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Mealybugs & Mealybug Look-Alikes of the Southeastern United States

This publication was produced and distributed by USDA-CSREES Integrated Pest Management •Centers in cooperation with the National Plant Diagnostic Network, APHIS, the National Plant Board, and the Land Grant Universities. USDA-CSREES Critical Needs grant 2005-1763-02 ‘Early Detec-tion and Rapid Response System for the Pink Hibiscus Mealybug through Education, Training, and Implementation of Integrated Pest Management’, by J. VanKirk, G. Wisler, R. Hammerschmidt, and S. Ratcliffe, funded graphic design and printing for this publication. Content development was funded through the USDA-CSREES Southern Plant Diagnostic Network, the University of Florida, Institute of Food & Agricultural Sciences, and the Florida Department of Agriculture & Consumer Services, Division of Plant Industry.
Publication Information
For further information regarding the development of this document, contact Amanda Hodges at •[email protected](352)392-1901ext.199
Susan Ratcliffe• , University of Illinois, NCIPMC, Project Coordinator
Scott Martin• , University of Illinois, NCIPMC, Graphic Design
321INCHES5 64321MM

Amanda Hodges• , SPDN, University of Florida, Institute of Food & Agricultural Sciences, Ento-mology & Nematology Department
Gregory Hodges• , Florida Department of Agriculture & Consumer Services, Division of Plant Industry
Lyle Buss• , University of Florida, Institute of Food & Agricultural Sciences, Entomology & Nema-tology Department
Lance Osborne• , University of Florida, Institute of Food & Agricultural Sciences, Mid-Florida Research & Education Center
Authors
Photo CreditsLyle Buss, University of Florida, Institute of Food & Agricultural Sciences, Entomology & •Nematology Department - 001c,001d,005,008,009,010,011,012,013,014a,014c,015,016,017,019a,019b,020a,021a,022a,022b,023b,026,027,028,029a,029b,030,031,032,034,036,037,038a,039,CoverPhoto
Florida Department of Agriculture & Consumer Services, Division of Plant Industry •001a,002,004,007,014b,018,020b,021b,023a,024,025,033,035,040
Lance Osborne, University of Florida, Institute of Food & Agricultural Sciences, Mid-Florida •Research & Education Center - 001b
Catharine Mannion, University of Florida, Institute of Food & Agricultural Sciences, Tropical •Research & Education Center - 003, 006
Mike Merchant, Texas A&M University, Texas AgriLife Extension Service - • 038b

Page 001Mealybugs (Family Pseudococcidae)
Page 022Cottony Cushion Scale & Related Species (Family Margarodidae)
Page 024Soft Scales (Family Coccidae)
Page 038Felt Scales (Family Eriococcidae)
Page 040Cochineal Scales (Family Dactylopiidae)
Mealybugs & Mealybug Look-Alikes of the Southeastern United States

What are mealybugs?Mealybugsareplantpeststhatfeedbypiercing-suckingmouthparts.Theyareclassifiedin•thefamilyPseudococcidae.Mealybugs,alongwithsoftscales(Coccidae)andarmoredscales(Diaspididae),arethemostcommonfamiliesofscaleinsects(Hemiptera:Coccoidea).
Approximately11ofthe21speciesofexoticscalesandmealybugsthathaveestablishedinthe•United States during the past 20 years are currently considered problematic pests. Four of these speciesaremealybugs:pinkhibiscusmealybug,miscanthusmealybug,papayamealybug,andvinemealybug.
Mostexoticscalesandmealybugsofconcernfeedonawiderangeofplantsspecies,andtheyare•capableofrapidpopulationexplosions.
Awarenessofcommonfieldcharacteristicspromotesearlydetectionforpestsofconcern.•
Mealybugs of the Southeastern United States
Why are mealybugs important?
How do you identify mealybugs?Thecommonname“mealybug”isderivedfromthefinepowderyto“mealy”waxthatcoversthe•body.Adultfemalemealybugsarewinglessandthebestlifestageforidentification.Adultmalemealybugsarewinged,buttheyarerarelyseenexceptwithspecificchemical-basedtrappingsystems.Mostadultfemalemealybugsaresmall(afewmillimeterstotypicallylessthan1/2inch),and a hand lens is useful for scouting. The immature or active “crawler” stage of mealybugs is typically less than half a millimeter and may be readily dispersed to new plants via wind currents.

Body shape, size, and color.•Thenumberofwaxfilamentsprotrudingfromthesideofthebody.•Presenceandlengthofwaxfilamentsattheendofthebody(i.e.,terminalwaxfilaments).•Colorofeggs(ifpresent).•Presenceofanovisac(awaxymasscoveringtheeggs).•Stripes on the body.•Coloroffluidswhencrushed.•
Thefollowingcharacteristicsareusefulforfieldidentification:
Becausemealybugsfeedonplantjuicestheyproducehoneydew.Honeydew(asugary,sticky•substance)promotesthegrowthofblacksootymold.Sootymoldmayinhibitplantphotosynthesis.Other signs of plant damage may include stunted plant growth, wilting, and crinkled leaves. Mealy-bugscommonlyfeedonnewplantgrowth.Thepinkhibiscusmealybugalsoinjectsatoxicsalivawhile feeding, and a characteristic known as “bunchy top” is often seen on its preferred host plant, hibiscus.
Plant damage
Chemical and biological control options may be available for some mealybug and scale insect •species.Ageneralwebsitefocusingonmealybugsisavailableathttp://mrec.ifas.ufl.edu/LSO/Mealybug.htm.Biologicalcontroloptionsoftenincludespecies-specificparasitoids,andchemi-cal recommendations may vary depending on pest species or local recommendations. Your local countyextensionserviceisthebestsourceforinformationregardingchemicalcontrolsinyourarea.
General mealybug integrated pest management

Both biological and chemical control options are available for pink hibiscus mealybug. Once this •pest has established in the landscape, biological control options are most often recommended. The most common natural enemies of pink hibiscus mealybug include the mealybug destroyer, Cryptolaemus montrouzieeri, and the parasitic wasps, Anagyrus kamali and Gyranusoidea indica. Incontrast,chemicaloptionsmaybetheonlychoiceinacommercial/nurserysetting.
General mealybug integrated pest management
Pink hibiscus mealybug is considered a regulatory pest of concern; subsequently, there is a zero tol-•erance for shipment of infested plant material. Please refer to the Pink Hibiscus Mealybug website http://mrec.ifas.ufl.edu/lso/pinkmealybug.htmforthelatestup-to-dateinformationonpinkhibiscusmealybug management.
Mealybugsinthisdeckhavebeenclassifiedasthefollowing:•
Warning! This deck is not a comprehensive listing of all mealybugs. Although useful as a •fieldscreeningtool,fieldidentificationisnotdefinitivefornewcounty,host,state,orconfi-dentialrecords.Slidemountingofspecimensandidentificationbyaspecialistisnecessaryforspecies-levelconfirmation.
Warning
Exotic:NotnativetotheUnitedStates,buthasinvaded.1. Native:NativetotheUnitedStates,orduetoitswidespreaddistributionandlong-termestablish-2. ment in the United States, its origin is uncertain.

001001E X O T I C
Maconellicoccus hirsutus
Pink Hibiscus Mealybug
Common Hosts
Known Southeastern Distribution
Field RecognitionBodycolorpink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopinkincolor.Ovisacsarepresentcoveringpinktoorange eggs. Feeding from pink hibiscus mealybug can cause twisted or distorted foilage. High populations may result in leaf drop.
Over 200 known hosts occur, but the most common host detected to-date is Hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern U.S. if estab-lished populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.
EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

Pink Hibiscus MealybugMaconellicoccus hirsutus
Bunchy Top

001001E X O T I C
Maconellicoccus hirsutus
Pink Hibiscus Mealybug
Common Hosts
Known Southeastern Distribution
Field RecognitionBodypink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopink.Ovisacsarepresentcoveringpinktoorangeeggs.Feeding from pink hibiscus mealybug can cause twisted or distorted foil-age. High populations may result in leaf drop.
More than 200 known hosts occur, but the most common host detected to date is hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.
EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

Pink Hibiscus MealybugMaconellicoccus hirsutus
Droplet of Red Body Fluid

001001E X O T I C
Maconellicoccus hirsutus
Pink Hibiscus Mealybug
Field Recognition
Known Southeastern Distribution
Common Hosts
Bodypink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopink.Ovisacsarepresentcoveringpinktoorangeeggs.Feeding from pink hibiscus mealybug can cause twisted or distorted foil-age. High populations may result in leaf drop.
More than 200 known hosts occur, but the most common host detected to date is hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.
EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

Pink Hibiscus MealybugMaconellicoccus hirsutus
White Wax on Abdomen

001001E X O T I C
Maconellicoccus hirsutus
Pink Hibiscus Mealybug
Field Recognition
Known Southeastern Distribution
Common Hosts
Bodypink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopink.Ovisacsarepresentcoveringpinktoorangeeggs.Feeding from pink hibiscus mealybug can cause twisted or distorted foil-age. High populations may result in leaf drop.
More than 200 known hosts occur, but the most common host detected to date is hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.
EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

Pink Hibiscus MealybugMaconellicoccus hirsutus Eggs

Common Hosts
Known Southeastern Distribution
Field Recognition
Jatropha, Hibiscus, and Plumeria.
Established in Florida.
Bodyyellow,2-3mmlong,withmanylateral(side)waxfilaments.Ovisacspresentwithyellowtogreenishyelloweggs.Waxpatternonbodylackinganystripesonitsuppersurface(i.e.,dorsum).
Papaya MealybugParacoccus marginatus
E X O T I C002002

Papaya MealybugParacoccus marginatus
Yellow Immatures

Field Recognition
Common Hosts
Known Southeastern Distribution
Jatropha, Hibiscus, and Plumeria.
Established in Florida.
Bodyyellow,2-3mmlong,withmanylateral(side)waxfilaments.Ovisacspresentwithyellowtogreenishyelloweggs.Waxpatternonbodylackinganystripesonitsuppersurface(i.e.,dorsum).
Papaya MealybugParacoccus marginatus
E X O T I C002002

Papaya MealybugParacoccus marginatus

Field Recognition
Common Hosts
Known Southeastern Distribution
N A T I V E003003
Wide host range, may vary in local areas but common on Croton, Coleus, and Hibiscus.
Considered cosmopolitan in greenhouses and interior landscapes throughout the southeastern United States. Outdoor populations estab-lished in Florida.
Bodygrayunderneaththewaxysurfaceandwithmanylateralwaxfila-ments. Body has two darker lengthwise strips on the body surface and size is about 3 mm long. Ovisacs present with yellow eggs.
Madeira MealybugPhenacoccus madeirensis

Madeira MealybugPhenacoccus madeirensis Lengthwise Dark Stripes
Many Lateral Wax Filaments

Common Hosts
Field Recognition
Known Southeastern Distribution
004004N A T I V E
Phenacoccus solenopsis
Solenopsis Mealybug
Bodygray,about4mmlong,withmanylateralwaxfilaments.Ovisacsnotpresent.Waxontopofbodysurfaceappearsasacottonymass,horizontaldarkstripesatthetipoftheendofthebody(i.e.,theabdomenoppositeofthefeedingmouthparts).
Florida and Mississippi.
Most common on Hibiscus and other malvaceous hosts.

Solenopsis MealybugPhenacoccus solenopsis
Dark Stripes

Common Hosts
Known Southeastern Distribution
Field Recognition
005005E X O T I C
Ferrisia virgata
Striped Mealybug
Widehostrange,commononcopperleaf(Acalypha), Alternanthera, silverbuttonwood(Conocarpus),andHibiscus.
Alabama, Florida, Georgia, Louisiana, Mississippi.
Bodygray,about4-5mmlong,withoutside(lateral)wasfilaments.Two“thick”waxfilamentsarisingfromtipofabdomen.Ovisacsnotpresent.Twodarklengthwisestripesontopsurfaceofbody.Thincrystal-likefila-ments protruding from sides and top of body.

Striped MealybugFerrisia virgata Crystal-Like Filaments
Two Filaments Protruding from Abdomen

Common Hosts
Known Southeastern Distribution
Field Recognition
006006
Widehostrange,commononCitrus.SPECIALNOTE:ThePacificmealybug(Planococcus minor)cannotbeseperatedinfieldfromcitrusmealybug.TherearenoknownpopulationsofPacificmealybugoccurringintheUnitedStates.ThePacificmealybugisalsoknownasthe passionvine mealybug.
Common on ornamental plants in interior landscapes and greenhouses. Does occur outdoors in Florida.
Body color light yellow to grayish yellow, about 3 mm long, with many side(lateral)filaments.Ovisacspresent,eggsyellow.Onecentralizeddarkstripeondorsum(topsurfaceofbody).
Planococcus citri
Citrus MealybugE X O T I C

Citrus MealybugPlanococcus citri

Common Hosts
Known Southeastern Distribution
Field Recognition
007007E X O T I C
Pseudococcus longispinus
Longtailed Mealybug
Body gray in adults, about 3 mm long, yellowish in immature stages, withmanylateralwaxfilaments.Fourlongfilamentsarisingfromtipof abdomen, middle pair being very long. One centralized dark stripe located in center of abdomen. No ovisac.
Common species found in interiorscapes and greenhouses. Does occur outdoors in Florida.
Wide host range, common on Sago palms in Florida.

Longtailed MealybugPseudococcus longispinus
Lengthwise Dark Stripes
Four Anal Wax Filaments

Common Hosts
Field Recognition
Known Southeastern Distribution
008008E X O T I C
Polyphagous species, most common in Florida on tropical fruit.
Not common, does occur outdoors in Florida.
Bodylightorangetopink,about3mmlong,withmanylateralfilaments.Twolongfilamentsarisingfromtipofabdomen.Bodyfluidsbrownish.Ovisac present, but not entirely covering body.
Pseudococcus jackbeardsleyi
Jack Beardsley Mealybug

Jack Beardsley MealybugPseudococcus jackbeardsleyi
Two Anal Filaments

Known Southeastern Distribution
Common Hosts
Field Recognition
009
Polyphagous species, common on cacti.
Not common, occurs in Florida.
Bodypinktolightpurple,about3mmlong,withmanyside(lateral)filaments.Verysimilar in appearance to Jack Beardsley mealybug, but body tends to be more pinkish.Bodyisoval.Twolonganalfilamentsarisingfromtheabdomen.Nostripe on top surface of body. Eggs light colored, gray to yellow. Ovisac present, but not entirely covering body.
Pseudococcus viburni
Obscure Mealybug009N A T I V E

Obscure MealybugPseudococcus viburni

Field Recognition
Known Southeastern Distribution
Common Hosts
010010E X O T I C
Literature reports that this mealybug can occur in cacti. However, in Florida the most common hosts are Portulaca and Alternathera species.
Established in Florida.
Bodyredtopink,about3mmlong,withoutlateralfilaments.Bodyfluidsreddish. Oval to round-shaped body. No ovisac, but produces large amountsofwax.Feedingtendstoclusteratnodalregionsofplants.
Hypogeococcus pungens

Hypogeococcus pungens
Adult Female

Common Hosts
Known Southeastern Distribution
Field Recognition
011
Polyphagous species, common in Florida on roots of many of the palms.
Florida and Louisiana.
Bodygray,about2-3mmlong,withmanylateralwaxfilaments.Nostripes on body. Occurs both above and below ground.
Dysmicoccus brevipes
Pineapple MealybugE X O T I C011

Pineapple MealybugDysmicoccus brevipes

Field Recognition
Known Southeastern Distribution
Common Hosts
012012N A T I V E
Oracella acuta
Acute Mealybug
Restrictedtothefollowingpines:loblolly,longleaf,shortleaf,andVirginiapines.
Florida, Georgia, Kentucky, Louisiana, Mississippi, North Carolina, SouthCarolina,Texas,andVirginia.
Bodyredtopink,about3mmlong,withoutside(lateral)waxfilaments.Generallyfoundbothunderneathbarkandonneedlesofhosts.

Acute MealybugOracella acuta

Field Recognition
Known Southeastern Distribution
Common Hosts
013013E X O T I C
Noxious Bamboo Mealybug
Restricted to bamboos.
Common species, occurring throughout Southeastern region where bamboo is grown.
Adultbodybrown,about2-3mmlong;immaturestages(i.e.,crawlers)yellow. Generally found at the nodal regions of various bamboos. Sooty moldoccurringatthenodalregionsandlongwaxfilamentsarising from nodal areas is a common symptom.
Antonina pretiosa

Noxious Bamboo MealybugAntonina pretiosa
Long Wax Filaments

Field Recognition
Known Southeastern Distribution
Common Hosts
014Nipaecoccus nipae
Polyphagous species, most common on palms.
Florida and Louisiana.
Bodymaroontored,about3mmlong,withfewside(lateral)waxfila-ments.Dorsalwaxfilamentsoccurring.Bodyroundtooval.Noovisacproduced, males common.
Coconut MealybugE X O T I C014

Coconut MealybugNipaecoccus nipae

Field Recognition
Known Southeastern Distribution
Common Hosts
014Nipaecoccus nipae
Polyphagous species, most common on palms.
Florida and Louisiana.
Bodymaroontored,about3mmlong,withfewside(lateral)waxfila-ments.Dorsalwaxfilamentsoccurring.Bodyroundtooval.Noovisacproduced, males common.
Coconut MealybugE X O T I C014

Coconut MealybugNipaecoccus nipae
Male Puparia

Field Recognition
Known Southeastern Distribution
Common Hosts
014Nipaecoccus nipae
Polyphagous species, most common on palms.
Florida and Louisiana.
Bodymaroontored,about3mmlong,withfewside(lateral)waxfila-ments.Dorsalwaxfilamentsoccurring.Bodyroundtooval.Noovisacproduced, males common.
Coconut MealybugE X O T I C014

Coconut MealybugNipaecoccus nipae

Field Recognition
Known Southeastern Distribution
Common Hosts
015015E X O T I C
Most commonly encountered on sugarcane but Andropogon and Miscanthus grasses also may be hosts.
Established in Florida.
Bodypinktored,about4mmlong,bodyelongatedandlackinglateralwaxfilaments.Usuallyfoundonstalksorbeneathsheathsonbladesofhost plants.
Pink Sugarcane MealybugSaccharicoccus sacchari

Pink Sugarcane MealybugSaccharicoccus sacchari

Field Recognition
Known Southeastern Distribution
Common Hosts
016016E X O T I C
Varies,butgenerallypolyphagous.SPECIALNOTE:Therearemanyspecies within both Rhizoecus and Ripersiella. All may be considered pestsandsomemayrequirequarantineactions.Expertidentificationisrequired.
Varies depending on species but many are common throughout the Southeastern region.
Verysmallmealybugs(1to2mminlength),bodywhitetoyellow-ishwhite,lackingside(lateral)waxfilaments.Rootswithinfestationsofgroundmealybugsgenerallyhaveareasofwhitewaxpresentandmealybugs may be visible with use of a hand lens.
Ground MealybugsRhizoecus & Ripersiella
N A T I V E&

Ground MealybugsRhizoecus & Ripersiella

E X O T I C Field Recognition
Known Southeastern Distribution
Common Hosts
017Vryburgia amaryllidis
Lily Bulb Mealybug017
Commonlyinterceptedonbulbplants(lilybulbmealybug)andjadeplants(short-leggedmealybug).
Smallmealybugs(2-3mmlong),redtopurple.Lackingside(lateral)waxfilaments.Twothickwaxfilamentsarisingfromtipofabdomen.
Not established in Southeastern region, but established in California.

Lily Bulb MealybugVryburgia amaryllidis

Field Recognition
Known Southeastern Distribution
Common Hosts
018
Commonlyinterceptedonbulbplants(lilybulbmealybug)andjadeplants(short-leggedmealybug).
Not established in Southeastern region, but established in Arizona and California.
Vryburgia brevicruris
Short-Legged Mealybug
Smallmealybugs(2-3mmlong),redtopurple.Lackingside(lateral)waxfilaments.Twothickwaxfilamentsarisingfromtipofabdomen.
018E X O T I C

Short-Legged MealybugVryburgia brevicruris

Field Recognition
Known Southeastern Distribution
Common Hosts
019019N A T I V E
Most common host is fakahatchee and muhlygrass.
Established in Florida.
Bodygraywithwhitewax,about2-3mmlong,somewaxseemstobefilamentous.Nolateralwaxfilamentsproduced.
Stemmatomerinx acircula

Stemmatomerinx acircula

Field Recognition
Known Southeastern Distribution
Common Hosts
019019N A T I V E
Most common host is fakahatchee and muhlygrass.
Established in Florida.
Bodygraywithwhitewax,about2-3mmlong,somewaxseemstobefilamentous.Nolateralwaxfilamentsproduced.
Stemmatomerinx acircula

Stemmatomerinx acircula
Ovisac
Body
Wax FilamentsCrawler

Field Recognition
Known Southeastern Distribution
Common HostsBamboo.
Established in Florida.
Bodygrayishpink,about3mmlong,withlargeamountsofwhitewaxvisibleonhostplant.Bodywithfewside(lateral)waxfilaments.Noovisacproduced.SPECIALNOTE:Thisspeciesisfoundunderneath sheaths of bamboo. It can be a severe pest.
020020E X O T I C
Palmicultor lumpurensis

Palmicultor lumpurensisAdult
Crawlers

Field Recognition
Common Hosts
Known Southeastern Distribution
Bamboo.
Established in Florida.
Bodygrayishpink,about3mmlong,withlargeamountsofwhitewaxvisibleonhostplant.Bodywithfewside(lateral)waxfilaments.Noovisacproduced.SPECIALNOTE:Thisspeciesisfoundunderneath sheaths of bamboo. It can be a severe pest.
020020E X O T I C
Palmicultor lumpurensis

Palmicultor lumpurensis
Infested Bamboo

Field Recognition
Known Southeastern Distribution
Common Hosts
021021E X O T I C
Various palms.
Established in Florida.
Bodyreddishbrowntopink,about3mmlong,withside(lateral)waxfilaments.Noovisacproduced.
Palmicultor browni

Palmicultor browni

Field Recognition
Known Southeastern Distribution
Common Hosts
021021E X O T I C
Various palms.
Established in Florida.
Bodyreddishbrowntopink,about3mmlong,withside(lateral)waxfilaments.Noovisacproduced.
Palmicultor browni

Palmicultor browni

Known Southeastern Distribution
Common Hosts
022022E X O T I C
Icerya genistae
Superficiallyresemblesthecottonycushionscale.Bodylightorangetopink,legsblack.Bodymaybecoveredwithawhitewax.Longovisacproduced(3-5xbodylength)andhelderecttohorizontalfrombody.
Established in Florida.
Wide host range, commonly collected on legumes.
Field Recognition

Icerya genistae

Known Southeastern Distribution
Common Hosts
022022E X O T I C
Icerya genistae
Superficiallyresemblesthecottonycushionscale.Bodylightorangetopink,legsblack.Bodymaybecoveredwithawhitewax.Longovisacproduced(3-5xbodylength)andhelderecttohorizontalfrombody.
Established in Florida.
Wide host range, commonly collected on legumes.
Field Recognition

Icerya genistae
Immatures

023023E X O T I C Field Recognition
Known Southeastern Distribution
Common HostsPolyphagous species, common on Citrus and Pittosporum.
A common species. Distribution scattered throughout the Southeastern region at different periods.
Bodyisorangewithsomewhitewaxpresentontopsideofbody.Legsareblack.Producesagroovedovisacthatisaboutthesamesize(or2x)asthebodyandgenerallytapersdowntowardthehostsubstrate.
Cottony Cushion ScaleIcerya purchasi

Cottony Cushion ScaleIcerya purchasi
Ladybugs Feeding on Cottony Cushion Scale

023023E X O T I C Field Recognition
Known Southeastern Distribution
Common HostsPolyphagous species, common on Citrus and Pittosporum.
A common species. Distribution scattered throughout the Southeastern region at different periods.
Bodyisorangewithsomewhitewaxpresentontopsideofbody.Legsareblack.Producesagroovedovisacthatisaboutthesamesize(or2x)asthebodyandgenerallytapersdowntowardthehostsubstrate.
Cottony Cushion ScaleIcerya purchasi

Cottony Cushion ScaleIcerya purchasi
Immatures

Field Recognition
Known Southeastern Distribution
Common Hosts
024
Maples, hollies, and dogwoods.
Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.
Body light brown to yellowish brown with young adults, dark brown on olderfemales.Body2.5-4.5mmlong.Ovisacproducedandgenerally found on underside of leaves.
Pulvinaria acericola
Cottony Maple Leaf ScaleN A T I V E024

Cottony Maple Leaf ScalePulvinaria acericola

Field Recognition
Known Southeastern Distribution
Common Hosts
025Pulvinaria ericicola
Azalea, lowbush blueberry and rusty lyonia.
Alabama, Florida, and Virginia.
Bodyredtoyellowandcoveredinglassywax.Body1.5-3.5mmlong.Ovisacproducedand2-4xaslongasbody.
Cottony Azalea Scale025N A T I V E

Cottony Azalea ScalePulvinaria ericicola

Field Recognition
Known Southeastern Distribution
Common Hosts
026E X O T I C026 Pulvinaria psidii
Green Shield Scale
Wide host range, common on Pittosporum, Citrus, Ficus species, and mango.
Alabama, Florida, Georgia, and Mississippi.
Adultfemalelightgreen,body2.0-4.5mmlong,ovisacproducedandwithoutnoticeablegrooves,approximately3xlengthofbody.Bodycoloris usually green but may seem somewhat brown depending on the host plant.

Green Shield ScalePulvinaria psidii
Ants Tending Scales for Honeydew
Ovisac

Field Recognition
Known Southeastern Distribution
Common Hosts
027E X O T I C027 Pulvinaria urbicola
Urbicola Soft Scale
Body of adult female brown to green, body 1.5 - 3.0 mm long, ovisac producedandwithoutnoticeablegrooves,approximately1.5-3xbodylength. Occurs on foliage but also near root line.
Alabama,Florida,Louisiana,andTexas.
Common on seagrape, guava, and Alternanthera species.

Urbicola Soft ScalePulvinaria urbicola
Grooved Ovisac
Brown Body

Field Recognition
Known Southeastern Distribution
Common Hosts
028N A T I V E028
Neopulvinaria innumerabilis
Cottony Maple Scale
Largescale(3-6mmlong)withcolorpatterndifferingdependingonhost, often blending in with host bark. Ovisac produced, but grooves areusuallynotclearlyvisible.Ovisac2xbodylength.Generallyfoundon woody portions of host plant material.
Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, Tennessee, and Virginia.
Polyphagous, common on dogwoods, maples and pecans.

Cottony Maple ScaleNeopulvinaria innumerabilis
Ovisac
Female Body

Common Hosts
Known Southeastern Distribution
Field Recognition
029E X O T I C029 Philephedra tuberculosa
Philephedra Scale
Largescale(3-6mmlong).Youngfemalesaregreensimilartogreenshieldscale.Ovisacsarelong(3-5xbodylength)andusuallyhavea“wet” appearance.
FloridaandTexas.
Wide host range but common on Acalypha, Codiaeum, and Conocarpus species.

Philephedra ScalePhilephedra tuberculosa
Ovisac

Field Recognition
Known Southeastern Distribution
Common Hosts
029E X O T I C029 Philephedra tuberculosa
Philephedra Scale
Largescale(3-6mmlong).Youngfemalesaregreensimilartogreenshieldscale.Ovisacsarelong(3-5xbodylength)andusuallyhavea“wet” appearance.
FloridaandTexas.
Wide host range but common on Acalypha, Codiaeum, and Conocarpus species.

Philephedra ScalePhilephedra tuberculosa
Green Female Body
Immature Scales

Field Recognition
Known Southeastern Distribution
Common Hosts
030E X O T I C030 Protopulvinaria pyriformis
Pyriform Scale
Body varies from tan to green to brown. Body about 2.5 mm long, pyriform. Ovisac produced, small and located beneath the scale insect. Long triangular anal plates visible in middle of top surface of body. Found on foliage of hosts.
Alabama, Florida, Georgia, Louisiana, North Carolina, South Carolina, Texas,andVirginia.
Wide host range, commonly found on Pittosporum, Ficus species, Schefflera, and tropical fruit.

Pyriform ScaleProtopulvinaria pyriformis
Ovisac
Triangular Anal Plates

Common Hosts
Known Southeastern Distribution
Field Recognition
031E X O T I C031 Milviscutulus mangiferae
Mango Shield Scale
Very similar to that of pyriform scale, body yellowish green, body size 2 - 3.5 mm long, pyriform. Ovisac produced and located beneath adult female. Found on foliage of hosts.
FloridaandTexas.
Polyphagous species, commonly found on Citrus, Ficus, Gardenia, jasmine and tropical fruit.

Mango Shield ScaleMilviscutulus mangiferae

Common Hosts
Known Southeastern Distribution
Field Recognition
032E X O T I C032 Ceroplastes ceriferus
Indian Wax Scale
Bodyred,coveredinathickwhitewetwax,normallywithananteriorprojecting “horn.” Body size 3-12 mm long. Found on woody portions of host plants.
Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.
Wide host range, common on hollies, camellias, and magnolias.

Indian Wax ScaleCeroplastes ceriferus

Field Recognition
Known Southeastern Distribution
Common Hosts
033E X O T I C033 Ceroplastes rusci
Fig Wax Scale
Similar in appearance to barnacle scale. Body red, covered in a thick pinkishwhitewax.Waxcoveringseemsplatedfromlateralview.Body2-3 mm long. Occurring on foliage and woody portions of host plant material.
Florida.
Wide host range, most commonly collected on Ficus species.

Fig Wax ScaleCeroplastes rusci
Plated Wax Cover

Field Recognition
Known Southeastern Distribution
Common Hosts
034E X O T I C034 Ceroplastes cirripediformis
Barnacle Scale
Bodymaroontored,coveredinathickbuff-coloredwax.Withoutananterior projecting horn. Appearing “plated” from lateral view. Body 1-6 mm long. Immatures maroon. Found on woody portions of host.
Alabama, Florida, Georgia, Louisiana, Mississippi, North Carolina, SouthCarolina,andTexas.
Wide host range, common on Ficus species, Pittosporum, hollies and Citrus.

Barnacle ScaleCeroplastes cirripediformis
Maroon Immature Stages

Field Recognition
Known Southeastern Distribution
Common Hosts
035E X O T I C035 Ceroplastes dugesii
Duges Wax Scale
Bodyred,coveredinthickgrayishwhitetobuffcoloredwaxcover.Body3-8 mm long. Found on woody portions of host plants.
Florida.
Wide host range, common on Bursera and Piper species.

Duges Wax ScaleCeroplastes dugesii

Field Recognition
Known Southeastern Distribution
Common Hosts
036E X O T I C036 Ceroplastes floridensis
Florida Wax Scale
Bodyred,coveredinathickgrayishtopinkishwhitewax.Generallyflat.Body2-4mmlong.Immaturesoftenresemblesmallwhitestars.Foundon twigs and foliage of hosts.
Alabama, Florida, Georgia, Louisiana, Mississippi, North Carolina, SouthCarolina,Texas,andVirginia.
Wide host range, common on hollies, Schefflera, Pittosporum, cycads, Citrus, and tropical fruit.

Florida Wax ScaleCeroplastes floridensis
Adult Female
Immatures

Field Recognition
Known Southeastern Distribution
Common Hosts
037E X O T I C037 Ceroplastes rubens
Red Wax Scale
Bodyred,coveredinathickredtopinkwax.Body2mmlong.Waxpentagonal in dorsal view. Occurring on foliage of host plants.
Florida.
Wide host range, common on Citrus and other tropical fruit.

Red Wax ScaleCeroplastes rubens

038
Common Hosts
Known Southeastern Distribution
Field RecognitionN A T I V E038 Eriococcus azaleae
Azalea Bark Scale
Body dark red to purple. Eggs and immature crawler stages pink. Bodyapproximately3-4mmlong,ovisacproducedandisashinywhite(almostsilvery).Generallyfoundinnodalareasor“forks”oftwigsandbranches.
Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.
Wide host range but most commonly collected on Azalea.

Azalea Bark ScaleEriococcus azaleae

038
Common Hosts
Known Southeastern Distribution
Field RecognitionN A T I V E038 Eriococcus azaleae
Azalea Bark Scale
Body dark red to purple in color. Eggs and immature crawler stages pink.Bodyapproximately3-4mmlong,ovisacproducedandisashinywhite(almostsilvery).Generallyfoundinnodalareasor“forks”oftwigsand branches.
Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.
Wide host range but most commonly collected on Azalea.

Azalea Bark ScaleEriococcus azaleae
Adult Female
Pink Immatures
Adult Female

039N A T I V E039 Eriococcus quercus
Oak Eriococcin
Field RecognitionBodyisdarkredtopurple.Bodyapproximately3-4mmlong,ovisacproducedandisashinywhite(almostsilvery).
Known Southeastern DistributionAlabama,Florida,Georgia,Louisiana,Mississippi,Texas,andVirginia.
Common HostsOaks.

Oak EriococcinEriococcus quercus

040Dactylopius confusus040
N A T I V E
California Cochineal Scale
Field RecognitionBodymaroontopurple,coveredwithsticky-stringywax.Bodyfluidsred. Found on pads of cacti.
Known Southeastern DistributionFlorida,Georgia,andTexas.
Common HostsOpuntia cacti.

California Cochineal ScaleDactylopius confusus