population the only instance seen of this colour Style colour varies mainly in regard to the intensity and extent of reddish coloration in the apical third the lemonshyperianth variant has the style light green throughout

Variation of this scale in tlower colour is not uncommon in Grevillea and comparable variation has heen observed in the closely related species G alpino and in the less-closdy related G arenaria seIlS sir among others

AFFINITIES

From comparative morphology G cetala is apparently m()st closely related to G alpina and G clI)sopwca Tahle I indicates some key diagnostic character statcs tor the three species It is possihle that G celata is a rdatively rccent and partially stabiliscd product of hyhridization hetween thcse ll

species It is now geographically isolated from the nearest known occurrences of hoth heing c 120 km from (i alpin a (on the lower northern slopes of the Victorian Alps ahovc Mt Beauty where forming hybrid often root-suckering swarms with G lanigera) and c 70 km from C cllI)SUpW( (which appears to have its easterly limit in the area of Bullock I-kat Creek Road off the Dargo Road) G ceta ( allowing for the polymorphisms noted) hreeds true from seed

Note regarding the distribution of C chrysophaea

cGilJjvray (1 93271) mentions TallaroQk and Merton (IS

localities for that species however from both field work and an inspection of herbarium material this author suggests that it is G alpilla which occurs at these localities

Both these related taxa are variable for certain characters especially for flower colour in C alpina as well as perianth size and shape as it is in G clIysophaea In these two species variability of these characters tends to be between not within populations

Root-suckering is not reliably recorded in either C alpilla or G chrysophaea McGillivray s (1993270) reference tll occasional root-suckering in the latter species results trom his inclusion of G celaa within it and occasional reports of root-suckering in the former always appear to involvc hybridization or intergrading with G lanigera

DISTRIBUTION

Known only from Victoria from the Colquhoun Statc Forest in central eastern Gippsland east and southeast of Bruthen

HABITAT AND ECOLOGY

The species grows on orange-red capping siliceous sands of apparently Tertiary age with low humus levels and on and around small low-relief Dovonian-Silurian granite outcrops in granitic sand The species is apparently absent from the black high-humus Tertiary sands that often abut the preferred soils Not all granite outcrops in the area carry the species and it is apparently absent from more southerly outcrops of the same granite formation at lower altitudes these however are more fire-prone and heavily colonised by denser taller species such as Kunzea ericoides and PomadeTis spp

Known populations of G ceala range in elevation from 240 to 290 m as1

November 1997 5 Grevillea Study Group NO 4R

The forests in which Grevilleacelata is found are classifiable as dry sclerophyll with Eucalyptus and Acacia being the dominant genera Proteaceae is also well rcpresen ted and there is a broad undcrstorev of small to medium and groundshycovering selerophyllous plants by a number of gc~era

REPRODUCTION

Recent observations (Molyneux Sept and Oct 1993) of areas burnt the previous summer show new growth from root -suckers and this appears to be a common means of regeneration in all populations possibly as a selected reshysponse to high fire frequencies Mature fruits and seedlings are also not uncommon

Pollination is probably both ornithophilous (While-naped Honeyeatcrs Melilhreplus IUllallis Veillot were observed in

the vicinity) and entomophilous (numbers of unidentified native bees were seen)- The flowers produce copious neetar It seems probable that ant-mediated seed transport and hurial is significant for reproduction in the case of G celaa

CONSERVATION STATUS

Currently known populations are scattered thruugh and confined to Colquhoun Slate Forest The area is subject t() rotational logging and wild and controlled tires A conservation code of 2Vi is recommended following Briggs and Leigh (1989) meaning that the species is of restricted distribution with a total range of less than 1 00 kin should be considered vulnerable and is inadequately npresented in conservation reserve (since the Slate Forest is a multiple-usc regime with the possibility of significant disturbance)

Table 1 Comparison of G celata with related species

Character G celnta G alpina G chr)sophaea

Habit Shrub root suckering shrub single-stemmed not shrub singlc-stemmed not root-suckering not root-suckering not lignotuberou~ lignotubernus

Leaves 15-58 mm long 4-18 mm 3-30 mm long 08-10 mm wide 9-55 mm long 3-21 mm wide size wide

venation brochididromous not brochididromous mostly brochididromous obvious obvious only midvein obvious

Conflorescence simple or branching up simple or branching up to 2 simple or branching up to 2 to three times 2 to 8 times 2 to many flowers times 2 to 12 lowers flowers

Floral bracts triangular or broadly triangular or linear c05-4 mm subobovate acute or linear linear c 18-25 mm long long 03-08 mm wide glabrous c18-3 mm long 08-15 mm 12-14 mm wide inside except for tip often wide tomentose inside for c glabrous inside except for persisten l to late bud top one third deciduous when tip deciduous when buds development buds c 3 mm long c 3 mm long

Flower colour red amp yellow red amp red amp yellow red amp while primarily golden or with green white pink amp white yellow amp white pink amp white patches around dorsal suture apricot amp white often yellow orange mauve oflen suffusing into base suffused or lemon in one collection

Nectary conspicuous angled at c conspicuous plusmn perpendicular not conspicuous angled 45deg to 45deg to pedicel arcuate to pedicel arcuate mostly pedicel shod often thick occasionally convex linguiform and tapering en tire pulvinus-like or broadly margin entire or irregular or tridendate regularly curving v-shaped nectary hairs present no nectary hairs present into di~ated perianth nectary sometimes c 05-13 mm high c 13-25 mm high hairs presen t sometimes c 15-25 mm wide 025 mm-O4 18-25 mm wide 02-07 12-19 mm high 04-25 mm mm thick mm thick wide 02-04 mm thick

Perianth positioned both dorsally less dense in the backward less dense dorsally than internal beard amp ventrally with long facing dorsal hairs the ventral ventrally both facing backward

white backward directed ones denser and + erect c 4 c 4 mm across 25 mm deep hairs c 5 mm across 2 mm across 2 mm deep and and 25 mm from base of mm deep and 2-3 mm 25-4 mm from base of perianth perianth from base of perianth

Pistil 18-25 mOl long 10-205 mm long 15-22 mm long

Grevillea Study Group NQ 48 6 November 1997

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 2: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

Prugress RCP Llft

h~ Giant

Gian A t m Plant Sa e 18-19 April 1998 - Mt Annan Botanic Garden

The organi cr will aL 0 have AhritJinul painting anu w r~~ by Pro Hart and Norman Linu ay l(lr~aIL Rcpr luuLl llln

u tUl11rl Plant ule 1 hI held al the Euucalion eL hing of the b ter featuring nuuL (1111111 ill 1 1 ~1 ) he e nt n 01 the Ml nnan Botanic Gardens un 1 -1~ April availahle for less tha n ~2()O While VilLI rnu rcasunahl y

1991- ha hoen confirmed rhL sale will not ht nstriltl~d qULly the r levance 01 thesl wnrk lail -sunJ the mduls tll re ilk a alth lugh it i h pet tha l ) wiue ran1C 111 wtre all Austra lian-born Ant III C11l11~ lhlluJ) (ilJp he on alt Al l genera ar welcomt The pTU ced~ will hen hl wi ll benefit from any pr (it q ak 1 1m lIs1 lryll1g It Ithe stud group whilth dl ~pcnd the money n c n crvaLion interest someone to hold an npLn air inl-las lmg In COIIshyprojects a~ weB as ho rticultural and tff anomie research junctjnn with lht cxhibl t 10 10 the genus Grcvlllea I halc managld to find an organiser

We hav two olher roum~ and kilhen ) uilabk 1lfC di~shyto run an art 1l1O in i ne l)f tht hUJrdlng~ u ing lhe (heme of nati e plant and lantiscaptls II YIJU arc a painter l)r play talk and auulcl-visurth v ill I~ f1 lu ku IILilli

lhe re be anyone willmg It) mlJunl ~1kn W SOl cnne who jy ami would hk LO exhihil work cUI-lll c Jl~plt )1

tm sail please contact me un (02) 9543 2242 Banksia and other proteaceueJ

The fo lio v in t pTllgramml i bullIf au IT1l11 11

Displays amp Activities Thl Gr vilka Study Group rh le)graphlc and cut-l1 wer di play I Grevll1ea

middot) lmiddot) ) Cut tl Wr uispla 01 Banksia and other proteaceae

Merv I1 odge Demnnstnltinn and Idlk Prupagali)n b graJung

d tarr iolt l Dernl nstraUon and Talk Propagation by ullin l 2 lIIustn t u lId ltIlk Tht Grcvil lea Gardlo

He ather Clarkt Grevilleas at llumcr Region Botanic G ardl o

Peter O lde (tU lk ) 1 The genus Grevl11ea 2 Rare amp Endangered G revilleas

erv Hodge ell Marri l)ll tiue dlif1I y f Grtvilleas

R Brown IIlawarra Grevillea Park i ) Banksia Audl -vi tJaJ

Mount Annan Stalt Laning lrom S(ratdl (dcmonitrat inn and talk I

Mount Annan taft Guided Walks through tht nursery ancl coll cct r)t1

Mount Ann an tafi GUIded walk lhmugh lh Botanic Gardens

The ve nue is superh and the eaLlhr Wlll Ol wllnderful I t i ju t tbe righl time to bu~ plunts and plant thelll out

Seed Bank by Judy Smith

The seed bmk h ilS gone inLcrnatioD II I stnt some s cd to Germany ia Ray Brown an d the internet - cry e iling

Any donations of seed belp La rnakt a bette r seed bank or everyone to beneshyfit T h nk you lu our nly U nors this year - Bill Lovett and Ian MiLcheLl

To ob tain free seed please send a selfshyaddressed envelop with n70cent stamp to Judy Smith 15 Cromddlt Lreel Mortdale 2223

FREE SIO

G Ql[Jleniifolia

bUllksii red (gT Y leaf shru b form ) tIdicherw lla jomsonii jWliperill(l (red upright ) lollglStya

macieayana pelrophiloid rivu mis Ihelena1fl i rcll a

triloba

CUTTING EXCHANGE

Cutting matcl ial ii avlriahk to iinanclal mem crs only from

Dav Ma~on Box l)1 Co aki 2471

Hundrellli of species available

Pkasc contac t me 1 n1JV lu vc thl plant au rcqulrt The cn~ t is

650 p yab k with orde r which CO I rs lht co 1 ( r p 1 I ing and

relurn VIa lf ss Pns t

Grevi llea Study Group NQ 4 2 l1vembcr ll)l)7

New Species of Grevi llea P ade October 7997

Sincc puhllcati on ofV ul I of the G rcvillea Boo k (Dcccm her 1l)l)4) Jescriptions of 4 new speci s (C celato W M Molyshyneux G n1aCClliCheol1ii G 1 K igherv amp RJ Cranfie ld G oxyollilw ssp oryo lliiza Makins(~n and G rhyolitica ssp rhvoirica Makinson IT -ranking of 1 ubspecies to species ( G p nlicoicics R H I) and descriptions of three new subshyP L c ICS ((J ()JIII If7orl ssp jecLllua G 0 J( IIIW 11]7 fea rishyIlata C rrolill ca ssp lcmivestilO) have heen published

T hi now brings the tn tal numhe r of accepted peci es to 3-1 8 The Flora Ll f Australia treatment of revillca by Bob llakin son wi ll be published probably in 1998 and this ill incluJ e lt1 num he r of additional new specie

Ithough it is not nrt cl in that all specis accepted b O ldc amp larrilltt will be acccr Lcd in th e n ora treatme nt I am u 1nl idcnt th at thl I-C wil l only be minm changes It is likely lh t the number uf acce pted spcli cs after the Flora trLatshyn1Lnl will excel lfl()

1~95 ( ircllicli cdu( (sec separatc article this newsct ter) 11)96 Grn illca I7ll1cclllcheof1ii

n ahb ni alcd d ~~cnptlo n of l71[ cculchcOllii was in shy1

cluded 111 Ncwsle tkr No 46

Commentary The re is no douht as to the distin ll1 (111 1 1

thi s species through its fidJIc-s hapcd ampleXicaul le aves wh ich are unlike any o ther in the genus My upi nion thilt this spccies belongs in our Group 14 ()ne III its closest relatives bcing G ripicola) has been lllnfirrmd hv Iiuits sent to me from a cultivated spccimen hy IIiabeth (l l1r [Cl

The fruits cxhihit an oh lique ly trullcal ed hI witl lt 1 I

ridging a major point separating this grlllip lrul1l our CirUllI 16 which incluJes c mllnglesiuitcs Apart iru l1l its kdll

G l71occll lcizeonii differs fwm C (l~csi( id(s 111 Its Il 111 ~ 1

pistils and the shape of its fruit s

A me ndme nts to the 0 amp M Kcv arc

Kev to Groups P 19 1 Insert 3ia Pistil IN mm long icdves nllt pltllldurifml1l (il (l u[1 I[i Iia Pistil 2() I1lm IOllg 1c1VlS PII1 J Lll ilp lTll (r l Illj I

K v to Species rruup 14 ( I =O)

iter I inse rt the f()lllWI1g bill

5a Leaves amplcxilaul ( CCIIc(ril ii Sa Leaves not amp lexiearl

(10 he cunl il1ucdj

co Wee Jasper Grevillea

The future of the end angered Grevillea iaJpicula which is restricted to th e Wee Jc sp r area about 40 km north wes t of Canberra IS looking much brighte r following the comb ined efforts of a small group of lanJholde rs and conshyse rva tion agenci s

The plant is the focus orth Wee Jasp rGrevill a R ecovery Team which invo lv s four landowners th local school teacher and representatives from the oc iety fo r G rowing Australian Plan t the G oobarra and ra-G oodradigbee Re shyserve Trust Burr inJuc Sta te Recreat ion Are a N W ashytiona I Parks amp Wildlife Se rvice th Biodiversity G roup of Environme nt Australia and the NSW Departmen t of Land and Water Conservation

There are less than 250 mature plan t in the wild which are subjec t to grazing by dom tic stock fe ral goats and blackbe rry and swe t briar weed infesta tion Many of the plants onl y survive in precipitous Inaccessible 10 ations

Rece ntly all signific t c llonics o f Grevillea iasp icula wen fenced off from graLing animals nd extensive control of weed infes tations has b en comple ted A nnual monitoring of the populationswi ll provide important information bout the effectiveness of recovery aclions and in particu lar the recruitment of eedlings

The cost to datt is $18000

~()urce J Briggs Dan thon ia 6( 1) June 1997 reprint fr 1m O n the Brink 9 May 1997

RVA ION

Grevilea iaspicula

ovembcr 1997 I G rev illc a StuJy G roup ltgt 4N

Grevillea ceata (PROTEACEAE) A NEW SPECIES FROM CENTRAL EASTERN GIPPSLAND VICTORIA

wM Molyneux PO Box 386 Yarra Glen Victoria 3775

ABSTRACT

~middot1olJ)eux v M Crelillea celata (Proteaceae) a new species from central eastern Gippsland Victoria Muelleria 8(3) 311-316 (1995)- Grcvilfea cclata is dcscribed and named and distinguished from G alpina Lindley and G chrysophaca F Mueller ex Meisner species with which it has been conshyfused by arious au thors Morphological varia tion affini tics ecology and conservation status are discussed

INTRODUCTION

Sevcral authors have noted the existence of a distinctive (jrcliflco in the Bruthen region assigning it to either G alpirw Lindley (eg Willis 197246) or to G cllIysophaca F 1ueller ex Meisner (eg cGillivray 1993 270-1 ) or to both taxa (Costcrmans 198 1 162 in circumscription and maps) The taxon has been informally known for some years as G sp nov Nowa Nowa from a nearby locality

The author has made observations of this taxon in the field since 197R observing especially its modes of regeneration and polymorphism in t10wer morphology and t10wer colour and in leaf shape and size These observations together with detailed measurements of live and herbarium material ol all three species arc detailed below and provide evidence for recognition of this taxon as a distinct species Some diagnostic features are not readily apparent from dried hershybarium specimens and their label data this difficulty probshyably underlies the uncertainty of diagnosis and assignmen t of this taxon in the past

Polymorphism within the taxon does exist with some apshyparently unstable character states it is possible that this species is of relatively recent hybrid origin The most typical form has red and yellow flowers and type material has been scleeted from a large population of this form as being the most representative

Definition of morphological characters and states follows that of McGillivray (1993)

TAXONOMY

Grevillea celata Molyneux sp nov

G alpillae Lindley et cluysophaea F Mueller ex Meisner affinis sed surculis radicibus cont1orescentiis magis ramosis forma amplitudine stato nectarii et pistilis longioribus saepe differt

TYPUS Victoria East Gi[lpsland Botanicil Rql( lll Colquhoun State Forest 1315 km cast north cast aillng Dclttd Horse Creek Road form the turnoff on the Bruthen-Buch111 Road c55 km south cast ()j BrullKn ~vLlp rc-I IldlllhL k R422 840306 13 Oct ISJ93 WM M(IIl(Ux (HU10Ir)l MEL ISOTYPI AD BRL CAN 13 K NSV IL](1I I)

Upright and open to low Clnd dense slzruh ()A-IS Il1 Lti l suckering from roots branchlets shortly tome 11 tuse II

mixed white and fawn hairs Juvenile le(Hs pink lH 111 1~ soon becoming green Adult leavcs mostly spread ing (11 rlti re 1 ascending shortly petiola te simple and en ti re (1hlung -L 1shyliptical to broadly ellipt ical or broadly linear often sUhCilIlVe upwards in eross seetiun ( 15- )2()-44( -51) mm long (4-) 7 -IS mm wide petiole (l5-15 mm long hase lttenualL 11L softly mucronate to apiculate margin~ irregularly wavy rLshycurved to loosely revolute seldom obscuring the lower surshyface upper surface tuberculate with a scattering or shurt hairs dull green or yellow-glecn lower surfacL tOI11LmiddotntlSL the hairs mainly white with an admixture of lightly krshyruginous ones hairs variably straight or twisted VL11lti(111 brochididromous obvious on upper surface less Sll helc11 due to indumentum Conflorescellces mainly terminal LliUshy

ally on short lateral branchlcts or occasionally axillar) (111

older wood floral rachis apically decurved or deflcxed llr rarely straight simple or up to 3-branched with no c()mmon peduncle (2-)4-8-t1owered forming a loose clustercentrifushygal (basipetal) peduncles (1- )2-3( -7) mm I()ng rachis 1-5 mm long both with an open indumentum of short mixLd hairs the hairs mainly white with some lightlv lcrruginuul

- - - shy

bracts triangular to broadly linear 11-25 mm long 12-14 mm wide tomentose outside with white hairs overlain hI

more erect lightly ferruginous hairs cm and ncar the mclrgins and mid-line glabrous inside except along the margins dl1Li

at the tip deciduous when buds are c 3 mm long fediccll 4-9 mm long with white and lightly ferruginous hairs (tlw proportion variab le ) pedicels ascending at c 35deg-45deg tll the rachis Torus oblique at 250-40deg more or less square in plan vicw (ie distinctly angled below the tepa I sutures) 17-22 mm across Perianth often tardily decidullus dilatcd at the base obliquclyoblong to ovate often with prunounced ribbing of the tepal mid-lines especially below the curve (3-)48-7(-8) mm across outer surface with an open short irregular indumentum of white and lightly ferruginous hairs inner surface glabrous in the basal 2-3 mm then densely bearded for c 2 mm towards the curve with retrorse white hairs distributed evenly on the dorsal and ventral tepals then with scattered white hairs above the the beard limb obliquely ovoid 2-3 mm long 22-4 mm wide tepals (nften only the dorsal pair) variably horned with a short villous appendage (sometimes scarcely apparent) dorsal tepals (13-)14-16(-18) mm long NectalY conspicuous at c 45deg to the pedicel arcuate-rectangular to oblong in plan view 13shy25 mm long 18-25 mm wide 02-07 mm thick margin entire or irregular sometimes concave and cupping the base of the ovary Pistil lR-20( -25)mm long uVCily sessife 2-25

Grevillca Study Circup Q 48 4 NllVemher 1ljlJ7

mm long villous ovules attached at about the midpoint hetween the basal and medial positions style dorsally villous in the lower half with mixed white and ferruginous hairs and with the distrihution and density of indumentum someshywhat variahle ventral side glabrous or with a few scattered hairs style-end lateral Pollen-presenter elliptical to almost round 28-4 mm long 2-32 mm wide face shallowly concave to slightly convex with the margins often thickened stigma slightly and distatly off-centre Fruits follicular obliquely Oblong-ellipsoid to ovoid-ellipsoid 12-14 mm long 5-8 mm wide tomentose surface beneath indumentum usually granulose and irregularly longitudinally ribbed style pershysistent pericarp c 025 mm thick suture opening to 4-6 mm wide opcned fruits often retained until following flowshycring season Seeds narrowly elliptical c 9 mm long 3 mm widc 19 mm thick outer surface convex irregularly rugose and granulate inner face more or less flat seed curved inwards to hoth ends each end with a prominent pulvinus 09-15 mm long elaiosome irregularly triangular c 2 mm long c 13 mm wide at base

SPECIMENS EXAMINED (6 CITED)

ETYMOLOGY

The specific epithet is from the Latin cclatus hidden or concealed within referring to the -previQus nfus ion of the taxon with both G alpina and G cllIysophaea

VARIATION

Flowercolour variation appears to be a feature of the species and is detailed under Flower Colour (below) all the colour variations there noted except the lemon-yellow occur in all populations surveyed The species also shows considshyerable variation ( especially as compared with its presumed closest relatives - sec under Affinities below) in flower size particularly in perianth width

FLOWERING PERIOD

Flowering has been observed from July to Decemher and in late January and in February

FLOWER COLOUR

Typically the perianth is red in the basal half shading to yellow from the eurve to the apex of the tepa Is and the style is green at the base shading to pink or cherry-red in the apical third with a green style-end Type material was selected as having flowers of this colour pattern as being the most representative of ( and apparently dominant in) all populations observed

Flower colour is however variable even within populations that appear to be wholly root-suckering and therefore probshyably clonal An example of the latter is the population (measshyuring c 40m 2) from which the Type material was selected within this population perianth colour may be red basally and yellow apically as above or (respectively) red and white pink and white or apricot and white A single plant with plain lemon-yellow perianths was found about 100m from

the Type population the only instance seen of this colour Style colour varies mainly in regard to the intensity and extent of reddish coloration in the apical third the lemonshyperianth variant has the style light green throughout

Variation of this scale in tlower colour is not uncommon in Grevillea and comparable variation has heen observed in the closely related species G alpino and in the less-closdy related G arenaria seIlS sir among others

AFFINITIES

From comparative morphology G cetala is apparently m()st closely related to G alpina and G clI)sopwca Tahle I indicates some key diagnostic character statcs tor the three species It is possihle that G celata is a rdatively rccent and partially stabiliscd product of hyhridization hetween thcse ll

species It is now geographically isolated from the nearest known occurrences of hoth heing c 120 km from (i alpin a (on the lower northern slopes of the Victorian Alps ahovc Mt Beauty where forming hybrid often root-suckering swarms with G lanigera) and c 70 km from C cllI)SUpW( (which appears to have its easterly limit in the area of Bullock I-kat Creek Road off the Dargo Road) G ceta ( allowing for the polymorphisms noted) hreeds true from seed

Note regarding the distribution of C chrysophaea

cGilJjvray (1 93271) mentions TallaroQk and Merton (IS

localities for that species however from both field work and an inspection of herbarium material this author suggests that it is G alpilla which occurs at these localities

Both these related taxa are variable for certain characters especially for flower colour in C alpina as well as perianth size and shape as it is in G clIysophaea In these two species variability of these characters tends to be between not within populations

Root-suckering is not reliably recorded in either C alpilla or G chrysophaea McGillivray s (1993270) reference tll occasional root-suckering in the latter species results trom his inclusion of G celaa within it and occasional reports of root-suckering in the former always appear to involvc hybridization or intergrading with G lanigera

DISTRIBUTION

Known only from Victoria from the Colquhoun Statc Forest in central eastern Gippsland east and southeast of Bruthen

HABITAT AND ECOLOGY

The species grows on orange-red capping siliceous sands of apparently Tertiary age with low humus levels and on and around small low-relief Dovonian-Silurian granite outcrops in granitic sand The species is apparently absent from the black high-humus Tertiary sands that often abut the preferred soils Not all granite outcrops in the area carry the species and it is apparently absent from more southerly outcrops of the same granite formation at lower altitudes these however are more fire-prone and heavily colonised by denser taller species such as Kunzea ericoides and PomadeTis spp

Known populations of G ceala range in elevation from 240 to 290 m as1

November 1997 5 Grevillea Study Group NO 4R

The forests in which Grevilleacelata is found are classifiable as dry sclerophyll with Eucalyptus and Acacia being the dominant genera Proteaceae is also well rcpresen ted and there is a broad undcrstorev of small to medium and groundshycovering selerophyllous plants by a number of gc~era

REPRODUCTION

Recent observations (Molyneux Sept and Oct 1993) of areas burnt the previous summer show new growth from root -suckers and this appears to be a common means of regeneration in all populations possibly as a selected reshysponse to high fire frequencies Mature fruits and seedlings are also not uncommon

Pollination is probably both ornithophilous (While-naped Honeyeatcrs Melilhreplus IUllallis Veillot were observed in

the vicinity) and entomophilous (numbers of unidentified native bees were seen)- The flowers produce copious neetar It seems probable that ant-mediated seed transport and hurial is significant for reproduction in the case of G celaa

CONSERVATION STATUS

Currently known populations are scattered thruugh and confined to Colquhoun Slate Forest The area is subject t() rotational logging and wild and controlled tires A conservation code of 2Vi is recommended following Briggs and Leigh (1989) meaning that the species is of restricted distribution with a total range of less than 1 00 kin should be considered vulnerable and is inadequately npresented in conservation reserve (since the Slate Forest is a multiple-usc regime with the possibility of significant disturbance)

Table 1 Comparison of G celata with related species

Character G celnta G alpina G chr)sophaea

Habit Shrub root suckering shrub single-stemmed not shrub singlc-stemmed not root-suckering not root-suckering not lignotuberou~ lignotubernus

Leaves 15-58 mm long 4-18 mm 3-30 mm long 08-10 mm wide 9-55 mm long 3-21 mm wide size wide

venation brochididromous not brochididromous mostly brochididromous obvious obvious only midvein obvious

Conflorescence simple or branching up simple or branching up to 2 simple or branching up to 2 to three times 2 to 8 times 2 to many flowers times 2 to 12 lowers flowers

Floral bracts triangular or broadly triangular or linear c05-4 mm subobovate acute or linear linear c 18-25 mm long long 03-08 mm wide glabrous c18-3 mm long 08-15 mm 12-14 mm wide inside except for tip often wide tomentose inside for c glabrous inside except for persisten l to late bud top one third deciduous when tip deciduous when buds development buds c 3 mm long c 3 mm long

Flower colour red amp yellow red amp red amp yellow red amp while primarily golden or with green white pink amp white yellow amp white pink amp white patches around dorsal suture apricot amp white often yellow orange mauve oflen suffusing into base suffused or lemon in one collection

Nectary conspicuous angled at c conspicuous plusmn perpendicular not conspicuous angled 45deg to 45deg to pedicel arcuate to pedicel arcuate mostly pedicel shod often thick occasionally convex linguiform and tapering en tire pulvinus-like or broadly margin entire or irregular or tridendate regularly curving v-shaped nectary hairs present no nectary hairs present into di~ated perianth nectary sometimes c 05-13 mm high c 13-25 mm high hairs presen t sometimes c 15-25 mm wide 025 mm-O4 18-25 mm wide 02-07 12-19 mm high 04-25 mm mm thick mm thick wide 02-04 mm thick

Perianth positioned both dorsally less dense in the backward less dense dorsally than internal beard amp ventrally with long facing dorsal hairs the ventral ventrally both facing backward

white backward directed ones denser and + erect c 4 c 4 mm across 25 mm deep hairs c 5 mm across 2 mm across 2 mm deep and and 25 mm from base of mm deep and 2-3 mm 25-4 mm from base of perianth perianth from base of perianth

Pistil 18-25 mOl long 10-205 mm long 15-22 mm long

Grevillea Study Group NQ 48 6 November 1997

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 3: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

New Species of Grevi llea P ade October 7997

Sincc puhllcati on ofV ul I of the G rcvillea Boo k (Dcccm her 1l)l)4) Jescriptions of 4 new speci s (C celato W M Molyshyneux G n1aCClliCheol1ii G 1 K igherv amp RJ Cranfie ld G oxyollilw ssp oryo lliiza Makins(~n and G rhyolitica ssp rhvoirica Makinson IT -ranking of 1 ubspecies to species ( G p nlicoicics R H I) and descriptions of three new subshyP L c ICS ((J ()JIII If7orl ssp jecLllua G 0 J( IIIW 11]7 fea rishyIlata C rrolill ca ssp lcmivestilO) have heen published

T hi now brings the tn tal numhe r of accepted peci es to 3-1 8 The Flora Ll f Australia treatment of revillca by Bob llakin son wi ll be published probably in 1998 and this ill incluJ e lt1 num he r of additional new specie

Ithough it is not nrt cl in that all specis accepted b O ldc amp larrilltt will be acccr Lcd in th e n ora treatme nt I am u 1nl idcnt th at thl I-C wil l only be minm changes It is likely lh t the number uf acce pted spcli cs after the Flora trLatshyn1Lnl will excel lfl()

1~95 ( ircllicli cdu( (sec separatc article this newsct ter) 11)96 Grn illca I7ll1cclllcheof1ii

n ahb ni alcd d ~~cnptlo n of l71[ cculchcOllii was in shy1

cluded 111 Ncwsle tkr No 46

Commentary The re is no douht as to the distin ll1 (111 1 1

thi s species through its fidJIc-s hapcd ampleXicaul le aves wh ich are unlike any o ther in the genus My upi nion thilt this spccies belongs in our Group 14 ()ne III its closest relatives bcing G ripicola) has been lllnfirrmd hv Iiuits sent to me from a cultivated spccimen hy IIiabeth (l l1r [Cl

The fruits cxhihit an oh lique ly trullcal ed hI witl lt 1 I

ridging a major point separating this grlllip lrul1l our CirUllI 16 which incluJes c mllnglesiuitcs Apart iru l1l its kdll

G l71occll lcizeonii differs fwm C (l~csi( id(s 111 Its Il 111 ~ 1

pistils and the shape of its fruit s

A me ndme nts to the 0 amp M Kcv arc

Kev to Groups P 19 1 Insert 3ia Pistil IN mm long icdves nllt pltllldurifml1l (il (l u[1 I[i Iia Pistil 2() I1lm IOllg 1c1VlS PII1 J Lll ilp lTll (r l Illj I

K v to Species rruup 14 ( I =O)

iter I inse rt the f()lllWI1g bill

5a Leaves amplcxilaul ( CCIIc(ril ii Sa Leaves not amp lexiearl

(10 he cunl il1ucdj

co Wee Jasper Grevillea

The future of the end angered Grevillea iaJpicula which is restricted to th e Wee Jc sp r area about 40 km north wes t of Canberra IS looking much brighte r following the comb ined efforts of a small group of lanJholde rs and conshyse rva tion agenci s

The plant is the focus orth Wee Jasp rGrevill a R ecovery Team which invo lv s four landowners th local school teacher and representatives from the oc iety fo r G rowing Australian Plan t the G oobarra and ra-G oodradigbee Re shyserve Trust Burr inJuc Sta te Recreat ion Are a N W ashytiona I Parks amp Wildlife Se rvice th Biodiversity G roup of Environme nt Australia and the NSW Departmen t of Land and Water Conservation

There are less than 250 mature plan t in the wild which are subjec t to grazing by dom tic stock fe ral goats and blackbe rry and swe t briar weed infesta tion Many of the plants onl y survive in precipitous Inaccessible 10 ations

Rece ntly all signific t c llonics o f Grevillea iasp icula wen fenced off from graLing animals nd extensive control of weed infes tations has b en comple ted A nnual monitoring of the populationswi ll provide important information bout the effectiveness of recovery aclions and in particu lar the recruitment of eedlings

The cost to datt is $18000

~()urce J Briggs Dan thon ia 6( 1) June 1997 reprint fr 1m O n the Brink 9 May 1997

RVA ION

Grevilea iaspicula

ovembcr 1997 I G rev illc a StuJy G roup ltgt 4N

Grevillea ceata (PROTEACEAE) A NEW SPECIES FROM CENTRAL EASTERN GIPPSLAND VICTORIA

wM Molyneux PO Box 386 Yarra Glen Victoria 3775

ABSTRACT

~middot1olJ)eux v M Crelillea celata (Proteaceae) a new species from central eastern Gippsland Victoria Muelleria 8(3) 311-316 (1995)- Grcvilfea cclata is dcscribed and named and distinguished from G alpina Lindley and G chrysophaca F Mueller ex Meisner species with which it has been conshyfused by arious au thors Morphological varia tion affini tics ecology and conservation status are discussed

INTRODUCTION

Sevcral authors have noted the existence of a distinctive (jrcliflco in the Bruthen region assigning it to either G alpirw Lindley (eg Willis 197246) or to G cllIysophaca F 1ueller ex Meisner (eg cGillivray 1993 270-1 ) or to both taxa (Costcrmans 198 1 162 in circumscription and maps) The taxon has been informally known for some years as G sp nov Nowa Nowa from a nearby locality

The author has made observations of this taxon in the field since 197R observing especially its modes of regeneration and polymorphism in t10wer morphology and t10wer colour and in leaf shape and size These observations together with detailed measurements of live and herbarium material ol all three species arc detailed below and provide evidence for recognition of this taxon as a distinct species Some diagnostic features are not readily apparent from dried hershybarium specimens and their label data this difficulty probshyably underlies the uncertainty of diagnosis and assignmen t of this taxon in the past

Polymorphism within the taxon does exist with some apshyparently unstable character states it is possible that this species is of relatively recent hybrid origin The most typical form has red and yellow flowers and type material has been scleeted from a large population of this form as being the most representative

Definition of morphological characters and states follows that of McGillivray (1993)

TAXONOMY

Grevillea celata Molyneux sp nov

G alpillae Lindley et cluysophaea F Mueller ex Meisner affinis sed surculis radicibus cont1orescentiis magis ramosis forma amplitudine stato nectarii et pistilis longioribus saepe differt

TYPUS Victoria East Gi[lpsland Botanicil Rql( lll Colquhoun State Forest 1315 km cast north cast aillng Dclttd Horse Creek Road form the turnoff on the Bruthen-Buch111 Road c55 km south cast ()j BrullKn ~vLlp rc-I IldlllhL k R422 840306 13 Oct ISJ93 WM M(IIl(Ux (HU10Ir)l MEL ISOTYPI AD BRL CAN 13 K NSV IL](1I I)

Upright and open to low Clnd dense slzruh ()A-IS Il1 Lti l suckering from roots branchlets shortly tome 11 tuse II

mixed white and fawn hairs Juvenile le(Hs pink lH 111 1~ soon becoming green Adult leavcs mostly spread ing (11 rlti re 1 ascending shortly petiola te simple and en ti re (1hlung -L 1shyliptical to broadly ellipt ical or broadly linear often sUhCilIlVe upwards in eross seetiun ( 15- )2()-44( -51) mm long (4-) 7 -IS mm wide petiole (l5-15 mm long hase lttenualL 11L softly mucronate to apiculate margin~ irregularly wavy rLshycurved to loosely revolute seldom obscuring the lower surshyface upper surface tuberculate with a scattering or shurt hairs dull green or yellow-glecn lower surfacL tOI11LmiddotntlSL the hairs mainly white with an admixture of lightly krshyruginous ones hairs variably straight or twisted VL11lti(111 brochididromous obvious on upper surface less Sll helc11 due to indumentum Conflorescellces mainly terminal LliUshy

ally on short lateral branchlcts or occasionally axillar) (111

older wood floral rachis apically decurved or deflcxed llr rarely straight simple or up to 3-branched with no c()mmon peduncle (2-)4-8-t1owered forming a loose clustercentrifushygal (basipetal) peduncles (1- )2-3( -7) mm I()ng rachis 1-5 mm long both with an open indumentum of short mixLd hairs the hairs mainly white with some lightlv lcrruginuul

- - - shy

bracts triangular to broadly linear 11-25 mm long 12-14 mm wide tomentose outside with white hairs overlain hI

more erect lightly ferruginous hairs cm and ncar the mclrgins and mid-line glabrous inside except along the margins dl1Li

at the tip deciduous when buds are c 3 mm long fediccll 4-9 mm long with white and lightly ferruginous hairs (tlw proportion variab le ) pedicels ascending at c 35deg-45deg tll the rachis Torus oblique at 250-40deg more or less square in plan vicw (ie distinctly angled below the tepa I sutures) 17-22 mm across Perianth often tardily decidullus dilatcd at the base obliquclyoblong to ovate often with prunounced ribbing of the tepal mid-lines especially below the curve (3-)48-7(-8) mm across outer surface with an open short irregular indumentum of white and lightly ferruginous hairs inner surface glabrous in the basal 2-3 mm then densely bearded for c 2 mm towards the curve with retrorse white hairs distributed evenly on the dorsal and ventral tepals then with scattered white hairs above the the beard limb obliquely ovoid 2-3 mm long 22-4 mm wide tepals (nften only the dorsal pair) variably horned with a short villous appendage (sometimes scarcely apparent) dorsal tepals (13-)14-16(-18) mm long NectalY conspicuous at c 45deg to the pedicel arcuate-rectangular to oblong in plan view 13shy25 mm long 18-25 mm wide 02-07 mm thick margin entire or irregular sometimes concave and cupping the base of the ovary Pistil lR-20( -25)mm long uVCily sessife 2-25

Grevillca Study Circup Q 48 4 NllVemher 1ljlJ7

mm long villous ovules attached at about the midpoint hetween the basal and medial positions style dorsally villous in the lower half with mixed white and ferruginous hairs and with the distrihution and density of indumentum someshywhat variahle ventral side glabrous or with a few scattered hairs style-end lateral Pollen-presenter elliptical to almost round 28-4 mm long 2-32 mm wide face shallowly concave to slightly convex with the margins often thickened stigma slightly and distatly off-centre Fruits follicular obliquely Oblong-ellipsoid to ovoid-ellipsoid 12-14 mm long 5-8 mm wide tomentose surface beneath indumentum usually granulose and irregularly longitudinally ribbed style pershysistent pericarp c 025 mm thick suture opening to 4-6 mm wide opcned fruits often retained until following flowshycring season Seeds narrowly elliptical c 9 mm long 3 mm widc 19 mm thick outer surface convex irregularly rugose and granulate inner face more or less flat seed curved inwards to hoth ends each end with a prominent pulvinus 09-15 mm long elaiosome irregularly triangular c 2 mm long c 13 mm wide at base

SPECIMENS EXAMINED (6 CITED)

ETYMOLOGY

The specific epithet is from the Latin cclatus hidden or concealed within referring to the -previQus nfus ion of the taxon with both G alpina and G cllIysophaea

VARIATION

Flowercolour variation appears to be a feature of the species and is detailed under Flower Colour (below) all the colour variations there noted except the lemon-yellow occur in all populations surveyed The species also shows considshyerable variation ( especially as compared with its presumed closest relatives - sec under Affinities below) in flower size particularly in perianth width

FLOWERING PERIOD

Flowering has been observed from July to Decemher and in late January and in February

FLOWER COLOUR

Typically the perianth is red in the basal half shading to yellow from the eurve to the apex of the tepa Is and the style is green at the base shading to pink or cherry-red in the apical third with a green style-end Type material was selected as having flowers of this colour pattern as being the most representative of ( and apparently dominant in) all populations observed

Flower colour is however variable even within populations that appear to be wholly root-suckering and therefore probshyably clonal An example of the latter is the population (measshyuring c 40m 2) from which the Type material was selected within this population perianth colour may be red basally and yellow apically as above or (respectively) red and white pink and white or apricot and white A single plant with plain lemon-yellow perianths was found about 100m from

the Type population the only instance seen of this colour Style colour varies mainly in regard to the intensity and extent of reddish coloration in the apical third the lemonshyperianth variant has the style light green throughout

Variation of this scale in tlower colour is not uncommon in Grevillea and comparable variation has heen observed in the closely related species G alpino and in the less-closdy related G arenaria seIlS sir among others

AFFINITIES

From comparative morphology G cetala is apparently m()st closely related to G alpina and G clI)sopwca Tahle I indicates some key diagnostic character statcs tor the three species It is possihle that G celata is a rdatively rccent and partially stabiliscd product of hyhridization hetween thcse ll

species It is now geographically isolated from the nearest known occurrences of hoth heing c 120 km from (i alpin a (on the lower northern slopes of the Victorian Alps ahovc Mt Beauty where forming hybrid often root-suckering swarms with G lanigera) and c 70 km from C cllI)SUpW( (which appears to have its easterly limit in the area of Bullock I-kat Creek Road off the Dargo Road) G ceta ( allowing for the polymorphisms noted) hreeds true from seed

Note regarding the distribution of C chrysophaea

cGilJjvray (1 93271) mentions TallaroQk and Merton (IS

localities for that species however from both field work and an inspection of herbarium material this author suggests that it is G alpilla which occurs at these localities

Both these related taxa are variable for certain characters especially for flower colour in C alpina as well as perianth size and shape as it is in G clIysophaea In these two species variability of these characters tends to be between not within populations

Root-suckering is not reliably recorded in either C alpilla or G chrysophaea McGillivray s (1993270) reference tll occasional root-suckering in the latter species results trom his inclusion of G celaa within it and occasional reports of root-suckering in the former always appear to involvc hybridization or intergrading with G lanigera

DISTRIBUTION

Known only from Victoria from the Colquhoun Statc Forest in central eastern Gippsland east and southeast of Bruthen

HABITAT AND ECOLOGY

The species grows on orange-red capping siliceous sands of apparently Tertiary age with low humus levels and on and around small low-relief Dovonian-Silurian granite outcrops in granitic sand The species is apparently absent from the black high-humus Tertiary sands that often abut the preferred soils Not all granite outcrops in the area carry the species and it is apparently absent from more southerly outcrops of the same granite formation at lower altitudes these however are more fire-prone and heavily colonised by denser taller species such as Kunzea ericoides and PomadeTis spp

Known populations of G ceala range in elevation from 240 to 290 m as1

November 1997 5 Grevillea Study Group NO 4R

The forests in which Grevilleacelata is found are classifiable as dry sclerophyll with Eucalyptus and Acacia being the dominant genera Proteaceae is also well rcpresen ted and there is a broad undcrstorev of small to medium and groundshycovering selerophyllous plants by a number of gc~era

REPRODUCTION

Recent observations (Molyneux Sept and Oct 1993) of areas burnt the previous summer show new growth from root -suckers and this appears to be a common means of regeneration in all populations possibly as a selected reshysponse to high fire frequencies Mature fruits and seedlings are also not uncommon

Pollination is probably both ornithophilous (While-naped Honeyeatcrs Melilhreplus IUllallis Veillot were observed in

the vicinity) and entomophilous (numbers of unidentified native bees were seen)- The flowers produce copious neetar It seems probable that ant-mediated seed transport and hurial is significant for reproduction in the case of G celaa

CONSERVATION STATUS

Currently known populations are scattered thruugh and confined to Colquhoun Slate Forest The area is subject t() rotational logging and wild and controlled tires A conservation code of 2Vi is recommended following Briggs and Leigh (1989) meaning that the species is of restricted distribution with a total range of less than 1 00 kin should be considered vulnerable and is inadequately npresented in conservation reserve (since the Slate Forest is a multiple-usc regime with the possibility of significant disturbance)

Table 1 Comparison of G celata with related species

Character G celnta G alpina G chr)sophaea

Habit Shrub root suckering shrub single-stemmed not shrub singlc-stemmed not root-suckering not root-suckering not lignotuberou~ lignotubernus

Leaves 15-58 mm long 4-18 mm 3-30 mm long 08-10 mm wide 9-55 mm long 3-21 mm wide size wide

venation brochididromous not brochididromous mostly brochididromous obvious obvious only midvein obvious

Conflorescence simple or branching up simple or branching up to 2 simple or branching up to 2 to three times 2 to 8 times 2 to many flowers times 2 to 12 lowers flowers

Floral bracts triangular or broadly triangular or linear c05-4 mm subobovate acute or linear linear c 18-25 mm long long 03-08 mm wide glabrous c18-3 mm long 08-15 mm 12-14 mm wide inside except for tip often wide tomentose inside for c glabrous inside except for persisten l to late bud top one third deciduous when tip deciduous when buds development buds c 3 mm long c 3 mm long

Flower colour red amp yellow red amp red amp yellow red amp while primarily golden or with green white pink amp white yellow amp white pink amp white patches around dorsal suture apricot amp white often yellow orange mauve oflen suffusing into base suffused or lemon in one collection

Nectary conspicuous angled at c conspicuous plusmn perpendicular not conspicuous angled 45deg to 45deg to pedicel arcuate to pedicel arcuate mostly pedicel shod often thick occasionally convex linguiform and tapering en tire pulvinus-like or broadly margin entire or irregular or tridendate regularly curving v-shaped nectary hairs present no nectary hairs present into di~ated perianth nectary sometimes c 05-13 mm high c 13-25 mm high hairs presen t sometimes c 15-25 mm wide 025 mm-O4 18-25 mm wide 02-07 12-19 mm high 04-25 mm mm thick mm thick wide 02-04 mm thick

Perianth positioned both dorsally less dense in the backward less dense dorsally than internal beard amp ventrally with long facing dorsal hairs the ventral ventrally both facing backward

white backward directed ones denser and + erect c 4 c 4 mm across 25 mm deep hairs c 5 mm across 2 mm across 2 mm deep and and 25 mm from base of mm deep and 2-3 mm 25-4 mm from base of perianth perianth from base of perianth

Pistil 18-25 mOl long 10-205 mm long 15-22 mm long

Grevillea Study Group NQ 48 6 November 1997

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 4: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

Grevillea ceata (PROTEACEAE) A NEW SPECIES FROM CENTRAL EASTERN GIPPSLAND VICTORIA

wM Molyneux PO Box 386 Yarra Glen Victoria 3775

ABSTRACT

~middot1olJ)eux v M Crelillea celata (Proteaceae) a new species from central eastern Gippsland Victoria Muelleria 8(3) 311-316 (1995)- Grcvilfea cclata is dcscribed and named and distinguished from G alpina Lindley and G chrysophaca F Mueller ex Meisner species with which it has been conshyfused by arious au thors Morphological varia tion affini tics ecology and conservation status are discussed

INTRODUCTION

Sevcral authors have noted the existence of a distinctive (jrcliflco in the Bruthen region assigning it to either G alpirw Lindley (eg Willis 197246) or to G cllIysophaca F 1ueller ex Meisner (eg cGillivray 1993 270-1 ) or to both taxa (Costcrmans 198 1 162 in circumscription and maps) The taxon has been informally known for some years as G sp nov Nowa Nowa from a nearby locality

The author has made observations of this taxon in the field since 197R observing especially its modes of regeneration and polymorphism in t10wer morphology and t10wer colour and in leaf shape and size These observations together with detailed measurements of live and herbarium material ol all three species arc detailed below and provide evidence for recognition of this taxon as a distinct species Some diagnostic features are not readily apparent from dried hershybarium specimens and their label data this difficulty probshyably underlies the uncertainty of diagnosis and assignmen t of this taxon in the past

Polymorphism within the taxon does exist with some apshyparently unstable character states it is possible that this species is of relatively recent hybrid origin The most typical form has red and yellow flowers and type material has been scleeted from a large population of this form as being the most representative

Definition of morphological characters and states follows that of McGillivray (1993)

TAXONOMY

Grevillea celata Molyneux sp nov

G alpillae Lindley et cluysophaea F Mueller ex Meisner affinis sed surculis radicibus cont1orescentiis magis ramosis forma amplitudine stato nectarii et pistilis longioribus saepe differt

TYPUS Victoria East Gi[lpsland Botanicil Rql( lll Colquhoun State Forest 1315 km cast north cast aillng Dclttd Horse Creek Road form the turnoff on the Bruthen-Buch111 Road c55 km south cast ()j BrullKn ~vLlp rc-I IldlllhL k R422 840306 13 Oct ISJ93 WM M(IIl(Ux (HU10Ir)l MEL ISOTYPI AD BRL CAN 13 K NSV IL](1I I)

Upright and open to low Clnd dense slzruh ()A-IS Il1 Lti l suckering from roots branchlets shortly tome 11 tuse II

mixed white and fawn hairs Juvenile le(Hs pink lH 111 1~ soon becoming green Adult leavcs mostly spread ing (11 rlti re 1 ascending shortly petiola te simple and en ti re (1hlung -L 1shyliptical to broadly ellipt ical or broadly linear often sUhCilIlVe upwards in eross seetiun ( 15- )2()-44( -51) mm long (4-) 7 -IS mm wide petiole (l5-15 mm long hase lttenualL 11L softly mucronate to apiculate margin~ irregularly wavy rLshycurved to loosely revolute seldom obscuring the lower surshyface upper surface tuberculate with a scattering or shurt hairs dull green or yellow-glecn lower surfacL tOI11LmiddotntlSL the hairs mainly white with an admixture of lightly krshyruginous ones hairs variably straight or twisted VL11lti(111 brochididromous obvious on upper surface less Sll helc11 due to indumentum Conflorescellces mainly terminal LliUshy

ally on short lateral branchlcts or occasionally axillar) (111

older wood floral rachis apically decurved or deflcxed llr rarely straight simple or up to 3-branched with no c()mmon peduncle (2-)4-8-t1owered forming a loose clustercentrifushygal (basipetal) peduncles (1- )2-3( -7) mm I()ng rachis 1-5 mm long both with an open indumentum of short mixLd hairs the hairs mainly white with some lightlv lcrruginuul

- - - shy

bracts triangular to broadly linear 11-25 mm long 12-14 mm wide tomentose outside with white hairs overlain hI

more erect lightly ferruginous hairs cm and ncar the mclrgins and mid-line glabrous inside except along the margins dl1Li

at the tip deciduous when buds are c 3 mm long fediccll 4-9 mm long with white and lightly ferruginous hairs (tlw proportion variab le ) pedicels ascending at c 35deg-45deg tll the rachis Torus oblique at 250-40deg more or less square in plan vicw (ie distinctly angled below the tepa I sutures) 17-22 mm across Perianth often tardily decidullus dilatcd at the base obliquclyoblong to ovate often with prunounced ribbing of the tepal mid-lines especially below the curve (3-)48-7(-8) mm across outer surface with an open short irregular indumentum of white and lightly ferruginous hairs inner surface glabrous in the basal 2-3 mm then densely bearded for c 2 mm towards the curve with retrorse white hairs distributed evenly on the dorsal and ventral tepals then with scattered white hairs above the the beard limb obliquely ovoid 2-3 mm long 22-4 mm wide tepals (nften only the dorsal pair) variably horned with a short villous appendage (sometimes scarcely apparent) dorsal tepals (13-)14-16(-18) mm long NectalY conspicuous at c 45deg to the pedicel arcuate-rectangular to oblong in plan view 13shy25 mm long 18-25 mm wide 02-07 mm thick margin entire or irregular sometimes concave and cupping the base of the ovary Pistil lR-20( -25)mm long uVCily sessife 2-25

Grevillca Study Circup Q 48 4 NllVemher 1ljlJ7

mm long villous ovules attached at about the midpoint hetween the basal and medial positions style dorsally villous in the lower half with mixed white and ferruginous hairs and with the distrihution and density of indumentum someshywhat variahle ventral side glabrous or with a few scattered hairs style-end lateral Pollen-presenter elliptical to almost round 28-4 mm long 2-32 mm wide face shallowly concave to slightly convex with the margins often thickened stigma slightly and distatly off-centre Fruits follicular obliquely Oblong-ellipsoid to ovoid-ellipsoid 12-14 mm long 5-8 mm wide tomentose surface beneath indumentum usually granulose and irregularly longitudinally ribbed style pershysistent pericarp c 025 mm thick suture opening to 4-6 mm wide opcned fruits often retained until following flowshycring season Seeds narrowly elliptical c 9 mm long 3 mm widc 19 mm thick outer surface convex irregularly rugose and granulate inner face more or less flat seed curved inwards to hoth ends each end with a prominent pulvinus 09-15 mm long elaiosome irregularly triangular c 2 mm long c 13 mm wide at base

SPECIMENS EXAMINED (6 CITED)

ETYMOLOGY

The specific epithet is from the Latin cclatus hidden or concealed within referring to the -previQus nfus ion of the taxon with both G alpina and G cllIysophaea

VARIATION

Flowercolour variation appears to be a feature of the species and is detailed under Flower Colour (below) all the colour variations there noted except the lemon-yellow occur in all populations surveyed The species also shows considshyerable variation ( especially as compared with its presumed closest relatives - sec under Affinities below) in flower size particularly in perianth width

FLOWERING PERIOD

Flowering has been observed from July to Decemher and in late January and in February

FLOWER COLOUR

Typically the perianth is red in the basal half shading to yellow from the eurve to the apex of the tepa Is and the style is green at the base shading to pink or cherry-red in the apical third with a green style-end Type material was selected as having flowers of this colour pattern as being the most representative of ( and apparently dominant in) all populations observed

Flower colour is however variable even within populations that appear to be wholly root-suckering and therefore probshyably clonal An example of the latter is the population (measshyuring c 40m 2) from which the Type material was selected within this population perianth colour may be red basally and yellow apically as above or (respectively) red and white pink and white or apricot and white A single plant with plain lemon-yellow perianths was found about 100m from

the Type population the only instance seen of this colour Style colour varies mainly in regard to the intensity and extent of reddish coloration in the apical third the lemonshyperianth variant has the style light green throughout

Variation of this scale in tlower colour is not uncommon in Grevillea and comparable variation has heen observed in the closely related species G alpino and in the less-closdy related G arenaria seIlS sir among others

AFFINITIES

From comparative morphology G cetala is apparently m()st closely related to G alpina and G clI)sopwca Tahle I indicates some key diagnostic character statcs tor the three species It is possihle that G celata is a rdatively rccent and partially stabiliscd product of hyhridization hetween thcse ll

species It is now geographically isolated from the nearest known occurrences of hoth heing c 120 km from (i alpin a (on the lower northern slopes of the Victorian Alps ahovc Mt Beauty where forming hybrid often root-suckering swarms with G lanigera) and c 70 km from C cllI)SUpW( (which appears to have its easterly limit in the area of Bullock I-kat Creek Road off the Dargo Road) G ceta ( allowing for the polymorphisms noted) hreeds true from seed

Note regarding the distribution of C chrysophaea

cGilJjvray (1 93271) mentions TallaroQk and Merton (IS

localities for that species however from both field work and an inspection of herbarium material this author suggests that it is G alpilla which occurs at these localities

Both these related taxa are variable for certain characters especially for flower colour in C alpina as well as perianth size and shape as it is in G clIysophaea In these two species variability of these characters tends to be between not within populations

Root-suckering is not reliably recorded in either C alpilla or G chrysophaea McGillivray s (1993270) reference tll occasional root-suckering in the latter species results trom his inclusion of G celaa within it and occasional reports of root-suckering in the former always appear to involvc hybridization or intergrading with G lanigera

DISTRIBUTION

Known only from Victoria from the Colquhoun Statc Forest in central eastern Gippsland east and southeast of Bruthen

HABITAT AND ECOLOGY

The species grows on orange-red capping siliceous sands of apparently Tertiary age with low humus levels and on and around small low-relief Dovonian-Silurian granite outcrops in granitic sand The species is apparently absent from the black high-humus Tertiary sands that often abut the preferred soils Not all granite outcrops in the area carry the species and it is apparently absent from more southerly outcrops of the same granite formation at lower altitudes these however are more fire-prone and heavily colonised by denser taller species such as Kunzea ericoides and PomadeTis spp

Known populations of G ceala range in elevation from 240 to 290 m as1

November 1997 5 Grevillea Study Group NO 4R

The forests in which Grevilleacelata is found are classifiable as dry sclerophyll with Eucalyptus and Acacia being the dominant genera Proteaceae is also well rcpresen ted and there is a broad undcrstorev of small to medium and groundshycovering selerophyllous plants by a number of gc~era

REPRODUCTION

Recent observations (Molyneux Sept and Oct 1993) of areas burnt the previous summer show new growth from root -suckers and this appears to be a common means of regeneration in all populations possibly as a selected reshysponse to high fire frequencies Mature fruits and seedlings are also not uncommon

Pollination is probably both ornithophilous (While-naped Honeyeatcrs Melilhreplus IUllallis Veillot were observed in

the vicinity) and entomophilous (numbers of unidentified native bees were seen)- The flowers produce copious neetar It seems probable that ant-mediated seed transport and hurial is significant for reproduction in the case of G celaa

CONSERVATION STATUS

Currently known populations are scattered thruugh and confined to Colquhoun Slate Forest The area is subject t() rotational logging and wild and controlled tires A conservation code of 2Vi is recommended following Briggs and Leigh (1989) meaning that the species is of restricted distribution with a total range of less than 1 00 kin should be considered vulnerable and is inadequately npresented in conservation reserve (since the Slate Forest is a multiple-usc regime with the possibility of significant disturbance)

Table 1 Comparison of G celata with related species

Character G celnta G alpina G chr)sophaea

Habit Shrub root suckering shrub single-stemmed not shrub singlc-stemmed not root-suckering not root-suckering not lignotuberou~ lignotubernus

Leaves 15-58 mm long 4-18 mm 3-30 mm long 08-10 mm wide 9-55 mm long 3-21 mm wide size wide

venation brochididromous not brochididromous mostly brochididromous obvious obvious only midvein obvious

Conflorescence simple or branching up simple or branching up to 2 simple or branching up to 2 to three times 2 to 8 times 2 to many flowers times 2 to 12 lowers flowers

Floral bracts triangular or broadly triangular or linear c05-4 mm subobovate acute or linear linear c 18-25 mm long long 03-08 mm wide glabrous c18-3 mm long 08-15 mm 12-14 mm wide inside except for tip often wide tomentose inside for c glabrous inside except for persisten l to late bud top one third deciduous when tip deciduous when buds development buds c 3 mm long c 3 mm long

Flower colour red amp yellow red amp red amp yellow red amp while primarily golden or with green white pink amp white yellow amp white pink amp white patches around dorsal suture apricot amp white often yellow orange mauve oflen suffusing into base suffused or lemon in one collection

Nectary conspicuous angled at c conspicuous plusmn perpendicular not conspicuous angled 45deg to 45deg to pedicel arcuate to pedicel arcuate mostly pedicel shod often thick occasionally convex linguiform and tapering en tire pulvinus-like or broadly margin entire or irregular or tridendate regularly curving v-shaped nectary hairs present no nectary hairs present into di~ated perianth nectary sometimes c 05-13 mm high c 13-25 mm high hairs presen t sometimes c 15-25 mm wide 025 mm-O4 18-25 mm wide 02-07 12-19 mm high 04-25 mm mm thick mm thick wide 02-04 mm thick

Perianth positioned both dorsally less dense in the backward less dense dorsally than internal beard amp ventrally with long facing dorsal hairs the ventral ventrally both facing backward

white backward directed ones denser and + erect c 4 c 4 mm across 25 mm deep hairs c 5 mm across 2 mm across 2 mm deep and and 25 mm from base of mm deep and 2-3 mm 25-4 mm from base of perianth perianth from base of perianth

Pistil 18-25 mOl long 10-205 mm long 15-22 mm long

Grevillea Study Group NQ 48 6 November 1997

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 5: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

mm long villous ovules attached at about the midpoint hetween the basal and medial positions style dorsally villous in the lower half with mixed white and ferruginous hairs and with the distrihution and density of indumentum someshywhat variahle ventral side glabrous or with a few scattered hairs style-end lateral Pollen-presenter elliptical to almost round 28-4 mm long 2-32 mm wide face shallowly concave to slightly convex with the margins often thickened stigma slightly and distatly off-centre Fruits follicular obliquely Oblong-ellipsoid to ovoid-ellipsoid 12-14 mm long 5-8 mm wide tomentose surface beneath indumentum usually granulose and irregularly longitudinally ribbed style pershysistent pericarp c 025 mm thick suture opening to 4-6 mm wide opcned fruits often retained until following flowshycring season Seeds narrowly elliptical c 9 mm long 3 mm widc 19 mm thick outer surface convex irregularly rugose and granulate inner face more or less flat seed curved inwards to hoth ends each end with a prominent pulvinus 09-15 mm long elaiosome irregularly triangular c 2 mm long c 13 mm wide at base

SPECIMENS EXAMINED (6 CITED)

ETYMOLOGY

The specific epithet is from the Latin cclatus hidden or concealed within referring to the -previQus nfus ion of the taxon with both G alpina and G cllIysophaea

VARIATION

Flowercolour variation appears to be a feature of the species and is detailed under Flower Colour (below) all the colour variations there noted except the lemon-yellow occur in all populations surveyed The species also shows considshyerable variation ( especially as compared with its presumed closest relatives - sec under Affinities below) in flower size particularly in perianth width

FLOWERING PERIOD

Flowering has been observed from July to Decemher and in late January and in February

FLOWER COLOUR

Typically the perianth is red in the basal half shading to yellow from the eurve to the apex of the tepa Is and the style is green at the base shading to pink or cherry-red in the apical third with a green style-end Type material was selected as having flowers of this colour pattern as being the most representative of ( and apparently dominant in) all populations observed

Flower colour is however variable even within populations that appear to be wholly root-suckering and therefore probshyably clonal An example of the latter is the population (measshyuring c 40m 2) from which the Type material was selected within this population perianth colour may be red basally and yellow apically as above or (respectively) red and white pink and white or apricot and white A single plant with plain lemon-yellow perianths was found about 100m from

the Type population the only instance seen of this colour Style colour varies mainly in regard to the intensity and extent of reddish coloration in the apical third the lemonshyperianth variant has the style light green throughout

Variation of this scale in tlower colour is not uncommon in Grevillea and comparable variation has heen observed in the closely related species G alpino and in the less-closdy related G arenaria seIlS sir among others

AFFINITIES

From comparative morphology G cetala is apparently m()st closely related to G alpina and G clI)sopwca Tahle I indicates some key diagnostic character statcs tor the three species It is possihle that G celata is a rdatively rccent and partially stabiliscd product of hyhridization hetween thcse ll

species It is now geographically isolated from the nearest known occurrences of hoth heing c 120 km from (i alpin a (on the lower northern slopes of the Victorian Alps ahovc Mt Beauty where forming hybrid often root-suckering swarms with G lanigera) and c 70 km from C cllI)SUpW( (which appears to have its easterly limit in the area of Bullock I-kat Creek Road off the Dargo Road) G ceta ( allowing for the polymorphisms noted) hreeds true from seed

Note regarding the distribution of C chrysophaea

cGilJjvray (1 93271) mentions TallaroQk and Merton (IS

localities for that species however from both field work and an inspection of herbarium material this author suggests that it is G alpilla which occurs at these localities

Both these related taxa are variable for certain characters especially for flower colour in C alpina as well as perianth size and shape as it is in G clIysophaea In these two species variability of these characters tends to be between not within populations

Root-suckering is not reliably recorded in either C alpilla or G chrysophaea McGillivray s (1993270) reference tll occasional root-suckering in the latter species results trom his inclusion of G celaa within it and occasional reports of root-suckering in the former always appear to involvc hybridization or intergrading with G lanigera

DISTRIBUTION

Known only from Victoria from the Colquhoun Statc Forest in central eastern Gippsland east and southeast of Bruthen

HABITAT AND ECOLOGY

The species grows on orange-red capping siliceous sands of apparently Tertiary age with low humus levels and on and around small low-relief Dovonian-Silurian granite outcrops in granitic sand The species is apparently absent from the black high-humus Tertiary sands that often abut the preferred soils Not all granite outcrops in the area carry the species and it is apparently absent from more southerly outcrops of the same granite formation at lower altitudes these however are more fire-prone and heavily colonised by denser taller species such as Kunzea ericoides and PomadeTis spp

Known populations of G ceala range in elevation from 240 to 290 m as1

November 1997 5 Grevillea Study Group NO 4R

The forests in which Grevilleacelata is found are classifiable as dry sclerophyll with Eucalyptus and Acacia being the dominant genera Proteaceae is also well rcpresen ted and there is a broad undcrstorev of small to medium and groundshycovering selerophyllous plants by a number of gc~era

REPRODUCTION

Recent observations (Molyneux Sept and Oct 1993) of areas burnt the previous summer show new growth from root -suckers and this appears to be a common means of regeneration in all populations possibly as a selected reshysponse to high fire frequencies Mature fruits and seedlings are also not uncommon

Pollination is probably both ornithophilous (While-naped Honeyeatcrs Melilhreplus IUllallis Veillot were observed in

the vicinity) and entomophilous (numbers of unidentified native bees were seen)- The flowers produce copious neetar It seems probable that ant-mediated seed transport and hurial is significant for reproduction in the case of G celaa

CONSERVATION STATUS

Currently known populations are scattered thruugh and confined to Colquhoun Slate Forest The area is subject t() rotational logging and wild and controlled tires A conservation code of 2Vi is recommended following Briggs and Leigh (1989) meaning that the species is of restricted distribution with a total range of less than 1 00 kin should be considered vulnerable and is inadequately npresented in conservation reserve (since the Slate Forest is a multiple-usc regime with the possibility of significant disturbance)

Table 1 Comparison of G celata with related species

Character G celnta G alpina G chr)sophaea

Habit Shrub root suckering shrub single-stemmed not shrub singlc-stemmed not root-suckering not root-suckering not lignotuberou~ lignotubernus

Leaves 15-58 mm long 4-18 mm 3-30 mm long 08-10 mm wide 9-55 mm long 3-21 mm wide size wide

venation brochididromous not brochididromous mostly brochididromous obvious obvious only midvein obvious

Conflorescence simple or branching up simple or branching up to 2 simple or branching up to 2 to three times 2 to 8 times 2 to many flowers times 2 to 12 lowers flowers

Floral bracts triangular or broadly triangular or linear c05-4 mm subobovate acute or linear linear c 18-25 mm long long 03-08 mm wide glabrous c18-3 mm long 08-15 mm 12-14 mm wide inside except for tip often wide tomentose inside for c glabrous inside except for persisten l to late bud top one third deciduous when tip deciduous when buds development buds c 3 mm long c 3 mm long

Flower colour red amp yellow red amp red amp yellow red amp while primarily golden or with green white pink amp white yellow amp white pink amp white patches around dorsal suture apricot amp white often yellow orange mauve oflen suffusing into base suffused or lemon in one collection

Nectary conspicuous angled at c conspicuous plusmn perpendicular not conspicuous angled 45deg to 45deg to pedicel arcuate to pedicel arcuate mostly pedicel shod often thick occasionally convex linguiform and tapering en tire pulvinus-like or broadly margin entire or irregular or tridendate regularly curving v-shaped nectary hairs present no nectary hairs present into di~ated perianth nectary sometimes c 05-13 mm high c 13-25 mm high hairs presen t sometimes c 15-25 mm wide 025 mm-O4 18-25 mm wide 02-07 12-19 mm high 04-25 mm mm thick mm thick wide 02-04 mm thick

Perianth positioned both dorsally less dense in the backward less dense dorsally than internal beard amp ventrally with long facing dorsal hairs the ventral ventrally both facing backward

white backward directed ones denser and + erect c 4 c 4 mm across 25 mm deep hairs c 5 mm across 2 mm across 2 mm deep and and 25 mm from base of mm deep and 2-3 mm 25-4 mm from base of perianth perianth from base of perianth

Pistil 18-25 mOl long 10-205 mm long 15-22 mm long

Grevillea Study Group NQ 48 6 November 1997

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 6: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

The forests in which Grevilleacelata is found are classifiable as dry sclerophyll with Eucalyptus and Acacia being the dominant genera Proteaceae is also well rcpresen ted and there is a broad undcrstorev of small to medium and groundshycovering selerophyllous plants by a number of gc~era

REPRODUCTION

Recent observations (Molyneux Sept and Oct 1993) of areas burnt the previous summer show new growth from root -suckers and this appears to be a common means of regeneration in all populations possibly as a selected reshysponse to high fire frequencies Mature fruits and seedlings are also not uncommon

Pollination is probably both ornithophilous (While-naped Honeyeatcrs Melilhreplus IUllallis Veillot were observed in

the vicinity) and entomophilous (numbers of unidentified native bees were seen)- The flowers produce copious neetar It seems probable that ant-mediated seed transport and hurial is significant for reproduction in the case of G celaa

CONSERVATION STATUS

Currently known populations are scattered thruugh and confined to Colquhoun Slate Forest The area is subject t() rotational logging and wild and controlled tires A conservation code of 2Vi is recommended following Briggs and Leigh (1989) meaning that the species is of restricted distribution with a total range of less than 1 00 kin should be considered vulnerable and is inadequately npresented in conservation reserve (since the Slate Forest is a multiple-usc regime with the possibility of significant disturbance)

Table 1 Comparison of G celata with related species

Character G celnta G alpina G chr)sophaea

Habit Shrub root suckering shrub single-stemmed not shrub singlc-stemmed not root-suckering not root-suckering not lignotuberou~ lignotubernus

Leaves 15-58 mm long 4-18 mm 3-30 mm long 08-10 mm wide 9-55 mm long 3-21 mm wide size wide

venation brochididromous not brochididromous mostly brochididromous obvious obvious only midvein obvious

Conflorescence simple or branching up simple or branching up to 2 simple or branching up to 2 to three times 2 to 8 times 2 to many flowers times 2 to 12 lowers flowers

Floral bracts triangular or broadly triangular or linear c05-4 mm subobovate acute or linear linear c 18-25 mm long long 03-08 mm wide glabrous c18-3 mm long 08-15 mm 12-14 mm wide inside except for tip often wide tomentose inside for c glabrous inside except for persisten l to late bud top one third deciduous when tip deciduous when buds development buds c 3 mm long c 3 mm long

Flower colour red amp yellow red amp red amp yellow red amp while primarily golden or with green white pink amp white yellow amp white pink amp white patches around dorsal suture apricot amp white often yellow orange mauve oflen suffusing into base suffused or lemon in one collection

Nectary conspicuous angled at c conspicuous plusmn perpendicular not conspicuous angled 45deg to 45deg to pedicel arcuate to pedicel arcuate mostly pedicel shod often thick occasionally convex linguiform and tapering en tire pulvinus-like or broadly margin entire or irregular or tridendate regularly curving v-shaped nectary hairs present no nectary hairs present into di~ated perianth nectary sometimes c 05-13 mm high c 13-25 mm high hairs presen t sometimes c 15-25 mm wide 025 mm-O4 18-25 mm wide 02-07 12-19 mm high 04-25 mm mm thick mm thick wide 02-04 mm thick

Perianth positioned both dorsally less dense in the backward less dense dorsally than internal beard amp ventrally with long facing dorsal hairs the ventral ventrally both facing backward

white backward directed ones denser and + erect c 4 c 4 mm across 25 mm deep hairs c 5 mm across 2 mm across 2 mm deep and and 25 mm from base of mm deep and 2-3 mm 25-4 mm from base of perianth perianth from base of perianth

Pistil 18-25 mOl long 10-205 mm long 15-22 mm long

Grevillea Study Group NQ 48 6 November 1997

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 7: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

Report on Guided Tours of HRBG Grevillea Garden August 1997

by Heather Clarke Hunter Region Botanic Gardens

Despite a slow start with bookings the tours were velY well patronised and after an initial nerve attack and a guest appearance on A C and 2NVR- 1 many interesting and inte rested people ame to vi si t

1~ format Va

I h ee ticket in a draw for a Gnvillea at the end of August

2 Grevillea historydiscove in 1770 etc - up to the present day whe re I was able to show some very recc[ltly discovercd revilleas

J Regional Grevilleas - section includes (1 mllcwlluala Banks first wllected G revillea

4 Diversity - followed by anecdotes un various plants from various widespread habitats

5 Landscaping - the challcnge of cmulating habitat as well as aesthetics of colour and form

6 Grevillcas rarc and endangered and potentially excellent as garden plants eg G f7exuosa - so easy to propagate and so beautiful in fluwer

7 AbofiglI1a an non-ationgin use 1n ~---~-~a~ o f p a ts- g 0 s e

striata and G slelio)otyra

The tour took longer than the proposed 20 minutes but almost without exception each group was attentive intershyested and keen to know more Groups varied from 5 to 25 people and includcd Welsh people who had never heard of a Grevillea before but were enthralled by my props of Bushmens Clothes Pegs and candelabras of G canshydelahroides To me the most pleasant group was some 20 visitors from the Friends of the a tional Botanic Gardens who I spoke to on a visit last year to ANBG in Canberra and urged to visit our Grevillea Garden (Beetwood) It was a great experience to face the public and has given me more confidence tor the future Study Group members Native Plants have been converted but I was pleasantly surprised at Queensland

Volume 2the interest in Grevillea created by my contact with people I only wish I could follow it up wi th sales of some appropriate

Keith Williams plants but labour constraints preclude this

Apology from the Editor In Newsletter 47 Rootstock Ramblings reference was made to using Grevillea barklyana as a r~otstock I t should have read G baileyan[ My apologies to Heather Clarke for this mistake

Items of Interest by HeatherClarke Hunter Region Botanic Gardens

bull The editorial un C rosmarinij)i(( type IllIm III Newsletter 47 was most interesting I knuw or its hiqory (I I ve historyl) but not details uf who when lH eX~Ill locatIon (in Scotland) I will happily propagate IS lllany of these as required for reintroduction Ours (Ire direct source from Sydncy R13C rvc impillred elch 111UP I ll

Guided Tours to gct in touch with Ille if they kIlll III

anyone from BathuI~t who might take ()n this rmiddotoject N i l

takers so far

bull G paluioia mentioned In NelVsletter 47 is aliv L (Jnl well in the HRBG It is gruwn frulll cuttings fmm MI Annan the source ui which is documented as bcill~ Heathcote (NSW) Identified by Peter Olde and ikn Marriot as G patuli(olia it seems to be a hardy plant in our climate and is thriving in many aspects in sunshade - showing long flowering potential Also showing th e sa po tential is sp Ij Llinmdfulia (Rutcemiddot ---llL--_

collected in October 1996 by P Olde [t is a lovel whilL fre e -tlowering plant that is ~asy l() propagate)

G striata

OFFICE BEARERS Leader Peter Olde 138 Fowler Road Illawong 2234 (02) 9543 2242

Treasurer and Newsletter Editor Christine G uthrie PO Box 275 Penshurst 2222 (02) 9579 3175

Curator of Living Collection amp Herbarium Ray Brown 29 Gwythir Avenue Bulli 2516 (042) 849216

Seed Bank Judy Smith 15 Cromda1e Street Mortdale 2223 (02) 9579 1455

Cuttings ExcllQuge Dave Mason Box 94 Coraki 2471 (066) 83 2583

November 1997 7 Grcvillea Study Group N9 4~

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997

Page 8: New letter - ANPSA · air \\inl:-las lmg . I In . COII projects, a~ weB as ho rticultural and . tff . anomie research junctjnn with lht: cxhibl t. 1010 . the genus, Grcvlllea. I ha'lc

ROPAGATIO The Great Grafting Debate by Richard Tomkin

Gin Gin Queensland Betorc ente ring into any form l) f debate 0 the suitability of Grcvillca robusta a a r o tstock I would love to bear from anyone who can advise me s Lo what is a tcn year scion and could they send me a list please

In ou r lalest catalogue - Jan uary 1997 - e list some 375 specIes cu[tivars and hybrids of which at least 50 u~e an interstoc k Why Becau~e it become - pre tty obvious that sciuns arc eith er compa tib or incompat ib le T he ones that drop o ff 12 mon Lhs la ter - yes that one Wilh a [1 the rootstock shoots - is either incompatible Or in my opinion batHy gra fted

Newsl tter 46 page 3 reported on a group vlsil to Fred and Joy McKews ell which P Ler Olde reported o n ca rly senesccnce and shoo ts fro m G robllsla rootsto ks In ewsshy[etter 47 page (j Mcrv Hodge rep a led Pett rs com ments but a[[owed that it appeared R ichard orn ki n had ove rshycome incurnpatihility problems hy u in a s ui table inshytcrstoc l-- fh ut th at it took too long to perfurm the two graft~ necessary

If the result is succe sfu l - long te rm - then I really don t mind that it took twice as long (prohahl 6()-)O scc shyuIlcb) Which hrings me to the real reason for writing (o the r than the I() yea r scions)

Shoots from G Robusta Rootstocks I laving u~ed G rohllsta exciusivd y for many ye ars exshyperi ence has led me tll believe tha t shoots will emerge from the rootstock when its p has nowhere to gu By this I mean that growth from below the graft can he expclled after

I PerllHllling the graft - where the iar has nowhe re to g() ~

) J1tcmiddot pOlling up -- when the surge 01 sap is too grcat lur the sma[1 sciun

3 After planting out - same as 2_ hut mml so if pruned to shape

4 Afte r heavy pruning - same as 1 amp 2

5 tter good rain - if it is after a dry spell its I amp 2 aga in

U you agree with 1 thruugh 5 albeit reluctant[y 1m Slime then it fo llows that

6 When the scion is incompatible and is unable t() accept the sap of the rootstock (nowhere to go) as in

H ving arrived at this point I come to 7 which will no do ubt cause all sorts of gasps oj disbelief and much shaking of head ~ d c Please rememher one thing - laquo f(l()htock has hu t one miss inn in lifc - to live

7 T he sc ion is comratihle hut thc graft union is incapab le of pe rforming to the nceds or the n)( lts toc k sap now - (nowhere to go)

If the graft is sho rt and stubhy roorly matched incom shypatible or generally had[y rerfu rmcd the ru()tslock will considcr the union to be a constrictionrcstrictiofl (II on II sort - as in 1 - and will sho()t tl) kee p itell ali c r it was content being a rt)ot stoc l-- it would IHI htlllr to shoot it wouldn t need to

or the 300+ grafted Grcvillca in our garden (1IHllC III gol) we have one that co ntinually hoots - il IH1l 1) 1

my early grafts that is short and stuhhy cracked ar()lIlHI

the edges and looks like a dried onion I

Before clumping G mOLlsta for having the duriacitv to ry and keep itself alive - rerhaps we should sec il he Ci11l1 is ind cd comratible and that it is grafte d properl

Please note that I am not - at this time - que s ti () llin ~

( mousla as a rootstock Thai wi[1 come nex t limc

FINANCIAL REPORT

SUl lSLT lp til 1I1S

1)1 Hwt il 1lS

ll l lCln cc Ull Ha nd

income

S175()O

4())(1

$2 1 ~i()(

16 IOLJ7

NOVEMBER 1997 Puhlishing

Postagc

Statione ry

$50220

Expenditure

~()()( I( I

I I A()

55 LJ()

$]7U()

[I d emss appc lr~ ill the hox your subscription of $500 is due Pc IS SLllll 1lt thllrlmiddotlS llITr Christine G uthrie PO Box 275 Penshursl 2222

1996 1997 PIcI Illltlh II cheq ues ravabk to the G rcvi[ica Study G roup

CJ CJ Grcvi[ica StuLly 1rour Nil 41 November 1997