Karoo Basin

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    Facultatea de Geologie si Geofizica

    Universitatea Bucuresti

    Geologie istorica

    Evolution of the Karoo

    sedimentary basin

    Autori:  Dobrin Sebastian 303B Olteanu Dan-Cristian 302A

     Ene Vlad 302A Voiculescu Alexandru 302B

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    Abstract 

    The main purpose of this paper is to present the evolution of the Karoo sedimentary basin,

    from its inception to it’s extinction. Situated in South Africa, it is a retroarc foreland basin developed

    in front of the Cape Fold Belt, wich is made up of rock formations from Late Carboniferous to Early

    Jurassic. In fact, it forms the thickest and stratigraphically most complete megasequence of several

    depositories of Permo-Carboniferous to Jurassic age in south-western Gondwana.

    Due to it being a foreland basin, there is a clear distinction between the southern and the

    northern part. The southern part is close to the CFB, and thus it is considered proximal, while the

    northern part is distal. A lack of correlation between formations can be observed, and also a thinning

    of the formations from south to north

    The climates in wich the deposition took places was quite different in Karoo’s history, andthus a shift from glaciar to deep water, shallow water and finally arid depositional system can be

    observed. Accomodation was also affected by different processes at different moments in Karoo’s

    history.

    The Karoo basin is famous for being rich in fossil-bearing rocks having quite an importance in

    dating some formations.

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    Tectonic settings

    The Karoo basins (fig 1) of south-central Africa evolved during the first-order cycle of supercontinent

    assembly and breakup of Pangea,under the influence of two distinct tectonic regimes sourced fromthe southern and northern margins of Gondwana.

    The southern tectonic regime was related to processes of subduction and orogenesis along the

    Panthalassan (palaeo-Pacific) margin of Gondwana, which resulted in the formation of a retroarc

    foreland system known as the ‘‘main Karoo’’ Basin, with the primary subsidence mechanisms

    represented by flexural and dynamic loading. This basin preserves the reference stratigraphy of the

    Late Carboniferous –Middle Jurassic Karoo time, which includes the Dwyka, Ecca, Beaufort and

    Stormberg lithostratigraphic units.

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    North of the main Karoo Basin, the tectonic regimes were dominated by extensional or

    transtensional stresses that propagated southwards into the supercontinent from the divergent

    Tethyan margin of Gondwana.

    Superimposed on the tectonic control on basin development, climatic fluctuations also left a mark on

    the stratigraphic record, providing a common thread that links the sedimentary fill of the Karoobasins formed under different tectonic regimes. As a general trend, the climate changed from cold

    and semi-arid during the Late Carboniferous –earliest Permian interval,to warmer and eventually hot

    with fluctuating precipitation during the rest of Karoo time.

    Due to the shifts in tectonic and climatic conditions from the southern to the northern margins of

     Africa during the Karoo interval,the lithostratigraphic character of the Karoo Supergroup also

    changes significantly across the African continent. 

    For this reason, the Karoo basins sensu stricto(fig.1), which show clear similarities with the main

    Karoo Basin of South Africa, are generally restricted tosouth-central Africa, whereas the Karoo-agesuccessions preserved to the north of the equator are distinctly different.

    The Cape and Karoo basins formed within the continental interior of Gondwana. Subsidence resulted

    from the vertical motion of rigid basement blocks and intervening crustal faults. Each basin episode a

    three-stage evolution consisting of crustal uplift, fault-controlled subsidence, and long periods of

    regional subsidence largely unaccompanied by faulting or erosional truncation. The large-scale

    episodes of subsidence were probably the result of lithospheric deflection due to subduction-driven

    mantle flow. (fig 2)

    The Karoo basin is a cratonic cover that mimics the underlying basement blocks. The Permian Ecca

    and lower Beaufort groups were deposited in a southward-deepening ramp syncline by extensional

    decoupling on the intra-crustal decollement. Reflection seismic and deep-burial diagenetic studies

    indicate that the Cape orogeny started in the Early Triassic. Deformation was partitioned into

    basement-involved strike-slip faults and thin-skinned thrusting. Uplift of the Namaqua basement

    resulted in erosion of the Beaufort cover.

    East of the Cape fold belt, contemporaneous subsidence and tilting of the Natal basement created a

    late Karoo transtensional foreland basin, the Stormberg depocentre. Early Jurassic tectonic resetting

    and continental flood basalts terminated the Karoo basin.

    2.  Accumulation of Karoo aged successions in Africa corresponds to the Pangean first-order cycle of

    supercontinent assembly and breakup. 

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    The tectonic regime during Karoo time was defined by

    compression and accretion along the southern margin

    of Gondwana coeval with extension propagating into

    the supercontinent from its Tethyan margin

    (Wopfner, 1994, 2002). This unique combination of

    tectonic stresses sourced from the convergent and

    divergent margins of Gondwana resulted in the

    formation of different basin types across Africa, with

    accommodation generated by tectonic and dynamic

    loads in the south, and riftingto the north. The

    extensional field in central and northern Africa during

    Karoo time is explained by the updoming caused by

    the self-induced Pangean heat anomaly that followed

    the onset of supercontinent assembly (Wopfner,

    1990, 1994; Veevers and Powell, 1994). Tensionalregimes initiated during that time resulted in the

    formation of the early Tethyan spreading centre, and

    continued to govern the Karoo deposition until the

    breakup of Gondwana in the Middle Jurassic. Starting

    with the onset of the Late Carboniferous, tensional

    stresses propagated gradually to the south from the

    Tethyan margin,controlling the deposition of Karoo

    sediments in grabens and subsequent rift structures.

    The age of the extensional structures in southern

    Africa is thus inferred to be younger than in central

    and northern Africa (e.g., Bordy and Catuneanu,

    2002c).

    Fig 2. Model evolution of Cape and Karoo

     basins

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    Discussion and conclusions

    The formation of the Karoo-age basins of Africa took place during the Late Paleozoic –Early Mesozoic

    interval,when the Pangea supercontinent reached its maximum extent.Tectonism was the primarycontrol on accommodation in the Karoo basins, with subsidence mechanisms (fig.3) ranging from

    flexural in the south, in relation to processes of subduction and orogenesis along the palaeo-Pacific

    margin,to extensional in the north, propagating southwards from the divergent Tethyan margin. The

    interplay of these tectonic mechanisms, combined

    with the influence exerted by the inherent structures

    of the underlying Precambrian basement, resulted in

    the formation of discrete depozones that follow

    regional tectonic trends. Sedimentation patterns in

    these Karoo basins were further influenced by a shift

    in climatic regimes, from early cold conditions during

    the Late Carboniferous –earliest Permian interval, to

    warmer and eventually hot climates with fluctuating

    precipitation during the rest of Karoo time. The

    climatic background provided a common thread for

    the sedimentary fill of all Karoo basins, which resulted

    in the development of similar depositional trends

    across much of south-central Africa in spite of the

    change in tectonic regimes.

    The lacuna which followed the coalescence of

    Pangea led to extensive denudation and

    peneplanation of central and eastern African cratonic

    regions. The first release of heat from the self-induced

    Pangean heat anomaly (Veevers et al., 1994a,b)

    caused updoming, followed by rifting in the eastern

    African and Malagasy region. Rust (1975) and

    subsequently Tewari and Veevers (1993) suggested

    that the shape of Pangean basins (like the Karoo

    basins) was largely determined by the structure of the

    basement.

    Syneclisic type Karoo basins formed by flexure and

    thermal sagging over isotropic, mainly Archean

    basement (e.g., Congo Craton), whereas linear, east

    African type basins developed 246 O. Catuneanu et al.

    / Journal of African Earth Sciences 43 (2005) 211 –253

    by fracturing over anisotropic, mainly Proterozoic fold

    belts (Wopfner, 1993, 1994). Similarly, Radelli (1975)that

    syn-depositional faulting within the MorondavaBasin followed old zones of weaknesses within the

    underlying Mozambique Fold Belt. More recently, Visser and Praekelt (1996) suggested a similar

    Fig 3. Geodynamic history of the Cape

    and Karoo basins.

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    model for the breakup between East Africa and Madagascar/India.Whilst the concept of rift

    directions being inherited from tectonic trends within the underlying basement holds true in a

    general way, there are a number of cases where Karoo rifts cut right across older fold structures. One

    of the best examples is the Ruhuhu Basin, the rift structures of which cut the fold fabric of the

    Palaeoproterozoic Ubendian Metamorphic Complex almost at a right angle (Wopfner and Diekmann,

    1996)

    Although the dominant trend of the eastern African and Malagasy graben structures is northeast,

    there are a number of shorter structures with northerly or even northwesterly trends. Some of them

    may be simple accommodation faults, but the great thickness of sediments accumulated against

    some of them suggests a pull-apart mechanism. It may be concluded therefore that East

    African/Malagasy rifts were not just simple tensional systems but that they were governed by a

    transtensional stress field with a clear left-lateral component (Wopfner et al., 1993).North of Kenya,

    at the point where later in the Jurassic the northwest trending, failed rift of the Anza Trough

    developed (Reeves et al., 1986), the Karoo rift system deviated from the trend of the Mozambique

    Belt altogether and presumably continued northeastward to southern Oman.Such a continuation is

    indicated not only by various palaeobiological assemblages (see Wopfner et al., 1993;Wopfner, 1994,

    1999, 2002 for references) but also by the relationship of the glacigene deposits of southern Oman to

    the Huqh-Haushi uplift (Lee, 1990). Permian pillow lavas overlain by radiolarian cherts in the Oman

    Mountains show that here the intracratonic rift of the Malagasy Trough merged with the

    Neotethyan Ocean.In summary it can be stated that the formation of the Karoo rift basins of eastern

    Africa resulted from left lateral transtension between India/Madagascar and Africa.

    Although the onset of rifting was triggered by an anomalous heat accumulation under the insulating

    blanket of the vast continental crust, transtension was the controlling mechanism for the

    establishment of rift directions. The forces were created by the opening of Neotethys combined with

    the right-lateral rotation between Gondwana and Laurasia (Veevers et al., 1994a). The release of

    heat from the self-induced heat anomaly is considered the cause for the sagging and the formation

    of the syneclise-type basins on the western side of African Gondwana.At the same time with the

    manifestation of dominantly extensional or transtensional tectonic regimes across much of

    Gondwana, the evolution of the southern part of the supercontinent was primarily linked to the

    formation of the Pan-Gondwanian fold-thrust belt, which provided the supracrustal load for the

    flexural subsidence recorded in the main Karoo Basin. Flexural loading, coupled with dynamic

    subsidence related to the process of subduction beneath Gondwana, conferred the main Karoo Basin

    of southern Africa unique characteristics that set it apart from all other Karoo-age basins of Africa.This retroarc foreland basin preserves the reference stratigraphy for the Late Carboniferous – Middle

    Jurassic interval of Gondwana, which includes the regionally correlatable Dwyka, Ecca, Beaufortand

    Stormberg lithostratigraphic units.

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    Distribution of the main lithostratigraphic units

    Fig.4. Distribution of the main lithostratigraphic units of the Main Karoo Basin

    Dwyka Group

    The initiation of sedimentary processes of the Dwyka Group was estimated at about 300 Myr

    (Moscovian) after a 30 Myr sedimantation break (Visean). The Dyka facies (tillites, cyclically grading

    upwards into finer-grainde clastic rocks) indicates a glacial environment, with deposition from both

    grounded and floating ice. The trend in which grounded and floating ice is separated is from north,

    respectively south, althought alternate deposition from grounded and floating ice has been recorded

    throughout the Karoo Basin. The foredeep accumulated about 800 m of silt-dominated marinediamictites with dropstones (Fig. 5) derived from the floating ice. In the south part of the basin the

    thickness of the fining-upward cycles varies from 60 to 100 metres featuring an uniform character

    and lateral continuity of the layers, suggesting that deposition from floating ice was taking place

    within a large marine basin, each cycle displaying a transition from terrestrial to subaqueous

    moraines at the base, to glaciolacustrine shales at the top. A general feature of the foredeep

    succession is the uniform character and lateral continuity of the diamictite layers, suggesting

    deposition from suspension in a relatively low energy environment, a large marine basin. Processes

    of resedimentation of the initial fallout deposits by debris flows have also been documented in this

    glacio-marine environment.

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    Fig.5. Dropstone

    Contrasting to the Dwyka succesion in the south, in the

    north, the lateral correlation of the facies were very difficult

    to comprehend due to the irregularity in thickness and

    complex facies relationships of the succession. Despite this,

    two finning-upwards cylces have been identified, each of

    them comprising a basal massive till, a result of continental

    glaciation, grading upwards into a stratified terminal zone

    deposited from floating ice. Another interesting aspect is

    that in the north part the top glacial-formed layers are

    younger than the southern top of the Dwyka group,

    meaning that the continental glaciation could have last

    longer in the north due to a higher altitude than in thesouth where the climate was warmer because of the

    marine environment during the Early Permian deglaciation.

    As the result of deglaciation of the northern zone of the

    basin, the uppermost layer of Dwyka succesion is

    compound by coal-bearing fluviodeltaic sequences. 

    In the north, the limit is represented by ravinement surface generated during the marine

    transgression of the Ecca Sea. In the south, a precise boundary between Dwyka and Ecca Groups is

    difficult to trace as the transition between glaciomarine and marine environments is gradual, the

    percentage of dropstones gradually deacreases upwards in parallel with the disintegration of the

    floating ice. 

    Ecca Group

    The term Ecca was first used to descrise argillaceous sedimentary strata exposed in the Ecca pass, in

    the Eastern Cape Province, South Africa. The Ecca group occurs between the Late Carboniferous

    Dwyka Group and the Late Permian-Middle Triassic Beaufort Group, encompassing the Permian time

    slot of the Karoo lithologies. Its maximum thickness is located in the southern part of the Karoo

    basin, in the foredeep, being of aproximately 3000 m. Most age determinations are based of fossil

    wood biostratigraphy and palynology. The absolute ages are generally in agreement with the 290 Ma

    aged inferred from palynomorphs. The Ecca group is mainly composed of mixed clastic sediments

    with some minor carbonates.

    The Ecca Group is diachronous, meaning that the distal region is younger than the proximal region,

    making the Volkrust Fm. correlative with the lowermost Adelaide Subgroup – Koonap Formation (Fig.

    6). With that being known, the age of the proximal region of the Ecca Group is from Artinskian to

    Kazanian, and in the distal region, the Ecca Group begins in Kungurian and finishes somewhere in

    Tatarian (Fig. 6). In the proximal region, the sedimentary processes was dominated by deep marine

    conditions, forming Bouma sequences in Collingham Formation. The Collingham Formation is made

    up of alternating siltstone and shale with yellowish layers of up. It is interpreted as a distal submarine

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    fan facies associated with pelagic sedimentation and wind-blown interbedded volcanic ash. The age

    is Late Permian, probably Ufimian.

    The Fort Brown Formation hasbeen formed in a shallow water,

    coresponding with a regressive

    marine environment in the Late

    Permian. This formation is

    represented by greenish-grey shale

    with subordinate sandstone

    becoming more prominent

    upwards. It comprises two

    coarsening-upward-sequences.

    Both are several 100 m thick. Their

    basal parts consist of dark pelites

    and rhythmic alternating sand- and

    mudstones layers of a prodelta.

    They pass over into sandy distal

    bar and distributary mouth bar

    deposits of the deltafront. The age

    of the Fort Brown Formation is

    Late Permian.

    Fig. 6. Lithostratigrahy of the Karoo

    Supergroup, along the profile

    shown on Fig. 4

    In the northern part of the basin, the Pietermaritzburg Fm. consists mainly of shales accumulated in

    a moderate to deep marine environment, of Kungurian age. The Vryheid Formation consist of

    fluviodeltaic deposition, containing the only nonmarine sedimentary deposits of the Ecca Group.Contrary to this, the correlation with the proximal Ecca Group is made with Whitehill Formation,

    apparently the deepest marine environment from this period, being recorded at Kungurian age. The

    Volkrust Fm. is accumulated in a deep to shallow marine environment, and is made up

    predominantly of dark shales with intercalations of fine-grained sandstone. The age of this formation

    is estimated as Umifian to Tatarian. The transgression of the Ecca Sea over the Vryheid Fm. formed

    an uncomformable boundary between the fluviodeltaic facies and the marine transgressive facies of

    Volkrust Fm.

    The history of the Ecca formation is quite hard to understand due to the complexity of structures

    and depositional enviromanets. After the Dwyka glaciation, the main facies types are meltwater rain-deposits, glaciofluvial and glaciolacustrine deposits, found in the norther Karoo, with deep-water

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    marine sedimentation in the foredeep. This is still in the underfilled settings of the basin, where the

    foredeep accumulates pelagic to gravity flow sediments. Accomodation is obtained by dynamic

    subsidence wich outpaces the rates of flexural uplift, leading to the lowering of the peripheral bulge

    below the sea level and the manifestation of the basin-wide transgressions of the interior seaway. At

    a higher frequency level, fluctuations in sediment supply and in the balance between the rates of

    flexural tectonics and dynamic loading may result in forced regressions on the side of the basin that

    is subject to flexural uplift, coeval with transgressions or normal regressions of the opposite shoreline

    of the interior seaway.

    The latter is the case with the age-equivalent Ripon and Vryheid formations in the Karoo

    Basin, when submarine fans in the foredeep formed at the same time as the progradation of fluvial –

    deltaic sequences over the forebulge. The filled foreland stage is represented by the Fort Brown and

    the Volkrust Formations, when the deep-water environment specific to the foreland is replaced by a

    shallow-marine environment. Also the forebulge is lowered below sea leve wich means that the

    dynamic subsidence is outpacing the rates of flexural uplift, which in turn implies base level rise

    across the entire foreland and sedimentation rates that are within the range of variaton of the rates

    of base level rise. 

    The boundary between Ecca Group and Adelaide Subgroup is comformable, both in proximal and

    distal regions.

    Beaufort Group

    The term Beaufort “Beds” was first introduced to describe

    sedimentary rocks of the lower part of the Karoo series at

    Beaufort West in South Africa, but has been expanded toinclude a wider range of fluviatile deposited Permo-Triasic

    rocks in the main Karoo. The absolute age of the Beaufort

    Group is not preciselly placed but the wealth of fossils

    tetrapods has allowed biostratigraphic subdvisions into eight

    biozones which have allowed the correlation with other

    better dated fossil bearing successions in the world.

    These rocks cover about 200000 square kilometers,

    making up about 20% of the local surface area of South

    Africa, with a maximum thickness of 7000 m in the

    foredeep, thinning rapidly northwards. Beaufort Group

    strata consist predominantly of mudstones and siltstones

    with subordinate lenticular and tabular

    Fig.7. Beaufort shale

    sandstones deposited by a variety of fluvial systems (Fig. 7).The boundary with the underlying Ecca

    Group is transitional and diachronous, recording a gradual change from deltaic to fluvial depositional

    systems. The climates had become semi-arid with seasonal rainfall. The presence of desert rose-

    gypsum, dessication cracks, palustrine carbonate beds and pedogenic carbonate horizons is a direct

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    indicator of a semi-arid region. Lower Beaufort sediments came from different margins of the Karoo

    basin, giving rise to pebbly sandstones in the north and fine-grained sandstones elsewhere. The

    Beaufort Groups is divided into two major subgrops : the Adelaide and Tarkastad Subgroups.

    Adelaide Subgroup

    By the time the Adelaide Subgroup had formed the nonmarine conditions of sedimentation were

    established in all Karoo Basin. The proximal facies of the Adelaide Subgroup includes the Koonap,

    Middleton and Balfour Formations being the correspondent for Tatarian age and the distal facies is

    represented by Normandien Fm. – Upper Tatarian – Lower Scythian (Fig. 6). The base of the Adelaide

    Subgroup is conformable with the lower Ecca Group in both distal and proximal facies, due to the

    transition marine-nonmarine environment; however the subgroup in its lower part is diachronous.

    The Koonap Formation has been formed in high-energy systems (braided river) grading upwards

    innto lower energy meandering systems. It is made up of greenish silty mudstones and sandstones.

    The Middleton Fm. composed by a finning-upward succesion made up of marron and greenish-grey

    mudstones interbedded with sandstones and is deposited in low-energy meandering and lacustrine

    systems. The Contact between the Middleton and the overlying Balfour Fm. is unconformable due to

    the changes in the facies and sedimentation energy rate, from low-energy meandering facies to high-

    energy braided facies. Similar to the lower sequence of Koonap and Middleton Formations, the

    Balfour Fm. presents changes in the depositional environment fom braided rivers grading upwards

    into meadnering systems. It also represents a finnin-up sequence, bounded by subaerial

    unconformities both at the top and at the base. They are composed of yellowish and bluish-greenish-grey sandstones interbedded with dark mudstones, and their age is Tatarian to early Scythian. The

    presence of calc-alkaline volcaniclastic detritus and ‘‘cherts’’ of tuffaceous origin (Ho-Tun, 1979)

    suggests that the provenance rocks in the southwest may have included an active andesitic volcanic

    chain located on the eastern side of the Andean Cordillera in South America and West Antarctica.

    The Normandien Formation is also the result of a fluviatile system, but showing evidence of wide

    semiarid floodplains, being made up of interbedded sandstones and mudstones deposited by

    meandring streams with channels flanked by this semiarid floodplains. It correlates with the upper

    part of the Balfour Fm., and therefore the age is tartarian-Early Scythian.

    Tarkastad Subgroup

    The Tarkastad proximal facies is composed of the Katberg and Burgersdorp formations. The distal

    facies is made up of the Driekoppen and Verkykerskop formations. It's age is Scythian to early

    Anisian.

    The Katberg is made up of thick, laterally extensive, light olive grey, coarse-grained sandstones,

    composed of transverse and longitudinal bar macroforms, which are internally structuredpredominantly by horizontal and trough cross-stratification. It unconformably overlays the Balfour

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    Formation. It was deposited in a shallow braided environment with pulsatory discharge. Abandoned

    channel fills are also present and are represented by thin sequences of red-olive yellow mudstones.

    The Burgersdrorp Formation conformably overlies the Katberg Formation and consists

    predominantly of thick fining-upward units of laterally inextensive, olive grey, fine to medium

    grained sandstones overlain by red-maroon coloured siltstones and mudstones. These fining-upwardsequences are thought to represent mixed-load meandering river and floodplain deposits and

    preserve a fauna assignable to the Cynognathus Assemblage Zone. The source area is composed

    predominantly of granitic, metamorphic and alkaline volcanic rocks.

    The Verkykerskop Fomation consists predominantly of thin, laterally extensive, medium to fine

    grained sandstones dominated by transverse bar macroforms wich are internally structured by

    planar cross-bedding. It is the northern equivalent of the Katberg formation. The Driekoppen

    formation is composed of thin fine-grained channel sandstones, internally structured by horizontal

    stratification, overlain by thick, massive to diffusely laminated siltstones and mudstones. They are

    believed to represent suspended-load-dominated meandering river deposits.

    The Tarkastad Subgroup may be regarded as a single finning-upward sequence : sandstone

    dominated braidplain deposits at the base, grade upwards into mudstone-dominated floodplain

    deposits associated with meandering river systems.

    The Beaufort Group marks the begging of the overfilled basin stage of the Karoo basin. Overfilled

    foreland systems are dominated by non-marine environments, and reflect stages in the evolution of

    the basin when sediment supply outpaces the available accommodation. The accommodation in this

    case is only achieved through tectonic processes.

    Stormberg Group

    A major stratigraphic gap, corresponding to the late Anisian-Ladinian interval separates these strata

    from underlying Tarkastad. In contrast to the other groups, there are no proximal or distal facies.

    Nevertheless, the entire group may be considered a distal facies for the Stormberg Group did not

    extend to the Cape Fold Belt. Furthermore, studies show that Dwyka, Ecca and Beaufort rocks are the

    sediment sources for the Stormberg Group, reworking this older Karoo sequences. The base-Moltenoangular unconformity, well developed in many basins, indicates a significant tectonic event across

    the region to usher in Stormberg sedimentation.

    The sequence discussed here is constituted by tree different formations: Molteno, Elliot and

    Clarens, from Middle Triassic to Middle Jurassic. (Fig. 6) 

    The Molteno Fm. is composed by two coarsening upward sequences that formed deposits

    interpreted as the fills of abandoned channel tracks and within ponded bodies of water on the

    braidplain. The formation is composed predominantly of tabular sheets of medium to coarse grained

    sandstone internally structured by horizontally and cross-stratified macroforms. Siltstone, mudstone

    and coal deposits also occur but are far less abundant. The Bamboesberg Member is dominated byolive grey fine to medium grained sandstones, internally structured

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     equally by horizontal and trough cross-stratification. The overlying Indwe Sandstone Member is also

    dominated by sandstone, internally structured predominantly by trough cross-stratification. Other

    distinctive features of the two members include the presence of dm-scale clasts

    of Witteberg quartzites within the sandstones (especially in the uppermost Bamboesberg and Indwe

    Sandstone members), and the large (mm-to cm-scale) crystals of feldspar within the Indwe sand-

    stones (Turner, 1975; Christie, 1981). The Transitional Member also coarsens upwards with the

    contact between the Molteno and overlying Elliot Formation marked by

    the top of the uppermost coarse sandstone. This contact also marks a sharp palaeontological break

    in that it coincides with the first occurrence of fossils assignable to the lowermost biozone of the

    Elliot Formation, the Euskelosaurus Assemblage Zone.

    The Elliot Formation (Norian –Early Jurassic, Fig. 8) is dominated by floodplain mudstones with

    subordinate channel and crevasse splay sandstones, probably representing a meandering system. In

    addition to the fluvial sediments, an aeolian loessic dust component is apearing on the upper part of

    the formation, increasing in the top, showing a transition not only in sedimentation processes, but

    also in the climate change, a more arid tipe of climate. The occurrence of aeolian sandstones starts as

    m-scale thick intercalations in the upper part of the Elliot Formation, before the definitive

    establishment of the aeolian environment, showing cyclical changes in climate or aeolian sediment

    input in the transition interval between the Elliot and Clarens environments.

    Fig.8. Elliot and Clarens Formations

    In the Clarens Fm. (Early to Middle Jurassic, Fig.6) consists of cream or yellow fine grained

    sandstones, sandy siltstones and mudstones with subordinate coarse-grained sandstones. Theclimate discussed previously intensified in the first period, and then a moderate tipe of climate with a

    more abundent rainfall takes place. Considering the stratigraphic conformity and the gradual

    character of the transition between the fluvial-dominated (Elliot) and the aeolian-dominated

    (Clarens) environments, the Elliot and Clarens formations may be taken together as one coarsening-

    upward sequence. The upper contact with the lavas suggests synchronous desert sedimentation and

    early volcanism, particularly in the Huab and main Karoo basin.

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    Fig.9 Idealized structure and process of the evolution of theforeland system which is influenced by cycles of orogenic

    loading and unloading

    Drakensberg

    The Karoo Igneous Province consists of several remnant floodbasalt accumulations, sills and vast

    dyke swarms . From the geochemical stratigraphy it appears that the tholeiitic lavas onceformed an extensive carapace centred on the 1800 m thick Lesotho remnant. 40Ar/39Ar dates on the

    flood basalts and intrusives record a 185 –180 Ma duration of magmatism. Continental flood basalts

    are commonly associated with continental breakup, such as the extensional basins that developed

    along the margin of the Namaqua plate. Synrift sediments of Kimmeridgian age have been drilled in

    the Outeniqua basin and Middle Jurassic age in the conjugate North Falkland basin. Magmatism and

    the start of extension were more or less contemporaneous.

    Interpretation

    A crucial issue in the analysis of

    the Karoo Basin is the

    identification of the flexural

    foredeep and forebulge

    characteristics, as well as the

    migration trends through time.

    The stages of orogenic loading

    and unloading represent animportant influence on the

    migration of the hinge line

    between phases of

    sedimentation, controlling also

    the subsidence and the uplift of

    the foreland system flexural

    profile (Fig.9)

    Flexural processes combined with

    sedimentation determine theposition of the basin depocentre

    in the proximal region during

    orogenic loading, and in the distal

    region during orogenic unloading.

    The out of phase base-level

    changes between the proximal

    and the distal regions result in

    contrasting stratigraphies (Fig.6),

    which may explain the

    stratigraphic hinge lines in

    Dwyka, Ecca and Beaufort

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     Fig.10 Variation of the stratigraphic hinge line through time

    Groups (Fig.10). The hinge line

    shifted towards the craton

    during the Late Carboniferous-

    Permian interval, and back

    towards the orogen during the

    Triassic-Middle Jurassic. As

    contrasting stratigraphies

    develop in response to ut of

    phase base-level changes

    between the proximal and the

    distal regions of the foreland

    system, the pattern of hinge

    line migration reflects a

    correspondig migration of the

    flexural foredeep andforebulge.

    The data of the studies of the

    Karoo Basin suggest that the

    cratonward migration of the

    foreland system (Late

    Carboniferous-Permian)

    occured during a time of

    orogenic loading in the CapeFold Belt. Although the first

    order orogenic loading lasted

    until the end of the Middle

    Triassic, which was continued

    with a low-rate shift in the

    same direction during the Late

    Triassic-Middle Jurassic stage

    of first order orogenic

    unloading. The figures.... offer

    a precise and detailed image

    of the variation of the changes

    that have been occured in

    sedimentation, relanshionship

    between proximal and distal

    regions depending on the

    flexural characteristics and

    paleoclimatic trend of a

    specific region. The

    progradation of the foreland

    system is attributed to the

    Fig.11 Schematic model for the evolution of the Karoo

    sedimentar basin

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    progradation of the orogenic fron during stages of orogenic loading. For the Late Carboniferous-

    Permian interval, when the progradation of the Karoo foreland system took place, evidence for the

    gradual advance of the CFB against the basin is supplied by the Dwyka, Ecca and lower Beaufort

    strata that are incorporated within the orogenic structures in the proximity of the orogenic front. The

    geology of the CFB indicates that no further progradation of the orogenic front occured after the end

    of the Permian, although the first-order orogenic loading continued until the end of the Middle

    Triassic. The subsequent low-rate retrogradation of the foreland system during the Late Triassic-

    Middle Jurassic stage of first-order orogenic unloading is attributed to the erosion of the orogenic

    front leading to a slow retrogradation of the centre of weight in the orogenic belt. Such stages of

    low-rate retrogradation of the foreland system due to the erosion of the Fig. Schematic model for the

    evolution of the Karoo sedimentary basin orogenic front probably took place during all quiescence

    stages of the CFB first order cycle.

    The depocentre of the Karoo basin is alternately the foredeep, during orogenic loading, and the

    foresag during which the retrogradation of the orogenic load due to the erosion of the orogenic front

    may cauze an orogenward migration of the foresag. The depocentre of the Karoo basin is alterately

    the foredeep during orogenic loading, and the foresag, during orogenic unloading. In Fig.11 , time

    step (1) suggest the coeval deposition of marine and nonmarine Dwyka facies, as well as the debut of

    the Ecca Sea (lower Prince Albert Formation) restricted to the southern part of the basin, which

    explains the diachroneity of

    the Dwyka-Ecca contact.

    The Ecca-Beaufort contact

    is also diachronous, which

    is explained by the

    persistence of the Ecca Seawithin the foresag area

    coeval with the debut of

    fluvial aggradation in the

    more proximal region

    during time-step (5).The

    basin is underfilledduring

    time-steps (1)-(4), with

    deep marine

    sedimentation, it reaches afilled phase during time

    step (5), with shallow

    marine=nonmarine

    sedimentation and evolves

    into an overfilled phase

    during time-step (6), with

    fully nonmarine

    sedimentation across the

    entire basin.

    Fig.11 Schematic model for the evolution of the Karoo -

    sedimentary basin (continuation)

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      In the period Triassic-Middle Jurrasic the orogenward migration of the peripheral bulge/sag may

    either be attributed to the retrogradation of the orogenic load due to the erosion of the orogenic

    front during times of quiescence. The basin is overfilled with fully nonmarine sedimentation during

    the stages presented at Fig. . As a function of the relative position between the equilibrium drainage

    profile and the flexural profile, areas of sedimentation, bypass or erosion are separated within the

    basin.

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    Paleontology

    Fossil Potential:

    * Dwyka Formation: a glacial deposit (tillite) containing no fossils.

    * Ecca Group: in this southern part of the basin, some fossil wood and fragmentary plant material is

    found. In addition, rare fish fossils have also been found (Jubb and Gardiner, 1975), as well as trace

    fossils in the form of burrows, feeding track and fish-fin drag marks at the interface of bedding

    planes.

    * Beaufort Group: these predominantly terrestrial sediments have, throughout South Africa, yielded

    a large number of vertebrate fossils in the form of amphibians, early primitive reptiles (the

    captorhinids), mammal-like reptiles (therapsids), and fish. Minor freshwater invertebrates (molluscs)

    and plant fossils have also been recovered. For the most part, however, the fossils found in the

    Beaufort sediments are rare, particularly in the lowermost part of the succession, just above the

    Ecca- Beaufort contact.

    The Beaufort Group is subdivided into eight biozones (Fig.12) (Rubidge, 1995) on the basis.

    Fig. 12 Beaufort Group

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    -The Eodicynodon Assemblage Zone:

    Eodicynodon (Fig. 13 ) was a therapsid reptile of the Capitanian stage (266-260 million years ago) of

    the Permian Period. It measured about 30 cm. (11.8 inches) long. The length of the skull was 9 cm.

    (3.5 inches).

    Fig. 13 Eodicynodon reconstruction

    -The Tapinocephalus Assemblage Zone:

    Tapinocephalus (Fig.14) is a genus of large herbivorous dinocephalian that lived during the Middle

    Permian Period 250 million years ago. These stocky, barrel-bodied animals were characterised by a

    massive bony skull roof and short weak snout. It is thought that, like the rest of the members of its

    family, the animals engaged in head-butting intraspecific behavior, possibly for territory or mates.

    The fossil remains (skull and postcranial elements) of Tapinocephalus are known from the Lower,Middle, and Upper part of the Tapinocephalus Assemblage Zone ( Capitanian age) of the Lower

    Beaufort Beds of the South African Karoo. Only the type species, T. atherstonei is now considered

    valid for this genus.

    In life, these animals were over 3 meters (10 ft) in length and weighed around 1.5 to 2 metric tonnes

    (1.6 to 2 short tons), making them among the largest animals of their time.They died off in the

    Triassic-Jurassic extinction about 200 million years ago, which opened the door to dinosaur

    domination of the earth.

    http://en.wikipedia.org/wiki/Therapsidhttp://en.wikipedia.org/wiki/Capitanianhttp://en.wikipedia.org/wiki/Capitanianhttp://en.wikipedia.org/wiki/Faunal_stagehttp://en.wikipedia.org/wiki/Faunal_stagehttp://en.wikipedia.org/wiki/Million_years_agohttp://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Genushttp://en.wikipedia.org/wiki/Herbivorehttp://en.wikipedia.org/wiki/Dinocephaliahttp://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Family_%28biology%29http://en.wikipedia.org/wiki/Agonistic_behaviorhttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Skullhttp://en.wikipedia.org/wiki/Postcraniahttp://en.wikipedia.org/wiki/Tapinocephalus_Assemblage_Zonehttp://en.wikipedia.org/wiki/Capitanianhttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Karoohttp://en.wikipedia.org/wiki/Type_specieshttp://en.wikipedia.org/wiki/Metrehttp://en.wikipedia.org/wiki/Foot_%28length%29http://en.wikipedia.org/wiki/Metric_tonnehttp://en.wikipedia.org/wiki/Short_tonhttp://en.wikipedia.org/wiki/Short_tonhttp://en.wikipedia.org/wiki/Metric_tonnehttp://en.wikipedia.org/wiki/Foot_%28length%29http://en.wikipedia.org/wiki/Metrehttp://en.wikipedia.org/wiki/Type_specieshttp://en.wikipedia.org/wiki/Karoohttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Capitanianhttp://en.wikipedia.org/wiki/Tapinocephalus_Assemblage_Zonehttp://en.wikipedia.org/wiki/Postcraniahttp://en.wikipedia.org/wiki/Skullhttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Agonistic_behaviorhttp://en.wikipedia.org/wiki/Family_%28biology%29http://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Dinocephaliahttp://en.wikipedia.org/wiki/Herbivorehttp://en.wikipedia.org/wiki/Genushttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Million_years_agohttp://en.wikipedia.org/wiki/Faunal_stagehttp://en.wikipedia.org/wiki/Capitanianhttp://en.wikipedia.org/wiki/Therapsid

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    Fig. 14 Tapinocephalus reconstruction

    -The Pristerognathus Assemblage Zone :

    Pristerognathus (Fig.15) a genus of  Therocephalian mammal-like reptile, whose fossils have been

    found in that structure. These animals were roughly cat-sized, and are characterized by long, narrow

    skulls with large canines. They are likely to have preyed on smaller therapsids and millerettids of the

    time.

    Fig. 15 Pristerognathus reconstruction

    http://en.wikipedia.org/wiki/Pristerognathushttp://en.wikipedia.org/wiki/Pristerognathushttp://en.wikipedia.org/wiki/Therocephaliahttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Therapsidshttp://en.wikipedia.org/wiki/Millerettidshttp://en.wikipedia.org/wiki/Pristerognathushttp://en.wikipedia.org/wiki/Pristerognathushttp://en.wikipedia.org/wiki/Millerettidshttp://en.wikipedia.org/wiki/Therapsidshttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Therocephaliahttp://en.wikipedia.org/wiki/Pristerognathus

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    -The Tropidostoma Assemblage Zone is a geological stratum and a faunal zone of the Beaufort

    Group, of the South African Karoo. The name refers to Tropidostoma, a genus of  dicynodont

    mammal-like reptile, whose fossils have been found in that structure.

    The Cistecephalus Assemblage Zone The name refers to Cistecephalus, a genus of small burrowing,

    mole-like reptile, whose fossils have been found in that structure.

    Cistecephalus (Fig. 16) was a small, specialised, burrowing dicynodont, possibly with habits similar to

    a modern mole. The head was flattened and wedge-shaped, the body short, and the forelimbs very

    strong, with similarities in structure to the forelimb of modern burrowing mammals.Cistecephalus is

    so far known from the Cistecephalus Assemblage Zone of the South African Karoo, as well as from

    Zambia and India. Cistecephalus was about 33 centimetres (13 in) in length.

    Fig. 16a A trace of Cistecephalus

    Fig. 16b Cistecephalus reconstruction

    http://en.wikipedia.org/wiki/Geologyhttp://en.wikipedia.org/wiki/Stratumhttp://en.wikipedia.org/wiki/Biozonehttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Karoohttp://en.wikipedia.org/wiki/Tropidostomahttp://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Cistecephalushttp://en.wikipedia.org/wiki/Reptilehttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Mole_%28animal%29http://en.wikipedia.org/wiki/Cistecephalus_Assemblage_Zonehttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Karoohttp://en.wikipedia.org/wiki/Zambiahttp://en.wikipedia.org/wiki/Indiahttp://en.wikipedia.org/wiki/Indiahttp://en.wikipedia.org/wiki/Zambiahttp://en.wikipedia.org/wiki/Karoohttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Cistecephalus_Assemblage_Zonehttp://en.wikipedia.org/wiki/Mole_%28animal%29http://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Reptilehttp://en.wikipedia.org/wiki/Cistecephalushttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Tropidostomahttp://en.wikipedia.org/wiki/Karoohttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Beaufort_Grouphttp://en.wikipedia.org/wiki/Biozonehttp://en.wikipedia.org/wiki/Stratumhttp://en.wikipedia.org/wiki/Geology

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    The Dicynodon Assemblage Zone :

    The name refers to Dicynodon (Fig. 17), a genus of  mammal-like reptile, that flourished in the

    Permian Period and whose fossils have been found in that structure. Is a type of  herbivorous

    mammal-like reptile that flourished during the Permian Period. This animal was toothless, except for

    prominent tusks, hence the name. It probably cropped vegetation with a horny beak, much like atortoise, while the tusks may have been used for digging up roots and tubers.

    Dicynodon was a medium sized and advanced member of the Dicynodont group. It had an average

    length of 1.2 metres (3.9 ft), although size differed among species. Its fossil remains have been found

    in sediments of latest Permian age in South Africa, Tanzania, Russia, and China.

    Fig. 7 Dicynodon reconstruction

    The Lystrosaurus Assemblage Zone The name refers to Lystrosaurus, a genus of  mammal-like reptile, 

    a dominant life form of the Early Triassic Period, whose fossils have been found in that structure.

    Lystrosaurus ( Fig. 18) was a genus of Late Permian and Early Triassic Period dicynodont therapsids, 

    which lived around 250 million years ago in what is now Antarctica, India, and South Africa. Four to

    six species are currently recognized.Being a dicynodont, Lystrosaurus had only two teeth, a pair of

    tusk-like canines, and is thought to have had a horny beak that was used for biting off pieces of

    vegetation. Lystrosaurus was a heavily-built, herbivorous animal, approximately the size of a pig. The

    structure of its shoulders and hip joints suggest that Lystrosaurus moved with a semi-sprawling gait. 

    The forelimbs were even more robust than the hindlimbs, and the animal is thought to have been apowerful digger that nested in burrows.

    Lystrosaurus was by far the most common terrestrial vertebrate of the Early Triassic, accounting for

    as many as 95% of the total individuals in some fossil beds. It has often been suggested that it had

    anatomical features that enabled it to adapt better than most animals to the atmospheric conditions

    that were created by the Permian –Triassic extinction event and which persisted through the Early

    Triassic—low concentrations of  oxygen and high concentrations of  carbon dioxide. However recent

    research suggests that these features were no more pronounced in Lystrosaurus than in genera that

    perished in the extinction or genera that survived but were much less abundant than Lystrosaurus.

    http://en.wikipedia.org/wiki/Dicynodonhttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Herbivorehttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Lystrosaurushttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Triassichttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Genushttp://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Triassichttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Therapsidhttp://en.wikipedia.org/wiki/Antarcticahttp://en.wikipedia.org/wiki/Indiahttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Tuskhttp://en.wikipedia.org/wiki/Canine_toothhttp://en.wikipedia.org/wiki/Herbivorehttp://en.wikipedia.org/wiki/Pighttp://en.wikipedia.org/wiki/Terrestrial_locomotionhttp://en.wikipedia.org/wiki/Early_Triassichttp://en.wikipedia.org/wiki/Permian%E2%80%93Triassic_extinction_eventhttp://en.wikipedia.org/wiki/Permian%E2%80%93Triassic_extinction_eventhttp://en.wikipedia.org/wiki/Permian%E2%80%93Triassic_extinction_eventhttp://en.wikipedia.org/wiki/Oxygenhttp://en.wikipedia.org/wiki/Carbon_dioxidehttp://en.wikipedia.org/wiki/Carbon_dioxidehttp://en.wikipedia.org/wiki/Oxygenhttp://en.wikipedia.org/wiki/Permian%E2%80%93Triassic_extinction_eventhttp://en.wikipedia.org/wiki/Early_Triassichttp://en.wikipedia.org/wiki/Terrestrial_locomotionhttp://en.wikipedia.org/wiki/Pighttp://en.wikipedia.org/wiki/Herbivorehttp://en.wikipedia.org/wiki/Canine_toothhttp://en.wikipedia.org/wiki/Tuskhttp://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/South_Africahttp://en.wikipedia.org/wiki/Indiahttp://en.wikipedia.org/wiki/Antarcticahttp://en.wikipedia.org/wiki/Therapsidhttp://en.wikipedia.org/wiki/Dicynodonthttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Triassichttp://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Genushttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Triassichttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Lystrosaurushttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Herbivorehttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Period_%28geology%29http://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Dicynodon

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    Fig. 18 Lystrosaurus

    The Cynognathus Assemblage Zone. The name refers to Cynognathus, a genus of  eucynodontian

    mammal-like reptile, whose fossils have been found in that structure.

    Cynognathus (Fig. 19) was a heavily built animal, and measured around 1 metre (3.3 ft) in body

    length. It had a particularly large head, 30 centimetres (1.0 ft) in length, with wide jaws and sharp

    teeth. Like hind limbs were placed directly beneath the body, as in mammals, but the fore-limbs

    sprawled outwards in a reptilian fashion.

    The dentary was equipped with differentiated teeth that show this animal could effectively process

    its food before swallowing. The presence of a secondary palate in the mouth indicates that

    Cynognathus would have been able to breathe and swallow simultaneously.

    The lack of ribs in the stomach region suggests the presence of an efficient diaphragm: an important

    muscle for mammalian breathing. Pits and canals on the bone of the snout indicate concentrations of

    nerves and blood vessels. In mammals, such structures allow hairs (whiskers) to be used as sensory

    organs.

    Fig. 19 Cynognathus

    http://en.wikipedia.org/wiki/Cynognathushttp://en.wikipedia.org/wiki/Eucynodontiahttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Dentaryhttp://en.wikipedia.org/wiki/Secondary_palatehttp://en.wikipedia.org/wiki/Diaphragm_%28anatomy%29http://en.wikipedia.org/wiki/Nervehttp://en.wikipedia.org/wiki/Whiskershttp://en.wikipedia.org/wiki/Whiskershttp://en.wikipedia.org/wiki/Nervehttp://en.wikipedia.org/wiki/Diaphragm_%28anatomy%29http://en.wikipedia.org/wiki/Secondary_palatehttp://en.wikipedia.org/wiki/Dentaryhttp://en.wikipedia.org/wiki/Fossilhttp://en.wikipedia.org/wiki/Mammal-like_reptilehttp://en.wikipedia.org/wiki/Eucynodontiahttp://en.wikipedia.org/wiki/Cynognathus

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    Fun Fact

    The Permian –Triassic (P –Tr) extinction event, informally known as the Great Dying, was an

    extinction event that occurred 251.4 million years ago, forming the boundary between the Permian

    and Triassic geologic periods. It was the Earth's most severe extinction event, with up to 96% of all

    marine species and 70% of  terrestrial vertebrate species becoming extinct It is the only known mass

    extinction of  insects. 

    Distinct assemblages of paleosols above and below the Permian –Triassic boundary in the Karoo

    Basin of South Africa are evidence for reorganization of ecosystems following this greatest of all

     mass

    extinctions. The Permian –Triassic boundary is recognized from the last appearance of Dicynodon and

    from a series of negative excursions in the isotopic composition of carbon within therapsid

     tusks,

    pedogenic carbonate nodules, and organic matter.

    Causes of the end-Permian mass extinction 

    The changes in fluvial style in the Karoo Basin at the end of the Permian were initially thought to

    be triggered by a pulse of thrusting in the southerly source area, which brought about rapid

    progradation of a large sandy braided fan system (the Katberg Fm.) into the central parts of the basin

    .Ward suggested an alternative that better fits the observed palaeoclimatic changes. They proposed

    that the switch in fluvial regime was a consequence of the extinction event rather than the cause.

    The global aridification and con comitant de-vegetation of the continental interiors caused unstable

    channel banks and increased run-off that is reflected by the onset of gullying of the floodplains and

    the switch from high to low sinuosity channel pattern.

    Stable isotopic analyses of pedogenic nodules and teeth taken from embedded fossils through

    the extinction interval at the Bethulie section revealed an anomalous negative excursion within the

    boundary laminites.This anomaly is interpreted as having been caused by a relatively rapid release of

    methane or CO2

    into the atmosphere over a period of some 150 kyr. The greenhouse effect from these emissions,

    whether they originated from the Siberian volcanism or oceanic overturn , could have caused the

    observed global aridification of continental interiors .

    Recovery fauna

    Only four genera (31%) survived the extinction: the dicynodont Lystrosaurus and the

    therocephalian genera Tetracynodon,Moschorhinus and Ictidosuchoides. However, none of the

    therocephalian genera survived into the succeeding Early Triassic Katberg Formation, because they

    died out in the upper Palingkloof some 160 kyr after the main extinction event during what could be

    interpreted as a second pulse of extinctions amongst the ‘survivor fauna’.Out of the ten taxa

    recovered from Earliest Triassic

    http://en.wikipedia.org/wiki/Extinction_eventhttp://toolserver.org/~verisimilus/Timeline/Timeline.php?Ma=251.4http://en.wikipedia.org/wiki/Permianhttp://en.wikipedia.org/wiki/Triassichttp://en.wikipedia.org/wiki/Geologic_periodhttp://en.wikipedia.org/wiki/Marine_biologyhttp://en.wikipedia.org/wiki/Specieshttp://en.wikipedia.org/wiki/Terrestrial_ecoregionhttp://en.wikipedia.org/wiki/Vertebratehttp://en.wikipedia.org/wiki/Extinctionhttp://en.wikipedia.org/wiki/Insectshttp://en.wikipedia.org/wiki/Insectshttp://en.wikipedia.org/wiki/Extinctionhttp://en.wikipedia.org/wiki/Vertebratehttp://en.wikipedia.org/wiki/Terrestrial_ecoregionhttp://en.wikipedia.org/wiki/Specieshttp://en.wikipedia.org/wiki/Marine_biologyhttp://en.wikipedia.org/wiki/Geologic_periodhttp://en.wikipedia.org/wiki/Triassichttp://en.wikipedia.org/wiki/Permianhttp://toolserver.org/~verisimilus/Timeline/Timeline.php?Ma=251.4http://en.wikipedia.org/wiki/Extinction_event

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    strata, directly above the PTB, seven taxa originate within 30 m of the boundary. The first new taxon

    to appear almost immediately above the boundary is the archosauriform Proterosuchus .

    This taxon is closely followed by the amphibian Micropholis and the procolophonoid

    ‘Owenetta’kitchingorum (soon to be placed in a new genus, the dicynodonts Lystrosaurus murrayi

    and L.declivis, and the cynodonts Thrinaxodon l iorhinus and Galesaurus. The amphibian, Lydekkerina., the procolophonoid)

    Procolophon and the therocephalian Scaloposaurus appear stratigraphically higher in the Katberg

    Formation, with Lydekkerina appearing firstat approximately 37 m above the boundary.

    Procolophonwas previously regarded as originating just abovethe PTB , but our collecting has

    repeatedly shown this taxon makes its first appearance approximately 60m above the PTB,

    thereafter it then occurs in abundance.

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    Conclusions

    Due to the flexural behaviour of the foredeep and forebulge of the basin, controlled by the orogenic

    loading and unloading processes. As a result, the depocentre of the foreland system alternatedbetween the depositional foresag, during orogenic unloading, and the depositional foredeep, during

    orogenic loading, so the orogenic tectonism played a crucial role in creating the stratigraphic

    arhitecture. The first-order CFB orogenic cycle icludes a Late Carboniferous-Middle Triassic orogenic

    loading stage followed by a Late Triassic-Middle Jurassic unloading stage. The Karoo foreland system

    migrated towards the craton during the Late Carboniferous-Permian, and back towards the orogen

    during the Triassic-Middle Jurassic. The cratonward migration of the foreland system is controlled by

    the progradation of the orogenic front during orogenic loading. The retrogradation of the centre of

    weight in the orogenic belt is probably attributed to the erosion the orogenic front.

    Bibliography

    The Karoo basins of south-central Africa - O. Catuneanu a,*, H. Wopfner b, P.G. Eriksson c, B.

    Cairncross d, B.S. Rubidge e, R.M.H. Smith f, P.J. Hancox e

    Tectonic evolution of the Cape and Karoo basins of South Africa - Anthony Tankard a,*, HermanWelsink b, Peter Aukes c, Robert Newton d, Edgar Stettler e

    The recovery of terrestrial vertebrate diversity in the South African Karoo Basin after the end-

    Permian extinction - Roger Smith *, Jennifer Botha

    Fluvial style variations in the Late Triassic –Early Jurassic Elliot formation, main Karoo Basin, South

    Africa - Emese M. Bordy *, P. John Hancox, Bruce S. Rubidge

    Reciprocal flexural behavior and contrasting stratigraphies: a new basin development model for the

    Karoo retroarc foreland system, South Africa – O. Catuneanu, P. J. Hancox, B. S. Rubidget

    Evolution of the Karoo basin - ??

    Wikipedia.org

    Different geology sites for small questions.