A N N A L E S H I S T O R I C O - N A T U R A L E S M U S E I N A T I O N A L I S H U N G A R I C I

Tomus 53. P A R S Z O O L O G I C A 1961.

New and Little Known Species of Braconidae (Hymenoptera) in the Carpathian Basin

B y J . PAPP, V e s z p r é m

I t is characteristical of the Hymenoptera literature of the last decades that the authors treat an entire group systematically, geographically, or systematico-geographically. The litera­ture on Braconidae is backward in this summarizing work and that is explained by the unsettled system. The taxonomical position of many species of Braconidae are uncertain, their descrip­tion are incomplete or short, and their distribution are scarcely known, — all these aggravate the edition of synthetizing works. The difficulty is increased by the fact that the braconid fauna of the Palaeartic Region is also unknown, or, respectively, in the last years a lot of species were described from that territory. However, the unequivocal systematization and the detailed descrip­tions of the species known up to the present is still necessary to form a solid opinion about the discovered species for the sake of further investigations and to avoid the description of species only to be synonymized in the future. Motivated by this reflection, I minutely charac­terize some species and publish new data of distribution in my treatise. The female of Chryso-pophtorus chrysopimaginis Goid. is new to science ; I append hereby its description.

Concurrently, I wish to acknowledge my most sincere thanks to M. C a p e k (Banská Stiavnica), G . E . J . N i x o n (London), and P. S t a r y (Praha) for their kindness in sending me their reprints and the (original or completed) descriptions of some of the species.

Dendrosoter Wesm.

The genus Dendrosoter was erected by W e s ni a e 1 in 1835 on the basis of a sole species : he placed Bracon protuberans Nees in that genus. L a t e r M a r s h a l l , R a t z e b u r g , and R u s c h k a had described newDendrosoter species from the Palaearctic region. I n 1862 F ö r s t e r created the genus Caenopachys, relegating to it Bracon hartigi R a t z . F a h r i n g e r (1934 : 128— 129) and N u o r t e v a (1957 : 53) acknowledged Caenopachys on the one hand, while on the other M u e s e b e c k (1938 : 282) and N i x o n (1939 : 7) emphasize after P i c a r d (1928) that the generic marks established by F ö r ­s t e r are unsatisfactory to further uphold the genus. W e know 11 Dendrosoter species from the Palaearctic region. Among them three species were found in the Carpathian B a s i n : D. hartigi R a t z . , D. middendorfi R a t z . , and D. protube­rans Nees. D. middendorfi R a t z , and D. protuberans Nees are deposited in the Collection of the Hungarian Natura l His tory Museum (Budapest) , D. hartigi R a t z , was reported by S t a r y (1957 : 285) from Slovakia.

Dendrosoter middendorfi R a t z . (F ig . 1—2)

9 — H e a d , viewed from above, approximately square, proportion of width and length 1 : 0,75—0,8. Head not widened behind oculi. Face finely rugulose. Ocul i not large. Antenna shorter than body (antenna of specimen

deficient, so number of joints unaccountable). Pedicel swollen, further joints always longer than broad, joints pubescent. On vertex two crest-shaped protu­berances besides oculi, their axis slightly divergent, transversely and roughly scrobiculated (fig. 27, i n T e l e n g a 1941: 73). A fine striation between protuberances. Occiput margined, finely rugulose, rugosity behind oculi indis­tinct. Temples totally smooth and polished over mandibles. Mandibels somewhat protruding. L a b i a l palpi 3-, maxi l lary palpi 6-jointed. Head rusty brown, crest-shaped protuberances yellow. Apex of mandibles black. Antenna also rusty brown, terminally (from 4—6. joint) black. Only face and mandibles hairy .

Pronotum striated. Mesonotum finely rugulose, two notauli, — meeting on posterior quarter, — divide mesonotum into three lobes, rugosity somewhat stronger along notauli. Central-anterior lobe abruptly declining towards prono­tum. Praescutellar furrow, with 7—9 crenulae. Scutel lum also finely rugidose. Propodeum posteriorly and gradually, roughly rugose. According to N i x o n (1939 : 6) areola (of propodeum) sometimes distinct, sometimes indistinct. O n single Carpathian specimen only traces of areola. Propleura rugulose. Upper angle of mesopleura (toward mesonotum) finely striate, otherwise smooth and shiny together with ventral side. Thorax largely blackish brown. Tegulae brown. Thorax not hairy, in contradiction to N i x o n (1939 : 6) . — Wing from nervus basalis fumous. Rat io of length and width of stigma 1 : 0,26. 2. cubital cell broader than that of D. protuberans Nees (fig. 1). — Legs pubescent, prepon­derantly yellowish brown.

Abdomen oval, proportion of length and width 1,6 : 1. tergite 1,7—1,8 times longer than broad (according to N i x o n only 1,5, 1 9 3 9 : 6 ) . F i r s t tergite, apart from anterior quarter, strongly striate. O n anterior quarter, a pair of convergent ridges. Base of 2(—3). tergite striate, central heart-shaped areola closed (fig. 2), otherwise — together with further tergites and every sternite — smooth and shiny. Abdomen not pubescent, 1—2. tergites brown, other parts of abdomen dark brown. Ovipositor as long as abdomen, brown, terminally black.

Length : 3,5—3,8 m m . çf — Deviation from female : sculpture somewhat finer ( F a h r i n g e r

1934 : 139), stigma of hind wing 6 times longer than wide (fig. 12, in N i x o n 1939 : 8), posterior edge of 3—7. tergites strongly excised (Abb. 1/A, in N u o r -t e v a 1957 : 51), length : 2 , 5 - ^ , 5 m m ( N o u r t e v a 1957 : 53).

D i s t r i b u t i o n : North- and Central-Europe, the Soviet Union, A s i a Minor, India . U p to the present collected only in S lovakia in the Carpathian Sasin ( S t a r f 1957 : 285, Capek 1960 : 206). New locality : D luha ( = D l h á ) , S l o v a k i a , 1901, 1 ? , leg. G a m m e l .

Dendrosoter protuberans Nees (F ig . 3—4)

Ç — Head , viewed from above, spuare, proportion of width and length 1 : 0,7—0,8. H e a d as broad as thorax, not widened behind ocelli. F a c e finely rugulose : irregularly striate beneath pedicel and over clypeus. Oculi not large, size as in D . middendorfi R a t z . Antenna almost as long as body, 29—31-jointed. 1. joint swollen, further joints cylindrical and gradually reduced in length, last joint, however, also longer than broad. E v e r y joint pubescent. O n vertex two crest-shaped protuberances besides oculi, transversely and roughly scrobiculate, axis of protuberances divergent. Between protuberances almost wholly smooth

and shiny. Occiput curvo-striated. Temples almost totally smooth and polished, contrary to D . middendorfi R a t z . Occiput margined. L a b i a l palpi 3-, maxil lar palpi 5-jointed. — Head rusty brown, lower part of face and crest-shapde protuberances yellow. Pedicel yellow, 4—6. joints of antenna brown, from 6. jo in t blackish brown. Hairs of face and mandibles long.

F i g . 1—2. Dendrosoter middendorfi Ratz . $ ; Fore right wing (1), 1—2. tergite (2). — F i g . 3—4. Dendrosoter protuberans Nees 9 '•> Fore right wing (3), 1—2. tergite (4). —• F ig . 5—6.

Microtypus trigonus Nees ; Mesonotum and scutellum (5), Propodeum (6).

Mesonotum abruptly connating selliform pronotum. Mesonotum divided into three lobes by two deep notauli meeting on posterior quarter. Praescutellar furrow with 7 crenulae. Sculpture of scutellum pointlike. Propodeum finely reticulate, posteriorly with three areas. Propleura finelv rugulose. Upper part of mesopleura strongly rugose, otherwise smooth and shiny. T h o r a x blackish brown ; along notauli , scutellum, and beneath tegulae only brown. Tegulae yellow. Mesonotum and pleurally thorax hairy . Wing fumous from nervus basalis. Special marks are form of stigma and 2. cubital cell. Rat io of length and width of stigma 1 : 0,32. 2. cubital cell narrow (fig. 3) .— Legs preponderantly yellow, pubescent.

Abdomen elongate-oval, proportion of length and width 1,8 : 1. F i r s t tergite twice longer than broad. I t s anterior quarter almost totally smooth, otherwise roughly and irregularly reticulate. Base of 2(—3). tergite striate, central heart-shaped areola open (fig. 4). Other surface o f that tergite together wi th further tergites and every sternite smooth and shiny. Abdomen dispersely pubescent, brown, 1. tergite blackish brown. Ovipositor somewhat longer than abdomen, brown, terminally b lack.

Length : 3,8 m m . çf — Deviat ion from female : 3. joint of maxil lar palpi broad (fig. 31, in

T e 1 e n g a 1941 : 79), sculpture somewhat more rough and extensive, posterior edge of 3—7. tergites strongly excised. Length : 4,5—6 m m .

D i s t r i b u t i o n : Europe , Caucasus, West A s i a , Localities of the Car­pathian specimens were reported by S z é p l i g e t i (1899 : 98), S t a r y (1957 : 286) and C a p e k (1960 : 206). New localities : Budapest , 2 çf and 1 Q, leg. Ú j h e l y i ; P á p a , 22 May 1958, 1 çf, leg. T ó t h .

Microtypus trigonus Nees (Fig . 5—8)

9 — Head, viewed from above, transverse, proportion of width and length 1 : 0,55. Occiput edged. Ocul i large, silver, not pubescent. On lower part of face oblique and deep incision on each side, centrally and beneath pedicel concentric rugulosity around a tubercle. Otherwise face, together with other surfaces of head, smooth and polished. Ocelli eminent, sitting in corner of triangles-haped tubercle. Maxillary palpi 5-, labial palpi 3-jointed. Antenna as long as bodv, 37—40-jointed. Hairs of head long, erect on face and papli , otherwise adpres-sed. H e a d brownish yellow, black beneath occiput, palpi yellow. Antenna blackish brown.

O n center of pronotum a crenulated transverse furrow, otherwise smooth and shiny. Mesonotum and scutellum smooth and polished, pubescent. Notauli deep, crenulate (fig. 5). Along central line and lateral side of propodeum rugose, upper angle grooved (fig. 6). L a t e r a l and ventral side pubescent, smooth and shiny. Thorax black. Pronotum, mesonotum and scutellum brownish yellow. — Wing hyaline. 2. cubital cell of fore wing three-sided, 2. cubital vein indistinct and interstitial with first abscissa of radius (fig. 7). Submedial cell of hind wing large (fig. 2, in N i x o n 1950 : 455). — Pubescence of legs dense, colour brownish yellow, tarsi dark.

Abdomen elongate-elliptic, proportion of length and width 2,5 : 1. 1. tergite 2,2 times longer than broad, reticulate, anterior well protruding (fig. 8). Center of base on 2. tergite finely rugulose. Abdomen smooth and shiny from 3. tergite, together with sternites. Abdomen shaggy, black, 1—2(—3) sternites light. Proportion of length of ovipositor and abdomen 3 : 2.

Length : 5 mm. çf — Deviation from female : antenna 39—41-jointed and somewhat

longer than body, face and central line of vertex black, pro- and mesonotum brown, coxae almost totally black. Length : 5,5 m m .

D i s t r i b u t i o n : Central-Europe. Collected in two Pannonicai locali­ties in the Carpathian B a s i n : Ócsa , t u r j á n i erdő , 18. September 1952, 1 9» leg. S z ő c s ; Apajpuszta , on Reseda lutea, 1. October 1952, 1 f , leg. M ó c z á r and B a j á r i . New for the fauna of the Carpathian Basin (Hungary) .

R e m a r k s : C a p e k (1960 : 210) reports about the new host ( Acrobasis sp.) from Slovakia. M. wesmaeli R a t z . , however, is synonymous with M. trigonus Nees according to T e 1 e n g a (1935 : 273), consequently, C a p e k's data refers to that species.

Syntretus vernalis Wesm. (Fig . 9—11)

Q — Head , viewed from above, squat ; ratio of length and width 1 : 0,8— 0,9. Ocul i middle-sized, their portion towards face prominent. Temples not broad. F a c e smooth and shiny, two point-form incisions over clypeus, finely rugulose under pedical. Long setae on sxiture between clypeus and face, hairs of face fine and short. Antenna longer than half of body : 2 mm, 21 — 25-jointed,

Fig. 7—8. Microtypus trigonus Nees; Fore right wing (7), 1. tergite (8). — Fig . 9—11. Syntretus vernalis Wesm. ; Mesonotum and scutellum (9), Fore right wing (10), 1. tergite (11).

— Fig . 12. Syntretus elegáns Ruthe ; Mesonotum and scutellum.

pubescent. 1. and 3. joints three times longer than wide, 2. joint almost square, from 4. joint single joints longer than broad and finely grooved. Vertex , occiput, and temples smooth and polished, occiput margined. Ocelli at equal distance from each other, protuberant. H e a d yellow, eyes dark, 1—4(—5). joints of antenna light yellow, from 5 — 6. joints dark. Triangle formed by ocelli eventually brown.

Thorax at tegulae somewhat more narrow than head. Pronotum finely wrinkled above, smooth laterally. Notauli shallow, together with scutellum smooth and shiny. Praescutellar furrow straight, uncrenulated (fig. 9). Mesono­tum ribbed besides scutellum. Propodeum smooth and polished, with five areas, finely rugulose along ridges which separate areas. Central posterior area concave. Mesopleura and ventral side smooth and shiny. Metapleura reticulate. Thorax yellow. Pronotum and propodeum often brown, areas of propodeum generally dark. — Wing transparent. Submedial vein indistinct. 1. cubital and discoidal cell confluent. Stigma yellow, ratio of length and width 1 : 0,3 (fig. 10). — Legs slender, light yellow, densely pubescent. 1. trochanter 4 times longer than 2. Claws dark.

Abdomen laterally compressed. 5. segment as wide as thorax. 1. segment strongly petiolate, proportion of length and greatest posterior width 1 : 0,13 ; spiracles on base and behind middle (fig. 11). 2. and 3. segments connated. E n t i r e abdomen smooth and polished, yellow. 1. segment and sometimes hind segments brown. Ovipositor sheath short : only a quarter of abdomen, dark brown.

Length : 3—3,2 mm. rf — Deviat ion from female : thorax dorsallv and abdomen totally blackish

brown, 2 — 3. segments light. Length : 2,5 — 2,8 m m .

D i s t r i b u t i o n : Belg ium, Eng land , F in land . New for the fauna of the Carpathian Bas in (Hungary and Roumania) . Localities : B u d a p e s t — H ű v ö s ­v ö l g y , 8 May 1918, 1 9> leg. B i r ó ; B u d a p e s t — H ű v ö s v ö l g y , 8 June 1918, 1 Q and 2 rf, leg. B i r ó ; ó s e b e s h e l y , 16 J u n e 1913, 1 9 » i e g - B i r ó ; V á c — T u d ó s d o m b , in pratis humidis, 2 June 1929, 1 9 •> l e g- B i r ó .

Syntretus vernalis var . rugosus var . n.

Differing from nominate form by the following marks :

1. 1. tergite rugulose posteriorly behind spiracles. — 2. Thorax dorsally brown, notauli, however, yellow. — 3. Ridges on thorax dorsally black. — 4. A b ­domen dorsally brown, ventrally yellow.

Local i ty : Budapest , 1919, leg. B i r ó .

Syntretus e legáns Ruthe (Fig . 12 — 14)

9 çf — Head transverse, squat (as seen from above), ratio of width and length 1 : 0,6—0,7. Ocul i small. Temples convex. Ocelli on triangular black tubercle. Base of triangle of ocelli greater than its two sides. Clypeus separated b y linear incision from face, incision ending in little hollow. Face equally reti-ciilate. Vertex punctate between pedicel and ocelli. Temples finely and dispersely punctate along oculi. Occiput smooth and polished. Antenna short, 2,2—2,4 m m , approximately as long as half of body. Antenna 29—30-jointed, joints striated, pubescent, joints from 5. to last but one scarcely longer than wide. Head yellow. A black spot below margin of occiput. Apex of mandibles brown. 1 — 3. joints of antenna yellow, from 4. to 10—12. gradually darkening, from 10—12. brown. Head , especially face, pubescent.

Thorax at tegulae more narrow than head. E n t i r e thorax hairy. Pronotum transversely rugulose. Anterior half of mesonotum punctate, punctation sparse and rough towards scutellum. Feeble dentation in center of punctated surface. Posterior half of mesonotum together with scutellum smooth and shiny. Prae-scutellar furrow spacious, crenulate (fig. 12). Metanotum besides scutellum rough­ly ribbed. Propodeum roughly wrinkled, concave five-sided area well separated by ridges. Pro- and metapleura totally rugulose, mesopleura punctate along edges, all pleurae polished. Thorax ventral ly smooth and shiny. E n t i r e thorax yellow. Pronotum, metanotum (besides scutellum) and propodeum black. — W i n g hyaline. Submedial vein indistinct. 1. cubital and discoidal cell < onfluent, interstitial meeting of 1. cubital and recurrent vein pale. Length and width of yellowish transparent stigma 1 : 0,45—0,5 (fig. 13). — Legs slender, yellow, pubes­cent. 1. trochanter twice (occasionally 3 times) longer than 2. Only claws brown.

Abdomen laterally compressed, slender than thorax. Length and width of petiolate 1. segment 1 : 0,2—0,25. Spiracles on each end of segment. Posterior half of 1. tergite striated (fig. 14). F u r t h e r segments smooth and shiny. 2—3. segments confused. 1. segment black, subsequent segments yellow. Ovipositor one third of abdomen, brown.

Length : 5 mm, sometimes 3,5 m m .

D i s t r i b u t i o n : North-Germany, F in land . F i r s t collected by Wachs-mann at P á p a in the Carpathian B a s i n ( S z é p l i g e t i 1899 : 101). F u r t h e r localities : B o r o s j e n ő , 27 A p r i l 1927, 1 Q, leg. D i ó s z e g h y ? ; Ka locsa , 17 A p r i l 1893, 1 $ , leg. T h a l h a m m e r ; T o k a j , 4 May 1881, leg. B i r ó ? ; ( S á t o r a l j a - ) Ű j h e l y , 1 May 1881, 1 çf, leg. B i r ó ? .

Chrysopophtorus chrysopimaginis Goid. , Q nova (F ig . 15—17)

H e a d rounded, six-angled, as seen frontally ; transverse when viewed from abo\e , ratio of width and length (measured between pedicles) 1 : 0,4—0,5. Oculi large, protruding. Temple convex. Mandibles long and thin. Maxi l lary

F i g . 13—14. Syntretus elegáns Ruthe ; Fore right wing (13), 1. tergite (14). — Fig . 15—17* Chrysopophtorus chrysopimaginis Goid. ; Mesonotum and scutellum (15), Fore right wing (16),

Lateral view of abdomen (17).

palpi 6—, labial palpi 3—jointed. Ha ir s of palpi long. Edge of clypeus curved, end of its margin like a point, sporadically punctated and shiny. F a c e centrally wi th dense, around oculi with disperse, punctation, weekly polished. Vertex finely punctate and slightly bright between ocelli and pedicel. Ocelli sit on apex of ecruilateral triangle, scarcely eminent. Temples and occiput sparsely punctate, shiny. Occiput edged, concave when viewed from above. Antenna 25-jointed. 1. joint swollen, 2. joint short, 3—4. joints 4—5 times longer than broad. 5—7. joints gradually shortening, 7—25. joints hardly longer than broad. Antenna finely striated from end of 5. jo int , two opposite setae sitting at end of every jo int . Whole antenna with adpressed pubescence. H e a d light yellow. Mandibles brown, palpi translucent yellow, oculi greenish si lvery, ocelli brown. Antenna light yellow, from 7—8. joint brownish fumous.

Thorax more slender than head, ratio of width of head and thorax (at tegulae) 1 : 0,7—0,8. Length and height of thorax 1 : 0,8—0,9. Pronotum selli-form, finely striated. Mesonotum vertically connated to pronotum. Notauli lineately impressed, curved and wi th corners posteriorly. Mesonotum finely rugulose, feebly wrinkled before scutellum, whole thorax dim. Praescutellar furrow spacious and deep with 6 crenulae. Scutel lum sparsely punctate and polished (fig. 15). Anterior half of metanotum ribbed, posterior half smooth and shiny. Propodeum reticulate, honeycomb-like, opaque. Posterior center of propodeum concave, and shghtly, irregularly rugulose. Thorax light yellow, posterior half of propodeum dark. — Wings hyaline. Stigma brown. Central v e m of first abscissa of radius, first cubital vein and a short abscissa of nervus media ( L a n h a m 1951) normal, other veins obsolescent. Radius curved. 2. cubital cell 5-sided (fig. 16). — Legs light yellow, slender, pubescent. Coxae swollen. Claws brown.

Abdomen somwehat longer than head and thorax together. Abdomen late­ral ly compressed, 1. segment petiolate, 2—3. segments confused, protruding downwards and overlapping further segments. Whole abdomen shiny and smooth,

29 Természettudományi Múzeum Évkönyve

with scattered hairs, yellow, eventually 2—3. segments light brown. Hypopy-gium prominent. Ovipositor sheath almost as long as abdomen, ovipositor shor­ter than half of abdomen.

Length : 4 m m .

Deviat ion from male : number of joints of antenna, sculpture of face and mesonotum, colour of body.

D i s t r i b u t i o n : I ta ly—Bologna ( G o i d a n i c h 1948 : 86), Czecho­slovakia^—Brno (C a p e k 1957 : 1). New for the fauna of the Carpathian Bas in (Hungary and Roumania) . Localities : B o r o s j e n ő ( = Borosienu, Roumania) , 21 A p r i l 1927, leg. D i ó s z e g h y , 1 9 (Allotype, in Natural His tory Museum Budapest) ; Ő r s z e n t m i k l ó s , 15 September 1915. leg. S a j ó , 1 9* (in Natural His tory Museum Budapest) .

R e m a r k s : Right antenna of allotype missing form 13. joint. Head of paratype and right fore leg wanting.

G o i d a n i c h (1948 : 86—88) classified Chrysopophtorus chrysopimaginis as a new genus and species in the subfamily Euphorinae. On the basis of S c h m i e d e k n e c h t ' s system (1930 : 348), this genus may be a distinct subfamily near Holorimorphinae. G o i d a n i c h , however, on the basis of M u e s e b e c k's arguments (1936 : 5) considers it as one of the genera of the subfamily Euphorinae. I f the assumption is valid, — namely that Euphorinae are wel l characterized by the two following features : a) 1. segment ofabdomen petiolate, h) segments 2. and 3. fused, we may accept the systematic relegation of Chrysopophtorus chrysopimaginis Goid. as outlined above.

Bibliography : C a p e k , M, : Zwei bemerkenswerte Braconiden aus der Sammlung von Prof. S n o f l á k (Acta Soc. E n t . Cech., 54. 1957, p. 1). — C a p e k, M. : Verzeichnis der Parasiten, die aus schädlichen Insekten an V U L H in B a n s k á Stiavnica erzogen wurden. Teil I . Wirte der Brackwespen — Braconidae (Hymenoptera) (Vedecké Práce , 1960, p. 199 — 212). — F a h r i n ­g e r , J . : Opuscula braconologica, I I — I I I . (Wien, Verlag F . Wagner, I I , 1934, pp. 594, I I I . , 1937, pp. 520). — G o i d a n i c h , A. : Materiah per 10 studio degli Imenotteri Braconidi V I . (Boll . I s t . Univ. Bologna, 17, 1948, p. 83—92). — H e 11 é n, W . : F ü r die Fauna Finnlands neue Braconiden (Hym.) I . (Not. E n t . , 18, 1938, p. 108—114). — H e 11 é n, W. : F ü r die Fauna Finnlands neue Braconiden (Hym.) I I . (Not. E n t . , 25, 1946, p. 130 — 137). — L a n h a m , U . N . : Review of the wing venetion of the higher Hymenoptera (suborder Clistrogastra), and speculations of the higher phylogeny of the Hymenoptera (Ann. E n t . Soc. Am. , 44, 4, 1951, p. 614—628). — M o c s á r y, S. : Hymenoptera (in Fauna Regni Hungáriáé, 1897, p. 1 — 113). — M u e s e b e c k, C. F . W . : The genera of parasitic wasps of the Braconid subfamily Eupho­rinae, with a review of the Nearctic species (United States Dept. Agr. Miscellaneous Publica-tions No. 241, 1936, p. 1—36). — M u e s e b e c k, C. F . W. : The gennus Dendrosoter Wesmael in the United States (Hymenoptera: Braconidae) (Proc. E n t . , Soc. Wash., 40, 1938, p. 281 — 288). — N i x o n G. È. J . : New species of Dendrosoter from India and Africa ( H y m . , Braco­nidae) (Proc. Roy. E n t . Soc. London, 8, 1939, p. 1—11) — N i x o n, G. E . J . : New Indian Braconidae bred from Lepidopterous defoliators (Hymenoptera) (Ann. Mag. Nat. Hist. . 12, I I I , 1950, p. 453—474). — N u o r t e v a , M. : Zur Kenntnis des parasitischen Hymenopteren der B o r k e n k ä f e r Finnlands (Ann. E n t . Fennici , 23, 1957, p. 47—71) — P i c a r d , F . : Les variations des caractères sexuels secodares chez les Braconides des genres Caenopachys Foerst. et Dendrosoter Wesm. (Bull . Soc. Zool. F r . , 53, 1928, p. 50—56). — R u t h e , J . F . : Prodro-mus einer Monographie der Gattung Microtonus Wesm. (Stettin. E n t . Zeit., 17, 1856, p. 289 — 308). — S c h m i e d e k n e c h t , O . : Die Hymenonteren Nord- und Mittel-Europas (Jena, G. Fischer Verlag, 1930, pp. 1062). — S t a r y, P . : Notes on the Braconidae (Hym.) of Czecho­slovakia I V . (Part 1) (Acta Soc. E n t . Cech., 54, 1957, p. 277 — 292 ). — S z é p l i g e t i , G y. : Magyarországi Braconidák (Rov. Lapok, 6, 1899, p. 98 — 1 0 2 ) . — T e 1 e n g a, N . A . : Neue und weniger bekannte palearktische Braconiden (Hym.) (Arb. physiol. angew. E n t . Berlin, 2, 1935, p. 271—275). — T e R n g a , N . A . : Braconidae (in Faune de l ' U R S S , Hymenoptera, 5, No. 3, 1941, pp. 456). — W e s m a e l , C. : Monographie des Braconides de Belgique (Bru­xelles, M. Hayez, 1835, pp. 252 et 166).