Flowers and Inflorescences

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Flowers and Inflorescences

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    FLOWERS AND

    INFLORESCENCES

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    Flowers

    The flower is the reproductive organ of the

    angiosperm plants. It is the most important

    organ for the plant classification and

    identification.

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    Dicot. And Monocot Flowers

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    Parts of the Flower

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    Parts of Flowers

    Perianth:

    When the sepals and petals are not differentiated

    It is called perianth ( units are called tepals,sepalloid or petalloid ).

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    Parts of the Flower

    - Pedicle

    - Receptacle( if elongated it is called Hypanthium as in clove ).

    - Calyx [ sepals, polysepalous (separate sepals) or gamosepalous

    (united sepals)] - Corolla [petals, polypetalous (separate petals) or gamopetalous

    (united petals)]

    - Perianth ( tepals when the calyx and corolla areundifferentiated).

    - Androecium [stamens (filament and anthers), free or joined] - Gynaecium ( Pestil)[carpels (ovary, style and stigma) free or

    joined] .

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    Adhesion in Calyx and Corolla

    Gamosepalous calyx:

    Gamopetalous corolla:

    http://images.google.com/imgres?imgurl=http://www.maltawildplants.com/FABC/Pics/AGYFT/T/AGYFT-FlowerScan2.jpg&imgrefurl=http://www.maltawildplants.com/FABC/Anagyris_foetida.php&usg=__ph8xNlsfWvXvj3ZIB6RVgGDpP4Q=&h=120&w=142&sz=12&hl=en&start=2&sig2=Nx9oBSj7lvjVDYraIAWbMA&tbnid=KJThNI9cVZ4SYM:&tbnh=79&tbnw=94&ei=495VSdeMNdCs-gbA-cjUDw&prev=/images%3Fq%3DGamosepalous%2Bcalyx%26gbv%3D2%26hl%3Den%26safe%3Dactive%26sa%3DGhttp://lh6.ggpht.com/_bGVNrAX-Iqc/RqDz0yRi7NI/AAAAAAAABm0/rMR8Oa50oIg/Gentianella_germanica_5.jpghttp://images.google.com/imgres?imgurl=http://lh5.ggpht.com/_bGVNrAX-Iqc/RqD5biRi9TI/AAAAAAAAB24/u2Zzp5r5bPI/5_Symphytu,Officinale.jpg&imgrefurl=http://picasaweb.google.com/lh/photo/MulJUnJyXZDf-qJM48MOIg&usg=__2GPT_T8oJkptH1hB7djsT12tjXw=&h=1224&w=800&sz=8&hl=en&start=5&sig2=iiguyYO0CAKH6D-OOoEeuQ&tbnid=x1ALrhiojGZZIM:&tbnh=150&tbnw=98&ei=Gd5VSd_nDIeQ-gaus6DIDw&prev=/images%3Fq%3DGamopetalous%2Bcorolla%26gbv%3D2%26hl%3Den%26safe%3Dactive%26sa%3DG
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    Androecium Consists of stamens. Each stamen consists of a filament

    and an anther( formed of two lobes open by slits andlined by a lignified fibrous layer and containing thepollen grains ).

    Stamens may be united either by filaments or byanthers.

    - Stamens united by filaments:

    Monadelphous, diadelphous, tetradelphousetc.

    - Stamens united by anthers ( syngenesious ):Didynamous, tetradynamous,..etc.

    Stamens may be situated over the inner side of petals andin this case called Epipetallous.

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    Adhesion of Androecium

    United by filament:

    Monadelphous, diadelphous.

    United by anthers (syngenesious):

    Diadynamous, Teradynamous.

    United with the petals:

    Epipetalous

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    Gynaecium(Pistil)

    It consists of one or more carpels (ovary, style andstigma).

    Carpels may be free (apocarpous) or united

    (syncarpous, note the number of locules; mono,bi,etc ).

    The ovary (containing the ovules) may be superior (in

    hypogenous flower) or inferior (in epigenous flower).

    The ovary contains one or more ovules carried on

    placenta which may be: Axile placentation

    Parietal placentation

    The stigma usually bifid and have papillosed surfacewh ?

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    Gynaecium and Placentation

    Free Carpels (Apocarpous).

    Fused Carpels (Syncarpous).

    Axile placentation. Parietal placentation.

    Free central placentation.

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    Types of flowers

    According to the presence or absence of any of

    the gynaecium or the androecium the flower

    may be:

    Hermaphrodite ( contains both the gynaecium

    and androecium.

    Unisexual:

    - Pistillate (contains gynaecium only)

    - Staminate (contains androecium only)

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    Types of Flowers

    According to the position of the floral parts in

    comparison to the gynaecium the flower may be

    described as:

    - Hypogenous (A1).

    - Perigenous(A2).

    -

    Epigenous(A3).

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    Types of Flowers

    According to giving two equal parts if divided in

    any direction the flower may be described as:

    Regular (actinomorphic) ( giving two equal parts

    when divided at any direction).

    Zygomorphic ( giving two equal parts only if

    divided in one direction).

    This is usually represented in the floral diagram.

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    Anatomy of Flowers

    The Pedicel has the stem structure.

    The bract, calyx and corolla have the leaf structure(epidermis of corolla often colored and has papillosed

    or striated cuticle). The stigma usually have papillosed epidermis.

    The anthers contain fibrous layer characteristic for theflowers.

    Of first importance for the existence of flowers is thepresence of pollen grains. The shape , size and wallstructure of them are characteristic for each flower.

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    Characteristic Elements Of

    Powdered Flowers

    Pollen grains:

    Shape, exine,

    germinal pores and

    furrows.

    Fibrous layer of anther ( lignified appear beaded in s. v.).

    Papillosed epidermis of stigma.

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    Inflorescence

    It consists of a group of flowers arranged in certain

    order.

    The main axis of the inflorescences is called Rachis

    Types of inflorescences:

    - Racemose.

    - Cymose.- Mixed ( racemes of cymes as in clove).

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    Types of Inflorescences

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    Example of Flower Medicinal Drugs

    Clove.

    Echinacea.

    German Chamomile.

    Roman Chamomile.

    Santonica.

    Pyrethrum.

    Calendula.

    Arnica.

    Saffron.

    Safflower.

    Hibiscus.

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    Clove

    Botanical origin:

    It is the dried flower buds ofEugenia

    caryophyllusfamilyMyrtaceae.

    Geographical Source:

    Madagascar, Indonesia, Brazil and smaller

    amounts in Seri Lanka and Tanzania.

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    Photo Image of Clove

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    Microscopical Charaters A)&B) Two types of clove.

    C) Fruit (mother Clove).

    D) Clove stalk.

    E)L.S. in Clove.

    F) T.S. in Hypanthium.

    G) Stamen.

    H) Petal (s.v.). I) Filament.

    J) Fibrous layer.

    K) Oil gland.

    L) Epidermis. of hypanthium.

    M) Parenchyma. Of hypanthium. N) Phloem fibers.

    O) Pollen grains

    P) Sclereids of mother Clove.

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    Clove PowderKey Elements:

    -Trianguler pollen grains.

    -Schizolysigenous oil

    glands.

    -Fibrous layer of anther.

    - Cluster crystals of

    calcium oxalate.

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    Active Constituents

    Volatile oil (14-20%), the main constituent of

    which is the phenolic compounds eugenol,

    isoeugenol and acetyl eugenol, in addition to

    sesquiterpenes(- and -phellandrene).

    Hydrolysable tannin.

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    Therapeutic Uses

    Stimulant and aromatic flavoring and antiseptic.

    The oil has local anesthetic effect ( used for

    treatment of toothache).

    The sesquiterpenes in oil of clove are cited to be

    potential anticarcinogenic compounds.

    Eugenol is used for the preparation of vanillin.

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    Compositae Flower Head

    (Sunflower inflorescence)

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    Ray ( Ligulate ) Floret

    It is pistillate.

    Corolla is strip-like consisting of 5 united petals.

    It has an inferior ovary and a bifid stigma,sometimes a bifid style.

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    Disc( Tubular ) Floret

    It is hermaphrodite.

    Corolla is tubular consisting of 5 united petals

    showing 5 teeth.

    It has an inferior ovary and a bifid stigma.

    The androecium is joined by anther

    (synginaceous anther).

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    Roman Chamomile

    Botanical origin:

    It is the dried flower heads ofAnthemis nobilis

    familyCompositae.

    Geographical Source:

    It is cultivated in south England, Belgium,

    France, Germany, Hungary and Poland and

    others.

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    Roman chamomile

    Photo-image

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    Chamomile Powder

    Anthemis nobilis

    Key Elements:

    - Pollen grains ( spherical with

    spiny exine ).

    - Fibrous layer of anther.- Papillosed stigma.

    - Compositae glandular trichomes

    ( biseriate[2-4 cells] stalk, bicellularbiseriate head ).

    - Non glandular trichomes.

    - Cluster crystals of calcium

    oxalate.

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    Active Constituents

    Volatile oil (0.4-1%) the main constituent of

    which is n-butyl angelate, and azulene (gives the

    oil blue color).

    Sesquiterpene lactones.

    Apigenin and luteolin (flavonoids).

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    Therapeutic Uses

    For the treatment of dyspepsia (in the form of

    infusion).

    In shampoo preparations.

    In poultice preparation for treatment of some

    skin diseases.

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    German Chamomile

    Botanical origin:

    It is the dried flower heads ofMatricaria recutita

    family Compositae.

    Geographical Sourse:

    It is native to and cultivated in southern and

    eastern Europe such as Germany and Hungery.

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    Photo-image of German Chamomile

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    Roman and German Chamomile

    A,B)Roman Ch.

    C,E)German Ch.

    D)Ray floret of

    Germa German Ch.1)Disc floret

    2)Ray floret

    3)Palae

    4)Receptacle

    5)bract of

    involucre

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    Difference between German and

    Roman Chamomile

    Roman Chamomile German Chamomile

    *Very few disc florets * One whorl ray florets

    and abundant ray and abundant discflorets (double capitulum) florets.

    * Solid receptacle * Hollow receptacle.

    *There are palae (thin * No palae are found.membranous bracts)

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    Powdered Matricaria

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    Active Constituents

    Volatile Oil (0.4%)blue in color due to the

    presence of chamazulene which is a

    sesquiterpene lactone found in the oil together

    with other sesquiterpenes (bisabolol andfarnesene.

    Apigenin flavonoid.

    Coumarins.

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    Therapeutic Uses

    Antispasmodic for digestive disorders.

    Anti-inflammatory in skin preparations.

    It has an ulcer-protective properties (due to thebisabolol content).