B.v. gueorguiev annotated catalogue of the carabid beetles of albania coleoptera carabidae v. 64...

ANNOTATED CATALOGUEof the

Carabid Beetlesof Albania

(Coleoptera: Carabidae)by

B.V.Guéorguiev

This catalogue summarizes all available literature recordsand a lot of new material revised or determined by theauthor in various European museums and private collec-

tions. At present the carabid fauna of Albania can be consideredas well-studied. The catalogue includes references to or data for543 species, 103 genera and 31 tribes of Carabidae. Several dozentaxa of specific or supraspecific rank are added for the first timeto the fauna of either the Balkan Peninsula or Albania. A moderatetaxonomic contribution based on a study of types, topotypes orseries of previously problematic species or subspecies fromAlbania and the neighbouring countries is also presented; as aresult, 38 various taxonomic changes (new synonyms, revalida-tions, new statuses etc.) are introduced. All species are listed withdetailed localities and references. The book is addressed to spe-cialists in coleopterology and entomology, biogeographers andnature conservationists, collectors and all ground beetle lovers.

Without doubt, this excellent catalogue, compiled with dili-gence and knowledge, will be an important tool for anyone,

not only specialists on Balkan Carabidae, and will improve ourinsights into the systematics and distribution of carabid beetles.The hope of the author to place Albania with this catalogue sideby side to Bulgaria (certainly the best-known regional fauna ofCarabidae in the Balkan Peninsula, catalogued in 1995 by thesame author and his father) is surely fulfilled. David Wrase

Pensoft Series Faunistica 64

”A

nnotated Catalogue of the C

arabid Beetles of A

lbania(C

oleoptera: Carabidae)

“

Annotated Catalogue of the

Carabid Beetles of Albania

(Coleoptera: Carabidae)

Borislav V. Guéorguiev

2 ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV

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SOFIA–MOSCOW

2007

Annotated Catalogue of the

Carabid Beetles of Albania

(Coleoptera: Carabidae)

BORISLAV V. GUÉORGUIEV


Pensoft Publishers

Geo Milev Str. 13a, Sofia 1111, Bulgaria

[email protected]

www.pensoft.net

ANNOTATED CATALOGUE OF THE CARABID

BEETLES OF ALBANIA (COLEOPTERA: CARABIDAE)

Borislav V. Guéorguiev

First published 2007

ISBN 978-954-642-295-8

Pensoft Series Faunistica No 64

ISSN 1312-0174

© PENSOFT Publishers

All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or

transmitted in any form by any means, electronic, mechanical, photocopying, recording or

otherwise, without the prior written permission of the copyright owner.

Printed in Bulgaria, February 2007

Editor: Ivailo Stoyanov

Scientific readers: Augusto Vigna Taglianti & David W. Wrase

Contents

Introduction 7

Background of the study 7

Aims of the study 8

Short description of the studied area 8

Treatment of the taxa 10

Material and depositories of the collections 10

Acknowledgments 11

Abbreviations 13

Topographic treatment of some localities 14

Systematic part 16

Results 187

Conclusions 188

Taxonomic structure of the Albanian Carabidae (Table 1) 189

Appendix 193

References 205

Index 224


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Introduction

The ground beetle fauna of Albania is not very well known taking into account the

fact that an overall investigation based upon studied material and critical survey on

the literature has never been made. So far, our knowledge of this fauna is still largely

due to Apfelbeck’s well-known monograph (Apfelbeck, 1904) and other works of the

same author published in the period 1901-1930 (Apfelbeck, 1901a; 1901b; 1902a;

1903; 1904; 1905; 1906; 1907a; 1907b; 1909; 1911; 1916; 1917; 1918a; 1918b; 1918c;

1919a; 1919b; 1930). Later, important faunistic contributions were made also by Csiki

(1940), Schatzmayr (1943b), and Hieke et al. (1986). Dealing with the Bulgarian ground

beetles, Hieke & Wrase (1988) mentioned 296 species (Nebriinae – 14, Cicindelinae –

5, Carabinae – 10, Omophroninae – 1, Brachininae – 10, Siagoninae – 1, Scaritinae –

12, Trechinae – 58, Harpalinae – 185) for Albania without detailed records. For polit-

ical reasons, in the period 1945-1990 not many faunistic records were published, and

relatively few materials from the country were available in the collections. Since 1990

collecting ground beetles there became possible again. Recently, 363 species (Nebrii-

nae – 10, Cicindelinae – 8, Carabinae – 18, Omophroninae – 1, Brachininae – 11,

Siagoninae – 1, Scaritinae – 20, Apotominae – 1, Trechinae – 55, Harpalinae – 238)

have been mentioned for Albania in the most recent catalogue of Palaearctic Co-

leoptera by Löbl & Smetana (2003).

Examination of the available collections added information about 55 new species

for the country plus 119 species omitted in the last work as well as many new faunistic

data for other species. The subsequent critical survey of the literature data brought to

the conclusion that 543 species of Carabidae live in Albania. Moreover, some taxo-

nomic changes and a lot of comments for many taxa have been made.

Background of the study

My interest in the Albanian Carabidae has appeared under the stimulus of my late

father, the well-known Bulgarian coleopterologist Vassil Borissov Guéorguiev (*1935

– †1996), immediately after the printing of the catalogue of the Bulgarian Carabidae

(Guéorguiev & Guéorguiev, 1995). The insufficient level of exploration of the cara-

bid fauna of the country at this time (because of the strong and prolonged political

isolation and lack of regional specialists), along with relatively diverse and fresh mate-

rial collected by Bulgarian zoologists in Albania, were the principal reasons for start-

ing this investigation. The complicated orography of this country, particularly the

complex relief, determines a great diversity of zonal (seacoast, lowland, hilly, forest,

subalpine, alpine) and intrazonal (riparian, stony, limestone) habitats, that could favor

INTRODUCTION 7


a high diversity of the focused animal groups – the ground beetles. Finally, it was

necessary to sort out some interpretation of toponyms in the region, which were

incorrectly treated in the past (for example, some authors have erroneously cited

records or have referenced old collection material as being from Albania).

Aims of the study

The present study pursues two objectives, and the first one is the accomplishment of

faunistic inventory of the Carabidae from Albania. This process includes discovery

of all faunistic publications for Albania and identification or partial revision of the

collections (for unpublished and published specimens, respectively) and their regis-

tration. The result of these activities is the improvement of the faunistic knowledge

(species diversity and distribution) about the ground beetles not only on the territory

of Albania, but also on the Balkan Peninsula, and in Europe on a large scale. The

second objective of the study is taxonomical – examination of type material and

revision of the status of some doubtful taxa from Albania and its adjacent regions

(Montenegro, the Kosovo Region of Serbia, Macedonia and Northwestern Greece),

which will contribute to the systematics of the group.

Short description of the studied area

Albania (indigenous name Shqipëri) is situated in the southwestern part of the Bal-

kan Peninsula and is meridionally 300-400 km long. The country occupies about 28,748

km2, and its maximal width is about 150 km. Albania is bordered in northwest with

Montenegro, in northeast and east with the Kosovo Region of Serbia and Mace-

donia, and in south with Greece. In the west, Ionic Sea divides Albania from the

Apennine Peninsula where the strait Otranto is the close water area between both

lands. One third of the country, mainly located in the western part, lies lower than

300 m a.s.l., the territory at altitude between 300 and 1000 m is almost 40 % of the

country, while that above 1000 m a.s.l. is about 30 %. The mountains situated in the

east of Albania are part of the Dinaric mountainous system and they represent rela-

tively young and high massifs. The highest top is Korabi Peak (2764 m) located on the

east state border. The main Albanian rivers, Drin, Shkumbin, Semani, Vijosë, flow in

direction northwest - southeast.

The situation of the country in the Mediterranean area and in a zone of subtropical

climate determines the principal features of the flora and fauna. The summer is dry and

warm with average temperatures in July between 24 and 25 C°, the autumn and winter

are mild and rainy. In Albania falls one of the most abundant rainfalls in Europe –

between 1000 and 1800 mm per year in the western lowlands, and to 2500 mm in the

mountains. The seasonal distribution of the precipitations is very uneven – in the sum-

Map of Albania

MAP OF ALBANIA 9


mer the rainfall is just about 1/10 from the annual sum, and most plains and hilly parts

of the country are characterized with heavy summer droughts. The days with average

temperature more than 10 C° are about 300 (in the seacoast plains) and favor the devel-

opment of Mediterranean flora and fauna. The evergreen sclerophilous Mediterranean

forests are developed in the western part of Albania up to altitude of 400-500 m and in

some other regions – higher. These forests consist of maquises (mosaically distributed

in the country) where Olea sp., Quercus coccifera, Q. ilex, other Quercus spp., Laurus nobilis,

Phyllirea angustifolia, Pistacia lentiskus, Juniperus oxicedrus and other dominate. The following

higher zone is occupied by xeric oak-hornbeam forests. The two zones are ones mostly

suffering from human activities – in many places those forests are destroyed and, in are

replaced by farmlands or secondary xeric formations including mostly Paliurus spina-

cristi. The belt of the mesophilous oak-hornbeam and beech forests is developed at

altitude from 900-1000 up to 1600-1800 m a.s.l. This vegetation is more abundant in the

northern parts of Albania, where the mesophilous forests form the upper forest bor-

der. Normally in Northern Albania the coniferous communities are above the beech

forests, but elsewhere inversions in the distribution of the deciduous and coniferous

forests, owing to local humidity conditions, may occur. The orophytic (subalpine and

alpine) grass belt starts from 1900-2000 m a.s.l. It may be noted that the intrazonal

riverside communities consist mainly of Platanus orientalis.

Treatment of taxa

The arrangement of the taxa followed the order used in the recent Palaearctic Cata-

logue (Löbl & Smetana, 2003), with only two exceptions: genus Sinechostictus (Bembi-

diini) is regarded as separate genus, according to the recent studies on the phylogeny

of Trechitae (Grebennikov, 1997; Grebennikov & Maddison, 2005), and the arrange-

ment of Tachyini follows Sciaky & Vigna Taglianti (2003). Synonyms are included if

they consider taxa from Albania.

Material and depositories of the collections

So far, faunal records for Albania were found in 279 publications, not all of them

including original or detailed data. The main part of the studied materials are kept in

three large Central European museums, which preserve the most diverse collections of

ground beetle species from the Balkan Peninsula, as well as type material from the

region. Common feature of these collections is that the principal part of the Albanian

material had been collected before 1942. Both the most diverse and most numerous

materials collected from the country are these in the Museum für Naturkunde der Hum-

boldt Universität in Berlin. There, the collection of A. Bischoff is the most outstanding

one; other notable collections are these of E. v. Oertzen and H. Hopp, V. Apfelbeck

and A. Winneguth, A. Penther and H. Zerny (most part), H. Priesner (most part), L.

Mader, C. Lona, J. Müller. F. Hieke, G. Müller-Motzfeld and L. Behne (Hieke et al.,

1986), as well as the materials collected in the second half of XX century by H. Peks

and S. Peks. The second important depository is that in the Naturhistorisches Museum

Wien, Vienna, Austria. From aspect of the species diversity the materials from Albania

there are comparable with these in Berlin. The most significant ground beetle collec-

tions in Vienna are these of V. Apfelbeck and his laboratory assistant A. Winneguth, E.

Csiki, A. Penther and H. Zerny, H. Priesner, L. Mader (most part), A. Bischoff and A.

Winkler (most part). Additionally, small parts of the collections of Matzenauer, C. Ravas-

ini and C. Lona, J. Müller, L. Weirather and J. Meschnigg are also housed in Vienna.

Another place of important carabid collection is the Hungarian Natural History

Museum, Budapest. The most important collections here are these of V. Apfelbeck

and A. Winneguth, E. Csiki (most part) and Xh. Murraj. Other small collections housed

in Budapest are those of M. Grabowski, L. Mader, C. Ravasini and C. Lona, A. Win-

kler and A. Bischoff. Recently, Fehér et al. (2004) also collected material of Carabidae

from Albania that is currently in process of preparation.

An important place as depository of type material from the Balkans is the collec-

tion of the Deutsches Entomologisches Institut in Müncheberg, Germany; some of

the types deposited there have been reviewed. Materials from one American and two

Balkan museums have been examined too. The most substantial collection among

three is that housed in the National Museum of Natural History in Sofia. It has been

collected by Bulgarian expeditions to Albania after 1992 and at present it is contain-

ing about 160 species. The other two collections are: i/ the one in the Museum of

Natural Sciences, Tiranë (Albania), which in part was identified and published by

Murraj (1968), and ii/ that in the Carnegie Museum of Natural History, Pittsburgh

(USA), consisting of recently collected material of 57 species.1

Acknowledgments

The most important part of the material was examined in the course of the author’s

work under the European Union-funded Integrated Infrastructure Initiative “Syn-

thesys”, which is actually the prime cause for the completion of this work (Applica-

tions DE-TAF-725, AT-TAF-758 and HU-TAF-817). For the work with the collec-

1Except the aforementioned collections, there are other four European museums that may house more

or less large number of Carabidae from Albania. At the present time a significant part of the former

collection of V. Apfelbeck is preserving in the Zemaljski muzej Bosne i Hercegovine in Sarajevo

(Bosnia Herzegovina). The Museo Civico di Storia Naturale in Trieste (Italy) and the Museo Civico di

Storia Naturale in Milano (Italy) house the most part of the materials collected by the Italian expeditions

in Albania before the Second World War. The first one is remarkable with the collections of C. Ravasini,

C. Lona and J. Müller, and the second one - with that of A. Schatzmayr. Most probably, the collection

of the Narodni Museum in Prague (Czech Republic) also contains materials from Albania.

ACKNOWLEDGEMENTS 11


tions and the materials on loan I am indebted to Prof. V. Andoni (MNST), Dr. P.

Beron (NMNHS), Dr. R. Booth (BMNH), Mr. R. Davidson (CMNH), Mr. B. Jaeger

(ZMHU), Dr. A. Popov (NMNHS), Dr. H. Schönmann (NMW), Dr. Gy. Szel

(HNHM), Dr. M. Uglih (ZMHU) and Dr. L. Zerche (DEI).

With pleasure, I record my deep appreciation to the colleagues who contributed

this study giving their unpublished data (marked with the paragraphs “Material in-

cluded” or “Additional material included”): in the first place Prof. Dr. A. Vigna Taglianti

(Università di Roma”La Sapienza”, Roma, Italy) as well as Mr. P. Bulirsch (Prague,

Czech Republic), Dr. F. Hieke (ZMHU), Mr. S. Hristovski (Institute of Biology, Sko-

pje, Republic of Macedonia), Mr. B. Jaeger (ZMHU), Prof. Xh. Murraj (Tiranë, Alba-

nia) and Mr. D.W. Wrase (Berlin, Germany). It is worth mentioning that, Prof. Dr. A.

Vigna Taglianti told me that several years ago he began the preparation of a checklist

of the Carabidae of Albania, being still not completed. Generously, he entrusted to

me the draft of this work which included a moderate number of records collected

from the Italian expeditions to Albania carried out during the last 15 years. Dr. F.

Hieke made valuable comments and improvements in the part about genus Amara, as

well as kindly provided me the data of both unavailable papers and papers in press.

The following colleagues also revised and improved different parts of the catalogue:

Dr. P. Bonavita (Bembidiini; Università di Roma”La Sapienza”, Roma, Italy), Mr. P.

Bulirsch (Scaritinae), Mr. J. Gebert (Cicindelinae; Rohne, Germany), Dr. P.M. Giachi-

no (Speluncarius, Tapinopterus; Settore Fitosanitario Regionale, Torino, Italy), Mr. B.

Jaeger (Stenolophina), Mr. P. Neri (Bembidiini; Forlí, Italy), Dr. B. Kataev (Harpalini;

ZISP), Mr. L. Picciau (Speluncarius, Tapinopterus; Museo Regionale de Scienze Naturali,

Torino), Dr. L. Toledano (Bembidiini; Museo Civico di Storia Naturale, Verona, Ita-

ly), Prof. Dr. A. Vigna Taglianti (whole draft, and especially Bembidiini) and Mr. D.W.

Wrase (whole draft, and especially Harpalini).

For the determination of material, I thank Dr. D. Fedorenko (Dyschirius; Moscow

Pedagogical State University, Moscow, Russia), Dr. F. Hieke (Amara), Dr. B. Kataev

(Anisodactylus, Harpalus, Ophonus, Trichotichnus, Acupalpus, Bradycellus), both Mr. G. Ledoux

and Mr. Ph. Roux (Nebria; Paris, France), Mr. P. Neri (Bembidion), Mr. M. Pavesi (Syn-

tomus, Olisthopus; MSNM, Italy) and Mr. D.W. Wrase (Notiophilus, Harpalus, Ophonus,

Parophonus, Microlestes).

Special thanks are due to Prof. Dr. A. Vigna Taglianti, Mr. D.W. Wrase, and Mr. I.

Stoyanov (Central Laboratory of General Ecology, Sofia, Bulgaria), who made care-

ful reviews and proposed important changes of the earlier versions of the manu-

script on which this paper is based. Willingly, again Mr. I. Stoyanov helped me with

the revision of the English text. Prof. Dr. A. Casale (Università di Sassari, Sassari)

kindly allowed me the reproduction of his drawing of Abanotrechus beroni, and Dr. P.

Stoev (NMNHS) gave me the basis of the map.

I owe deep gratitude to my late farther for his strong and enthusiastic support in the

initial stages of the work, as well as to my mother and my wife for their tolerance.

Abbreviations

I. Abbreviations of collections studied upon the European Union-funded In-

tegrated Infrastructure Initiative “Synthesys” by the author

DEI Deutsches Entomologisches Institut, Müncheberg, Germany

HNHM Hungarian Natural History Museum (Magyar Természettudományi Múzeum),

Budapest, Hungary

MNHUB Museum für Naturkunde der Humboldt Universität zu Berlin, Bereich Zoo-

logisches Museum, Berlin, Germany

NMW Naturhistorisches Museum Wien, Vienna, Austria

II. Abbrevuiations of other collections studied by the author

BMNH The Natural History Museum, London, United Kingdom

CMNH Carnegie Museum of Natural History, Pittsburgh, USA

MNST Museum of Natural Sciences, Tirana, Albania

NMNHS National Museum of Natural History, Sofia, Bulgaria

III. Abbreviations of collections studied by other colleagues

MCG Museo Civico di Storia Naturale “Giacomo Doria”, Genoa, Italy

MHNG Muséum d’Histoire Naturelle, Geneva, Switzerland

MNHP Muséum National d’Histoire Naturelle, Paris, France

MSNM Museo Civico di Storia Naturale di Milano, Milano, Italy

MTKD Museum für Tierkunde, Dresden, Germany

NMPC National Museum Prague (Národni muzeum v Praze), Prague, Czech Re-

public

ZISP Zoological Institute, St. Petersburg, Russia

ZSM Zoologische Staatssammlung, Munich, Germany

cAVT private collection A. Vigna Taglianti, Roma, Italy

cBJ private collection B. Jaeger, Berlin, Germany

cFC private collection F. Cassola, Rome, Italy

cLR private collection G. Ledoux & P. Roux, Paris, France

cPB private collection P. Bulirsch, Prague, Czech Republic

cPN private collection P. Neri, Forlí, Italy

cSH private collection S. Hristovski, Skopje, Macedonia

cWH private collection W. Heinz, Schwanfeld, Germany

cWR private collection D.W. Wrase, Berlin, Germany

cXM private collection Xh. Murraj, Tiranë, Albania

ABBREVIATIONS 13


IV. Other abbreviations and symbols used in the systematic part of the catalogue

! taxa still not well defined

? taxa with doubtful or questionable occurrence in Albania

[ ] taxa that most probably inhabit Albania; they were recorded from neighbouring

region

# taxa incorrectly indicated for Albania, and therefore excluded from the catalogue

f. female specimen/s

m. male specimens/s

s. specimen/s

Topographic treatment of some localities

(first are the correct treatments of toponyms, then their most frequent variants in the

examined references and collections)

Buenë River = Bojana River

Çukall Mt. = Cukali; Cucale

Dukati = Fush-Dukati; Fushes Dukati; Dukat-Fshat

Durrës = Durazzo

Gjalicë Mt. or Gjalicë e Lumes = Montes Gjallica Ljums; Djalica Ljums

Gjirokastër = Argirokastro

Gostilë = Köstil

Ibë (? Ibë e Postmë; ? Ibë e Sipërme) = Iba

Karbunara e Madhe = Karbunara

Kukës = Kukësi

Lezhë = Alessio

Liqen i Shkodrës = Shkodër Lake

Llogora = Logora; Lungara

Lura Mt. = Kunora e Lurës

Lurë e Vjetër = Lurja

Lushnje = Lushnia

Maja e Rrabës = “Raba Massiv”

Mal i Dajtit Mt. = Dajti Mt.; Dajt Massif

Mal i Shenjt Mt. = Mal i Shêit

Maranajt Mt. = Maranaj

Nangë = Nangat

North Albanian Alps = Prokletija

Peshtrik Mt. = Paštrik; Pashtrik

Rapshë = Rapša; Rapsa

Shengjin = San Giovanni di Medua; Medua

Shëndelli Mt. = Shendeli; Shen Ili

Shkëlzen Mt. = Shkölsen; Skölsen

Shkodër = Scutari

Stan-Karbunarë = Star Karbunara

Tejas = Radomir (village in East Albania, Korab Mt.)

Tepelenë = Tepelene

Tërpan = Terpan

Theth = Thethi

Vermosh = Vermosa

Vijosë River = Vijosa River

Vlorë = Valona; Avlona

Zepë Mt. = Zebia; Zebja

TOPOGRAPHIC TREATMENT OF SOME LOCALITIESEDITORIAL 15


Family CARABIDAE Latreille, 1802

Subfamily NEBRIINAE Laporte, 1834

Tribe NEBRIINI Laporte, 1834

1. Genus Eurynebria Ganglbauer, 1891

Notes. New genus for the fauna of Albania.

1.1. Eurynebria complanata (Linnaeus, 1767)

Material studied. “Apfelb. Albanien”, 4 s. (MNHUB); “Albania: Valona.” / “complanata orientalis Typus!

Apfb.”, 1 s. (HNHM); “Valona. Albania” / “Typus! Apf.”, 1 s. (HNHM); “Valona” / “v. albanica

Cotypus! Apfb”, 4 s. (HNHM); “Südalbanien Kopstein” / “Eurynebria complanata var. albanica Apf.”, 2

s. (NMW); “Alban. Wght. Valona” / “Eurynebria complanata var. albanica Apf.”, 8 s. (NMW); “Alban.

Wght. Valona”, 4 s. (MNHUB); “Albanien Avlona 08” / “E. complanata albanica Coll. Apfelbeck”, 4 s.

(HNHM); “Valona Albanien V.1908 H. Hopp”, 2 s. (MNHUB); “Ruskuli Alb., Mader” / “Eurynebria

complanata var. albanica Apf.”, 2 s. (NMW); “Shëng… 25.X.1965 kol. Murraj”, 2 s. (MNHUB).

Additional material included (studied A. Vigna Taglianti). Shën Pjeter, Durrës District,

29.V.2004, 1 f., A. Paparistos leg. (cAVT).

Notes. Eurynebria complanata orientalis Apfelbeck, E. complanata albanica and E. complanata var.

albanica Apfelbeck are nomen nudum. The species was recorded hitherto only for European

Turkey on the Balkans (Farkač & Janata, 2003: 78). New species for the fauna of Albania.

2. Genus Leistus Frölich, 1799

Subgenus Leistus s.str.

2.1. Leistus fulvibarbis Dejean, 1826

2.1.1. Leistus fulvibarbis fulvibarbis Dejean, 1826

References. Leistus fulvibarbis: Apfelbeck, 1904: 50 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);

Schatzmayr, 1943b: 94 (Stan-Karbunarë); Perrault, 1986: 58, fig. 3; Hieke & Wrase, 1988: 21;

Perrault, 1991: 198, fig. 22.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Skutari Bojana”, 6 s. (NMW;

MNHUB); “Apfelb. Alban. spt. IV.-V.’05”, 1 s. (NMW); “Albania leg. Bischoff 1932” / “Logara

Juni”, 1 s. (MNHUB); “leg. Bischoff Albania 1934” / “Logara”, 1 s. (MNHUB); “Shen Ili” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”,

1 s. (MNHUB); Tiranë Park, 1-9.IX.1997, 7 s., R. Ward leg. (CMNH).

Notes. Farkač & Janata (2003: 82) have omitted L. fulvibarbis for Albania. Here, the species is

confirmed for the country.

SYSTEMATIC PART

COLEOPTERA: CARABIDAE: NEBRIINAE 17

2.2. Leistus nitidus (Duftschmid, 1812)

References. Leistus nitidus: Perrault, 1986: 58, fig. 3; Perrault, 1991: 198, fig. 23.

Material studied. “Albania leg. Bischoff 1936” / “Shkëlzen”, 1 s. (MNHUB).

Notes. Perrault displayed the species range as including North Albania (Perrault, 1986;

Perrault, 1991). In the second work he noted also a single locality on the map. First detailed

record for Albania.

Subgenus Pogonophorus Latreille, 1802

2.3. Leistus apfelbecki Ganglbauer, 1891

Material included (studied A. Vigna Taglianti). Valbona, Lugu i Cirkut Mt., 2000 m,

16.VII.1993, 1 m., L. Latella leg. (cAVT).

Notes. New species for the fauna of Albania.

2.4. Leistus magnicollis Motschulsky, 1866

References. Leistus magnicollis: Reitter, 1884: 36; Apfelbeck, 1904: 50; Apfelbeck, 1907a: 498,

501 (Mal i Shenjt Mt.: top region, 1500 m); Hieke & Wrase, 1988: 21; Perrault, 1992: 21, fig. 11.

Material studied. “Oroši (Ptr.) Alban. spt.”, 2 s. (NMW); “Merdita Oroschi”, 2 s. (HNHM);

“Alban. Exp.’18 Pashtrik 4-14.7.”, 1 s. (NMW); “Albania leg. Bischoff 1935” / “Koritnik”, 1 s.

(MNHUB); “Albania leg. Bischoff 1935” / “Llogora”, 3 s. (MNHUB); “Giallica” / “Albania

leg. Bischoff 1936”, 9 s. (MNHUB); “Albania leg. Bischoff 1936” / “Shkalla Haruse” / “Korab”,

1 s. (MNHUB); “Cafa e Sharit Iballe” / “Albania leg. Mueller 1939”, 7 s. (MNHUB); “Maja e

Shpatit Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë,

1000 m, 9.V.1995, 2 s., L. Penev leg. (NMNHS); Petrela, 15 km NE Tiranë, 300 m, 9.V.1995, 1

s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).

Notes. Farkač & Janata (2003: 83) have omitted L. magnicollis for Albania. Here the species is


2.5. Leistus parvicollis Chaudoir, 1869

References. Leistus parvicollis: Perrault, 1992: 18-19, 22, fig. 12; Farkač & Janata, 2003: 83.

Material studied. “Oroši (Ptr.) Alban. spt.”, 1 s. (NMW); “Alban. spt. Maranai”, 1 s. (HNHM);

“Alban. Exped. Kula Ljums 15-20.VIII.’18”, 1 s. (NMW); “Albania leg. Bischoff 1935” /

“Cukale”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 1 s. (MNHUB); “Bashtrik”

/ “Albania leg. Bischoff 1936”, 7 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”, 7 s.

(MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania leg. Bischoff

1936” / “Mal i Krujes”, 2 s. (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Mal i

Dajtit”, 1 s. (MNHUB); “Mali i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB);

Gjirokastër, 6.V.1994, 1 s., P. Stoev leg. (NMNHS).

Notes. For the present, the differentiation of L. parvicollis from L. spinibarbis s.l. bases only on

the ratio of the maximal width of head and the width of pronotum base (Perrault, 1992: 19).

First detailed records for Albania.


2.6. Leistus rufomarginatus (Duftschmid, 1812)

References. Leistus rufomarginatus: Apfelbeck, 1907a: 498 (Oroshi); Hieke & Wrase, 1988: 21;

Farkač & Janata, 2003: 84.

Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.-V.’05”, 5 s. (NMW); “Albania

leg. Bischoff 1932” / “Mal i Dajit”, 1 s. (MNHUB); “Albania leg. Bischoff 1932” / “Tomor-

Buranj Juli 1932”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 1 s. (MNHUB).

2.7. Leistus spinibarbis (Fabricius, 1775)

2.7.1. Leistus spinibarbis rufipes Chaudoir, 1843 [= punctatus Reitter, 1885]

References. Leistus spinibarbis rufipes: Apfelbeck, 1904: 49 (Vlorë; Berat); Perrault, 1992: 18, 22

(fig. 12); Farkač & Janata, 2003: 84. Leistus spinibarbis: Anonymous, 1907: 279; Hieke & Wrase,

1988: 21. Leistus spinibarbis punctatus: Apfelbeck, 1909: 297 (Vlorë).

Material studied. “Latif Skutari”, 2 s. (NMW); “Skutari Mesi” / “Apfelb. Alban. spt. IV.-V.’05”,

12 s. (NMW); “Merdita Munela”, 4 s. (NMW); “Alban. spt. Maranai”, 9 s. (HNHM; MNHUB);

“Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 9 s. (NMW); “Albania leg. Bischoff 1932” / “Ige-

Ezren”, 1 s. (MNHUB); “Albania leg. Bischoff 1932” / “Kjori”, 2 s. (MNHUB); “leg. Bischoff

Albania 1934” / “Mal i Krues”, 10 s. (MNHUB); “Giallica” / “Albania leg. Bischoff 1936”, 1 s.

(MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 24 s. (MNHUB); “Mal i Dajit” / “Albania

leg. Bischoff 1936”, 11 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 23 s. (MNHUB);

“Maja e Policanit” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albania Sauk” / “23.IV.1958

leg. Xh. Murraj”, 10 s. (HNHM); Mal i Dajtit Mt., 10-11.V.1965, 4 s. (MNST).

Notes. Except the material studied above I have seen more specimens in MNHUB, which

remained unregistered.

3. Genus Nebria Latreille, 1802

Subgenus Alpaeus Bonelli, 1810

Notes. The systematic of the Balkan species from this group is complex and still not well

settled. Comparison carried out between specimens from all the subspecies of Nebria aetolica

Apfelbeck, 1901 s.l. in HNHM revealed that some of them are more closely related each other

than to others. The subspecies have vicarious distribution in the alpine zones of the high

mountains of South Albania, Southwestern Macedonia, North and Central Greece. Details in

the pronotum form manifested that N. aetolica galicica, N. aetolica macedonica, N. aetolica papingutica

and N. aetolica rambouseki are more similar each other than any of them to the remaining

subspecies. For the reason to display more natural relations in this group the following

nomenclatorial acts are proposed:

Nebria (Alpaeus) macedonica Mařan, 1938, stat. rest. of Nebria (Alpaeus) aetolica macedonica Mařan, 1938

Nebria (Alpaeus) macedonica galicica Horvatovich, 1974, new assessment of Nebria (Alpaeus)

aetolica galicica Horvatovich, 1974

Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974, new assessment of Nebria (Alpaeus)

aetolica papingutica Horvatovich, 1974

Nebria (Alpaeus) macedonica rambouseki Mařan, 1938, new assessment of Nebria (Alpaeus) aetolica

rambouseki Mařan, 1938


3.1. Nebria attemsi Apfelbeck, 1908

References. Nebria attemsi: Bänninger, 1925: 272; Mařan, 1938: 51 (Korab Mt.); Csiki, 1940:

210 (Korab Mt.); Ledoux & Roux, 1992: 258; Farkač & Janata, 2003: 85.

Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 40 s. (HNHM); “Akad. Balk.

Exp. Csiki 1918” / “Albania Mts. Korab”, more than 70 s. (HNHM); “Alban. Exped. Korab, 1918,

24-31.VII.”, more than 80 s. (NMW); “Albania leg. Bischoff 1932” / “Tomor”, 1 s. (MNHUB);

“Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Albania leg. Bischoff 1935” /

“Derzë”, 2 s. (MNHUB); “Albania leg. Bischoff 1935” / “Korab”, more than 50 s. (MNHUB).

Notes. Apfelbeck published three descriptions of this species (Apfelbeck, 1908; Apfelbeck,

1917; Apfelbeck, 1930), the oldest one is correctly treated as valid.

3.2. Nebria bosnica Ganglbauer, 1889

3.2.1. Nebria bosnica jezercensis Horvatovich, 1974

References. Nebria bosnica jezercensis: Horvatovich, 1974b: 90-91 (Maja e Jezercë Mt., loc. typ.);

Huber, 2001: 78 (Maja e Jezercë Mt.); Farkač & Janata, 2003: 85 (Maja e Jezercë Mt.).

3.2.2. Nebria bosnica kulti Mařan, 1939

References. Nebria apfelbecki kulti: Mařan, 1939a: 20-21 (Shkëlzen Mt., loc. typ.). Nebria bosnica:

Csiki, 1940: 210 (Shkëlzen Mt.). Nebria bosnica kulti: Horvatovich, 1974b: 90 (Shkëlzen Mt.);

Huber, 2001: 78 (Shkëlzen Mt.); Farkač & Janata, 2003: 85 (Shkëlzen Mt.).

Material studied. “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 5 s. (HNHM); “Shkëlzën”

/ “Albania leg. Bischoff 1936”, 1 syntype (HNHM); “Albania leg. Bischoff 1936” / “Shkëlzën”,

15 s. (MNHUB).

3.2.3. Nebria bosnica prokletiensis Horvatovich, 1974

References. Nebria apfelbecki: Apfelbeck, 1918c: 164 (North Albanian Alps: Grebeni near

Vermosh); Mařan, 1939a: 20 (North Albanian Alps: Grebeni near Vermosh). Nebria bosnica

prokletiensis: Horvatovich, 1974b: 91 (North Albanian Alps, loc. typ., state attribution

unspecified); Farkač & Janata, 2003: 85.

Material studied. “Prokletija Gbt. Dr. Penther”, 1 paratype (HNHM).

Notes. N. bosnica apfelbecki Ganglbauer, 1891 inhabits only mountains in Eastern Bosnia

Herzegovina and Western Montenegro.

3.2.4. Nebria bosnica sturanyi Apfelbeck, 1906

References. Nebria sturanyi: Apfelbeck, 1906: 1662-1663 (Maranajt Mt., loc. typ.); Apfelbeck,

1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1918c: 160 (Maranajt Mt.);

Bänninger, 1925: 270; Mařan, 1939a: 20 (Maranajt Mt.); Pretner, 1978: 146, 166. Nebria bosnica

sturanyi: Horvatovich, 1974b: 89 (Maranajt Mt.); Huber, 2001: 77-78 (Maranajt Mt.); Farkač &

Janata, 2003: 85 (Maranajt Mt.).

Material studied. “Latif Alban spt.”, 1 s. (HNHM); “Alban. spt. Maranai”, 4 s. (MNHUB);

“Maranai Alban.-occ.” / “Coll. Apfelbeck”, 1 s. (HNHM); “Maranai G.” / “Apfelb. Alban. spt.

11.V.05”, 3 s. (HNHM); “Apfelb. Alban. spt. IV.-V.05” / “Maranai” / “sturanyi Apfelb.”, 15 s.

(NMW); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 6 s., P. Beron & B.

Petrov leg. (cLR; NMNHS).


? Nebria bosnica tresnjevikensis Horvatovich, 1974

Notes. This geographical race inhabits the Montenegrin part of the North Albanian Alps and

may live in the northernmost alpine spot in Albania too. Huber (2001: 81, Abb. 3) showed

map where the subspecies is shown to live in the country in spite of lack of exact locality.

? Nebria dahli (Duftschmid, 1812)

? Nebria dahli montenegrina Apfelbeck, 1904

References. Nebria dahli velebitica: Apfelbeck, 1904: 56 (“Sv. Troice bei Prisren”). Nebria dahli:

Horion, 1941: 81.

Notes. The locality “Sv. Troice bei Prisren” (Apfelbeck, 1904) is situated on the northern slopes of the

Shar Planina Mountains, in the Kosovo Region of Serbia. Probably Horion (1941) followed the same

record. N. dahli may live in Albania since it occurs in the massif of Durmitor (Apfelbeck, 1904 sub N.

dahli montenegrina), and N. dahli velebitica Heyden, 1884 inhabits only in Croatia (Farkač & Janata, 2003: 85).

3.3. Nebria ganglbaueri Apfelbeck, 1905

3.3.1. Nebria ganglbaueri ganglbaueri Apfelbeck, 1905

References. Nebria ganglbaueri: Apfelbeck, 1905: 240 (Albania / Kosovo border: Koritnik Mt.,

alpine belt, loc. typ.); Apfelbeck, 1907b: 636 (Koritnik Mt., alpine belt, loc. typ.); Bänninger,

1925: 270. Nebria ganglbaueri ganglbaueri: Horvatovich, 1975: 272 (Koritnik Mt.); Farkač & Janata,

2003: 86 (Koritnik Mt.).

Material studied. “Latif Koritnik” / “ganglbaueri Apfelb.”, 2 syntypes (NMW).

Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b),

the older one is correctly treated as valid.

3.3.2. Nebria ganglbaueri gyalicaensis Horvatovich, 1975

References. Nebria ganglbaueri: Csiki, 1940: 210 (Gjalicë Mt.). Nebria ganglbaueri gyalicaensis:

Horvatovich, 1975: 275-276 (Gjalicë Mt., loc. typ.); Farkač & Janata, 2003: 86 (Gjalicë Mt.).

Material studied. “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 holotype

and 18 paratypes (HNHM); “Alban. Exp. 18. Gjalica Ljums 17-26.VI.”, 1 paratype (NMW);

“Alban. Exp. 18. Gjalica Ljums 17-26.VI.”, 16 s. (NMW).

3.3.3. Nebria ganglbaueri korabensis Horvatovich, 1975

References. Nebria ganglbaueri: Csiki, 1940: 210 (Korab Mt.). Nebria ganglbaueri korabensis:

Horvatovich, 1975: 274-275 (Korab Mt., loc. typ.); Farkač & Janata, 2003: 86.

Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 holotype and 22

paratypes (HNHM); “Alban. Exped. Korab 1918 24-31.VII.”, 23 s. (NMW); “Albania leg.

Bischoff 1935” / “Korab”, more than 140 s., incl. 27 paratypes (MNHUB).

Notes. Farkač & Janata (2003: 86) noted this subspecies for ex-Yugoslavia (Serbia and Montenegro).

In fact its type locality is the Korab Mt. lying on the border between Albania and Macedonia.

3.3.4. Nebria ganglbaueri merditana Apfelbeck, 1906

References. Nebria merditana: Apfelbeck, 1906: 1661-1662 (Merdita: Munella Mt., loc. typ.);

Apfelbeck, 1907a: 500, 503 (Munella Mt.); Bänninger, 1925: 270; Mandl, 1964: 33-36;

Horvatovich, 1974a: 359. Nebria ganglbaueri merditana: Horvatovich, 1975: 272-273 (Munella

Mt.); Farkač & Janata, 2003: 86 (Munella Mt.).


Material studied. “Albania”, 1 s. (DEI); “Albania Merdita”, 1 s. designated incorrectly as

syntype of Nebria ganglbaueri ditana Apfelbeck (DEI); “Merdita Munela” / “Coll. Apfelbeck”,

5 s. (HNHM); “Merdita Munela”, 1 s. designated incorrectly as syntype of Nebria ganglbaueri

ditana Apfelbeck (DEI); “Merdita Munela”, 9 s. (MNHUB); “Merdita Munela” / “Vgth. 1906

Merdita”, 14 s. (NMW).

Notes. Apfelbeck, 1906 described this taxon from two closely situated mountains (Munella

Mt. and Zepë Mt.) situated in the area of Merdita. Also, he gave no exact “loc. typ.”. Later on,

Horvatovich (1975) attributed the specimens from Zepë Mt. to the ssp. zebiaensis, so that “loc.

typ.” of the ssp. merditana should be Munella Mt. Nebria ganglbaueri ditana Apfelbeck is either

nomen nudum or incorrect spelling of the name merditana. In the second case the two specimens

in DEI may be syntypes.

3.3.5. Nebria ganglbaueri zebiaensis Horvatovich, 1975

References. Nebria merditana: Apfelbeck, 1906: 1661-1662 (Merdita: Zepë Mt.); Apfelbeck,

1907a: 500, 503 (Zepë Mt.). Nebria ganglbaueri zebiaensis: Horvatovich, 1975: 273-274 (Zepë

Mt., loc. typ.); Farkač & Janata, 2003: 86 (Zepë Mt.).

Material studied. “Merdita Zebia” / “Coll.Apfelbeck”, 1 holotype and 4 paratypes (HNHM);

“Merdita Zebia” / “Vgth. 1906 Merdita”, 2 s. (DEI).

Notes. Further study on the validity of this race (with respect to the previous one) is needed.

3.4. Nebria germarii Heer, 1837

! Nebria germarii ssp.

Material studied. “Albania leg. Bischoff 1935” / “Maja e Jezeroe” / “Nebria!! wahrscheinlich”

/ “N. germari ssp. n. det. Dr. Horvatovich”, 1 s. (MNHUB).

Notes. New species for the fauna of Albania. For the present the subspecies status of the

local population is not defined, but it probably tends to ssp. durmitorensis Apfelbeck, 1904.

3.5. Nebria macedonica Mařan, 1938

3.5.1. Nebria macedonica galicica Horvatovich, 1974 [= korabica Horvatovich, 1974]

References. Nebria aetolica galicica: Horvatovich, 1974a: 360-361 (loc. typ. in Macedonian part

of Galičica Mt.); Ledoux & Roux, 1992: 257 (Korab Mt.); Farkač & Janata, 2003: 84. Nebria

aetolica korabica: Horvatovich, 1974a: 361 (loc. typ. in Macedonian part of Korab Mt.).

Material studied. “Korab”, 1 paratype of Nebria aetolica korabica (HNHM).

3.5.2. Nebria macedonica papingutica Horvatovich, 1974 [= tomoricensis Horvatovich, 1974]

References. Nebria aetolica tomoricensis: Mařan, 1945: 56 (Tomor Mt.); Horvatovich, 1974a:

364-365 (Tomor Mt., loc. typ. of tomoricensis; Tomor Mt.: Abbas Ali). Nebria aetolica papingutica:

Horvatovich, 1974a: 363-364 (Maja e Papingut Mt., loc. typ. of ssp. papingutica; Maja e Policanit

Mt.); Ledoux & Roux, 1992: 257 (Maja e Papingut Mt.; Tomor Mt.); Farkač & Janata, 2003: 84.

Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 4 s., including 3

paratypes of Nebria aetolica tomoricensis (HNHM; NMW); “Tomor Monte Toik” / “Albania

1922 Ravasini, Lona”, 1 paratype of Nebria aetolica tomoricensis (HNHM); “Tomor: Tomor Alban.

mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 holotype and 4 paratypes of Nebria aetolica

tomoricensis (HNHM); “Tomori Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 5


paratypes of Nebria aetolica tomoricensis (NMW); “Tomori Toik Alban. mer.” / “Winkler, Lona,

Bischoff, V.1931”, 1 s. (NMW); “Albania leg. Bischoff 1932” / “Tojk Tomor”, 10 paratypes of

Nebria aetolica tomoricensis (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Maja e

Papingut”, 1 holotype and 26 paratypes of Nebria aetolica papingutica (HNHM; MNHUB);

“Albania leg. Bischoff 1937” / “Maja e Policanit”, 2 paratypes of Nebria aetolica papingutica

(HNHM); “Albania leg. Bischoff 1937” / “Mal i That”, 2 s. (HNHM).

[Nebria macedonica rambouseki Mařan, 1938]

Notes. This geographical race was described from the Macedonian part of the Jablanitsa Mt.

(Mařan, 1938: 57; Mařan, 1945: 56). So far there is no sure record from Albania.

Subgenus Boreonebria Jeannel, 1937

3.6. Nebria rufescens (Strøm, 1768)

3.6.1. Nebria rufescens rufescens (Strøm, 1768) [= gyllenhali (Schönherr, 1806)]

References. Nebria gyllenhali: Hieke & Wrase, 1988: 22. 12. Nebria rufescens rufescens: Farkač &Janata, 2003: 88.

Material studied. “Albania leg. Bischoff 1935” / “Shkëlzen”, 4 s. (MNHUB).

Notes. First detailed record for Albania.

Subgenus Eunebria Jeannel, 1937

3.7. Nebria jockischi Sturm, 1815

3.7.1. Nebria jockischi jockischi Sturm, 1815

References. Nebria jockischi: Hieke & Wrase, 1988: 22.

Material studied. “Albania leg. Bischoff 1936” / “Gjallica”, 3 s. (MNHUB); “Albania leg.

Bischoff 1936” / “Korab”, 8 s. (MNHUB).

Notes. Farkač & Janata (2003: 90) have omitted N. jockischi for Albania. The species is here

confirmed for the country with first detailed records.

Subgenus Nebria s.str.

3.8. Nebria brevicollis (Fabricius, 1792)

References. Nebria brevicollis: Apfelbeck, 1904: 54 (Velipoja; Vlorë); Apfelbeck, 1909: 297

(Vlorë); Schatzmayr, 1943b: 95 (Mal i Krujes; Lushnje; Stan-Karbunarë; Berat); Hieke & Wrase,

1988: 22; Farkač & Janata, 2003: 92. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlorë).

Material studied. “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 5 s. (HNHM); “Albania

Sauk” / “2.V.1958 leg. Xh. Murraj”, 3 s. (HNHM); “Albania Dajti” / “2.6.1959 leg. Xh. Murraj”,

1 s. (HNHM); Tiranë, 10.V.1961, 1 s. (MNST); Mal i Dajtit Mt., 10-11.V.1965, 3 s. (MNST);

Tiranë Park, 8.III.1998, 54 s., shaded lake edge under leaves and wet, rotten wood, R. Ward

leg. (CMNH; NMNHS); “Albania Tomor Mts. Kajala e Tomorit, 26.V.2004, K. Harmos & D.

Muránya”, 1 s. (HNHM).


Additional material included (studied A. Vigna Taglianti). Tiranë, Mal i Dajtit Mt.,

7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelenë District), valley

of Vijosë River, 300-360 m, 18.IV.2003, 3 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT);

Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 1 m., 2 f., P. Crucitti & C.

Cavalier leg. (cAVT).

Notes. Except the material studied above I have seen additionally more than 20 s. from different

localities in MNHUB.

3.9. Nebria kratteri Dejean, 1831

3.9.1. Nebria kratteri kratteri Dejean, 1831 [= valonensis Apfelbeck, 1904]

References. Nebria kratteri: Apfelbeck, 1904: 52 (South Albania); Apfelbeck, 1907a: 498, 501,

503 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1928: 46; Holdhaus, 1956: 7 (Mal i

Dajtit Mt.); Magistretti, 1965: 79; Ledoux & Roux, 1990: 94. Nebria kratteri valonensis: Apfelbeck,

1904: 55 (Vlorë); Bänninger, 1925: 274; Mařan, 1939a: 19 (Vlorë); Jeanne, 1974: 66. Nebria

kratteri ssp.: Schatzmayr, 1943b: 94 (cave “Stere Gropa e Vasos” near Dukati). Nebria kratteri

kratteri: Farkač & Janata, 2003: 92.

Material studied. “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);

“Mal-i-Dajtit Tiranë” / “Albania Bischoff 1931”, 2 s. (DEI); “Tomor: Kulmak Albania mer.”

/ “Winkler, Lona, Bischoff, V.1931”, 5 s. (NMW); “Logara juni” / “Albania Bischoff 1932”, 1

s. (DEI); “Tomor-Buranj juli 1932” / “Albania Bischoff 1932”, 1 s. (DEI).

Notes. The record “Chani Driskos bei Jannina” (Apfelbeck, 1904: 55) refers to Northwestern

Greece. The locality at Mal i Shenit Mt. (Apfelbeck, 1907a), representing the northernmost

known spot in the species range, needs verification. In MNHUB, I have seen more than 100

specimens from different localities, which remained unregistered.

# Nebria salina Fairmaire & Laboulbène, 1854

References. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlorë).

Notes. This Atlantic species does not inhabit the Balkan Peninsula, so that the record for it

referred to N. brevicollis.

Tribe NOTIOPHILINI Motschulsky, 1850

4. Genus Notiophilus Dumeril, 1806

4.1. Notiophilus aestuans Dejean, 1826 [= pusillus C.R. Waterhouse, 1833]

References. Notiophilus aestuans: Csiki, 1940: 212 (Gjalicë Mt.). Notiophilus pusillus: Hieke &

Wrase, 1988: 24.

Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania,

Mts. Gyalicalums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Pashtrik 1918

29.V.-4.VI.”, 2 s. (NMW); “Mal i Krujes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” /

“Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).

Notes. Bousquet & Barševskis (2003: 96) have omitted N. aestuans for Albania. Here it is confirmed

for the country. The record “Peristeri-Gebirge bei Jannina” (Apfelbeck, 1904: 61 sub N. pusillus)

refers to Greece.


4.2. Notiophilus aquaticus (Linnaeus, 1758)

References. Notiophilus aquaticus: Hieke & Wrase, 1988: 24.

Material studied. “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Korab”

/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB).

Notes. Bousquet & Barševskis (2003: 96) have omitted the species for Albania. Here it is


4.3. Notiophilus biguttatus (Fabricius, 1779)

References. Notiophilus biguttatus: Csiki, 1940: 212 (Gjalicë Mt.); Hieke & Wrase, 1988: 25.

Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

“Albania leg. Bischoff 1935” / “Korab”, 3 s. (MNHUB); “Albania leg. Bischoff 1935” / “Maja

e Jezercoë”, 3 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 3 s. (MNHUB); “Albania

leg. Bischoff 1936” / “Shkëlzën”, 3 s. (MNHUB); “Albania leg. Mueller 1939” / “Kunora

Drunit Cucale”, 8 s. (MNHUB); “Albania leg. Mueller 1939” / “Maja e Shpatit Iballe”, 1 s.

(MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron & B. Petrov

leg. (NMNHS); Lushnje, Divjaka National Park, 10.V.1995, 1 s., S. Golovach, B. Petrov, P.

Stoev leg. (NMNHS).



4.4. Notiophilus danieli Reitter, 1897

References. Notiophilus danieli: Apfelbeck, 1904: 64 (Vlorë); Ganglbauer, 1904: 648; Apfelbeck,

1909: 297 (Vlorë); Magistretti, 1965: 84; Hieke & Wrase, 1988: 25.

Material studied. Butrinti, 4.VI.1995, 1 s., G. Blagoev leg. (NMNHS).



4.5. Notiophilus germinyi Fauvel, 1863 [= hypocrita Putzeys, 1866]

References. Notiophilus hypocrita: Hieke & Wrase, 1988: 24.

Material studied. “Alban. Exp.’ 18 Pashtrik 4-14.7.”, 3 s. (NMW); “Koritnik” / “Albania leg.

Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB);

“Gjallica” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).



4.6. Notiophilus palustris (Duftschmid, 1812)

References. Notiophilus palustris: Csiki, 1940: 212 (Gjalicë Mt.).

Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM).



COLEOPTERA: CARABIDAE: CICINDELINAE 25

4.7. Notiophilus rufipes Curtis, 1829

References. Notiophilus rufipes: Apfelbeck, 1904: 64 (Velipoja near Lezhë); Hieke & Wrase,

1988: 25.

Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomor-

Buranj Juli 1932” / “Albania Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936

leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania leg. Mueller

1939”, 2 s. (MNHUB); “Maja e Shpatit Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB);

Tiranë Park, 1-9.IX.1997, 2 s., R. Ward leg. (CMNH).



4.8. Notiophilus substriatus C.R. Waterhouse, 1833

References. Notiophilus substriatus: Csiki, 1940: 212 (Kula Ljums; foot of Gjalicë Mt.); Hieke

& Wrase, 1988: 25.

Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

“Logara Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Burelle” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Murrsi” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal

i Krujes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Tiranë Park, 22-31.X.1997, 1 s., R.

Ward leg. (CMNH); Tiranë Park, 1-9.IX.1997, 1 s., R. Ward leg. (CMNH); Tiranë Park, 12-

18.XII.1997, 1 s., R. Ward leg. (NMNHS).



Subfamily CICINDELINAE Latreille, 1802

Tribe CICINDELINI Latreille, 1802

5. Genus Calomera Motschulsky, 1862

5.1. Calomera fischeri (M. F. Adams, 1817)

5.1.1. Calomera fischeri fischeri (M. F. Adams, 1817)

References. Cicindela fischeri: Apfelbeck, 1907a: 496, 501 (Kiri River near Mesi); Csiki, 1940:

209 (Kula Ljums); Hieke & Wrase, 1988: 11.

Material studied. “Albania Kula Ljums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

Shëngjin, 16.V.1963, 1 m. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev

leg. (NMNHS).

Notes. Puchkov & Matalin (2003: 100) have omitted the species for Albania. Here it is



5.2. Calomera littoralis (Fabricius, 1787)

5.2.1. Calomera littoralis nemoralis (Olivier, 1790)

References. Cicindela lunulata var. nemoralis: Apfelbeck, 1894: 546. Cicindela lunulata: Murraj,

1968: 33 (Golem; Suhagora; Theth; Tomor Mt.). Lophyridia nemoralis nemoralis: Mandl, 1981:

150; 154. Calomera littoralis nemoralis: Puchkov & Matalin, 2003: 100.

Material studied. Golem, 19.V.1958, 2 m. (MNST); Golem, 24.III. 1961, 1 m., 1 f. (MNST);

Mal i Dajtit Mt.: Shkall Priskë, 30.VI.1960, 1 m. (MNST); Shëngjik, 20.VII.1961, 1 m. (MNST);

Shëngjik, 16.V.1963, 2 m., 2 f. (MNST); Laguna Patog, 18.IX.1993, 4 s., S. Beshkov leg.

(NMNHS); Velipoja, 19.IX.1993, 5 s., S. Beshkov leg. (NMNHS).

Notes. Part of the material cited by Murraj (1968) was found and revised. Calomera lunulata

(Fabricius, 1781) is a valid species with primarily North African range, in Europe it lives only

in Sicily (Puchkov & Matalin, 2003: 101).

[genus Cephalota Dokhtouroff, 1883]

[subgenus Taenidia Rivalier, 1950]

[Cephalota circumdata (Dejean, 1822)]

[Cephalota circumdata circumdata (Dejean, 1822)]

Notes. Most probably, the species inhabits the southern seacoast of Albania (see also Gebert,

1999: 30, Abb. 1.2.).

6. Genus Cicindela Linnaeus, 1758

Subgenus Cicindela s.str.

6.1. Cicindela campestris Linnaeus, 1758

6.1.2. Cicindela campestris olivieria Brullé, 1832

References. Cicindela campestris: Apfelbeck, 1904: 6 (Velipoja near Lezhë; Vlorë; Berat; Durrës);

Apfelbeck, 1907a: 496 (Kiri River near Mesi); Apfelbeck, 1909: 297 (Vlorë; Berat); Csiki, 1940:

209 (Gjalicë Mt.; Korab Mt.); Murraj, 1968: 33 (Tiranë; Lurë; Sauk; Pashaliman; Kulla e Lumës);

Hieke & Wrase, 1988: 9. Cicindela campestris pontica: Schatzmayr, 1943b: 93 (Karbunara e Madhe;

Lushnje; Tërpan; “Poggio Boschetto”). Cicindela campestris olivieria: Mandl, 1944: 7; 12; 13 (Tiranë;

Korab Mt.); Puchkov & Matalin, 2003: 104. Cicindela campestris campestris: Puchkov & Matalin,

2003: 103.

Material studied. Dukati, 7.V.1958, 1 m. (MNST); Golem, 19.V.1958, 1 f. (MNST); Golem,

8.VII.1965, 1 f. (MNST); Mali-me Gropa, 3.VII.1958, 1 f. (MNST); Tiranë, 4.V.1959, 1 m. (MNST);

Tiranë, 10.IV.1965, 1 f. (MNST); Sauk, 9.VII.1961, 1 f. (MNST); Sauk, 11.IV.1964, 2 m. (MNST);

Sauk, 30.10.1963, 1 f. (MNST); Lurë, 18.VII.1961, 1 m. (MNST); Ibë, 24.IV.1962, 1 f. (MNST);

Mal i Dajtit Mt., 10.V.1965, 3 f. (MNST); North Albanian Alps, Maj’e Redohimes, 2400-2550 m,

29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m,

1.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m,

20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); Vlorë, 1.V.1994, 1 s., P. Stoev leg.

(NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS); North

Albanian Alps, near cave B 52, 1600 m, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS); Petrela 15

km NE Tiranë, 300 m, 9.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).

COLEOPTERA: CARABIDAE: CICINDELINAE 27

Additional material included (studied F. Cassola). Rapsh, 15.V.2002, 1 s., A. Paparistos leg.

(cFC); Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003,

1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cFC); Vasjar (Tepelenë District), 18.IV.2003, 1 m., P.

Crucitti & C. Cavalieri leg. (cFC).

Notes. By the lack of modern study, the intraspecific relations of C. campestris s.l. in the Balkan

Peninsula are obscure. For this reason all the data about it from Albania are referred to ssp.

olivieria provisionally. The ssp. pontica Fischer von Waldheim, 1828 inhabits only the Black Sea

Coast, so that the record for this race is referred to ssp. olivieria. Except the material studied

above I have also seen additional specimens from Gjallica Lums and the Korab Mt. in HNHM.

# Cicindela hybrida Linnaeus, 1758

References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlorë); Csiki, 1940: 209 (Kula Ljums); Hieke

& Wrase, 1988: 10.

Notes. I follow the view of Gebert (1995) and refer the literature records for C. hybrida to C.

monticola albanica.

6.2. Cicindela monticola Menetries, 1832

6.2.1. Cicindela monticola albanica Apfelbeck, 1909

References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlorë); Csiki, 1940: 209 (Kula Ljums); Hieke

& Wrase, 1988: 10. Cicindela hybrida albanica: Apfelbeck, 1909: 297 (Vlorë, loc. typ.; Berat, loc.

typ.); Apfelbeck, 1917: 219 (Kiri River near Mesi, loc. typ.; Merdita: Munella, loc. typ.; Vlorë,

loc. typ.); Apfelbeck, 1918a: 75 (Shkodër, loc. typ.; Orish, loc. typ.; Vlorë, loc. typ.); Apfelbeck,

1930: 161; Murraj, 1968: 33 (Golem; Suhagora; Tomor Mt.; Shëngjin); Cassola, 1973: 26. Cicindela

sahlbergi albanica: Gebert, 1995: 16 (Buenë River near Shkorër; Mesi; Vlorë; Merdita: Munella;

Berat). Cicindela monticola albanica: Puchkov & Matalin, 2003: 106.

Material studied. “Skutari Bojana” / “C. v. albanica Apfb. Coll. Apfelbeck”, 2 s. (HNHM);

“Albania Kula Ljums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania coll. Lichtn.”,

1 s. (HNHM); Shkumbi - Kavajes, 7.V.1958, 1 f. (MNST); Golem, 24.III.1961, 2 m., 3 f.

(MNST); Divjake, 18.VI.1964, 1 f. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G.

Blagoev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 2 s., B. Guéorguiev leg. (NMNHS);

Leskovac - Barmachi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).

Notes. Apfelbeck published four consecutive descriptions of the ssp. albanica (Apfelbeck, 1909;

Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930: 161), the oldest one is correctly treated as valid.

6.3. Cicindela sylvatica Linnaeus, 1758

6.3.1. Cicindela sylvatica sylvatica Linnaeus, 1758

References. Cicindela sylvatica sylvatica: Puchkov & Matalin, 2003: 107.

6.4. Cicindela transversalis Dejean, 1822

6.4.1. Cicindela transversalis transversalis Dejean, 1822 [= riparia Dejean, 1822]

References. Cicindela hybrida riparia: Apfelbeck, 1907a: 496 (Kiri River near Mesi).

Notes. Puchkov & Matalin (2003: 108) have omitted the species for Albania, therefore its

occurrence in Albania needs further confirmation.


7. Genus Cylindera Westood, 1831

Subgenus Cylindera s.str.

7.1. Cylindera germanica (Linnaeus, 1758)

7.1.1. Cylindera germanica germanica (Linnaeus, 1758)

References. Cicindela germanica: Hieke & Wrase, 1988: 10. Cylindera germanica germanica: Puchkov

& Matalin, 2003: 109.

7.1.2. Cylindera germanica muelleri (Magistretti, 1966)

References. Cylindera germanica muelleri: Puchkov & Matalin, 2003: 109.

Subgenus Eugrapha Rivalier, 1950

7.2. Cylindera arenaria (Füessly, 1775)

7.2.1. Cylindera arenaria viennensis (Schranck, 1781)

References. Cicindela arenaria: Hieke & Wrase, 1988: 11. Cylindera arenaria viennensis: Puchkov

& Matalin, 2003: 110.

7.3 Cylindera trisignata (Dejean, 1822)

7.3.1. Cylindera trisignata trisignata (Dejean, 1822)

References. Cicindela literata sic!: Murraj, 1968: 33 (Shëngjin). Cylindera trisignata: Cassola, 1973:

33, Fig. 2. Cylindera trisignata trisignata: Puchkov & Matalin, 2003: 111.

Material studied. Shëngjin, 16.V.1963, 4 m., 4 f. (MNST); Radona, Shkiela, 21.VI.1994, 8 s.,

B. Guéorguiev leg. (NMNHS).

Notes. Cicindela litterata Sulzer, 1776 is synonym of Cylindera arenaria. Revision of material

published by Murraj (1968) revealed that it is identical with C. trisignata.

8. Genus Myriochila Motschulsky, 1858


Subgenus Myriochila s.str.

Notes. New subgenus for the fauna of Albania.

8.1. Myriochila melancholica (Fabricius, 1798)

8.1. Myriochila melancholica melancholica (Fabricius, 1798)

Material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 8-

15.VII.2006, 1 s., M. Komnenov leg. (cSH).


COLEOPTERA: CARABIDAE: CARABINAE 29

Subfamily CARABINAE Latreille, 1802

Tribe CARABINI Latreille, 1802

9. Genus Calosoma Weber, 1801

Subgenus Calosoma s.str.

9.1. Calosoma inquisitor (Linnaeus, 1758)

9.1.1. Calosoma inquisitor inquisitor (Linnaeus, 1758)

References. Calosoma inquisitor inquisitor: Bousquet et al., 2003: 119.

Material studied. “Albania Rausa Dr. Grabowski”, 2 s. (HNHM); North Albanian Alps, Bogë,

1800-1900 m, 20-23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Mal i Dajtit Mt., 27.V.2000, 1 f.,

A. Paparistos leg. (cAVT).

Notes. First detailed records for Albania.

9.2. Calosoma sycophanta (Linnaeus, 1758)

References. Calosoma sycophanta: Murraj, 1968: 33 (Sauk; Voskopojë; Shkodër); Hieke et al.,

1986: 183 (Borshi near Vlorë); Bousquet et al., 2003: 120.

Notes. Due to misidentifications found in the collection of Xh. Murraj, his literature record

for C. sycophanta (Murraj, 1968) is included here with minimal reservation because of the easily

recognition of this species.

Subgenus Microcallisthenes Apfelbeck, 1918

9.3. Calosoma pentheri Apfelbeck, 1918

References. Calosoma pentheri: Apfelbeck, 1918c: 160-163 (eastern part of the North Albanian

Alps, 1700 m, loc. typ.); Bousquet et al., 2003: 121.

Material included (studied D.W. Wrase). “Prokletia-Geb. Alban., leg Popp”, 1 m. (cWR).

9.4. Calosoma relictum Apfelbeck, 1918 [= pentheroides Roubal, 1932]

References. Calosoma relictum var. pentheroides: Roubal, 1932: 19. Calosoma relictum: Csiki, 1940:

209 (Korab Mt., Tejas); Bousquet et al., 2003: 121.

Material examined. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 53 s. (HNHM);

“Korab-Gebirge Albanien coll. Winkler”, 2 s. (HNHM).

10. Genus Carabus Linnaeus, 1758

Notes. I accept Březina’s (1999) “well geographical and morphologically defined” subspecies

when dealing with the subgenera Megodontus, Procerus and Procrustes.


Subgenus Archicarabus Seidlitz, 1887

10.1. Carabus montivagus Palliardi, 1825

10.1.1. Carabus montivagus vellepiticus Hampe, 1850 [= sutomorensis Reitter, 1885]

References. Carabus montivagus m. sutomorensis: Breuning, 1933: 677 (Vlorë; Korab Mt.); Breuning,

1978: 194. Carabus montivagus: Hieke & Wrase, 1988: 18. Carabus montivagus vellepiticus: Březina,

1999: 25; Bousquet et al., 2003: 129; Turin et al., 2003: 198.

Subgenus Carabus s.str.

10.2. Carabus granulatus Linnaeus, 1758

10.2.1. Carabus granulatus interstitialis Duftschmid, 1812 [= aetolicus Schaum, 1857; miriditus

Apfelbeck, 1901]

References. Carabus granulatus var. miridita: Apfelbeck, 1901b: 121-122 (Lezhë, loc. typ.). Carabus

granulatus miridita: Apfelbeck, 1904: 33-34 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Buenë

River Outfall); Monzini & Angelini, 1997: 39. Carabus granulatus aetolicus: Apfelbeck, 1904: 34

(Durrës; Vlorë); Vigna Taglianti et al., 2001: 72-73 (Velipoja near Lezhë). Carabus granulatus

granulatus n. interstitialis: Breuning, 1933: 533 (Vlorë; Durrës); Breuning, 1978: 183. Carabus

granulatus granulatus m. merdita sic!: Breuning, 1933: 534 (Velipoja near Lezhë); Breuning, 1978:

184. Carabus granulatus interstitialis v. miridita: Arnold, 1939: 158; Březina, 1999: 8 (Velipoja near

Lezhë). Carabus granulatus m. miridita: Magistretti, 1965: 26. Carabus granulatus: Hieke & Wrase,

1988: 17. Carabus granulatus interstitialis: Bousquet et al., 2003: 136.

Material included (studied A. Vigna Taglianti). Vlorë, 10.IV.1999, 1 f., A. Paparistos leg. (cAVT).

Subgenus Chaetocarabus C.G. Thomson, 1875

10.3. Carabus intricatus Linnaeus, 1761

10.3.1. Carabus intricatus intricatus Linnaeus, 1761 [= montenegrinus Kraatz, 1876; chionobatus Born,

1918; weiratheri Breuning, 1934]

References. Carabus intricatus montenegrinus: Apfelbeck, 1904: 28 (Durrës). Carabus intricatus var.:

Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m). Carabus intricatus: Klaptocz, 1910:

399 (Theth); Horion, 1941: 42. Ruspoli, 1965: 8; Geisthardt, 1975: 7; Hieke & Wrase, 1988: 16;

Turin et al., 2003: 243 (North Albania). Carabus intricatus var. montenegrinus: Apfelbeck, 1918c: 160,

163 (North Albanian Alps, 1300-2000 m). Carabus intricatus chinobatus: Buresch & Kantardjieva,

1928: 84. Carabus intricatus intricatus m. montenegrinus: Breuning, 1934: 1043, 1045 (Merdita; Tiranë;

Durrës; Vlorë). Carabus intricatus intricatus n. weiratheri: Breuning, 1934: 1046 (Vlorë); Magistretti,

1965: 45 (Vlorë); Breuning, 1978: 234 (Vlorë). Carabus intricatus intricatus var. weiratheri: Březina,

1999: 53. Carabus intricatus intricatus: Bousquet et al., 2003: 138.

Material studied. Bizë, 27.VII.1966, 1 f. (MNST); North Albanian Alps, Bogë, 1400-1800 m,

1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100

m, 5-9.VI.1993, 1 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë,

1800-1900 m, 20-23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Razma, 1560 m,

12.VI.1995, 1 m., V. Sakalian leg. (NMNHS).


Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt.,

1900 m, 10.VII.1993, 1 m., L. Latella leg. (cAVT).

# subgenus Eucarabus Géhin, 1876

# Carabus catenulatus Scopoli, 1763

# Carabus catenulatus catenulatus Scopoli, 1763 [= catenatus Panzer, 1803; albanensis Géhin, 1885]

References. Carabus catenatus var. albanensis: Apfelbeck, 1904: 35.

Notes. Probably, the original record for this species from Albania is that in the description of

albanensis Géhin, 1885. Apfelbeck (1904: 35) has based indirectly on this record, indicating no

exact locality for the county. I cannot find the original description of this variation, and believe

the species does not inhabit Albania (see also Turin et al., 2003).

[subgenus Hygrocarabus C.G. Thomson, 1875]

[Carabus variolosus Fabricius, 1787]

[Carabus variolosus nodulosus Creutzer, 1799]

References. Carabus variolosus nodulosus: Breuning, 1926: 19; Breuning, 1928a: 113; Buresch &

Kantardjieva, 1928: 88. Carabus nodulosus: Turin et al., 2003: 240.

Notes. Actually, the record “Albanien: Ljuboten” (Breuning, 1926: 22) concerns the territory

of Republic of Macedonia (see also Breuning, 1935: 1169). The following authors had followed

this data mentioning the species for Albania. In spite of this, C. variolosus may inhabit the

marginal northern or northeastern regions of the country.

[subgenus Limnocarabus Géhin, 1876]

[Carabus clathratus Linnaeus, 1761]

[Carabus clathratus auraniensis J. Müller, 1903]

Notes. I cannot find any exact record for this species from Albania even though Turin et al.

(2003: 155) showed in the map it occurs there. This rare Carabus-species may inhabit the

country as Purkyně (1928) described the variation epirensis Purkyně, 1928 (synonym of C.

clathratus auraniensis) from the Greek Epirus.

Subgenus Megodontus Solier, 1848

10.4. Carabus caelatus Fabricius, 1801

10.4.1. Carabus caelatus caelatus Fabricius, 1801 [= sarajevoensis Apfelbeck, 1890; malissorum

Apfelbeck, 1919; metalkanus Apfelbeck, 1919]

References. Carabus caelatus sarajevensis sic!: Apfelbeck, 1907a: 498, 502 (Mal i Shenjt Mt.: top

region, 1500 m). Carabus caelatus: Apfelbeck, 1907a: 500; Apfelbeck, 1917: 220 (Merdita);

Apfelbeck, 1930: 162 (Merdita); Horion, 1941: 41; Březina, 1999: 73. Carabus caelatus malissorum:

Apfelbeck, 1919a: 226-267 (North Albanian Alps, Maja e Jezerzë, loc. typ.); Apfelbeck, 1919b:

45-46 (North Albanian Alps, Maja e Jezerzë, loc. typ.). Carabus caelatus caelatus n. metalkanus:

Breuning, 1935: 1297 (Merdita; Gjalicë e Lumes; Sarisaltik near Kruja; “Male Dojtitz”; Mal e

Krujes); Breuning, 1978: 258. Carabus caelatus caelatus m. malissorum: Breuning, 1935: 1298 (North

Albanian Alps; Maja e Jezercë); Breuning, 1978: 258. Carabus caelatus metalkanus: Arnold, 1939:

189. Carabus caelatus metalkanus v. malissorum: Arnold, 1939: 189. Carabus caelatus caelatus: Březina,

1999: 73; Bousquet et al., 2003: 163. Carabus caelatus sarajevoensis: Turin et al., 2003: 253.


Material studied. North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 2 m., P. Beron & B.

Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 3 m., P. Beron

& B. Petrov leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 1 m., 1 f., P.

Dioli leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 f., L. Latella leg.

(cAVT); Valbona, Kollatës Mt., 1800 m, 9.VII.1993, 1 f., F. Sirtori leg. (cAVT).

Notes. Apfelbeck published two description of the form malissorum (Apfelbeck, 1919;

Apfelbeck, 1920), the older one is correctly treated as valid.

10.4.2. Carabus caelatus dalmatinus Duftschmid, 1812 [= procerus Reitter, 1885]

References. Carabus caelatus dalmatinus: Apfelbeck, 1904: 25 (Velipoja near Lezhë); Wohlberedt,

1909: 127 (near Liqen i Shkodrës); Březina, 1999: 74; Bousquet et al., 2003: 163; Turin et al.,

2003: 253 (Velipoja; Lezhë). Carabus dalmatinus procerus: Apfelbeck, 1918c: 160 (North Albanian

Alps). Carabus caelatus var. procerus: Apfelbeck, 1918c: 163 (North Albanian Alps, 1300-2000 m).

Carabus caelatus dalmatinus n. procerus: Breuning, 1935: 1300 (Lezhë District); Breuning, 1978: 258.

Material included (studied A. Vigna Taglianti). Durrës, 13,VI.1998, 1 f., A. Paparistos leg. (cAVT).

Notes. The mentioning of the locality “Vilajet” (Turin et al., 2003: 253) is misunderstanding.

Actually this Turkish word is an equivalent to the English one “district”.

10.5. Carabus croaticus Dejean, 1826

10.5.1. Carabus croaticus durmitorensis Apfelbeck, 1904 [= ljubetensis Apfelbeck, 1917; babinjensis

Apfelbeck, 1919]

References. Carabus croaticus babinjensis: Apfelbeck, 1919a: 226 (North Albanian Alps, loc. typ.);

Apfelbeck, 1919b: 45 (North Albanian Alps, loc. typ.); Krätschmer & Drovenik, 1977: 215 (Abb.1);

Turin et al., 2003: 254 (Babinje Planina Mt.). Carabus croaticus ljubetensis n. ljubetensis: Breuning,

1935: 1291 (North Albanian Alps; Babinje Planina Mt.); Breuning, 1978: 257. Carabus croaticus

ljubetensis m. durmitorensis: Breuning, 1935: 1291 (Korab Mt.); Breuning, 1978: 257. Carabus croaticus:

Horion, 1941: 41; Tabaković-Tošić, 1988: 135; Březina, 1999: 74. Carabus croaticus durmitorensis:

Březina, 1999: 74 (North Albanian Alps); Bousquet et al., 2003: 163. Carabus croaticus durmitorensis

var. babinjensis: Březina, 1999: 74 (North Albanian Alps). Carabus croaticus durmitorensis var. ljubetensis:

Březina, 1999: 74. Carabus croaticus ljubetensis: Turin et al., 2003: 254.

Notes. Apfelbeck published three consecutive descriptions of Carabus croaticus ljubetensis

(Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930), the second one has long time been

accredited to be correct. However it is wrong and the original description must be assigned to

the oldest one. The same author gave also two descriptions of Carabus croaticus babinjensis (Apfelbeck,

1919a; Apfelbeck, 1919b), as for the present it is not clear which of both has priority.

10.6. Carabus violaceus Linnaeus, 1758

10.6.1. Carabus violaceus azurescens Dejean, 1826 [= rilvensis Kolbe, 1887; zabljakensis Eidam,

1927; shljebensis Mandl, 1985]

References. Carabus violaceus azurescens n. zabljakensis: Breuning, 1935: 1261 (Maranajt Mt.; Merdita);

Breuning, 1978: 254. Carabus violaceus azurescens m. zabljakensis: Štěrba, 1945: 152 (Maranajt Mt.;

Merdita). Carabus violaceus: Hieke & Wrase, 1988: 15. Carabus violaceus azurescens var. shljebensis:

Březina, 1999: 77 (North Albanian Alps). Carabus violaceus azurescens: Turin et al., 2003: 260.


10.6.2. Carabus violaceus dryas Gistl, 1857 [= merditanus Apfelbeck, 1917; shardagensis Apfelbeck,

1917; korabensis Csiki, 1944]

References. Carabus violaceus merditanus: Apfelbeck, 1917: 220 (Merdita: Zepë Mt. near Fandi,

loc. typ.); Apfelbeck, 1918a: 75-76 (Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1930: 162 (Merdita:

Zepë Mt., loc. typ.); Cleu, 1968: 28 (Merdita); Cleu, 1969: 167. Carabus violaceus azurescens m.

merditana: Breuning, 1935: 1262 (Vermosh; Zepë Mt. near Oroshi; Peshtrik Mt.; Gjalicë e

Lumes); Breuning, 1978: 254 (Zepë Mt. near Oroshi). Carabus violaceus zabljakensis v. merditanus:

Arnold, 1939: 186. Carabus violaceus var. shardagensis: Csiki, 1940: 210 (Gjalicë Mt.; Korab Mt.).

Carabus violaceus var. korabensis: Csiki, 1944: 50-51 (Korab Mt., loc. typ.). Carabus violaceus azurescens

m. merditanus: Štěrba, 1945: 152 (Vermosh; Zebë Mt.). Carabus violaceus shardagensis: Cleu, 1968:

28 (Korab Mt.); Cleu, 1969: 167 (Korab Mt.). Carabus picenus peristericus: Tarrier, 1974: 242

(Korab Mt.). Carabus violaceus dryas: Březina, 1999: 77; Bousquet et al., 2003: 165; Turin et al.,

2003: 260. Carabus violaceus dryas var. merditanus: Březina, 1999: 77 (Merdita: Zepë Mt.; Gjalicë

e Lumes). Carabus violaceus dryas var. korabensis: Březina, 1999: 77 (Korab Mt.).

Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m,

11.VII.1993, 1 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 2000 m, 14.VII.1993,

rests of 1 s., A. Gobetti leg. (cAVT).

Notes. Apfelbeck published three descriptions of both C. violaceus merditanus and C. violaceus

shardagensis, respectively in Serbo-Croatian, in Latin and in German (Apfelbeck, 1917; Apfelbeck,

1918a; Apfelbeck, 1930), the second ones have long time been accredited to be correct. However

it is incorrect and the original descriptions must be assigned to the oldest ones. A revision of

the type of the var. shardagensis Apfelbeck, 1917 is needed to elucidate to which subspecies,

azurescens or dryas, it belongs. Csiki (1944: 50, Fig. 1a) represented a figure of this variation

which let me assume that it belongs rather to the former subspecies.

# Carabus violaceus exasperatus Duftschmid, 1812 [= vlasuljensis Apfelbeck, 1894]

References. Carabus violaceus vlasuljensis: Apfelbeck, 1907a: 502 (Merdita); Apfelbeck, 1918c:

160, 163 (North Albanian Alps, Vermosh).

Notes. The easternmost known records of ssp. exasperatus come from Bosnia Herzegovina, so that

the literature data for it should be referred to some of both subspecies that really live in Albania.

Subgenus Oreocarabus Géhin, 1876

Notes. In my opinion, C. hortensis, C. neumeyeri and C. preslii represent separate species by

reason of its clear and constant differences in both the overall aspect of the pronotum and

the penis (Casale et al., 1982: 185, 190 – fig. 75, 195). Externally, the first species can be easily

distinguished from the other two in the more quadrate aspect of the pronotum (vs. subovate

or trapezium-shaped), the lateral border of pronotum posteriorly sinuate (vs. straight to slightly

sinuate) and in the lateral lobes of pronotum clearly horizontal or slightly oblique (vs. oblique).

Additionally C. neumeyeri has lateral borders of pronotum posteriorly more convergent than

those in C. hortensis. The following nomenclatorial acts are proposed: Carabus (Oreocarabus)

neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri Schaum, 1856;

Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii

Dejean, 1830 (see Casale et al., 1982: 194).


10.7. Carabus hortensis Linnaeus, 1758 [= rhodopensis Apfelbeck, 1904]

References. Carabus hortensis: Apfelbeck, 1918c: 163 (North Albanian Alps: Vermosh). Carabus

hortensis rhodopensis: Apfelbeck, 1918c: 164 (North Albanian Alps: Grebeni near Vermosh, 1100-

1750 m). Carabus hortensis hortensis: Breuning, 1933: 709 (Peshtrik Mt.; Vermosh); Bousquet et

al., 2003: 179.

Material included (studied A. Vigna Taglianti). Tropoia, Valle di Valbona, 900 m, 8-

16.VII.1993, 2 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 1900 m, 10.VII.1993, 1 f., L.

Latella leg. (cAVT); Valbona, Kollatës Mt., 1000 m, 14.VII.1993, 1 f., L. Latella leg. (cAVT).

10.8. Carabus neumeyeri Schaum, 1856 [= herzegowinensis Apfelbeck, 1904; pindicus Apfel-

beck, 1904; albanicola Breuning, 1965; hilleri Mandl, 1964]

References. Carabus hortensis neumeyeri: Apfelbeck, 1904: 42 (Velipoja); Apfelbeck, 1907a: 496

(Maranajt Mt.: between Vorfai Postme, 300 m, and Vorfai Siperme, 700 m); Apfelbeck, 1909:

297 (Berat); Schatzmayr, 1943b: 94 (Shendelliut); Březina, 1999: 46; Bousquet et al., 2003: 179.

Carabus hortensis neumeyeri n. neumeyeri: Breuning, 1933: 710 (Sarisaltik near Kruja; Vlorë;);

Breuning, 1978: 198. Carabus hortensis neumeyeri n. herzegowinensis: Breuning, 1933: 712 (Velipoja

near Lezhë). Carabus hortensis: Hieke & Wrase, 1988: 19. Carabus hortensis neumeyeri var. pindicus:

Březina, 1999: 46. Carabus hortensis neumeyeri var. albanicola: Březina, 1999: 46 (Llogora). Carabus

hortensis neumeyeri var. hilleri: Březina, 1999: 46 (Reza e Kanalit near Llogora). Carabus preslii

neumeyeri: Turin et al., 2003: 228.

Material studied. Kurbneshi, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); North

Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS);

North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., 2 f., P. Beron & B. Petrov leg.

(NMNHS).

10.9. Carabus preslii Dejean, 1830 [= jonicus Apfelbeck, 1904]

References. Carabus hortensis preslii m. jonica: Breuning, 1933: 713 (Gjirokastër); Breuning,

1978: 198.

Material studied. Pogyara, 1027 m, 4.VI.1994, 2 f., G. Blagoev leg. (NMNHS); Leskovac,

650 m, 2.VI.1995, 1 f., V. Sakalian leg. (NMNHS).

Notes. Bousquet et al. (2003: 179) has omitted the taxon for Albania, so that it is confirmed

here for the country.

Subgenus Pachystus Motschulsky, 1866

10.10. Carabus cavernosus I. Frivaldszky von Frivald, 1838

10.10.1. Carabus cavernosus cavernosus I. Frivaldszky von Frivald, 1838 [= sterbai Breuning, 1936]

References. Carabus cavernosus cavernosus: Breuning, 1933: 798 (Korab Mt.; Gjalicë e Lumes);

Guéorguiev et al., 1997: 8; Březina, 1999: 35; Bousquet et al., 2003: 181; Turin et al., 2003: 234.

Carabus cavernosus var. sterbai: Csiki, 1940: 210 (Korab Mt.). Carabus cavernosus n. cavernosus:

Breuning, 1978: 207. Carabus cavernosus: Hieke & Wrase, 1988: 15.


10.11. Carabus graecus Dejean, 1826

10.11.1. Carabus graecus morio Mannerheim, 1830 [= winkleri Breuning, 1933; neowinkleri Deuve, 1991]

References. Carabus graecus winkleri: Breuning, 1933: 802 (Maja e Tartarit Mt., 800 m, loc. typ.).

Carabus graecus n. winkleri: Breuning, 1978: 207 (Maja e Tartarit Mt.). Carabus graecus: Hieke & Wrase,

1988: 14; Březina, 1999: 36. Carabus graecus morio: Březina, 1999: 36; Bousquet et al., 2003: 182.

Carabus graecus morio var. winkleri: Březina, 1999: 36 (Maja e Tartarit Mt.). Carabus graecus morio var.

neowinkleri: Březina, 1999: 36 (Maja e Tartarit Mt.). Carabus graecus graecus: Turin et al., 2003: 236.

? subgenus Platycarabus A. Morawitz, 1886

? Carabus irregularis Fabricius, 1792

? Carabus irregularis ramanus Sokolář, 1909 [= bucephalus Kraatz, 1879]]

References. Carabus irregularis bucephalus: Turin et al., 2003: 249.

Notes. Turin et al. (2003) showed the species range reaching North Albania. According to

Březina (1999: 53) ssp. ramanus is known from the Bjelasièa Mt. in Northeastern Montenegro

not very far from the studied territory. The species may live in Albania.

Subgenus Procerus Dejean, 1821

10.12. Carabus gigas Creutzer, 1799

10.12.1. Carabus gigas gigas Creutzer, 1799 [= parnassicus Kraatz, 1884]

References. Carabus gigas parnassicus: Březina, 1999: 72. Carabus gigas gigas: Bousquet et al., 2003: 185.

Material studied. Malesia e Polisit Mt., 1850 m, near Sopoti, VI.1966, 1 s. (cXM).


Subgenus Procrustes Bonelli, 1810

10.13. Carabus coriaceus Linnaeus, 1758

10.13.1. Carabus coriaceus cerisyi Dejean, 1826 [= hopffgarteni Kraatz, 1877; subrugosus Kraatz,

1877; weiratherianus Breuning, 1936]

References. Carabus coriaceus cerisyi m. weiratheriana: Breuning, 1937: 1406 (Korab Mt.); Breuning,

1978: 270. Carabus coriaceus subrugosus v. weiratherianus: Arnold, 1939: 192. Carabus coriaceus var.

hopffgarteni: Csiki, 1940: 209 (Kula Ljums; Gjalicë Mt.; Korab Mt.). Carabus coriaceus subrugosus:

Hieke et al., 1986: 183 (Borshi near Vlorë). Carabus coriaceus: Hieke & Wrase, 1988: 13.

10.13.2. Carabus coriaceus excavatus Charpentier, 1825 [= rugosus Dejean, 1826; dalmaticus Géhin,

1885; joniensis Breuning, 1932; albanicus Breuning, 1943]

References. Carabus coriaceus rugosus: Apfelbeck, 1904: 20 (Velipoja; Berat; Vlorë); Apfelbeck,

1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 297 (Ardenica; Vlorë;

Pojani). Carabus coriaceus rugosus n. joniensis m. albanicus: Breuning, 1932: 20 (Maja e Tartarit Mt.,

loc. typ.); Mařan, 1949: 3 (Maja e Tartarit Mt.). Carabus coriaceus excavatus m. dalmatica: Breuning,

1937: 1403 (Velipoja; Kruja; Durrës; Merdita; Tiranë; Berat; Vlorë; south of Vlorë); Breuning,

1978: 269. Carabus coriaceus excavatus n. joniensis: Breuning, 1937: 1404 (Vlorë; Gjirokastër);

Magistretti, 1965: 55; Breuning, 1978: 269. Carabus coriaceus excavatus: Arnold, 1939: 191; Hieke


et al., 1986: 183 (Mal i Dajtit Mt.: Shkall Priskë); Březina, 1999: 62; Bousquet et al., 2003: 187;

Turin et al., 2003: 278. Carabus coriaceus excavatus dalmaticus: Schatzmayr, 1943b: 94 (Lushnje;

Karbunara e Madhe; Dukati, south of Vlorë). Carabus coriaceus joniensis: Březina, 1999: 62;

Bousquet et al., 2003: 187. Carabus coriaceus joniensis: var. albanicus: Březina, 1999: 62 (Maja e

Tartarit Mt.). Carabus coriaceus mediterraneus: Turin et al., 2003: 278.

Material studied. Sauk, 18.II.1960, 1 m. (MNST); Ibë, 3.VII.1962, 1 f. (MNST); Bizë,

27.VII.1966, 1 f. (MNST); Tiranë, 27.III.1968, 1 f. (MNST); Tiranë Park, 22-31.X.1997, 2 s.,

R. Ward leg. (CMNH); Tiranë Park, 1-4.I.1998, 2 s., R. Ward leg. (CMNH).


2250 m, 11.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 2000

m, 14.VII.1993, 1 f., A. Gobetti leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley

of Vijosë River, 300-360 m, 18.IV.2003, 2 m., P. Crucitti & C. Cavalieri leg. (cAVT).

Note: According to Turin et al. (2003) C. coriaceus mediterraneus Born, 1906 has transionian

distribution and lives in Albania.

Subgenus Tachypus Weber, 1801

10.14. Carabus cancellatus Illiger, 1798

10.14.1. Carabus cancellatus alessiensis Apfelbeck, 1901 [= hypsobius Apfelbeck, 1918]

References. Carabus cancellatus var. alessiensis: Apfelbeck, 1901b: 121 (Lezhë, loc. typ.). Carabus cancellatus

alessiensis: Apfelbeck, 1904: 30; 33 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Buenë River Outfall);

Breuning, 1933: 594 (Velipoja near Lezhë; Shkodër; Pulaj); Arnold, 1939: 161 (North Albania);

Breuning, 1978: 187 (Velipoja); Březina, 1999: 41 (Velipoja near Lezhë); Bousquet et al., 2003: 195;

Turin et al., 2003: 194. Carabus cancellatus subgraniger var. alessiensis: Bernau, 1913: 198 (Lezhë). Carabus

cancellatus emarginatus: Bernau, 1913: 200. Carabus cancellatus: Hieke & Wrase, 1988: 17.

Material studied. Shkodër, 14.V.1963, 2 m. (MNST).

Notes. The type locality of the form hypsobius Apfelbeck, 1918c: 163 is Vunšaj (in Albanian)

or Vusanje (in Serbo-Croatian) in the North Albanian Alps. It is situated in the Montenegrin

territory very closely to Albania. C. cancellatus emarginatus Duftschmid, 1812 is a valid subspecies

reaching to Bosnia Herzegovina in the east.

Subgenus Tomocarabus Reitter, 1896

10.15. Carabus convexus Fabricius, 1775

10.15.1. Carabus convexus dilatatus Dejean, 1826 [= chionophilus Apfelbeck, 1904; miliusi Breuning, 1927]

References. Carabus convexus var.: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500

m). Carabus convexus dilatatus: Breuning, 1933: 874 (Vlorë; Gjirokastër); Bousquet et al., 2003:

198; Turin et al., 2003: 230. Carabus convexus dilatatus n. miliusi: Breuning, 1933: 876 (North

Albanian Alps; Merdita); Breuning, 1978: 214. Carabus convexus miliusi: Arnold, 1939: 178. Carabus

convexus var. chionophilus: Csiki, 1940: 210 (Korab Mt.). Carabus convexus: Hieke & Wrase, 1988:

19. Carabus convexus dilatatus var. miliusi: Březina, 1999: 33.

Material studied. Mal i Dajtit Mt., 11.V.1965, 1 m. (MNST); North Albanian Alps, Maj’e

Radohimes, 1900-2200, 28.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian

Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS);

COLEOPTERA: CARABIDAE: ELAPHIRINAE 37

North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS);

North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 3 f., P. Beron & B. Petrov leg. (NMNHS).

? subgenus Trachycarabus Géhin, 1876

? Carabus scabriusculus Olivier, 1795

? Carabus scabriusculus bulgarus Lapouge, 1908

References. Carabus scabriusculus bulgarus: Turin et al., 1993: 42.

Notes. This species is questionable for Albania due to lack of exact records. On the other

side, it was quoted for the neighboring countries Montenegro and Serbia.

Tribe CYCHRINI Laporte, 1834

11. Genus Cychrus Fabricius, 1794

11.1. Cychrus semigranosus Palliardi, 1825

# Cychrus semigranosus semigranosus Palliardi, 1825

References. Cychrus semigranosus semigranosus: Häckel, 2003: 205.

Notes. This record is referred to the ssp. montenegrinus because the nominotypical race does

not populate Albania.

# Cychrus semigranosus balcanicus Hopffgarten, 1881

References. Cychrus semigranosus n. balcanicus: Mařan, 1940c: 133-134 (North Albanian Alps).

Notes. This East Balkan race does not inhabit Albania, therefore it is removed from the species list.

# Cychrus semigranosus montanus Winkler, 1939

References. Cychrus semigranosus montanus: Häckel, 2003: 205.

Notes. Winkler (1939: 199) described this subspecies on material from two different regions - the

Eastern Carpathians and Western Balkans. Mandl (1989: 69) designated lectotype and paralectotypes

for the specimens from the former region, and syntypes of montanus from the latter region were

assigned to C. semigranosus dinaricus Mandl, 1989 (“Cvrstnica”, loc. typ.). Presumably, the subspecies

mentioning for Albania (Häckel, 2003: 205) is based on incorrect interpretation of the locality

“Zljeb planina nördlich der Nord-albanischen Alpen” (Winkler, 1939: 199; Mařan, 1940c: 133;

Mandl, 1989: 68). In fact the Zljeb Mt. is situated in the Kosovo Region of Serbia.

11.1.1. Cychrus semigranosus montenegrinus Apfelbeck, 1904 [= albanicus Hopp, 1929; dinaricus Mandl, 1989]

References. Cychrus semigranosus semigranosus: Röschke, 1907: 255; Häckel, 2003: 205. Cychrus semigranosus:

Apfelbeck, 1907a: 498, 503 (Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 210 (Shkëlzen Mt.;

Korab Mt.). Cychrus semigranosus albanicus: Hopp, 1929: 2 (Tomor Mt, 2000 m, loc. typ.); Schatzmayr,

1943b: 94 (Tomor Mt.); Mandl, 1989: 69 (Peshtrik Mt.; Gjalicë Mt.; Çukall Mt.; Merdita); Häckel,

2003: 205. Cychrus semigranosus ssp.: Schatzmayr, 1943b: 93 (Shendelliut). Cychrus semigranosus montenegrinus:

Winkler, 1939: 200 (Llogora; Tomor Mt.; Kruja). Cychrus semigranosus n. montenegrinus: Mařan, 1940c:

128, 130-132, 135-136 (Llogora). Cychrus semigranosus montanus: Häckel, 2003: 205.

Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Alban.

Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Alban. Exp.’18. Pashtrik 4-14.7”, 3 s. (NMW);

“Cucale” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Giallica” / “Albania leg. Bischoff 1936”, 1

s. (NMW).


Notes. Winkler (1939) synonymized the ssp. albanicus with ssp. montenegrinus. Mařan (1940c)

accepted this point of view, but Mandl (1989) treated both as separate subspecies. Samples of

the ssp. montenegrinus, ssp. albanicus, as well as the holotype and paratypes of the ssp. dinaricus

have been studied in NMW. Differences between specimens from different localities exist

mostly in the sizes of the body and shape of the pronotum, but they also have occurred in the

limits of one and the same population. In reality characters that can support well-defined

geographical races in the Western Balkans, excluding the well-differentiated ssp. travnikanus

Apfelbeck, 1904 (type series in NMW) and the not studied ssp. jakupicensis Mařan, 1943, are

absent. Therefore the following synonymy is proposed: Cychrus semigranosus albanicus Hopp,

1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck, 1904; Cychrus semigranosus dinaricus

Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck, 1904. In MNHUB, I

have furthermore seen 69 s. from different localities, which remained unregistered.

Subfamily ELAPHRINAE Erichson, 1837

Tribe ELAPHRINI Erichson, 1837

12. Genus Elaphrus Fabricius, 1775

Subgenus Neoelaphrus Hatch, 1951

12.1. Elaphrus uliginosus Fabricius, 1792

References. Elaphrus uliginosus: Murraj, 1968: 33 (Bizë).

Material studied. “N 88”, 1 f. (MNST).

Notes. The record of Murraj (1968) is included here with minimal reservation because of his

correct determination of the female above. Goulet (2003: 207) omitted the species for Albania,

but herewith it is confirmed for the country.

Subfamily OMOPHRONINAE Bonelli, 1810

Tribe OMOPHRONINI Bonelli, 1810

13. Genus Omophron Latreille, 1802

Subgenus Omophron s.str.

13.1. Omophron limbatum (Fabricius, 1777)

References. Omophron limbatum: Hieke et al., 1986: 183 (Bizë near Shëngjergji); Hieke & Wrase,

1988: 23; Hůrka, 2003: 208.

Material studied. “Skutari Bojana”, 3 s. (NMW); “Oboti Bojana” / “Apfelb. Alban. spt. IV-V.05”,

1 s. (NMW); “Berat” / “Albania leg. Bischoff 1931”, 17 s. (MNHUB); “Ibe-Erzen” / “Albania leg.

Bischoff 1932”, 2 s. (MNHUB); “Tepelene” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Vojusa

COLEOPTERA: CARABIDAE: BRACHININAE 39

Dragot” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff

1937”, 1 s. (MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).

Subfamily BRACHININAE Bonelli, 1810

Tribe BRACHININI Bonelli, 1810

14. Genus Aptinus Bonelli, 1810

Subgenus Aptinus s.str.

14.1. Aptinus acutangulus Chaudoir, 1876

Material studied. North Albanian Alps, Bogë, 1400-1800 m, 1.6.1993, 1 f., P. Beron & B. Petrov

leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.6.1993, 1 m., 1 f., P. Beron & B.

Petrov leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 15 m., 14 f., P.

Dioli leg. (cAVT).


14.2. Aptinus bombarda (Illiger, 1800)

References. Aptinus bombarda: Apfelbeck, 1918c: 168 (North Albanian Alps, Vermosh, 1200

m); Hůrka, 1988: 290-291 (Shkëlzen Mt.); Hrdlička, 2003: 212.

Material studied. “Vermosa’14 Penther”, 1 m. (NMW); “Prokletija 1300-2000 m” / “Alban.

montenegr. Grenze, Penther’14”, 1 f. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban.

montenegr. Grenze, Penther’14”, 1 m. (NMW); “Maja e Jezerce Albania sept.”, 1 m. (NMW);

“Shkëlzën” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); North Albanian Alps, Bogë, 1400-

1800 m, 1.6.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS).


2000 m, 11.VII.1993, 2 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 1000 m,


Notes. A. bombarda occurs sympatrically with A. acutangulus in the North Albanian Alps (see

also Casale & Vigna Taglianti, 1983: 36; Hůrka, 1988: 289); the former lives at the same altitude

or higher than the latter. The study of the penis reveals that the male from Bogë has somewhat

differing apex compared with those from Vermosa and Maja e Jezercoë. The penis apex in the

latter is identical with the illustration of Casale & Vigna Taglianti (1983: 32, Fig. 8). Very likely

the population from the massif of Bogë forms a separate race.

14.3. Aptinus merditanus Apfelbeck, 1917

14.3.1. Aptinus merditanus merditanus Apfelbeck, 1917 [= granulipennis Apfelbeck, 1917]

References. Aptinus lugubris rufipes: Apfelbeck, 1907a: 499, 501 (Mal i Shenjt Mt.: top region,

1500 m). Aptinus merditanus: Apfelbeck, 1917: 227 (Merdita: Zepë Mt. near Fandi, loc. typ.);

Apfelbeck, 1918b: 89 (Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1930: 171-172


(Merdita: Zepë Mt. near Fandi, loc. typ.); Casale & Vigna Taglianti, 1983: 29, 39, 41 (Merdita:

Zepë Mt.; Shen Thanas Mt. near Vlorë); Hůrka, 1988: 293-294 (Mal i Shenjt Mt.; Zepë Mt.;

Mal i Dajtit Mt.; Mal i Krujës Mt.; Shen Ili; Maja e Tartarit Mt.; Cafa Kjorës; Tomor Mt.; Shen

Thanas Mt.; Llogora); Wrase, 1991: 18. Aptinus merditanus var. granulipennis: Apfelbeck, 1917:

227 (South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt.

near Joannina, loc. typ.); Apfelbeck, 1918b: 89 (South Albania: Shen Thanas Mt. near Vlorë,

loc. typ.; Northwestern Greece: Peristeri Mt. near Joannina, loc. typ.); Apfelbeck, 1930: 172

(South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt.

near Joannina, loc. typ.). Aptinus merditanus merditanus: Hrdlička, 2003: 212.

Material studied. “Merdita M. Schèit” / “Coll. Apfelbeck” / “merditanus Typ. Apf.”, 1 m. (HNHM);

1 s. from “Çafa Kjores”, 53 s. from “Llogora” and “Logara, 8 s. from “Maja e Tartarit”, 40 s. from

“Mal i Dajtit”, 1 s. from “Mal i Krujes”, 10 s. from “Shen Ili”, 1 s. from “Tomor” (MNHUB).

Notes. Apfelbeck published three consecutive descriptions for both A. merditanus and A.

merditanus var. granulipennis (Apfelbeck, 1917; Apfelbeck, 1918b; Apfelbeck, 1930) and the second

ones have long time been accredited to be valid. Actually this is incorrect and the original

descriptions must be assigned to the oldest ones. Aptinus lugubris rufipes (Apfelbeck, 1907a:

499) is nomen nudum (see also Hůrka, 1988: 292). In that case, Apfelbeck had most probably the

intention to introduce a name for A. merditanus earlier than the year of 1917.

15. Genus Brachinus Weber, 1801

Subgenus Brachinus s.str.

# Brachinus alexandri F. Battoni, 1984

References. Brachinus alexandri: Guéorguiev & Guéorguiev, 1995: 239.

Notes. Dealing with B. plagiatus data in the paper of Hùrka (1988), I noted that B. alexandri

was recorded by mistake for Albania (Guéorguiev & Guéorguiev, 1995). Most likely

Hrdlička(2003: 213) followed this mistake marking the species for the country. Here B. alexandri

is removed from the list of Albanian ground beetles.

15.1. Brachinus crepitans (Linnaeus, 1758) [= fallax Apfelbeck, 1904]

References. Brachinus crepitans: Apfelbeck, 1904: 348 (Vlorë; Berat); Apfelbeck, 1909: 298 (Vlorë;

Berat); Csiki, 1940: 222 (Kula Ljums; Gjalicë Mt.); Hieke & Wrase, 1988: 163; Hrdlička, 2003:

213. Brachinus crepitans ab. fallax: Apfelbeck, 1904: 348 (Vlorë); Csiki, 1940: 222 (Kula Ljums).

Material studied. “Avlona”, 1 s. (HNHM); “Han Holfit 5.-10. Mai” / “Alban. montenegr. Grenze,

Penther’14”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

“Alban-Exp.’18. Pashtrik, 4-14.7.”, 1 s. (NMW); “Dures” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB); “Lume i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Kara Alibey” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Iballe” / “Albania leg. Bischoff

1939”, 1 s. (MNHUB); “Albania Fush-Dukati” / “7.V.1958 leg. Xh. Murraj”, 1 s. (HNHM);

Vlorë District, Dukati Mt., 450 m, 11.V.1995, 1 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS);

Ljukove, 100 m, 5.VI.1995, 1 f., G. Blagoev leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 2

m., 1 f., D. Antonelli leg. (cAVT).

COLEOPTERA: CARABIDAE: BRACHININAE 41

15.2. Brachinus ejaculans Fischer von Waldheim, 1828

References. Brachinus ejaculans: Apfelbeck, 1904: 351 (Velipoja; Vlorë); Hieke & Wrase, 1988:

165; Hrdlička, 2003: 213.

Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB); “Miloti, Alb. Matzenauer”, 3

s. (NMW); “Shullaz” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Piqeros” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 11 s. (MNHUB);

“Rushkull” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB).

15.3. Brachinus elegans Chaudoir, 1842 [= ganglbaueri Apfelbeck, 1904]

References. Brachynus psophiae or ganglbaueri: Klaptocz, 1910: 399 (Shkodër); Schatzmayr, 1943b:

113 (Lushnje). Brachinus ganglbaueri: Hieke & Wrase, 1988: 164.

Material studied. “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Murthi” / “Albania

leg. Bischoff 1936”, 5 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

Notes. Hrdlička (2003: 213) has omitted the species for Albania. Here it is confirmed for the

country.

15.4. Brachinus plagiatus Reiche, 1868

References. Brachinus plagiatus: Apfelbeck, 1904: 350 (Velipoja; Vlorë); Horion, 1941: 353;

Schatzmayr, 1943b: 112 (Skrofotina); Hieke & Wrase, 1988: 165; Hùrka, 1988: 205 (Tiranë).

Material studied. “Avlona”, 6 s. (HNHM); “Durazzo”, 2 s. (HNHM); “Alban. Wgth. Valona”, 9

s. (NMW); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Rushkull” / “Albania leg.

Bischoff 1936”, 3 s. (MNHUB).


country.

15.5. Brachinus psophia Audinet-Serville, 1821

References. Brachinus psophia: Apfelbeck, 1904: 349 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);

Hieke & Wrase, 1988: 164; Hrdlička, 2003: 213. “Brachynus psophiae aut Ganglbaueri”: Schatzmayr,

1943b: 113 (Lushnje).

Material studied. “Avlona”, 1 s. (HNHM); “Alban. Wgth. Valona”, 2 s. (NMW); Radona,

Shkiela, 21.VI.1994, 1 m., B. Guéorguiev leg. (NMNHS).

Subgenus Brachynidius Reitter, 1919

15.6. Brachinus bodemeyeri Apfelbeck, 1904

References. Brachinus bodemeyeri: Apfelbeck, 1904: 352 (Vlorë); Hieke & Wrase, 1988: 166;

Hrdlička, 2003: 213.

Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB); “Alban. Wgth. Valona”, 4 s.

(NMW); “Dukati” / “ex Coll. Scheibel”, 1 s. (MNHUB).


15.7. Brachinus brevicollis Motschulsky, 1844 [= peregrinus Apfelbeck, 1904]

References. Brachinus peregrinus: Apfelbeck, 1904: 351 (Vlorë, loc. typ.); Reitter, 1919: 140;

Schatzmayr, 1935: 245; Schatzmayr, 1943b: 112 (Tërpan; Dukati; Lushnje; Vlorë); Hieke &

Wrase, 1988: 165. Brachinus brevicollis: Hrdlička, 2003: 213.

Material included (studied A. Vigna Taglianti). Qesarat, near Memaliaj (Tepelenë District),

valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).

15.8. Brachinus explodens Duftschmid, 1812 [= glabratus Dejean, 1824; strepens Fischer von

Waldheim, 1828]

References. Brachinus explodens: Apfelbeck, 1904: 353 (Vlorë; Berat); Apfelbeck, 1909: 298

(Vlorë; Berat); Csiki, 1940: 222 (Ura i Lopez); Schatzmayr, 1943b: 113 (Dukati); Hieke et al.,

1986: 190 (North Albanian Alps near Theth); Hieke & Wrase, 1988: 166; Hrdlička, 2003: 214.

Brachinus explodens ab. strepens: Csiki, 1940: 222 (Ura i Lopez; “Stičen” near Gjalicë Mt.). Brachinus

explodens ab. glabratus: Csiki, 1940: 222 (Kula Ljums; Ura i Lopez).

Material studied. “Avlona”, 2 s. (HNHM); “Skutari-Holcekai” / “Alban. montenegr. Grenze,

Penther’14”, 12 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM);

“Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Ura i

Lopez” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 8

s. (NMW); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Cerrik”

/ “16.IV.1958 leg. Xh. Murraj”, 7 s. (HNHM); Tiranë, 7.IV.1966, 1 s. (MNST); Vlorë, 1.V.1994,

2 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 2 s., G. Blagoev leg.

(NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 3 s., S. Golovach, B. Petrov, P. Stoev

leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-

4.I.1998 / 20-25.I.1998 / 1.III.1998 / 8.III.1998, 13 s., shaded lake edge under leaves and wet,

rotten wood, R. Ward leg. (CMNH).

Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near

Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti &

C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., M.

Komnenov leg. (cSH).

Notes. I have seen furthermore 160 s. from different localities in MNHUB, which remained

unregistered.

15.9. Brachinus sichemita Reiche & Saulcy, 1855

References. Brachinus explodens sichemita: Apfelbeck, 1904: 353 (Berat; Vlorë).

Notes. Hrdlička (2003: 214) has omitted the species for Albania, therefore its occurrence in

the country needs further confirmation.

Subgenus Brachynolomus Reitter, 1919

15.10. Brachinus immaculicornis Dejean, 1826

15.10.1. Brachinus immaculicornis immaculicornis Dejean, 1826

References. Brachinus immaculicornis: Apfelbeck, 1909: 298 (Vlorë).

COLEOPTERA: CARABIDAE: SIAGONINAE 43

Notes. Hrdlička (2003: 214) has omitted the species for Albania; therefore its occurrence in

the country needs further confirmation.

Subgenus Cnecostolus Reitter, 1919

15.11. Brachinus bayardi Dejean, 1831

References. Brachinus bayardi: Schatzmayr, 1943b: 112 (Mal i Krujes).

Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB);

“Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).


country.

15.12. Brachinus bipustulatus Quensel, 1806

References. Brachinus bipustulatus: Apfelbeck, 1904: 354 (Velipoja; Vlorë); Apfelbeck, 1909:

298 (Vlorë); Reitter, 1919: 135; Hieke & Wrase, 1988: 166; Hrdlička, 2003: 214.

Material studied. “Scutari” / “collectio Paganetti”, 1 s. (NMW); “Mustajbeg Velipoja Albania”,

1 s. (MNHUB); “Durazzo Apfelbeck”, 1 s. (HNHM); “Oboti Bojana”, 4 s. (HNHM); “Alban.

Wgth. Valona”, 1 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB);

“Myzeqe” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg.


15.13. Brachinus exhalans (P. Rossi, 1792)

References. Brachinus exhalans: Apfelbeck, 1904: 354 (Velipoja; Vlorë); Apfelbeck, 1909: 298

(Vlorë); Schatzmayr, 1943b: 112 (Lushnje; Mal i Krujes); Hrdlička, 2003: 214.

Material studied. “Avlona”, 3 s. (HNHM); “Avlona Albania mer.”, 1 s. (HNHM); “Albanien

Avlona v. Oertzen.”, 5 s. (MNHUB); “Durazzo Apfelbeck”, 2 s. (HNHM); “Alban. Wgth.

Valona”, 6 s. (NMW); “Murshi” / “Albania leg. Bischoff 1936”, 111 s. (MNHUB); Radona,

Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).

Species insertae sedis

15.14. Brachinus nigricornis Gebler, 1829 [= incertus Brullé, 1834]

References. Brachinus incertus: Apfelbeck, 1904: 350 (Velipoja near Lezhë); Apfelbeck, 1909:

298 (Vlorë); Hieke & Wrase, 1988: 165. Brachinus nigricornis: Hrdlička, 2003: 216.

Material studied. “Durazzo”, 1 s. (HNHM); “Albania Avlona”, 5 s. (HNHM); “Bojana

Mundung”, 3 s. (MNHUB); “Alban. Wgth. Valona”, 12 s. (NMW).


Subfamily SIAGONINAE Bonelli, 1813

Tribe SIAGONINI Bonelli, 1813

16. Genus Siagona Latreille, 1804

16.1. Siagona europaea Dejean, 1826

16.1.1. Siagona europaea europaea Dejean, 1826 [= depressa Bedel, 1887 (nec Fabricius, 1798)]

References. Siagona europaea: Apfelbeck, 1904: 358 (Vlorë); Schatzmayr, 1943b: 95 (Mal i Krujes;

Lushnje; Skrofotina); Hieke, 1981: 76; Hieke & Wrase, 1988: 27. Siagona depressa: Apfelbeck,

1909: 297 (Vlorë). Siagona europaea europaea: Baehr, 2003a: 218.

Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Krabe” /

“Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 2 s.

(MNHUB); “Liqen i Butrinten Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tirane”

/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1936”, 1 s.

(MNHUB); “Tirane” / “leg. Bischoff Albania 1937”, 2 s. (MNHUB).


Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., P. Crucitti

& C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., M.


? Siagona europaea oberleitneri Dejean, 1830

References. Siagona europaea var. oberleitneri: Ganglbauer, 1904: 648.

Notes. This race is endemic for the Ionic Islands. Taking into account the proximity of these

islands to South Albania further collectings are needed to determine the subspecies status of

the populations in the studied territory.

Subfamily SCARITINAE Bonelli, 1810

Tribe CLIVININI Rafinesque, 1815

Subtribe CLIVININA Rafinesque, 1815

17. Genus Clivina Latreille, 1802

17.1. Clivina collaris (Herbst, 1784)

References. Clivina collaris: Balkenohl, 2003: 219.

Material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 8-



COLEOPTERA: CARABIDAE: SCARITINAE 45

17.2. Clivina fossor (Linnaeus, 1758)

17.2.1. Clivina fossor fossor (Linnaeus, 1758)

References. Clivina fossor: Apfelbeck, 1904: 68 (Velipoja near Lezhë; Vlorë); Apfelbeck, 1909:

297 (Vlorë); Hieke et al., 1986: 184 (Shkodër); Hieke & Wrase, 1988: 27. Clivina fossor fossor:

Balkenohl, 2003: 219.

Material studied. “Scutary Bojana”, 3 s. (NMW); “Scutary i. See Ufer”, 2 s. (NMW); “Scutary

Vraka”, 3 s. (NMW); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mamurash” /

“Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB); Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet,

rotten wood, R. Ward leg. (CMNH).

17.3. Clivina laevifrons Chaudoir, 1842

References. Clivina laevifrons: Apfelbeck, 1904: 69 (Velipoja); Rambousek, 1912: 69; Hieke &

Wrase, 1988:28; Balkenohl, 2003: 220.

Material studied. “Scutary Bojana”, 1 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.”, 1 s. (NMW);

“Scutary i. See Ufer”, 2 s. (NMW); “Scutary Vraka”, 1 s. (NMW).

Subtribe REICHEINA Jeannel, 1957

! genus Dalmatoreicheia Magrini & Bulirsch, 2005

! Dalmatoreicheia sp.

Material studied. “Kruja, Alban., Mader”, 1 m. (NMW).

Notes. The specimen belongs to a new species.

18. Genus Reicheadella Reitter, 1913

Notes. The Albanian species of this subtribe are object of taxonomic work (Bulirsch &

Guéorguiev, in preparation).

Subgenus Reicheadella s.str.

18.1. Reicheadella bischoffi (Meschnigg, 1933)

References. Reicheia bischoffi: Meschnigg, 1933:172 (Tomor Mt. near Berat, loc. typ.); Turin,

1981: 72. Reicheadella bischoffi: Jeannel, 1957: 179; Casale et al., 1998: 95; Balkenohl, 2003: 222.

Material studied. “Tomor Juli 1932” / “Albania leg. Bischoff 1932”, 2 syntypes (MNHUB);

“Tomor: Tomor Alban. mer.” / “lg. Winkler Mai 1931” / “bischoffi Meschn. det. A. Winkler”,

2 m. (NMW).

! Reicheadella sp.

Material studied. “Albania mer. Liqueni i Butrintit, 1.V.1958; A. Smetana leg.”, 1 f. (NMPC).

Notes. The specimen belongs to a new species.


Subtribe DYSCHIRIINI W. Kolbe, 1880

19. Genus Dyschirius Bonelli, 1810

Subgenus Chiridysus Fedorenko, 1996

19.1. Dyschirius strumosus Erichson, 1837

References. Dyschirius strumosus: Balkenohl, 2003: 224.

Subgenus Dyschiriodes Jeannel, 1941

Species group “integer”

19.2. Dyschirius intermedius Putzeys, 1846

References. Dyschirus intermedius: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 78

(Oboti); Hieke & Wrase, 1988: 31; Fedorenko, 1993: 819; Balkenohl, 2003: 225.

Species group “nitidus”

19.3. Dyschirius agnatus Motschulsky, 1844 [= lucidus Putzeys, 1846]

References. Dyschirius lucidus: Hieke et al., 1986: 184 (Kula e Lumës near Kukës). Dyschirius

agnatus: Balkenohl, 2003: 224.


“Tepelene” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Durrës” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Bojana

Shkoder” / “Albania leg. Bischoff 1937”, 5 s. (MNHUB).

19.4. Dyschirius nitidus (Dejean, 1825)

19.4.1. Dyschirius nitidus nitidus (Dejean, 1825)

References. Dyschirius nitidus: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 63-64 (Berat);

Csiki, 1940: 212 (Kula Ljums); Horion, 1941: 98; Ieniștea, 1979: 224; Gryuntal, 1984b: 514;

Hieke & Wrase, 1988: 29.

Material studied. “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB).

Notes. Balkenohl (2003: 226) has omitted the species for Albania. Here it is confirmed for the country.

Species group “minutus”

19.5. Dyschirius aeneus (Dejean, 1825)

19.5.1. Dyschirius aeneus aeneus (Dejean, 1825)

References. Dyschirius aeneus: Apfelbeck, 1904: 72 (Velipoja near Lezhë); Apfelbeck, 1907a: 495 (Buenë

River); Müller, 1922b: 76; Hieke & Wrase, 1988: 31. Dyschirius aeneus aeneus: Balkenohl, 2003: 224.


Material studied. “Durrazo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Durres” /

“Albania leg. Bischoff 1932”, 1 s. (MNHUB).

19.6. Dyschirius apicalis Putzeys, 1846

References. Dyschirius apicalis: Apfelbeck, 1907a: 496 (Buenë River Outfall); Müller, 1922b: 77

(Durrës); Hieke & Wrase, 1988: 31; Balkenohl, 2003: 224.

Material studied. “Alban. Wgth. Valona”, 1 s. (HNHM); “Skutari Bojana” / “Mündg. Apf. 08.”,

3 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Liqen Butrintës” /

“Albania leg. Bischoff 1936”, 5 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS).

[Dyschirius auriculatus (Wollaston, 1867)]

Notes. An expected species for Albania due to its occurring in Montenegro (P. Bulirsch,

personal communication)

19.7. Dyschirius bacillus Schaum, 1857

19.7.1. Dyschirius bacillus arbensis J. Müller, 1911

References. Dyschiriodes bacillus arbensis: Fedorenko, 1996: 191 (Vlorë); Balkenohl, 2003: 224.

Dyschirius bacillus bacillus: Balkenohl, 2003: 224.

Material studied. Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).

Notes. According to Fedorenko (1996: 191) the specimen from Vlorë shows transitional

characters between the subspecies arbensis and lucanus (Focarile, 1959). P. Bulirsch (personal

communication) identified the specimen from Shkiela and referred it to spp. arbensis.

19.8. Dyschirius cylindricus (Dejean, 1825)

19.8.1. Dyschirius cylindricus hauseri A. Fleischer, 1898

References. Dyschirius cylindricus: Apfelbeck, 1907a: 496 (Buenë River Outfall).

Material studied. “Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Durres” /



19.9. Dyschirius laeviusculus Putzeys, 1846

References. Dyschirius laeviusculus: Hieke et al., 1986: 184 (Kula e Lumës near Kukës).

Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).

Notes. Balkenohl (2003: 226) has omitted the species for Albania. Here D. laeviusculus is

confirmed for the country. The locality Kula e Lumës outlines the southern limit of the

species distribution in the Balkan Peninsula.

19.10. Dyschirius luticola Chaudoir, 1850

19.10.1. Dyschirius luticola luticola Chaudoir, 1850

Material studied. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).



19.11. Dyschirius macroderus Chaudoir, 1850

19.11.1. Dyschirius macroderus macroderus Chaudoir, 1850

References. Dyschirius macroderus: Apfelbeck, 1907a: 496 (Buenë River Outfall).

Notes. Balkenohl (2003: 226) has omitted the species for Albania. Actually D. macroderus needs

further confirmation for the country.

19.12. Dyschirius minutus (Dejean, 1825)

19.12.1. Dyschirius minutus albanicus J. Müller, 1922

References. Dyschirius minutus albanicus: Müller, 1922b: 48, 82-83 (Berat, loc. typ.); Müller, 1937:

122; Focarile, 1959: 51; Guéorguiev & Guéorguiev, 1995: 71; Guéorguiev et al., 1997: 9. Dyschirius

punctatus albanicus: Müller, 1923: 107 (Osum River near Berat, loc. typ.); Drovenik & Peks, 1994:

25 (Berat); Drovenik & Peks, 1999: 29 (Berat). Dyschirius minutus: Balkenohl, 2003: 226.

Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Ura-Erzen”

/ “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff

1936”, 2 s. (MNHUB).

Notes. Müller published two descriptions of the subspecies albanicus. First he described D.

minutus albanicus (Müller, 1922), and next D. punctatus albanicus (Müller, 1923). Drovenik and

Peks (Drovenik & Peks, 1994; Drovenik & Peks, 1999) erroneously indicated the locality

Berat for Montenegro. Fedorenko (2001) confirmed the subspecific status of albanicus and his

point of view is followed here.

19.13. Dyschirius morio Putzeys, 1866 [= simulator J. Müller, 1922]

References. Dyschirius simulator: Müller, 1922b: 48, 90 (Berat, loc. typ.; Shkodër, loc. typ.);

Müller, 1923: 105-107 (Osum River near Berat, loc. typ.); Ieniștea, 1979: 220 (Berat); Hieke &

Wrase, 1988: 32. Dyschirius morio: Guéorguiev & Guéorguiev, 1995: 66; Balkenohl, 2003: 226.

Material studied. “Berat” / “Albania leg. Bischoff 1931”, 15 s. (MNHUB).

Notes. Müller published two description of D. simulator (Müller, 1922; Müller, 1923), the older

one is correctly treated as valid.

# Dyschirius punctatus (Dejean, 1825)

References. Dyschirius punctatus: Magistretti, 1965: 107; Balkenohl, 2003: 227.

Notes. A primarily West-Mediterranean species (Fedorenko, 1996: 161-162, Fig. 382) which

presence in Albania is highly doubtful. Likely the confusion about its presence there is due to

the view of Magistretti (1965: 106-107) who treated D. minutus as synonym of D. punctatus.

Species removed from the list of the Albanian Carabidae.

[Dyschirius pusillus (Dejean, 1825)]

Notes. An expected species for Albania as it was recently collected in North Greece (P. Bulirsch,

personal communication)


Species group “chalybeus”

19.14. Dyschirius chalybeus Putzeys, 1846

19.14.1. Dyschirius chalybeus gibbifrons Apfelbeck, 1899

References. Dyschirius gibbifrons: Apfelbeck, 1899: 290; Focarile, 1959: 48 (Shkodër; Vlorë).

Dyschirius chalybeus gibbifrons: Balkenohl, 2003: 225.

Material studied. “Alban. Wgth. Valona”, 7 s. (HNHM; MNHUB); “Skutari l. See-Ufer”, 9 s.

(HNHM; MNHUB); “Skutari 29.3.16”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”,

1 s. (MNHUB); “Liqen Butrintës” / “Albania leg. Bischoff 1936”, 10 s. (MNHUB); Tiranë

Park, 22.II.1998, 1 s., from grasses along small sluggish stream, R. Ward leg. (CMNH).

19.15. Dyschirius salinus Schaum, 1843

19.15.1. Dyschirius salinus striatopunctatus Putzeys, 1846

References. Dyschirius salinus: Apfelbeck, 1907a: 496 (Buenë River Outfall); Müller, 1922b: 72

(Durrës); Hieke & Wrase, 1988: 30. Dyschirius salinus striatopunctatus: Balkenohl, 2003: 227.

Material studied. “Berat” / “Albania leg. Bischoff 1931”, 14 s. (MNHUB); “Durrazo” /

“Albania leg. Bischoff 1931”, 21 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB): “Durrës” /


Subgenus Dyschirius s.str.


19.16. Dyschirius latipennis Seidlitz, 1867

Material included (studied P. Bulirsch). Labinoti-Fushe (Elbasan District), Shkumbinit River,

2.VII.2001, 4 s., Fencl & Moravec leg. (cPB).


Subgenus Eudyschirius Fedorenko, 1996

Species group “globosus”

19.17. Dyschirius globosus (Herbst, 1784)

References. Dyschirius globosus: Balkenohl, 2003: 229.

Species group “lafertei”

19.18. Dyschirius gracilis (Heer, 1837)

19.18.1. Dyschirius gracilis gracilis (Heer, 1837) [= lafertei Putzeys, 1846]

References. Dyschirius lafertei: Apfelbeck, 1907a: 495 (Buenë River).


Material studied. “Berat” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB).

Notes. Balkenohl (2003: 229) omitted this species for the Balkans in spite of at least three

localities from the region that were showed by Fedorenko (1996: 129, Fig. 355). P. Bulirsch

(personal communication) informed me that there is a reliable record of D. gracilis from

Northwestern Greece. The species is confirmed here for the fauna of Albania.

19.19. Dyschirius importunus Schaum, 1857

19.19.1. Dyschirius importunus importunus Schaum, 1857

References. Dyschirius importunus: Apfelbeck, 1904: 74 (Velipoja near Lezhë); Gryuntal, 1984a:

292; Hieke & Wrase, 1988: 33. Dyschirius importunus importunus: Balkenohl, 2003: 229.

Material studied. “Alban. Wgth. Valona”, 3 s. (HNHM).

Subgenus Paradyschirius Fedorenko, 1996

Species group “substriatus”

19.20. Dyschirius parallelus Motschulsky, 1844

19.20.1. Dyschirius parallelus ruficornis Putzeys, 1846

References. Dyschirius ruficornis: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 79

(Shkodër; Berat); Messner, 1964: 277; Hieke & Wrase, 1988: 31. Dyschiriodes parallelus ruficornis:

Fedorenko, 1996: 202 (Vijosë River, Tepelenë District). Dyschirius parallelus ruficornis: Balkenohl,

2003: 230.

Material studied. “Berat” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Vojusa Tepelene”


19.21. Dyschirius substriatus (Duftschmid, 1812)

19.21.1. Dyschirius substriatus priscus J. Müller, 1922

References. Dyschirius substriatus priscus: Müller, 1922b: 80 (Berat); Horion, 1941: 104; Ieniștea,

1979: 221; Hieke et al., 1986: 184 (Kula e Lumës near Kukës); Balkenohl, 2003: 230. Dyschirius

substriatus: Hieke & Wrase, 1988: 32. Dyschiriodes substriatus: Fedorenko, 1996: 204.

Tribe SCARITINI Bonelli, 1810

Subtribe SCARITINA Bonelli, 1810

20. Genus Scarites Fabricius, 1775

Subgenus Parallelomorphus Motschulsky, 1849

20.1. Scarites laevigatus Fabricius, 1792

References. Scarites laevigatus: Apfelbeck, 1904: 67 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke

& Wrase, 1988: 27; Balkenohl, 2003: 232.

COLEOPTERA: CARABIDAE: TRECHINAE 51

Material studied. “Durazzo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Himara Juni”

/ “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Liqen i Butrintes Vivari” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Valona” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); 2 km

N Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Shkodër, 12.VI.1993, 2 s.,

P. Beron & B. Petrov leg. (NMNHS).

20.2. Scarites terricola Bonelli, 1813

20.2.1. Scarites terricola terricola Bonelli, 1813

Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB);

“Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albania Sauk” / “16.VI.1958

leg. Xh. Murraj”, 1 s. (HNHM); Saranda District, Butrinti, 16.IV.1994, 1 s., S. Beshkov leg.

(NMNHS); Përmeti District, Pelombasi, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).


Subgenus Scallophorites Motschulsky, 1858

20.3. Scarites buparius (Forster, 1771)

References. Scarites buparius: Hieke et al., 1986: 184 (Vlorë).

Material examined. “Vom Alb. Amerik Institut…” / “Zweifellos Alb. Konvenienc”, 1 s.

(MNHUB); “S Albanien Vlora, Strand p.7.58 leg. Grebenscikov”, 1 s. (DEI).

Additional material included (studied A. Vigna Taglianti). Tiranë, botanical garden, 6.VI.1998,

1 s., A. Paparistos leg. (cAVT).


Subfamily APOTOMINAE LeConte, 1853

Tribe APOTOMINI LeConte, 1853

21. Genus Apotomus Illiger, 1807

21.1. Apotomus rufithorax Pecchioli, 1837

References. Apotomus rufithorax: Schatzmayr, 1943b: 99 (Mal i Krujes; Karbunara e Madhe);

Wrase, 1992: 93 (Durrës); Wrase, 2003a: 237.



m., 2 f., D. Antonelli leg. (cAVT).

21.2. Apotomus rufus (P. Rossi, 1790)

References. Apotomus rufus: Wrase, 1992: 77 (Lum i Tiranë).

Notes. Wrase (2003a: 237) has omitted the species for Albania, therefore it occurrence in the

country needs further confirmation.


Subfamily TRECHINAE Bonelli, 1810

Tribe BEMBIDIINI Stephens, 1827

Subtribe BEMBIDIINA Stephens, 1827

22. Genus Asaphidion Gozis, 1886

Notes. Hartmann has revised the material studied under this genus from HNHM.

? Asaphidion cyanicorne (Pandellé, 1867)

? Asaphidion cyanicorne cyanicorne (Pandellé, 1867)

Material studied. “Korab” / “A. cyanicornis cyanicornis det. H. Schweiger” / “Fundort wahrsch.

falsch!”, 1 m. with extracted genitalia (NMW); “Korab, Alb. Bischoff 1935” / “A. cyanicornis

cyanicornis det. H. Schweiger”, 1 m. with extracted genitalia (NMW).

Notes. Schweiger noted on the label that this locality might be untrustworthy. For the time

being his remark is taken into consideration despite A. cyanicorne has been recorded for Croatia

and Bulgaria (Mařan, 1933: 89; Marggi et al., 2003: 242).

22.1. Asaphidion flavipes (Linnaeus, 1761)

References. Asaphidion flavipes: Apfelbeck, 1904: 79 (Velipoja near Lezhë); Schatzmayr, 1943b:

95 (Tërpan); Hieke & Wrase, 1988: 42.

Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Mamuras Alban. Kruja Dr.

J. Müller”, 5 s. (NMW); “Argirocastro Albania Weirather”, 2 s. (NMW); “Lumi Bençes Tepelene,

Alb. m.”, 1 s. (NMW); “Berat Alb. Dr. Müller’18”, 1 s. (NMW); “Elbasan Alb., Mader”, 16 s.

(HNHM; NMW); “Tomor-Buranj Juli 1932” / “Albania leg. Bischoff 1932”, 12 s. (MNHUB);

“Bojana Shkoder 1934”, 8 s. (NMW); “Gjinokastrë Umg. Albania 1934 Bischoff ”, 1 ex. (NMW);

“Mal i Dajtit” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Berat” / “Albania leg. Bischoff 1936”, 5 s. (NMW; MNHUB); Saranda

District, Lyukove, 5.V.1994, 1 m., P. Stoev leg. (NMNHS); Përmeti District, Pelombasi,

2.VI.1994, 1 f., G. Blagoev leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995,

1 f., L. Penev leg. (NMNHS); Tiranë Park, 12-19.XI.1997, 1 f., R. Ward leg. (CMNH).

Notes. Marggi et al. (2003: 242) have omitted the species for Albania. Here A. flavipes is


22.2. Asaphidion nebulosum (P. Rossi, 1792)

22.1.1. Asaphidion nebulosum balcanicum Netolitzky, 1918

References. Asaphidion caraboides: Apfelbeck, 1904: 78 (Durrës). Asaphidion caraboides balcanicus:

Netolitzky, 1918: 215 (Buenë River near Shkodër; Berat); Schatzmayr, 1943b: 95 (Ura Zezes);

Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelenë; Bizë near Shëngjergji); Hieke &

Wrase, 1988: 41; Sakalian & Guéorguiev, 1997: 46; Marggi et al., 2003: 242. Asaphidion caraboides

var. balcanicus: Csiki, 1940: 212 (Kula Ljums; Gjalicë Mt.). Bembidion elegans: Murraj, 1968: 33

(Tomor Mt.).


Material studied. “Shkëlzën Alban.”, 3 s. (NMW); “Albania Buneševce” / “Akad. Balk. Exp.

Csiki 1917”, 14 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s.

(HNHM); “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 62 s. (NMW); “Skutari, Alb. Klenka”, 3

s. (NMW); “Vojusa – Shkoder 1934”, 1 s. (NMW); “Dajtit Alb. Bischoff 1935”, 8 s. (NMW);

“Cucale” / “Albania leg. Bischoff 1936”, 1 s. (NMW); Tomor Mt., 21.IX.1961, 2 m., 3 f., Xh.

Murraj leg. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).

Notes. Bonavita & Vigna Taglianti (2005) concluded that A. caraboides and A. nebulosum are

two distinct species, the first monotypic and living in Central Europe, the second polytypic

(with four subspecies) and living in South Europe and Anatolia. The record of Murraj (1968)

about Bembidion elegans is based on misidentification. Except the material mentioned above the

present author has seen further more than 40 s. in MNHUB, which remained unregistered.

22.3. Asaphidion rossii (Schaum, 1857)

References. Asaphidion rossii: Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelenë); Marggi

et al., 2003: 243.

Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Tiranë” / “Albania leg. Bischoff

1931”, 2 s. (NMW; MNHUB); “Bushek Alb. Elbasan Bischoff 1932”, more than 45 s. (NMW;

MNHUB); “Ibe Erzen” / “Albania leg. Bischoff 1932”, 2 s. (NMW; MNHUB); “Gjirakastrë

Ung. Albania 1937 Bischoff 1934”, 3 s. (NMW); “Dajtit, Albania Bischoff 1935”, 8 s. (NMW);

“Mal i Dajtit” / “Albania leg. Bischoff 1935”, 11 s. (MNHUB); “Berat” / “Albania leg. Bischoff

1936”, 2 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Vojusa –

Tepelene 1936”, 4 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);

“Lem i Kuri Ura Mes” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania

leg. Bischoff 1939”, 2 s. (MNHUB).

22.4. Asaphidion stierlini (Heyden, 1880)

References. Asaphidion flavipes: Csiki, 1940: 212 (Kula Ljums).

Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM).


23. Genus Bembidion Latreille, 1802

Subgenus Bembidion s.str.

? Bembidion humerale Sturm, 1825

References. Bembidion humerale: Murraj, 1968: 33 (Tomor Mt.).

Notes. The record of Murraj (1968) might be doubtful because of some misidentifications

found in his collection. Unfortunately, the cited material cannot be found.

? Bembidion quadrimaculatum (Linnaeus, 1761)

? Bembidion quadrimaculatum quadrimaculatum (Linnaeus, 1761) [= quadriguttatum (Fabricius, 1775)

nec Pontoppidan, 1763]

References. Bembidion quadrimaculatum: Csiki, 1940: 213 (Kula Ljums). Bembidion quadriguttatum

(Fabricius): Csiki, 1940: 213 (Kula Ljums).


Notes. In HNHM, I found single specimen identical with that cited by Csiki (1940) to belong

to B. quadripustulatum. The occurrence of B. quadrimaculatum in Albania is very probable due to

the distribution of this common species on the Balkans.

23.1. Bembidion quadripustulatum Audinet-Serville, 1821

23.1.1. Bembidion quadripustulatum quadripustulatum Audinet-Serville, 1821 [= quadriguttatum

(Olivier, 1795) nec Fabricius, 1775]

Material studied. “Skutari Bojana” / “Apfelbeck”, 1 s. (NMW); “Alban., Wgth. Valona” /

“Apfelbeck”, 1 s. (NMW); “Elbasan, Alb. 15.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Albania

Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Skutari, Alb. Klenka”, 3 s.

(NMW); “Elbasan Alb., Mader”, 1 s. (NMW); “Bojana - Ufer b. Shkoder, Alb.” / “Alb. Winkler

1934”, 1 s. (NMW); “Durazzo, Alb. Bischoff ”, 1 s. (NMW); Tiranë Park, 1.III.1998, 1 s., R.

Ward leg. (CMNH).


Subgenus Bembidionetolitzkya E. Strand, 1929

Notes. This is one of the most complex subgenera from the genus in the Balkans. Apparently,

the center of diversity of this subgenus in the Peninsula is situated in the northern half of

Albania and the adjacent territories of Montenegro and Kosovo. The taxonomical problems

with several yet not specified forms inhabiting the studied territory deserve special endeavour.

Species group “tibiale”

23.2. Bembidion rhodopense Apfelbeck, 1902

References. Bembidion rhodopense: Hieke & Wrase, 1988: 47 (Korab Mt.); Guéorguiev &

Guéorguiev, 1995: 106; Guéorguiev et al., 1997: 14; Sakalian & Guéorguiev, 1997: 46.

Material studied. “…Albanien”, 5 s. (NMW); “Shkëlzën Alb. Bischoff 1935”, 1 s. (NMW);

“Korab” / “Albania leg. Bischoff 1935”, 2 s. (NMW; MNHUB); “Korab” / “Albania leg.

Bischoff 1936”, 5 s. (NMW; MNHUB); “Korab, Alb. Guri Kuq Bischoff 1936”, 27 s. (NMW;

MNHUB).

Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for

the country.

23.3. Bembidion tibiale (Duftschmid, 1812)

References. Bembidion tibiale: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke & Wrase,

1988: 47.

Material studied. “Shkëlzën Alban.”, 1 s. (NMW); “Shkëlzën” / “Albania Bischoff 1935”, 9

s. (MNHUB).


the country.


Species group “complanatum-geniculatum”

23.4. Bembidion geniculatum Heer, 1837

23.4.1. Bembidion geniculatum geniculatum Heer, 1837

References. Bembidion geniculatum: Hieke & Wrase, 1988: 48.

Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Korab”

/ “Albania Bischoff 1935”, 1 s. (MNHUB); “Korab, Alb. Guri Kuq Bischoff 1936”, 2 s. (NMW).

Notes. Marggi et al. (2003: 245) have omitted the species for Albania. Here B. geniculatum is


23.5. Bembidion justinae Meschnigg, 1947

References. Bembidion justinae: Meschnigg, 1947: 94 (North Albanian Alps: Dedaj, 780 m, south of

Maja e Radohines Massif, near a little brook, loc. typ.); Turin, 1981: 81; Marggi et al., 2003: 245.

Notes. Finding and revising the type of B. justinae is needed to clarify its systematic position

and relationships with the related species.

? Bembidion relictum Apfelbeck, 1904

References. Bembidion relictum: Netolitzky, 1943a: 116; Hieke & Wrase, 1988: 47; Nitzu, 1995: 158.

Material studied. “Dr. Penther Prokletija” / “Sammlung Apfelbeck”, 1 f. (HNHM).

Notes. Hieke & Wrase (1988) and Nitzu (1995) noted this species for Albania. Obviously,

these authors have followed the record “Albanian (Drin, Orsk leg. Winkler)” (Netolitzky, 1943a:

116). Actually this locality is situated in Western Kosovo close to the border with Montenegro.

Due to lack of exact records B. relictum is still regarded as questionable for Albania; the material

studied above is without definite state attribution.

! Bembidion sp. (aff. complanatum Heer, 1937)

Material studied. “Jezerce K. Wgth.18”, 1 f. (NMW).

Notes. Netolitzky has identified the female above as B. complanatum Heer, 1837. I examined it

and think also that this specimen is externally most similar to B. complanatum. The female at

hand has pointed apex of elytra and a body-size of 6.6-6.7 mm. The specimen from Maja e

Jezerzë differs from B. justinae and B. relictum in the black antennomeres, excluding the reddish

scapus. The study of more male specimens, including such from the same locality, and their

comparison with specimens of B. complanatum, B. relictum and the type of B. justinae, could

establish the correct status of this form.

Species group “varicolor”

23.6. Bembidion conforme (Dejean, 1831)

References. Bembidion conforme: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Horion,

1941: 132.

Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW); Leskovac

- Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).

Notes. Marggi et al. (2003: 245) have omitted for Albania. Here it is confirmed for the country.


23.7. Bembidion meschniggi Netolitzky, 1943

References. Bembidion meschniggi: Netolitzky, 1943a: 113 (Dedaj, loc. typ.); Turin, 1981: 82;

Marggi et al., 2003: 246.

Material studied. “Albania Markdedaj Ing. Meschnigg” / “coll. Netolitzky” / “meschniggi n.

sp. det. Netolitzky” / “Type” / “Genital - Nr 458 präpariert G. Müller 1978”, 1 syntype

(NMW); “Albania Markdedaj Ing. Meschnigg” / “coll. Paul Meyer” / “Cotypus det. Netolitzky”

/ “Type” / “meschniggi n. sp. det. Netolitzky” / “meschniggi Cotypus anno 1996 revidit Miloš

Fassati”, 1 syntype (NMW); “Albania Markdedaj Ing. Meschnigg” / “coll. Netolitzky” /

“Cotypus det. Netolitzky” / “Type” / “meschniggi n. sp. det. Netolitzky” / “meschniggi Cotypus

anno 1996 revidit Miloš Fassati”, 1 syntype (NMW).

Notes. I consider B. meschniggi being a distinct species and most probably a sister taxon of B.

atrocaeruleum (Stephens, 1828), but further study should differentiate the former from the latter

and to prove its species status. There are in Albania two places with the name Dedaj (? =

Markdedaj), situated not far from each other. The first place is north of the Drin River (Shkodër

District), at the southern slopes of the North Albanian Alps, at 1012 meters altitude, whereas

the second one is south of the Drin River (Pukë District), at 500 meters altitude. For the

present one cannot specify what of both the type locality of B. meschniggi is.

23.8. Bembidion varicolor (Fabricius, 1803)

23.8.1. Bembidion varicolor varicolor (Fabricius, 1803) [= tricolor (Fabricius, 1801)]

References. Bembidion tricolor: Apfelbeck, 1904: 87 (Velipoja); Hieke & Wrase, 1988: 46.

Material studied. “Shkëlzën Alban.”, 2 s. (NMW); “Shkëlzën Alb. Alfons Bischoff ”, 1 s.

(NMW); “Shkëlzën” / “Albania Bischoff 1935”, 9 s. (MNHUB).


the country.

Species group “fasciolatum”

# Bembidion ascendens K. Daniel, 1902

References. Bembidion ascendens: Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Marggi

et al., 2003: 245.

Notes. Sure records for this species from the Balkan Peninsula are known from Slovenia,

Croatia, and Western Bosnia Herzegovina. Since no reliable records are known eastwards of

the city of Sarajevo, B. ascendens is removed from the list of the Albanian ground beetles. Most

likely, the literature data for it refer to B. brunoi.

23.9. Bembidion brunoi (Bonavita, 2001)

References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 468 (Kiri River near Mesi); Hieke et

al., 1986: 185 (Uji Ftohte, south of Tepelenë); Marggi et al., 2003: 245. Bembidion ascendens:

Hieke et al., 1986: 185 (Uji Ftohte near Tepelenë); Marggi et al., 2003: 245.

Material studied. “Skutari Mesi”, 1 f. (NMW); “Merdita Oroshi” / “Alban. spt. IV.V.’05”, 2

m., 4 f. (NMW); “Mesi Kiri Ufer” / “Albania leg. Bischoff 1931”, 1 m., 2 f. (MNHUB); 5 km

from Tepelenë, Drina River, 3.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS).


Notes. In both the shape and the internal structure of the median lobe this species seems

most related to B. fasciolatum. On the other hand, in the wider margin of the pronotum and

bigger size of the body, it resembles B. ascendens.

23.10. Bembidion concoeruleum Netolitzky, 1943

References. Bembidion coeruleum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Hieke &

Wrase, 1988: 47. Bembidion concoeruleum: Netolitzky, 1943a: 114 (Oroshi; Shkodër; Dukati); Marggi et

al., 2003: 245. Bembidion coeruleum concoeruleum: Hieke et al., 1986: 185 (Borshi near Vlorë; Uji Ftohte,

south of Tepelenë). Ocydromus concoeruleus: Bonavita & Vigna Taglianti, 1993: 84.

Material studied. “Orosi (Ptr.) Alban. spt.”, 5 s. (HNHM; NMW); “Skutari Mesi” / “Apfelb.

Alban. spt. IV.V.’05”, 1 s. (NMW); “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s.

(NMW); “Prokletija”, 1 s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze,

Penther’14”, 1 s. (NMW); “Dukati”, 1 s. (NMW); “Skutari, Alb. Klenka”, 4 s. (NMW); “Skutari,

Alb. Mader”, 1 s. (NMW); “Lumi, Benges Tepelene, Alb. m.”, 2 s. (NMW); “Argirokastron

Albania mai 1929 Weirather”, 6 s. (NMW); “Lumi Dukati Alban. mer.” / “Winkler, Lona,

Bischoff V.1931”, 1 s. (NMW); “Korab, Alb. Guri Kuq Bischoff 1936”, 2 s. (MNHUB); Tepelenë

District, Vijosë River, 2.VI.1994, 2 m., G. Blagoev leg. (NMNHS).

Notes. All previously published data for B. coeruleum Audinet-Serville, 1821 from the Balkans

refer to this species (see also Bonavita & Vigna Taglianti, 1993).

# Bembidion fasciolatum (Duftschmid, 1812)

References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i

Madh River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte near Tepelenë); Marggi et al.,

2003: 245.

Notes. I failed to find material of this species from the Balkan Peninsula in the collections,

excluding a single specimen from the locality “Travnik”. There are two places with this

name - one in Slovenia and one in Bosnia Herzegovina. B. fasciolatum is removed from the

list of the Albanian Carabidae. The records cited for it in the past most probably concern

either an unnamed species (the record near Oroshi) or B. brunoi (the records by Mesi River

and in South Albania).

! Bembidion sp. (ex. gruppo “fasciolatum”)

References. Bembidion fasciolatum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi);

Marggi et al., 2003: 245.

Material studied. “Orosi (Ptr.) Alban. spt.”, 6 s. (DEI; HNHM; MNHUB).

Notes. The specimens from DEI and HNHM have been identified as Bembidion merditanum

Apfelbeck as two of them from the first depository were incorrectly designated as syntypes

of this form. In fact, Bembidion merditanum Apfelbeck is nomen nudum. The series from Oroshi

may belong to separate species.

[subgenus Bracteon Bedel, 1879]

[Bembidion litorale (Olivier, 1790) [= elegans Germar, 1824]]

References. Bembidion elegans: Murraj, 1968: 33 (Tomor Mt.).

Notes. Murraj (1968) cited this species from the Tomor Mt. Among the material loaned from

MNST, I found five specimens from this locality bearing label data as the published ones and


identified by the author as B. elegans. In fact, all the specimens concern Asaphidion nebulosum

balcanicum. For the present B. litorale is treated as an expected species for Albania because it

inhabits most of the Balkan countries.

Subgenus Chlorodium Motschulsky, 1864

23.11. Bembidion splendidum Sturm, 1825

23.11.1. Bembidion splendidum pincum De Monte, 1957

References. Bembidion splendidum: Apfelbeck, 1907a: 495 (Buenë River).

Material studied. “Scutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 5 s. (NMW); “Vojusa

Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg.

Bischoff 1937”, 3 s. (NMW).


the country. According to Bonavita and Vigna Taglianti (2005) the Balkan populations of the

species belong to ssp. pincum.

Subgenus Diplocampa Bedel, 1896

23.12. Bembidion assimile Gyllenhal, 1810

References. Bembidion assimile: Apfelbeck, 1904: 111 (Velipoja); Hieke & Wrase, 1988: 57.

Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM).


the country.

23.13. Bembidion fumigatum (Duftschmid, 1812)

References. Bembidion fumigatum: Apfelbeck, 1904: 111 (Velipoja near Lezhë); Hieke & Wrase,

1988: 57.

Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Actually, further

confirmation for the country is needed.

Subgenus Emphanes Motschulsky, 1850

23.14. Bembidion axillare (Motschulsky, 1844)

23.14. Bembidion axillare occiduum Margi & Huber, 2001 [= rivulare (Dejean, 1831)]

References. Bembidion rivulare: Hieke & Wrase, 1988: 58.

Material studied. “Durrës Alban.”, 1 s. (NMW); “Albanien Ruskuli 24.IX.18 Dr. H. Priesner”,

1 s. (NMW); “Ruskuli Alb., Mader”, 1 s. (NMW); “Durazzo, Alb. Ravasini, Lona”, 2 s. (HNHM);

“Durrazo, Alb. Bischoff ”, 4 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 9 s.

(MNHUB); “Valona, Alb. Bischoff 1936”, 2 s. (NMW); Radona, Shkiela, 21.VI.1994, 3 s., B.

Guéorguiev leg. (NMNHS).

Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here B. axillare is



23.15. Bembidion azurescens Dalla Torre, 1877

23.15.1. Bembidion azurescens azurescens Dalla Torre, 1877

References. Bembidion azurescens: Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Kula e Lumës

near Kukës); Hieke & Wrase, 1988: 58. Bembidion azurescens azurescens: Marggi et al., 2003: 248.

Material studied. “Oboti Schirel” / “Apfelbeck”, 1 s. (NMW); “Berat, Alb. Ravasini, Lona”,

3 s. (NMW); “Tepelene” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Vojusa Tepelene” /

“Albania leg. Bischoff 1936”, 8 s. (NMW).

23.16. Bembidion latiplaga Chaudoir, 1850

23.16.1. Bembidion latiplaga latiplaga Chaudoir, 1850

References. Bembidion latiplaga: Apfelbeck, 1907a: 495 (Buenë River); Hieke & Wrase, 1988: 59.

Material studied. “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Oboti Schirei”

/ “Apfelbeck”, 1 s. (NMW); “Skutari Alb. Dr. J. Müller”, 1 s. (NMW); “Vojusa Tepelene” /

“Albania leg. Bischoff 1936”, 1 s. (NMW; MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff

1937”, 3 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb. Winkler 1937”, 1 s. (NMW); “Bojana-

Ufer b. Shkoder, Alb.” / “dch. Winkler XI 1942”, 1 s. (NMW); “Bojana Oboti, Alb.” / “ex coll.

Winkler 1948”, 1 s. (NMW); “Tirana Ungb. Albania” / “ex coll. Winkler 1948”, 2 s. (NMW).


the country.

23.17. Bembidion normannum Dejean, 1831

23.17.1. Bembidion normannum mediterraneum Csiki, 1928 [= meridionale Ganglbauer, 1891]

References. Bembidion normannum meridionale: Apfelbeck, 1904: 103 (Velipoja near Lezhë).

Bembidion normannum apfelbecki: Müller-Motzfeld, 1986a: 262 (Raskuli); Hieke & Wrase, 1988:

59. Bembidion normannum mediterraneum: Hieke & Wrase, 1988: 59; Marggi et al., 2003: 249.

Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM).

Notes. Hieke & Wrase (1988) have noted that both subspecies of B. normannum inhabit Albania.

However, I consider the material studied and reference data as belonging to ssp. mediterraneum

(see also Marggi et al., 2003: 249).

23.18. Bembidion tenellum Erichson, 1837

23.18.1. Bembidion tenellum tenellum Erichson, 1837

References. Bembidion tenellum: Csiki, 1940: 213 (Kula Ljums).

Material studied. “Skutari Vraka” / “Apfelbeck”, 1 s. (NMW); “Oboti Schiren” / “Apfelbeck”,

1 s. (NMW); “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 7 s. (HNHM); “Albania

Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” / “Akad.

Balk. Exp. Csiki 1918”, 7 s. (HNHM); “Skutari, Alb. Dr. J. Müller”, 3 s. (NMW); “Berat, Alb.

Ravasini, Lona”, 1 s. (NMW); “Liqen Butrintës Alb. 1936. Bischoff ”, 1 s. (NMW); “Bojana

Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb.

Winkler 1937”, 1 s. (NMW); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Tepelenë

District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 /

22.II.1998, 2 s., from grasses along small sluggish stream, R. Ward leg. (CMNH).



the country.

Subgenus Euperyphus Jeannel, 1941

23.19. Bembidion combustum Ménétries, 1832

23.19.1. Bembidion combustum combustum Ménétries, 1832

References. Bembidion combustum: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi);

Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 48. Bembidion

combustum combustum: Marggi et al., 2003: 249.

Material studied. “Albania, Berat”, 1 s. (NMW); “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”,

9 s. (NMW); “Oroši (Ptr.) Alban. spt.”, 11 s. (HNHM; MNHUB); “Berat, Alb. Ravasini, Lona”, 3 s.

(NMW); “Skutari, Alb. Klenka”, 1 s. (NMW); “Tiranë, Alb. Alf. Bischoff ”, 2 s. (NMW); “Cucale

Geb.” / “Albania leg. Bischoff 1931”, 1 s. (NMW); “Elbasan Juni” / “Albania leg. Bischoff 1932”,

1 s. (MNHUB); “Lum i Tiranës Alb. V.1934 Mjr. Strupi”, 2 s. (NMW); “Lum Erzen Alb. V.1934

Mjr. Strupi”, 2 s. (NMW); “Tiranë” / “lgt. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “lgt.

Bischoff Albania 1935”, 2 s. (MNHUB); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST).

23.20. Bembidion scapulare Dejean, 1831

23.20.1. Bembidion scapulare oblongum Dejean, 1831

References. Bembidion oblongum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi).

Bembidion scapulare oblongum: Netolitzky, 1943b: 17; Hieke & Wrase, 1988: 50.

Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW); “Berat Alb.

Dr. J. Müller’18”, 1 s. (NMW); “Elbasan, Alb. 4.IX.18. Dr. H. Priesner”, 1 s. (NMW); “Elbasan

Alb., Mader”, 3 s. (NMW); “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s.

(NMW); “Elbasan Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomorica” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 2 s. (NMW;

MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).


the country. Presently the subspecific status of the Balkan populations remains unsettled.

23.21. Bembidion testaceum (Duftschmid, 1812)

23.21.1. Bembidion testaceum testaceum (Duftschmid, 1812)

Material studied. “Berat, Alb. Ravasini, Lona”, 1 s. (NMW); “Bojana-Schkodër Alb., Bischoff

1934”, 1 s. (NMW).


Subgenus Metallina Motschulsky, 1850

23.22. Bembidion lampros (Herbst, 1784)

References. Bembidion lampros: Apfelbeck, 1904: 83 (Velipoja; Vlorë); Apfelbeck, 1909: 297

(Vlorë; Sop); Csiki, 1940: 212 (Gjalicë Mt.); Hieke & Wrase, 1988: 44; Marggi et al., 2003: 251.


Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

North Albanian Alps, Bogë, 1000-1100 m, 30.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS);

North Albanian Alps, Bogë, 1600 m, near cave B 52, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS);

Tiranë Park, 22-31.X.1997 / 1-9.IX.1997 / 8.II.1998 / 11.II.1998 / 15.II.1998 / 22.II.1998, 9

s., from grass and soil around tree trunks, R. Ward leg. (CMNH).

Notes. Except the specimens mentioned above, I have seen additionally nearly 100 s. in NMW

and MNHUB, which remained unregistered.

23.23. Bembidion properans (Stephens, 1828)

References. Bembidion lampros var. properans: Csiki, 1940: 212 (Kula Ljums). Bembidion properans:


Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Elbasan

Alb., Mader”, 1 s. (HNHM); “Lumi i Tiranës” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);

“Mal i Dajtit” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff

1935”, 4 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 2 s. (MNHUB);

Tiranë Park, 8.II.1998, 2 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).


the country. Except the specimens mentioned above, in NMW the present author has seen

other specimens, which remained not registered.

Subgenus Microserrullula Netolitzky, 1921

23.24. Bembidion quadricolle (Motschulsky, 1844) [= inserticeps Chaudoir, 1850; cyaneomi-

cans Mařan, 1933]

References. Bembidion inserticeps: Apfelbeck, 1907a: 495 (Buenë River). Bembidion quadricolle:

Hieke & Wrase, 1988: 42; Nitzu, 2003: 180.

Material studied. “Albania Scutari”, 1 s. (MNHUB); “Skutari Bojana”, 2 s. (HNHM); “Skutari

Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 5 s. (NMW; MNHUB); “Berat, Alb. Ravasini, Lona”,

1 s. (NMW); “Berat, Alb. Ravasini, Lona” / “cyaneomicans det. Paul Meyer”, 1 s. (NMW); “Berat”

/ “Albania leg. Bischoff 1931”, 1 s. (NMW); “Lumi Berati Alban. mer.” / “Winkler, Lona,

Bischoff, V.1931”, 1 s. (NMW); “Tepelene” / “Albania leg. Bischoff 1932”, 5 s. (NMW;

MNHUB); “Ura Erzen” / “Albania leg. Bischoff. 1932”, 2 s. (NMW); “Bojana Shkoder” /

“Albania leg. Bischoff 1934”, 2 s. (NMW).

Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here B. quadricolle is


Subgenus Nepha Motschulsky, 1864

23.25. Bembidion callosum Küster, 1847

23.25.1. Bembidion callosum callosum Küster, 1847 [= laterale Dejean, 1831; pseudolaterale Netolitzky,

1943a: 103]

References. Bembidion laterale: Apfelbeck, 1904: 108 (Velipoja). Bembidion laterale pseudolaterale:

Netolitzky, 1943a: 103.


Material included (studied P. Bonavita). “Maja e Tartarit” / “Albania leg. Bischoff 1936”, 1 f. (NMW).


the country.

23.26. Bembidion caucasicum (Motschulsky, 1844) [= substriatum Chaudoir, 1846]

References. Bembidion substriatum: Apfelbeck, 1907a: 501 (Merdita); Csiki, 1940: 213 (Shkëlzen

Mt.; Gjalicë Mt.). Bembidion caucasicum: Müller, 1937: 124; Hieke & Wrase, 1988: 56.

Material studied. “Merdita Munela”, 2 s. (NMW); “Albania Galica Lums” / “Akad. Balk.

Exp. Csiki 1917”, 4 s. (HNHM); “Albania, Mts Gyalicalums” / “Akad. Balk. Exp. Csiki 1917”,

1 s. (HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Tomor:

Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Gjallica, Alb. Bischoff ”,

1 s. (NMW); “Maja e Jezercë Albania Bischoff leg.”, 2 s. (NMW); “Korab, Guri Kua Alban.

Bischoff 35”, 5 s. (NMW); “Shkëlzen, Alb. Bischoff 1935”, 1 s. (NMW); “Maja e Policanit” /

“Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (NMW); “Korab” / “Albania leg. Bischoff

1936”, 1 s. (NMW); “Shkëlzen, Alb. Bischoff 5.8.1936”, 3 s. (HNHM; NMW); North Albanian

Alps, Maj’e Radohimes, 2200-2400 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-e-

Rubës, 2000 m, 8.VII.1993, 2 f., L. Latella leg. (cAVT).


the country. Besides the material studied above, in NMW and MNHUB I have seen other

numerous specimens from the massifs of the North Albanian Alps, Korab Mt., Koritnik Mt.,

Gjalicë Mt. and Maja e Policanit Mt.

23.27. Bembidion vseteckai Mařan, 1936

23.27.1. Bembidion vseteckai dissimile G. Müller, 1943

References. Bembidion genei illigeri: Müller, 1943: 13; Hieke et al., 1986: 185 (Uji Ftohte, south

of Tepelenë; Kula e Lumës near Kukës); Hieke & Wrase, 1988: 55. Bembidion illigeri: Marggi et

al., 2003: 253.

Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Elbasan,

Alb. Leop. Mader leg.”, 2 s. (NMW); “Kruja Alb. Leop. Mader”, 2 s. (NMW); “Terbaci Alban.

mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); Mal i Dajtit Mt., 6.V.1962, 6 s. (MNST).

Notes. Except the specimens studied above I have seen several additional ones in NMW,

which remained unregistered. Bonavita (2005), as well P. Bonavita and A. Vigna Taglianti

(personal communication), noticed that all records or literature data about B. genei illigeri

Netolitzky, 1914 from the southern parts of Balkan Peninsula concern B. vseteckai dissimile.

Subgenus Notaphemphanes Netolitzky, 1920

23.28. Bembidion ephippium (Marsham, 1802)

References. Bembidion ephippium: Hieke & Wrase, 1988: 46.

Material studied. “Valona, Albania”, 4 s. (NMW); “Durazzo Alb. Dr. J. Müller”, 5 s. (NMW);

“Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Durres” / “Albania leg. Bischoff

1932”, 9 s. (MNHUB).



the country with first detailed records.

Subgenus Notaphus Dejean, 1821

23.29. Bembidion varium (Olivier, 1795) [= rumelicum Apfelbeck, 1902; dyrrhachii De

Monte, 1947]

References. Bembidion varium: Apfelbeck, 1894: 547; Apfelbeck, 1904: 85 (Velipoja near Lezhë);

Hieke & Wrase, 1988: 46; Marggi et al., 2003: 254. Bembidion varium dyrrhachii: De Monte, 1947:

52 (Durrës, loc. typ.). Bembidion rumelicum: Guéorguiev & Guéorguiev, 1995: 100.

Material studied. “Bojana Oboti, Alb.”, 1 s. (NMW); “Oboti Schirei” / “Apfelbeck”, 1 s.

(NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 12 s. (MNHUB); “Durres” / “Albania leg.

Bischoff 1932”, 7 s. (MNHUB); “Drin Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW);

Radona, Shkiela, 21.VI.1994, 4 s., B. Guéorguiev leg. (NMNHS); Tiranë Park, 22.II.1998, 30

s., from grasses along small sluggish stream, R. Ward leg. (CMNH).

Notes. Except the material studied above the present author has seen more specimens in

NMW, which remained unregistered.

Subgenus Ocydromus Clairville, 1806

23.30. Bembidion aetolicum Apfelbeck, 1901

23.30.1. Bembidion aetolicum aetolicum Apfelbeck, 1901

References. Bembidion aetolicum: Netolitzky, 1943b: 12 (Tomor Mt.). Bembidion aetolicum

aetolicum: Müller-Motzfeld, 1996b: 164 (Tomor Mt.: Buranj; Tomor Mt.: Kulmak); Marggi

et al., 2003: 254.

Material studied. “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Tomor Buranj Alb.

10.4.1932 Alf. Bischoff ”, 4 s. (NMW); “Barshë-i-Vjetër Wildbach-Quellen Bischoff

21.VII.1940”, 2 s. (NMW).

23.31. Bembidion decorum (Zenker in Panzer, 1799)

23.31.1. Bembidion decorum decorum (Zenker in Panzer, 1799)

References. Bembidion decorum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i Madh

River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Lurë e Vjetër near

Lura Mt.); Hieke & Wrase, 1988: 49. Bembidion decorum decorum: Müller-Motzfeld, 1996b: 153

(Shkodër; Mal i Dajtit Mt.; Shkëlzen Mt.; Bushek near Elbasan; Tiranë; Zeza-Brüche near

Kruja; Lumi i Bençes near Tepelenë; Tomor Mt.: Kulmak; Miloti; Gjirokastër; Prespa Lake;

Cafa Stams; Kiri River near Mesi); Marggi et al., 2003: 255.

Material studied. Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS);

Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS).

Notes. Except the specimens studied above, I have seen several more ones in NMW, which



23.32. Bembidion siculum Dejean, 1831

23.32.1. Bembidion siculum smyrnense Apfelbeck, 1904

References. Bembidion siculum: Apfelbeck, 1907a: 497-498 (Maranajt Mt.: top region, 1500-1576

m; Fani i Madh River near Oroshi); Hieke et al., 1986: 185 (Lurë e Vjetër near Lura Mt.); Hieke

& Wrase, 1988: 49. Bembidion siculum smyrnense: Müller-Motzfeld, 1996b: 170 (Shkodër; Maje

Shbogut; Himara; Llogora; Radavac; Sarapta; Lura Mt.: Lurë e Vjetër); Marggi et al., 2003: 256.

Material studied. “Merdita Oroschi”, 3 s. (NMW); “Oroši (Ptr.) Alban. spt.”, 5 s. (HNHM);

“Winkler, Lona, Bischoff, V.1931” / “Terbaci Alban. mer.”, 1 s. (NMW); “Llogora” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Himara Juni” / “Albania leg. Bischoff 1932”, 2 s.

(MNHUB); Përmeti District, Pelombashi, 2.VI.1994, 3 s., G. Blagoev leg. (NMNHS).

Notes. Besides the material studied above, I have seen other more in NMW, which remained

unregistered.

Subgenus Ocyturanes Müller-Motzfeld, 1986

23.33. Bembidion balcanicum Apfelbeck, 1899

23.33.1. Bembidion balcanicum balcanicum Apfelbeck, 1899

References. Bembidion balcanicum: Müller, 1918b: 85 (Munella Mt.); Müller, 1926: 61 (Munella

Mt.); Csiki, 1940: 213 (Gjalicë Mt.; Korab Mt.); Horion, 1941: 136; Netolitzky, 1943a: 42

(Korab Mt.); Hieke & Wrase, 1988: 52. Bembidion balcanicum balcanicum: Marggi et al., 2003: 256.

Material studied. “Latif Koritnik”, 11 s. (HNHM; NMW); “Merdita Munela”, 1 s. (HNHM);

“Merdita Munela” / “Wgth 1906 Merdita”, 10 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” /

“Alban. montenegr. Grenze, Penther,’14”, 2 s. (NMW); “Albania Galica Lums” / “Akad. Balk.

Exp. Csiki 1917”, 1 s. (HNHM); “Albania, Mts. Gyalicalums” / “Akad. Balk. Exp. Csiki 1918”,

21 s. (HNHM); “Alban Exped. Gialica Ljums 17.-26.VI’18”, 2 s. (NMW); “Albania Mts. Korab”

/ “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Maja e Jezerce Albania Bischoff leg.”, 4 s.

(NMW); “Maja e Jezerce Alban. sept.” / “Alb. Winkler 1934”, 1 s. (NMW); “Albania Korab

1935 Bischoff leg.”, 1 s. (NMW); “Giallica, Alb. Bischoff 1935”, 3 s. (NMW); “Tomor - Tojk

Bischoff Alban.”, 3 s. (NMW); “Tomor - Tojk Bischoff 1935”, 3 s. (NMW); “Maja Policanit”

/ “Albania 1936 leg. Ramadan coll. Bischoff ”, 3 s. (NMW); “Shkelzën, Alb. Bischoff, 5.8.1936”

/ “Albania leg. Bischoff 1936”, 8 s. (HNHM; NMW).

Notes. Besides the material studied above, in NMW and MNHUB I have seen numerous

other specimens from the massifs of Shkëlzën Mt., Gjalicë Mt., Korab Mt. and Tomor Mt.,

which remained unregistered.

23.34. Bembidion gudenzii (Neri, 1981)

References. Bembidion gudenzii: Müller-Motzfeld, 1985: 225 (“Hochgebirge bei Valona (Epyrus)”).

Notes. Marggi et al. (2003: 256) have omitted B. gudenzii for Albania. Actually, the species

needs further confirmation for the country.

23.35. Bembidion pindicum Apfelbeck, 1901

References. Bembidion pindicum: Netolitzky, 1943a: 49 (Korab Mt.); Guéorguiev, 1999: 71.


Material studied. “Korab, Guri-Kua Alban. Bischoff 35”, 1 s. (NMW); “Tomor-Tojk Alban

1935 Bischoff leg.” / “pindicum var.? dtt. Netolitzky”, 1 s. (NMW); “Tomor - Tojk Alban. 1935

Bischoff leg.”, 2 s. (NMW).

Notes. Marggi et al. (2003: 257) have omitted the species for Albania. The specimens studied

have been found to be conspecific with three syntypes of B. pindicum labelled “Apfelbeck

Peristeri” / “(Peryphus) pindicum typ. Apf.” / “Typus” (NMW), therefore the species is here

confirmed for Albania.

23.36. Bembidion praeustum Dejean, 1831

23.36.1. Bembidion praeustum praeustum Dejean, 1831

References. Bembidion praeustum: Hieke et al. 1986: 186 (Uji Ftohte, soth of Tepelenë); Hieke

& Wrase, 1988: 52. Bembidion praeustum praeustum: Marggi et al., 2003: 257.

Material studied. “Dukati” / “Apfelbeck”, 1 s. (NMW); “Tomor: Kulmak Albania mer.” /

“Ravasini & Lona”, 1 s. (NMW); “Lumi i Bençes Tepelene, Alb. m.” / “Alb. Winkler”, 1 s.

(NMW); “Bushek b. Elbasan, Alb. Alf. Bischoff ”, 1 s. (NMW); “Maj’e Shbogut Alnania mer.” /

“Winkler, Lona, Bischoff, V.1931”, 2 s. (NMW); “Humara juni” / “Albania leg. Bischoff 1932”,

2 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB); “Maranaj, Alb. Bischoff,

1934”, 1 s. (NMW); “Lumi i Tiranës Alb. V.1934 Mjr. Strupi”, 1 s. (NMW); “Köleyrë” / “Albania

leg. Bischoff 1936”, 1 s. (NMW); “Boge ober d. Quelle” / “Albania leg. Mueller 1939”, 1 s.

(NMW); “Borshë-i-Vjetër Wildbach-Quellen Bischoff 21.VII.1940”, 8 s. (HNHM; NMW);

“Borshë, Alb. Bischoff 1940”, 1 s. (NMW); “Tirane, Alb. dch. Winkler 42”, 1 s. (NMW); Tomor

Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Përmeti District, Pelombasi, 2.VI.1994, 1 s., G.

Blagoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS);

Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS).

Notes. A part of the material in NMW has been determined as B. praeustum viridifluum J. Müller,

1929. However, I refer provisionally all the specimens studied to the nominotypical race.

23.37. Bembidion signatipenne Jacquelin du Val, 1852

References. Bembidion signatipenne: Hieke & Wrase, 1988: 53.

Notes. Marggi et al. (2003: 257) have omitted B. signatipenne for Albania. Actually, the species

needs further confirmation for the country.

23.38. Bembidion stolfai J. Müller, 1943

References. Bembidion stolfai: Müller, 1943: 12 (North Albanian Alps: Okol near Bogë, loc.

typ.; Maja e Jezercë); Netolitzky, 1943a: 41 (North Albanian Alps: Okol near Bogë; Maja e

Jezercë); Marggi et al., 2003: 257.

Material studied. “Maja e Jezerce Albania sept.” / “Alb. Winkler 1934” / “coll. Netolitzky”

/ “stolfai m. n. sp. det. J. Müller” / “Co-typus” / “Coll. Netolitzky”, 1 f. syntype (NMW).

Notes. The syntype of B. stolfai was studied and compared with specimens of B. reiseri reiseri

Apfelbeck, 1902 from Bosnia Herzegovina (including a syntype labelled “reis. 1900 Cvrstica”

/ “Testediolum bosnicum typ. Apf.” / “Reiseri” / “Typus”), topotypical material of B. reiseri vranense

Apfelbeck, 1902 and extensive material of B. balcanicum balcanicum. B. stolfai is distinct from the

other two species in the pronotum wider than long being narrower in the anterior part and in


the elytra having distinct transverse microsculpture. Besides, it differs from B. reiseri s.l. in the

wrinkles on outer basal fovea of pronotum clear and long (vs. reduced), the shoulders of

elytra distinct (vs. round) and the sides of elytra after middle parallel (vs. slightly round). B.

stolfai is also living sympatrically with B. balcanicum balcanicum in the massif of Maja e Jezercë.

Subgenus Odontium LeConte, 1848

23.39. Bembidion foraminosum Sturm, 1825

References. Bembidion foraminosum: Hieke & Wrase, 1988: 43.

Material studied. “Tiranë” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB).


the country with first detailed record.

23.40. Bembidion striatum (Fabricius, 1792)

References. Bembidion striatum: Netolitzky, 1918: Beilage zu Heft 4-6 (Shkodër; Oboti);

Geisthardt, 1975: 8. Bembidion striatul sic!: Panin, 1942: 567 (Shkodër).

Material studied. “Oboti Schirei”, 1 s. (NMW); “Skutari Bojana”, 5 s. (NMW); “Skutari

Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW).


the country.

Subgenus Omoperyphus Netolitzky, 1931

23.41. Bembidion hypocrita Dejean, 1831

23.41.1. Bembidion hypocrita illyricum Netolitzky, 1918

References. Bembidion hypocrita illyricum: Müller, 1918b: 89; Müller, 1926: 66; Netolitzky, 1943a: 122.

Material studied. “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Skutari, Alb. Mader”,

1 s. (NMW); “Argyrókastron, Alb. leg. Weirather”, 1 s. (NMW); “Mal i Dajtit” / “Albania leg.

Bischoff 1935”, 3 s. (NMW; MNHUB); “Shkelzën, Alb. Bischoff, 5.8.1936”, 1 s. (NMW);

“Bischoff 21.VII.1940 Börshë-i-Vjetër Wildbach-Quellen”, 7 s. (NMW); “Börshë-i-Vjetër Alb.

21.7.1940 Bischoff ”, 1 s. (MNHUB).



Subgenus Peryphanes Jeannel, 1941

23.42. Bembidion brunnicorne Dejean, 1831

23.42.1. Bembidion brunnicorne brunnicorne Dejean, 1831

References. Bembidion brunnicorne: Netolitzky, 1943a: 50; Csiki, 1940: 213 (Kula Ljums).

Material studied. “Albania Kula Ljums”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk.

Exp. Csiki 1918”, 7 s. (HNHM); “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Kruja,

Alban., Mader”, 1 s. (NMW); “Tomor: Kulmak Albania mer.”, 3 s. (NMW); “Mal i Dajtit” /


“Albania leg. Bischoff 1935”, 2 s. (HNHM; MNHUB); “Maranaj 1937” / “Albania bei Skutari”,

1 s. (NMW); “Börshë i Vjetër Wildbach-Quellen Bischoff, 21.VII.1940”, 4 s. (NMW).


the country. Besides the material listed above, I have seen additionally more than 70 s. from

different localities in MNHUB, which remained unregistered. All the material in the last museum

was identified as B. milleri.

23.43. Bembidion dalmatinum Dejean, 1831

23.43.1. Bembidion dalmatinum dalmatinum Dejean, 1831

References. Bembidion dalmaticum sic!: Anonymous, 1907: 286. Bembidion dalmatinum: Apfelbeck,

1907a: 498 (Fani i Madh River near Oroshi); Csiki, 1940: 213 (Gjalicë Mt.); Schatzmayr, 1943b:

95 (Dukati; Lushnje); Hieke & Wrase, 1988: 53. Chlaenius nigricornis: Murraj, 1968: 33 (Tiranë;

Tomor Mt.). Bembidion dalmatinum dalmatinum: Marggi et al., 2003: 258.

Material studied. Tomor Mt., 21.IX.1961, 9 s., Xh. Murraj leg. (MNST); Theth, 800-1200 m,

28.V.1993, 7 s., P. Beron & B. Petrov leg. (cPN; NMNHS); North Albanian Alps, Bogë, 1400-

1800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë,

1000-1100 m, 5-9.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District, Merkurth,

11.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); Përmeti District, Pelombashi, 2.VI.1994,

1 s., G. Blagoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 27 s., G. Blagoev

leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS);

Leskoec - Barmashi, 880 m, 2.VI.1995, 2 s., G. Blagoev leg. (NMNHS); 5 km Tepelenë, Drina

River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Ljukove near Vlorë, 6.VI.1995, 4 s., G.

Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH).

Notes. About the record of Murraj (1968) above see the notes under Chlaenius nigricornis.

Besides the material studied, the present author has seen numerous other specimens in NMW

and MNHUB, which remained unregistered.

23.44. Bembidion deletum Audinet-Serville, 1821

23.44.1. Bembidion deletum deletum Audinet-Serville, 1821

Material studied. “Borshë, Alb. Bischoff 1940”, 1 s. (NMW).


23.45. Bembidion grandipenne Schaum, 1862

23.45.1. Bembidion grandipenne grandipenne Schaum, 1862

References. Bembidion grandipenne: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi);


Material studied. “Oroschi (Ptr.) Alban. spt.”, 4 s. (HNHM; MNHUB); “Merdita Oroschi” /

“Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Albania Malčija” / “Akad. Balk. Exp. Csiki 1917”,

1 s. (HNHM); “Skutari Alb., Mader”, 1 s. (NMW); “Lumi i Bençes Tepelene, Alb. m.”, 2 s.

(NMW); “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Ibe-Erzen” / “Albania leg. Bischoff

1932”, 2 s. (NMW; MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB); “Shkëlzën, Alb. Bischoff 1935”, 1 s. (NMW); “Borshë-i-Vjetër Wildbach-Quellen

Bischoff 21.VII.1940”, 4 s. (NMW; MNHUB); “Borshë, Alb. Bischoff 1940”, 1 s. (NMW).



the country.

# Bembidion milleri Jacquelin du Val, 1852

# Bembidion milleri milleri Jacquelin du Val, 1852

References. Bembidion milleri: Hieke & Wrase, 1988: 54.

Notes. According to Müller-Motzfeld (personal communication in 1994) B. milleri does not

live in the Balkans. P. Bonavita and A. Vigna Taglianti (personal communication) stated that it

occurs only in Slovenia and probably in Serbia. Here the upper opinions are followed, though

Marggi et al. (2003: 258-259) noted the species from the Balkan Peninsula.

Subgenus Peryphus Dejean, 1821

23.46. Bembidion bualei Jacquelin du Val, 1852

23.46.1. Bembidion bualei bualei Jacquelin du Val, 1852 [= albanicum J. Müller, 1935]

References. Bembidion andreae: Apfelbeck, 1904: 92 (Durrës). Bembidion andreae albanicum: Müller,

1935: 152 (Berat, loc. typ.; Tiranë; Shkodër); Netolitzky, 1943a: 27 (Berat); Hieke et al. 1986:

186 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 51; Guéorguiev & Guéorguiev,

1995: 111; Guéorguiev et al., 1997: 15. Bembidion andreae bualei: Panin, 1942: 582 (Berat);

Schatzmayr, 1943b: 95 (Ura Zezes). Bembidion cruciatum albanicum: Marggi et al., 2003: 260.

Material studied. “Alban. Exped. Gjalica Ljuma 17.-26.VI.’18”, 6 s. (NMW); “Bojana Shkodër

Alb., Bischoff 1937” / “Albania leg. Bischoff 1937”, 1 s. (HNHM).

Notes. According to a recent study (Coulon, 2006) the Balkan populations of the species

related to B. cruciatum Dejean, 1831 belong to B. bualei bualei, and the form albanicum is a synonym

of the latter. Except the material studied above, I have seen more specimens from different

localities in NMW and MNHUB, which remained unregistered.

23.47. Bembidion femoratum Sturm, 1825

23.47.1. Bembidion femoratum femoratum Sturm, 1825

Material studied. “Scutari, Alb. Klenka”, 1 s. (NMW); “Bojana, Alb. Shkodër Bischoff 1934”,

31 s. (NMW); “Albania leg. Bischoff 1937”, 1 s. (NMW); “Bojana Shkodër Albania leg. Bischoff

1937”, 4 s. (NMW); 5 km from Tepelenë, Drina River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS).


23.48. Bembidion subcostatum (Motschuslky, 1850)

23.48.1. Bembidion subcostatum vau Netolitzky, 1913 [= javurkovae Fassati, 1944]

References. Bembidion subcostatum: Hieke & Wrase, 1988: 52. Bembidion subcostatum javurkovae: Hieke

et al. 1986: 186 (Borshi near Vlorë; Lurë e Vjetër near Lura Mt.); Marggi et al., 2003: 263.

Material studied. “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 4 s. (NMW); “Scutari Alb. Dr.

J. Müller”, 1 s. (NMW); “Berat, Alb. sch. Winkler”, 1 s. (NMW); “Tirana Alb. dch. Winkler 42”,

2 s. (NMW); “Bushek Juni” / “Berat Alban. mer.” / “Albania leg. Bischoff 1932”, 4 s. (NMW);

Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994,

1 s., B. Guéorguiev leg. (NMNHS).


Notes. Netolitzky identified the specimens in NMW as ssp. vau, ssp. javurkovae Fassati, 1944 or

ssp. spartanum Fassati, 1944. However, in a forthcoming paper these three taxa are synonymized

(Guéorguiev & Lobo, in press). In HNHM I have seen numerous additional specimens mostly

from the localities “Korab” and “Kula Lums”, and in MNHUB - other more than 200 s., as all

those data remained unregistered.

23.49. Bembidion tetracolum Say, 1823

23.49.1. Bembidion tetracolum tetracolum Say, 1823

References. Bembidion ustulatum (Linnaeus) (sic!): Csiki, 1940: 213 (Kula Ljums; Korab Mt.);

Murraj, 1968: 33 (Tomor Mt.).

Material studied. “Dukati” / “Apfelbeck”, 1 s. (NMW); “Merdita M. Scheit” / “Mali Scheit

Apfelbeck”, 1 s. (NMW); “Alban. Vermosa Penther”, 1 s. (NMW); “Kruja, Alban., Mader”, 4

s. (NMW); “Berat Alban. mer.”, 2 s. (NMW); “Albania leg. Bischoff 1936”, 2 s. (NMW);

“Kèlcyrë” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Schkoder” / “Albania leg.

Bischoff 1937”, 2 s. (NMW); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Ljukova,

100 m, 5.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS).

Notes. Marggi et al. (2003: 263) have omitted the species for Albania. Here it is confirmed

for the country. The record of Murraj (1968) is included here since the revision of one male

from the Tomor Mt., having the same label data as the published material, has been found

to be correct.

Subgenus Philochthus Stephens, 1828

23.50. Bembidion decolor Apfelbeck, 1911

References. Bembidion decolor: Apfelbeck, 1911: 210 (Buenë River; Vlorë); Apfelbeck, 1916:

340-341 (Buenë River; Vlorë); Netolitzky, 1943a: 63 (Elbasan; Kruja); Hieke & Wrase, 1988:

62; Marggi et al., 2003: 263.

Material studied. “Alban. Wgth Valona” / “Coll. Apfelbeck”, 1 m. (HNHM); “Albania Kula

Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).

Notes. Apfelbeck published two descriptions of this species, the first in Latin and Serbo-

Croatian and second one in Latin and German (Apfelbeck, 1911; Apfelbeck, 1916). The older

one is correctly treated as valid.

23.51. Bembidion guttula (Fabricius, 1792)

23.51.1. Bembidion guttula guttula (Fabricius, 1792)

References. Bembidion guttula: Hieke & Wrase, 1988: 61.

Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Tomor: Kulmak Alban. mer.” / “Alb.

Winkler leg.”, 1 s. (NMW); “Maje Tartarit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).




23.52. Bembidion inoptatum Schaum, 1857

References. Bembidion inoptatum: Apfelbeck, 1904: 112 (Velipoja near Lezhë; Vlorë); Apfelbeck,

1911: 210 (Buenë River; Vlorë); Apfelbeck, 1916: 341 (Buenë River; Vlorë); Csiki, 1940: 213

(Kula Ljums); Hieke & Wrase, 1988: 62.

Material studied. “Bojana Oboti, Alb.” / “ex coll. Winkler 1948”, 1 s. (NMW).


the country.

23.53. Bembidion iricolor Bedel, 1879

References. Bembidion iricolor: Apfelbeck, 1909: 297 (Vlorë).

Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Actually, it needs further

confirmation for the country.

23.54. Bembidion lunulatum (Geoffroy, 1785)

References. Bembidion lunulatum: Schatzmayr, 1943b: 96 (Stan-Karbunarë).

Material studied. “Elbasan Alb., Mader”, 4 s. (NMW); “Durazzo” / “Albania leg. Bischoff

1931”, 2 s. (NMW); “Tirana, Alb., dch. Winkler 42”, 1 s. (NMW); Tiranë Park, 22.II.1998, 1 s.,

from grasses along small sluggish stream, R. Ward leg. (CMNH).


the country.

23.55. Bembidion mannerheimii C.R. Sahlberg, 1827

Material studied. “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Tomor: Kulmak Albania

mer.” / “lg. Winkler mai 1931”, 5 s. (NMW).

Notes. New species for the fauna of Albania. Netolitzky has identified the material from the

Balkans either as “Bembidion neresheimeri” or as “cfr. neresheimeri”. I determined the material

from Albania as B. mannerheimii. According to D.W. Wrase (personal communication) B.

neresheimeri J. Müller, 1929 is a distinct species that also may inhabit the Balkan Peninsula;

Marggi et al. (2003: 264) have cited the latter only for Slovenia.

Subgenus Phyla Motschulsky, 1844

23.56. Bembidion tethys Netolitzky, 1926

References. Bembidion tethys: Focarile, 1959: 58; Hieke & Wrase, 1988: 61.

Material studied. “Elbasan Alb. 3.IX.18 Dr. H. Priesner”, 4 s. (NMW; MNHUB); “Elbasan

Alb., Mader”, 2 s. (NMW); “Tirana, Alb. Ravasini, Lona”, 1 s. (NMW); “Lum Erzen Alb.

V.1934 Mjr. Strupi”, 1 s. (NMW); Ljukove, 100 m, 5.VI.1995, 2 s., G. Blagoev leg. (NMNHS);

Tiranë Park, 1-9.XI.1997 / 23-28.II.1998, 3 s., R. Ward leg. (CMNH).


the country with first detailed records for the country.


Subgenus Princidium Motschulsky, 1864

23.57. Bembidion punctulatum Drapiez, 1820

23.57.1. Bembidion punctulatum punctulatum Drapiez, 1820

References. Bembidion punctulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke et al.,

1986: 185 (Uji Ftohte, south of Tepelenë; Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 44.

Material studied. “Tirane” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Lum i Tiranës”

/ “Albania leg. Bischoff 1935”, 2 s. (MNHUB); Tepelenë District, Drina River, 3.VI.1995, 1 s.,

G. Blagoev leg. (NMNHS).


the country. Besides the material studied above, the present author has seen some more

specimens in NMW, which remained unregistered.

Subgenus Talanes Motschulsky, 1864


23.58. Bembidion subfasciatum Chaudoir, 1850

Material studied. “Bojana - Umg. Pulaj” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Ruskuli

Alb., Mader”, 1 s. (NMW); “Liqen Butrintes” / “leg. Bischoff. 1936”, 3 s. (NMW).


Subgenus Terminophanes Müller-Motzfeld, 1998

Notes. New subgenus for the faunas of the Balkan Peninsula and Albania.

23.59. Bembidion terminale Heer, 1841

! Bembidion terminale ssp.

Material studied. “Albania, Alban Alps, Okol, 800 m, 15-20.VII.1996, leg. G. Holzinger”, 1 f. (HNHM).

Notes. New species for the faunas of the Balkan Peninsula and Albania. For the time being

the subspecific status of the local population remains unspecified. The specimen in question

has a somewhat different shape of the pronotum, compared with specimens from the Alps

and North Iran.

Subgenus Testedium Motschulsky, 1864

23.60. Bembidion bipunctatum (Linnaeus, 1761)

23.60.1. Bembidion bipunctatum nivale Heer, 1837

References. Bembidion bipunctatum var. pyritosum: Csiki, 1940: 212 (Shkëlzen Mt.; Gjalicë Mt.;

Korab Mt.). Bembidion bipunctatum: Hieke & Wrase, 1988: 45.

Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, more than 50 s.

(HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 3 s. (HNHM); “Albania Mts.


Korab” / “Akad. Balk. Exp. Csiki 1918”, 15 s. (HNHM); “Korab Preshk, Alb. Bischoff 1936”,

1 s. (HNHM).


Rubës, 2000 m, 8.VII.1993, 4 m., 1 f., L. Latella leg. (cAVT).


the country. Besides the material studied above, I have seen other numerous specimens from

different localities in NMW and MNHUB, which remained unregistered.

Subgenus Trepanes Motschulsky, 1864

23.61. Bembidion articulatum (Panzer, 1796)

References. Bembidion articulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Marggi et al.,

2003: 270.

Material studied. “Skutari Bojana”, 1 s. (NMW); “Skutari Alb. Dr. J. Müller”, 1 s. (NMW);

“Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Shkoder” / “Albania

leg. Bischoff 1937”, 3 s. (NMW).

23.62. Bembidion maculatum Dejean, 1831

23.62.1. Bembidion maculatum maculatum Dejean, 1831

Material studied. “Tirana, Alb. Ravasini, Lona”, 1 s. (NMW).


23.63. Bembidion octomaculatum (Goeze, 1777)

References. Bembidion octomaculatum: Apfelbeck, 1904: 110 (Velipoja near Lezhë); Apfelbeck,

1907a: 496 (Kiri River near Mesi); Hieke & Wrase, 1988: 60.

Material studied. “Alban. Wgth. Valona” / “Apfelbeck”, 1 s. (NMW); “Fieri, Albania a Semeni,

Müller”, 4 s. (NMW); “Skutari, Alb. Prof. Dr. Müller”, 2 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.”

/ “Alb. Winker 1934”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW).


the country.

24. Genus Ocys Stephens, 1829

24.1. Ocys harpaloides (Audinet-Serville, 1821)

References. Ocys harpaloides: Apfelbeck, 1904: 117 (Vlorë); Apfelbeck, 1909: 297 (Pasha Liman).

Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Scutari Alb., Mader”, 2 s. (NMW);

“Tiranë” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Borshë-i-Vjetër Wildbach-Quellen

Bischoff 21.VII.1940”, 1 s. (NMW).


the country.


25. Genus Sinechostictus Motschulsky, 1864

Subgenus Pseudolimnaeum Kraatz, 1888


25.1. Sinechostictus doderoi (Ganglbauer, 1891)

Material studied. “Tomor Buranj Alb. 10.7.1932 Alf. Bischoff ”, 1 s. (NMW); “Sckëlzën” /

“Albania Bischoff 1935”, 2 s. (NMW).


Subgenus Sinechostictus s.str.

25.2. Sinechostictus tarsicus (Peyron, 1858)

References. Bembidion elongatum var. effluviorum: Csiki, 1940: 213 (Kula Ljums). Bembidion elongatum

Dejean: Hieke & Wrase, 1988: 56.

Material studied. “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania

Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 6 s. (HNHM); “Miloti Alban. Matzenauer”, 1 s.

(NMW); “Elbasan Alb., Mader”, 1 s. (NMW); “Lumi Berati Alban. mer.”, 1 s. (NMW); “Lumi

Beratit Alban. mer. Lona. Winkler.”, 1 s. (NMW); “Llogora” / “Albania leg. Bischoff 1932”, 8

s. (MNHUB); “Llogora, Alb. Alf. Bischoff 2.VI.1932”, 3 s. (NMW); “Llogora” / “Albania leg.

Bischoff 1935”, 1 s. (NMW).

Notes. Marggi et al. (2003: 267 sub Bembidion elongatum tarsicum) have omitted the species for

Albania. Here it is confirmed for the country. Bonavita & Vigna Taglianti (2005) concluded

that in the Balkan Peninsula is present only S. tarsicus, a distinct species, while S. elongatus

(Ganglbauer, 1891) is a West European species. S. effluviorus (Peyron, 1858) is a separate species

confined only to the territory of Asian Turkey.

25.3. Sinechostictus millerianus (Heyden, 1883)

Material studied. “Shkëlzen” / “Albania leg. Bischoff 1936”, 1 s. (NMW).


Subtribe TACHYINA Motschulsky, 1862

26. Genus Lymnastis Motschulsky, 1862


26.1 Lymnastis galilaeus Piochard de la Brûlerie, 1876

Material studied. “Alban. Wgth. Valona”, 2 s. (MNHUB); “Mysege” / “Albania leg. Bischoff

1937”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB).



27. Genus Paratachys Casey, 1918

27.1. Paratachys bistriatus (Duftschmid, 1812)

References. Tachys bistriatus: Apfelbeck, 1904: 117 (Velipoja); Murraj, 1968: 33 (Vau i Dejës);

Hieke et al., 1986: 184 (Borshi near Vlorë); Hieke & Wrase, 1988: 62.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Mustajbeg Velipoja Albania”,

2 s. (HNHM); “Albania Miloti”, 3 s. (HNHM); “Miloti, Alb. Matzenauer”, 1 s. (HNHM);

“Elbasan Alb., Mader”, 7 s. (HNHM); “Lim i Tiranes” / “Albania leg. Bischoff 1932”, 2 s.

(MNHUB); “Burelle” / “Albania leg. Bischoff 1934”, 4 s. (MNHUB); “Mursi-Kara Alibey” /

“Albania leg. Bischoff 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1934”, 1 s.

(MNHUB); “Durres” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); Librazhd, Prenjas, 750

m, 7.V.1995, 1 s., L. Penev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 8.II.1998 / 28.II.1998,

54 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).

Notes. Kopecký (2003: 275) has omitted the species for Albania, so that it is here confirmed

for the country. The record of Murraj (1968) is included here with great reservation in view

of possible misidentification (it was found a male of Bembidion praeustum from the Tomor Mt.

determinated by Murraj as Tachys bistriatus).

27.2. Paratachys fulvicollis (Dejean, 1831)

References. Tachys fulvicollis: Apfelbeck, 1904: 118 (Velipoja near Lezhë); Hieke & Wrase,

1988: 63.

Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Tirane” / “Albania leg.

Bischoff 1932”, 2 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW);

Tiranë Park, 1.III.1998, 9 s., R. Ward leg. (CMNH).

Notes. Kopecký (2003: 275) has omitted the species for Albania. Here it is confirmed for the

country.

27.3. Paratachys micros (Fischer von Waldheim, 1828)

References. Tachys micros: Hieke & Wrase, 1988: 63.

Material studied. “Ibe Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

Notes. Kopecký (2003: 276) has omitted the species for Albania. Here P. micros is confirmed

for the country with first detailed record.

28. Genus Polyderis Motschulsky, 1862


28.1. Polyderis cardioderus (Chaudoir, 1850)

Material studied. Tiranë Park, 8.II.1998, 1 s., grass and soil around tree trunks, R. Ward leg.

(CMNH).



29. Genus Sphaerotachys J. Müller, 1926

29.1. Sphaerotachys haemorrhoidalis (Ponza, 1805)

References. Tachys haemorrhoidalis: Apfelbeck, 1904: 120 (Velipoja); Hieke et al., 1986: 184 (Uji

Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 65.

Material studied. “Bojana Shkoder”, 10 s. (NMW); “Medua, Alb. 10.5.914”, 2 s. (NMW);

“Elbasan. Alb. 29.VIII.18 Dr. H. Priesner”, 3 s. (MNHUB); “Berat Alb. Dr. Müller 18”, 2 s.

(NMW); “Skutari, Alb. Klenka”, 2 s. (NMW); “Elbasan Alb., Mader”, 7 s. (NMW); “Vorra,

Alban., Mader”, 5 s. (NMW); “Bushek Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Lumi i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg.

Bischoff 1933”, 3 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

Tiranë Park, 8.II.1998, 89 s., from grass and soil around tree trunks, R. Ward leg. (CMNH).

Notes. Kopecký (2003: 278) has omitted the species for Albania. Here it is confirmed for

the country.

30. Genus Tachys Dejean, 1821

30.1. Tachys scutellaris Stephens, 1828

References. Tachys scutellaris: Hieke & Wrase, 1988: 63.

Material studied. “Durazzo, Alb. Dr. J. Müller”, 9 s. (NMW); “Rusculi Alb., Mader”, 1 s.

(NMW); “Durres” / “Albania leg. Bischoff 1932”, 18 s. (MNHUB); “Humara Juni” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Liquen i Butrintes Vivari” / “Albania leg. Bischoff

1934”, 5 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 6 s. (MNHUB); “Liqen i

Butrintes” / “Albania leg. Bischoff 1936”, more than 30 s. (NMW).


country with first detailed records.

31. Genus Tachyta Kirby, 1837

Subgenus Tachyta s.str.

31.1. Tachyta nana (Gyllenhal, 1810)

References. Tachyta nana: Apfelbeck, 1904: 121 (Velipoja); Hieke et al., 1986: 184 (Bizë near

Shëngjergji); Hieke & Wrase, 1988: 66.

Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Ricavac 1300 m 25.VI.-

2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Mamuras Alban. Kruja Dr.

J. Müller”, 1 s. (NMW); “Kap Rodeni Alb., leg. Mader”, 1 s. (NMW); “Kruja, Alban., Mader”,

5 s. (NMW); “Llogora” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB).

Notes. Kopecký (2003: 277) has omitted the species for Albania. Here it is confirmed for

the country.


32. Genus Tachyura Motschulsky, 1862

Subgenus Tachyura s.str.

32.1. Tachyura diabrachys (Kolenati, 1845)

References. Tachys sexstriatus var. bisbimaculatus: Csiki, 1940: 214 (Kula Ljums). Tachys inaequalis:

Hieke et al., 1986: 184 (Borshi near Vlorë). Tachys diabrachys: Hieke & Wrase, 1988: 65.


“Elbasan, Alb. 4.IX.18 Dr. H. Priesner”, 1 s. (MNHUB); “Ibe Erzen” / “Albania leg. Bischoff

1932”, 10 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë”

/ “leg. Bischoff Albania 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff

1936”, 5 s. (NMW); “Drin Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW); Tepelenë

District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Librazhd, Prenjas, 750 m,

7.V.1995, 7 s., L. Penev leg. (NMNHS); Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev

leg. (NMNHS).


country. The most part of the literature data and older identifications of Tachys inaequalis

(Reitter, 1884, nec Kolenati, 1845), T. sexstriatus (Duftschmid, 1812) and T. bisbimaculatus

(Chevrolat, 1860) from the Balkans concern this species.

32.2. Tachyura parvula (Dejean, 1831)

References. Tachys parvulus: Hieke & Wrase, 1988: 64.

Material studied. “Elbasan Alb. 29.VII.18 Dr. H. Priesner”, 1 s. (NMW); “Elbasan Alb.

4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Skutari, Alb. Klenka”, 3 s. (NMW); “Skutari Alb.,

Mader”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB).

Notes. Kopecký (2003: 279) has omitted the species for Albania. Here T. parvula is confirmed

for the country with first detailed records.

# Tachyura sexstriata (Duftschmid, 1812)

Notes. I have not seen any specimen from this species from the Balkan Peninsula, excluding

Crete. The older records for it from the region refer to T. diabrachys.

Tribe PATROBINI Kirby, 1837

33. Genus Deltomerus Motschulsky, 1850

Subgenus Deltomerus s.str.

? Deltomerus malissorum Apfelbeck, 1918

References. Deltomerus malissorum: Zamotailov, 2003: 283.

Material studied. Montenegro. “Vunsaj, Alban. leg. Plec, 1914” / “Deltomerus mallisorum Typus!

Apfelb.” / “Typus”, 1 m. syntype (NMW); “Wgth. 1918 Kom VIII.”, 1 m., 2 f. (NMW); “Zljeb

Pl. Kulina” / “Montenegro leg. Bischoff 1931”, 1 m., 3 f. (MNHUB).


Notes. After the original description, only Zamotailov (2003: 283) noted this species for

Albania without any detailed records. However, the type locality, Vunšaj (in Albanian) or

Vusanje (in Serbo-Croatian) is situated in the Montenegrin section of the North Albanian

Alps very close to Albanian territory (Apfelbeck, 1918c: 160-161, 164-165). The massifs

of Kom Mt. and Zljeb Mt. also lie outside the investigated territory. Due to lack of any

reliable record from Albania D. malissorum is treated here as questionable. Very probably it

inhabits the country.

33.1. Deltomerus parumpunctatus Zamotailov, 1990

References. Deltomerus parumpunctatus: Zamotailov, 1990: 133, 135 (North Albanian Alps: “Čafa

Borit”, loc. typ.); Zamotailov, 1992: 56; Zamotailov, 2003: 283.

Material studied. “Wgth. 1918 Menina pl. N. alb. alp.” / “Deltomerus parumpunctatus m. paratype

V. N. Kurnakov det. 1970”, 1 m. designated by mistake as paratype (NMW); “Wgth. 1918

Menina pl.” / “Deltomerus parumpunctatus m. alotype V. N. Kurnakov det. 1970”, 1 f. designated

by mistake as paratype (NMW).

Notes. Zamotailov (1990: 135) have noted no paratypes in the description of D. parumpunctatus.

For this reason, the two specimens in NMW designated as paratypes should not be considered

as part of the type series.

Subgenus Paradeltomerus Apfelbeck, 1908

33.2. Deltomerus paradoxus Apfelbeck, 1908

Notes. Apfelbeck published three consecutive descriptions for both the subgenus and the

species (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest ones are correctly

treated as valid.

? Deltomerus paradoxus paradoxus Apfelbeck, 1908

References. Deltomerus paradoxus: Zamotailov, 1992: 56. Deltomerus paradoxus paradoxus:

Zamotailov, 2003: 283.

Notes. Hieke (1981: 80) stated by mistake that D. paradoxus is described from Albania.

In reality, the nominotypical form was described from the alpine zone of the Shar

Planina Mts. (Apfelbeck, 1907c; Apfelbeck, 1908). Dinchev & Atanasov (1998) stated

that only 1 % of the territory of the last mountain massif is situated in Albania. Actually

all specimens of the nominotypical race in the collections originate from Macedonia or

the Kosovo Region of Serbia. Due to lack of reliable data the subspecies is treated as

doubtful for Albania.

33.2.1. Deltomerus paradoxus korabensis J. Müller, 1936

References. Deltomerus paradoxus korabensis: Müller, 1936: 108-110 (Korab Mt., loc. typ.; Bjeshka

Matroxh); Zamotailov, 2003: 283. Deltomerus paradoxus: Zamotailov, 1992: 56.

Material studies. Nearly 15 s. from Korab Mt. (MNHUB).


Tribe POGONINI Laporte, 1834

34. Genus Pogonistes Chaudoir, 1871


34.1. Pogonistes convexicollis Chaudoir, 1871

Material examined. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

Notes. New species for the fauna of Albania. For the time being, the locality near Durrës is

the westernmost locality in the species range.

34.2. Pogonistes gracilis (Dejean, 1828)

Material examined. “Durres” / “Albania leg. Bischoff 1932”, 29 s. (MNHUB).

Notes. New species for the fauna of Albania. For the present, the locality near Durrës is one

of the easternmost finds in the species range.

35. Genus Pogonus Dejean, 1821

Subgenus Pogonus s.str.

35.1. Pogonus littoralis (Duftschmid, 1812)

References. Pogonus litoralis sic!: Apfelbeck, 1904: 149 (Velipoja); Hieke & Wrase, 1988: 67.

Material studied. “Alban. Wgth. Valona”, 2 s. (HNHM); “Latif Pulaj”, 1 s. (HNHM); “Latif

Pulaj” / “Latif Alban. sept.”, 5 s. (NMW).

Notes. Bousquet (2003a: 287) has omitted the species for Albania. Here it is confirmed for

the country. Besides the material studied above, in MNHUB I have seen more than 60 s. (most

of them come from the vicinity of Durrës), which remained unregistered.

? Pogonus luridipennis (Germar, 1823)

References. Pogonus luridipennis: Murraj, 1968: 33 (Kamëz).

Notes. The data of Murraj (1968) are doubtful because of possible misidentification (among

the materials loaned from MNST, I found a male of Bembidion combustum from the Tomor Mt.

which had been determinated by Murraj as P. luridipennis).

35.2. Pogonus riparius Dejean, 1828

References. Pogonus riparius: Apfelbeck, 1904: 149 (Velipoja near Lezhë; Shkodër); Hieke &

Wrase, 1988: 67; Bousquet, 2003a: 287.

Material studied. “Latif Pulaj”, 6 s. (HNHM); “Latif Pulaj” / “Latif Alban. sept.”, 7 s.

(NMW); “Merdita, Alb. Matzenauer”, 2 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”,

6 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB); “Liqen i Butrintes”

/ “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 3 s.

(MNHUB); “Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Valona” /



Tribe TRECHINI Bonelli, 1810

Subtribe PERILEPTINA Sloane, 1903

36. Genus Perileptus Schaum, 1860

Subgenus Perileptus s.str.

36.1. Perileptus areolatus (Creutzer, 1799)

36.1.1. Perileptus areolatus areolatus (Creutzer, 1799)

References. Perileptus areolatus: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Csiki, 1940: 214

(Kula Ljums); Hieke & Wrase, 1988: 34. Perileptus areolatus areolatus: Moravec et al., 2003: 288.


“Skutari Alb., Mader”, 1 s. (HNHM); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Leskovac - Barmashi,

880 m, 2.VI.1995, 4 s., G. Blagoev leg. (NMNHS).

Subtribe TRECHINA Bonelli, 1810

! genus Adriaphaenops Noesske, 1928

! Adriaphaenops sp.

References. Aphaenopsis sp.: Latella, 1990: 148 (North Albanian Alps: Mali i Thatë Mt., cave

“Shpella e Haxhisë”, 1720 m).

Material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-Thatë Mt., cave

“Shpella e Haxise”, 8.VII.1993, 1 f., L. Latella leg. (cAVT).

Notes. According to Vigna Taglianti (personal communication) the female mentioned above

belongs to a hitherto undescribed species.

37. Genus Albanotrechus Casale & V. B. Guéorguiev, 1994

37.1. Albanotrechus beroni Casale & V. B. Guéorguiev, 1994

References. Albanotrechus beroni: Casale & Guéorguiev, 1994: 418-420 (cave “Spela Merkurth”

near to village of Merkurth, Rreshen District); Moravec et al., 2003: 290.

38. Genus Duvalius Delarouzée, 1859

[subgenus Bicharotrechus Bokor, 1922]

[species group “pilifer”]

[Duvalius droveniki Magrini, 1998]

Notes. This species was described from the locality “Hajla Prokletije” in the Montenegrin

part of the North Albanian Alps (Magrini, 1998), very close to the Albanian border. D. droveniki

might live in Albania too.


# Duvalius maglicensis Winkler, 1933

References. Duvalius maglicensis: Turin, 1981: 112.

Notes. This species lives only in Bosnia Herzegovina and Montenegro (see also Moravec et

al., 2003: 299). Species deleted from the list of the Albanian ground beetles.

Subgenus Duvalius s.str.

Notes. The arrangement of the separate groups is in accordance with the works of Casale

(1983), Casale & Laneyrie (1983), Casale & Vigna Taglianti (1990) and Casale et al. (1996). The

specificity of the group of “fodori” is discussed in a coming paper (Guéorguiev, in preparation).

I. South Aegean groups

Species group “krueperi”

38.1. Duvalius didonnai Magrini, Vanni & Ingusio, 1996

References. Duvalius didonnai: Magrini et al., 1995a: 27-29 (cave “Shpella Lekpetes”, 1150 m, near to

Kurvelesh, loc. typ.; cave “Shpella e Kasarit”, 1100 m, near to Kurvelesh); Moravec et al., 2003: 302.

Species group “georgii”

38.2. Duvalius bischoffi Meschnigg, 1936

References. Duvalius bischoffi: Meschnigg, 1936: 48-49 (cave “Homberg-Höhle” at Maja e Tanasit

Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlorë); Schatzmayr, 1943b:

99 (Maja e Tanasit Mt.: cave “Homberg”); Turin, 1981: 110; Casale, 1983: 249; Casale & Laneyrie,

1982:199; Genest & Juberthie, 1994: 592 (cave “Homberg”, Maja e Tanasit Mt., south of

Vlorë); Moravec et al., 2003: 301.

38.3. Duvalius georgii (J. Müller, 1922)

References. Duvalius georgi: J. Müller, 1922a: 87-88 (hot pole in Tomor Mt., loc. typ.); Jeannel,

1928: 567-570 (Tomor Mt.: Toik, Kulmak); Scheibel, 1937a: 18, 20 (Tomor Mt.); Schatzmayr,

1943b: 98 (Tomor Mt.: Toik, Kulmak); Casale, 1983: 249; Casale & Laneyrie, 1982:199; Genest

& Juberthie, 1994: 592 (hot pole and cave at Tomor Mt., east of Berat); Moravec et al., 2003: 303.

Material studied. “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona” / “ex Coll. Bokor”,

1 f. (HNHM); “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona”, 3 s. (NMW); “Tomor:

Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor-Buranj juli

1932” / “Albania leg. Bischoff 1932”, 3 s. (NMW; MNHUB).

38.4. Duvalius muelleri Winkler, 1933

References. Duvalius muelleri: Winkler: 1933a: 75-76 (Shen Elias Mt., cave “Stere gropa e Vasos”,

800 m, south of Vlorë, loc. typ.); Scheibel, 1937a: 18, 20 (Shen Elias Mt., south of Vlorë);

Schatzmayr, 1943b: 98 (Shen Elias Mt., cave “Stere gropa e Vasos”); Turin, 1981: 112; Casale,


1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (cave at Shen Elias

Mt., south of Vlorë); Magrini et al., 1995b: 25 (Llogora Mt., cave “Stera Kullëxhive”, south of


Material studied. “Stere gropa de Vasos” / “Reza e Kanalit, Alban. m.” / “Winkler, Lona,

Bischoff, V.1931”, 8 syntypes (NMW); “Shen-E’Lias Juni 1932” / “Albania leg. Bischoff 1932”,

2 s. (MNHUB); “Shen-Elias” / “leg. Bischoff Albania 1934”, 1 f. (HNHM); “Shen-E’Lias Juni

1932” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB).

Species group “fuchsi”

38.5. Duvalius fuchsi Scheibel, 1937

References. Duvalius fuchsi: Scheibel, 1937a: 15-18, 20 (nameless cave near Llogora, south of

Vlorë, loc. typ.); Turin, 1981: 111; Casale, 1983: 249; Casale & Laneyrie, 1982: 199; Genest &

Juberthie, 1994: 592 (cave at Llogora, south of Vlorë); Moravec et al., 2003: 303.

Material studied. “Höhle in der Llogora” / “Albania leg. Bischoff 1935”, 1 syntype (MNHUB);

“Höhle i.-d. Llgora” / “Bischoff mai 1935”, 4 syntypes (MNHUB).

37.6. Duvalius pominii Schatzmayr, 1943

References. Duvalius pominii: Schatzmayr, 1943b: 96-97, 99 (cave “Stere e Vogel” and cave “Stere

e Madhe” near Dukati, south of Vlorë, loc. typ.); Turin, 1981: 112; Genest & Juberthie, 1994:

592 (cave “Stere e Vogel” and cave “Stere e Madhe” near Dukati); Moravec et al., 2003: 305.

38.7. Duvalius regiszogui Meschnigg, 1936

References. Duvalius regiszogui: Meschnigg, 1936: 46-47 (cave “Homberg-Höhle” Maja e Tanasit

Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlorë); Schatzmayr, 1943b:

99 (Maja e Tanasit Mt.: cave “Homberg”); Turin, 1981: 113; Casale, 1983: 249; Casale & Laneyrie,

1982:199; Genest & Juberthie, 1994: 592 (cave “Homberg”, Maja e Tanasit Mt., south of


II. North Aegean groups

Species group “leonhardi”

38.8. Duvalius sturanyi (Apfelbeck, 1904)

38.8.1. Duvalius sturanyi stenocephalus (Apfelbeck, 1918)

References. Trechus stenocephalus: Apfelbeck, 1918c: 164 (North Albanian Alps, Vermosh, 1100

m, loc. typ.). Duvalius leonhardi sturanyi: Jeannel, 1928: 548 (North Albanian Alps: Vermosh,

1100 m). Duvalius sturanyi stenocephalus: Scheibel, 1937a: 18-20 (North Albanian Alps: Vermosh,

Menina Planina Mt.); Pretner, 1978: 130, 135, 148, 167-168 (Vermosh; Menina Planina Mt.);

Moravec et al., 2003: 305. Duvalius leonhardi stenocephalus: Mařan, 1939c: 60 (Vermosh). Duvalius

sturanyi: Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (North Albanian Alps:

Menina Planina Mt. near Vermosh).


Material studied. “Vermoša 1100 m” / “Alban. montenegr. Grenze Penther’14” / “Typus” /

“Trechus stenocephalus Typus! Apfelb.”, 1 f. syntype of Trechus stenocephalus (NMW).

Notes. Jeannel (1928: 582-584) synonymized D. stenocephalus with D. leonhardi sturanyi, but later

Scheibel (1937a: 19-20) proved that the former is a subspecies of D. sturanyi. Further studies are

needed to test the status of D. sturanyi sturanyi and D. sturanyi stenocephalus, having in mind the different

forms of their aedeagus and copulatory pieces (Scheibel, 1937a). The same concerns the remote

populations of the former near Foča and from the Komovi Mt. as well as the status of D. sturanyi

matejkai Mařan, 1939 described from the Montenegrin section of the North Albanian Alps.

Species group “fodori”

Notes. The treatment of this species group is in accordance with a separate author’s paper in prepara-

tion. It treats Duvalius fodori Scheibel, 1937, D. strupii Scheibel, 1937 and D. vignai Casalei, 1983.

38.9. Duvalius strupii Scheibel, 1937

References. Duvalius strupii: Scheibel, 1937b: 282-283 (Korab Mt., 2000 m, loc. typ.); Schatzmayr,

1943b: 98 (Korab Mt.); Turin, 1981: 113; Casale & Laneyrie, 1982:191; Casale, 1983: 250

(Korab Mt.); Genest & Juberthie, 1994: 592 (Korab Mt.); Moravec et al., 2003: 305.

Material studied. “Korab” / “Albania leg. Bischoff 1936”, 17 specimens, incl. 12 syntypes (MNHUB).

Subgenus Euduvalius Jeannel, 1929

Species group “gogalai”

38.10. Duvalius zhalovi B. V. Guéorguiev, 2005

References. Duvalius zhalovi: Guéorguiev, 2005: 33-40 (cave “Peshtnik”, 1100 m near Ostreni

i Madh, loc. typ.).

39. Genus Neotrechus J. Müller, 1913

Species group “dalmatinus”

39.1. Neotrechus lonae (J. Müller, 1914)

39.1.1. Neotrechus lonae lonae (J. Müller, 1914)

References. Trechus lonae: J. Müller, 1914: 1003-1005 (Çukall Mt.: cave “Scpedha n’Giuka

Kunors”, loc. typ.; Çukall Mt.: cave “Scpedha n’Ciuk tugit Vukai”, loc. typ.). Neotrechus lonae:

Winkler, 1926: 86, 96 (caves in Çukall Mt.); Scheibel, 1936: 229 (Çukall Mt.); Scheibel, 1937a:

20 (Çukall Mt.). Neotrechus lonai sic!: Jeannel, 1928: 296-297 (Çukall Mt.: cave “Scpedha n’

giuka kunors” and cave “Scpedha n’ Ciuk tugit Zukali”); Schatzmayr, 1943b: 97 (Çukall Mt.:

cave “Scpedha n’ giuka kunors” and cave “Scpedha n’ Ciuk tugit Zukali”); Casale & Laneyrie,

1983: 180; Genest & Juberthie, 1994: 592 (caves in Çukall Mt., Rapshë). Neotrechus lonai sic!

lonai sic!: Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Çukall Mt.); Pretner, 1978: 147, 166

(Çukall Mt.). Neotrechus lonae lonae: Moravec et al., 2003: 315.


Material studied. “Cukali Gebirge N-Albanien Lona leg. 1914”, 4 s. (NMW); “Scpedha n’Giuka

Zukali Albanien Lona Giuno 1914”, 1 s. (DEI); “Cukale Albania sept.” / “Bischoff 1930 coll.

A. Winkler”, 5 s. (NMW); “Cucale Geb. Höhle von Vukaj” / “Albania leg. Bischoff 1931”, 1 s.

(MNHUB); “Shpella Kodra Çorit” / “Cukale Geb.” / “leg. Bischoff Albania 1934”, 3 s.

(MNHUB); “Shpella Kodra Carit Thep” / “Cukale” / “Albania leg. Bischoff 1935”, 8 s.

(MNHUB); “Shpella Luga Ìen Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 1 s.

(MNHUB); “Shpella Rasa-Kunors Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 4 s.

(MNHUB); “Shpella Çuka Vjit” / “Cucale” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB);

“Shpella Gollë” / “Cukale” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

Notes. N. lonae lonae is sympatric and syntopic with N. malissorum (J. Müller, 1914) in most of

the caves where they have been found. The report of this subspecies from the area of Rapshë

(Genest & Juberthie, 1994: 592) is erroneous since this region is populated by N. lonae atavus.

The nominotypical subspecies differs from both ssp. atavus and ssp. shkodrensis in the wider

and more rounded, almost not narrowed to apex, elytra.

39.1.2. Neotrechus lonae atavus J. Müller, 1934 [= bischoffi Meschnigg, 1936]

References. Neotrechus bischoffi: Meschnigg, 1936: 44-46 (North Albanian Alps: cave “Shpella

e lugstit, Hot-Brigje” near Rapshë, loc. typ.); Schatzmayr, 1943b: 98 (cave “Shpella e lugstit,

Hot-Brigje” near Rapshë); Casale & Laneyrie, 1983: 180. Neotrechus lonae atavus: Scheibel, 1936:

227-228 (cave “Shpella e lugstit, Hot Brgje” near Rapshë); Scheibel, 1937a: 20 (cave “Hot

Brigje”); Moravec et al., 2003: 315. Neotrechus lonai sic! atavus: Absolon & Mařan, 1947: 2;

Mařan, 1947: 11 (Rapshë); Pretner, 1978: 147, 166 (cave “Shpella e lugstit, Hot Brgje” near

Rapshë); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (cave “Shpella e Lugstit”,

cave “Hot Brigje” near Rapshë).

Material studied. “Shpella e Lugstit” / “Brigie Hot” / “leg. Bischoff Albania 1934”, 7 syntypes

of Neotrechus bischoffi (MNHUB); “Shpella e Lugstit” / “Brigie Hot” / “Albania leg. Bischoff

1935”, 11 syntypes of Neotrechus bischoffi (MNHUB).

Notes. Scheibel (1936) established the synonymy of N. bischoffi with N. lonae atavus. Regardless

of that, Schatzmayr (1943b) and Casale & Laneyrie (1983) listed the former as valid species.

The type localities of both N. lonae atavus and N. suturalis pretneri Scheibel, 1936 coincide and

are located in Montenegro very close to the Albanian border.

39.1.3. Neotrechus lonae shkodrensis Scheibel, 1936

References. Neotrechus lonae shkodrensis: Scheibel, 1936: 228-229 (cave “Shpella kufinit, Koplikut-

Hotit” near Ivanaj, north of Liqen i Shkodrës, loc. typ.); Scheibel, 1937a: 20 (Ivanaj); Moravec

et al., 2003: 315. Neotrechus lonai sic! shkodrensis: Absolon & Mařan, 1947: 2; Mařan, 1947: 11

(Ivanaj); Pretner, 1978: 147, 166 (cave “Shpella kufinit, Koplikut-Hotit” near Ivanaj); Casale &

Laneyrie, 1983: 180; Drovenik & Peks, 1994: 45; Genest & Juberthie, 1994: 592 (cave “Kufinit

et Kopikut, Hotit peés” near Ivanaj, north of Liqen i Shkodrës); Drovenik & Peks, 1999: 51.

Material studied. “Shpella Kufin Hot Koplicut Ivanaj” / “Albania leg. Bischoff 1935”, 4

syntypes (MNHUB); “Shpella Kufin Hot Koplicut Ivanaj” / “Albania leg. Bischoff 1937”, 4 s.

(MNHUB).

Notes. Drovenik & Peks (1994: 45) and Drovenik & Peks (1999: 51) indicated that the type

locality of this subspecies is situated in Montenegro. Actually it is located in Albania, very

close to the state border.


39.2. Neotrechus malissorum (J. Müller, 1914)

References. Trechus malissorum: Müller, 1914: 1005 (Çukall Mt.: cave “Scpedha n’ kunor

Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha n’ Ciuk tujit Vukai”, loc. typ.; Çukall Mt.: cave

“Scpedha e Bacit Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha n’ rasst’ kunors”, loc. typ.;

Çukall Mt.: cave “Scpedha n’ giuka kunors”, loc. typ.; Çukall Mt.: cave “Scpedha Nre Gegs

n’ gur t’ barzodhs”, loc. typ.). Neotrechus dalmatinus malissorum: Winkler, 1926: 86, 96 (caves in

Çukall Mt.); Scheibel, 1937a: 20 (Çukall Mt.). Neotrechus malissorum: Jeannel, 1928: 290-291

(Çukall Mt.: cave “Scpedha n’ kunor Zukali”, cave “Scpedha n’ Ciuk tujit Zukali”, cave

“Scpedha e bacit Zukali”; cave “Scpedha n’ rasst’ kunors”; cave “Scpedha n’ giuka kunors”;

cave “Scpedha Nre Gegs n’ gur t’ barzodhs Zukali”); Schatzmayr, 1943b: 98 (Çukall Mt.:

cave “Scpedha n’ kunor Zukali”, cave “Scpedha n’ Ciuk tujit Zukali”, cave “Scpedha e bacit

Zukali”; cave “Scpedha n’ rasst’ kunors”; cave “Scpedha n’ giuka kunors”; cave “Scpedha

Nre Gegs n’ gur t’ barzodhs Zukali”); Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Çukall

Mt.); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (Çukall Mt., cave “n’Kunor

Vukai”, cave “n’Ciuk tujit Vukai”, cave “Lacit Vukai”, cave “n’Rast’Kunors”, cave “n’Giuka

Kunors”); Moravec et al., 2003: 315.

Material studied. “Scpedha n’Giuka Zukali Albanien Lona Giuno 1914”, 6 s. (DEI); “Scpedha-

Nre Geg N’gur Bazzodha Zukali Albanien”, 1 s. (HNHM); “Scpedha-Nre Geg N’gur Barzodha

Zukali Albanien Lona Giugno 1914”, 4 s. (MNHUB); “Cukali Gebirge N-Albanien Lona leg.

1914”, 4 s. (NMW); “Albania m. Cucali C. Lona”, 2 s. (HNHM); “Cukali Albania sept.” / “leg.

Bischoff 1931”, 2 s. (NMW); “Shpellë Quku i Konors Vukaj Cucale” / “Albania leg. Bischoff

18.VI.31”, 2 s. (MNHUB); “Shpella Kodra Carit” / “Cukale Geb.” / “leg. Bischoff Albania

1934”, 2 s. (MNHUB); “Shpella Kodra Carit Thep” / “Cukale” / “Albania leg. Bischoff 1935”,

24 s. (NMW; MNHUB); “Shpella Luga Ìen Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”,

3 s. (MNHUB); “Shpella Rasa Kunors Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 3 s.

(MNHUB); “Shpella Gollë” / “Cukale” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).

Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-

Thatë Mt., cave “Shpella e Haxise”, 8.VII.1993, rests of 3 s., A. Gobetti leg. (cAVT).

39.3. Neotrechus suturalis (Schaufuss, 1864)

39.3.1. Neotrechus suturalis pentheri Winkler, 1926

References. Trechus suturalis: Apfelbeck, 1918c: 164 (North Albanian Alps: small cave near

Rapshë). Neotrechus dalmatinus suturalis var. pentheri: Winkler, 1926: 85-86, 96 (near Rapshë, loc.

typ.). Neotrechus dalmatinus pentheri: Scheibel, 1937a: 20 (Rapshë). Neotrechus suturalis pentheri:

Jeannel, 1928: 295 (North Albanian Alps: caves near Rapshë); Absolon & Mařan, 1947: 2;

Mařan, 1947: 11 (North Albanian Alps: Rapshë); Pretner, 1978: 146-147, 167 (Rapshë); Casale

& Laneyrie, 1983: 180; Drovenik & Peks, 1994: 46; Genest & Juberthie, 1994: 592 (North

Albanian Alps: caves at Rapshë); Drovenik & Peks, 1999: 53; Moravec et al., 2003: 316.

Material studied. “Alban. montenegr. Grenze, Penther’14” / “Höhle bei Rapsa, 727 m”, 15

syntypes of Trechus suturalis (DEI; MNHUB); “Albania Rapsa Dr. Grabowski”, 86 s. (HNHM);

“Höhlen bei Rapsa leg. Bischoff 1931”, 1 s. (DEI); “Höhlen bei Rapsa 20.VII.31” / “Albania

leg. Bischoff 1931”, 4 s. (MNHUB); “Rapsh” / “Albania leg. Bischoff 1935”, 40 s. (MNHUB).

Notes. Drovenik & Peks (1994: 46) and Drovenik & Peks (1999: 53) noted by mistake the

type locality in Montenegro. In fact, the village of Rapshë is situated in Albania.


40. Genus Trechus Clairville, 1806

Subgenus Trechus s.str.

Species group “quadristiatus”

40.1. Trechus obtusus Erichson, 1837

40.1.1. Trechus obtusus obtusus Erichson, 1837

References. Trechus obtusus: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m);

Hieke & Wrase, 1988: 35; Nonveiller et al., 1994: 9. Trechus obtusus obtusus: Jeannel, 1927: 309

(“Petrov planina”, 2000 m); Moravec et al., 2003: 337.

Material studies. “Latif Koritnik”, 1 s. (NMW); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).

40.2. Trechus quadristriatus (Schrank, 1781)

References. Trechus quadristriatus: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr,

1943b: 96 (Lushnje; cave “Stere e Madhe” near Dukati); Hieke et al., 1986: 184 (Uji Ftohte, south of

Tepelenë); Hieke & Wrase, 1988: 35; Nonveiller et al., 1994: 9; Moravec et al., 2003: 339.

Material studied. “Elbasan Alb., Mader”, 2 s. (HNHM); “Kruja, Alban., Mader”, 4 s. (HNHM);

“Sarisaltik, 1180 m Alb., leg. Mader”, 3 s. (HNHM); Tiranë Park, 22-31.X.1997 / 1-9.XI.1997

/ 23-28.XI.1997, 47 s., R. Ward leg. (CMNH).

Notes. Except the specimens studied above, in MNHUB the present author has seen additionally

about 210 s., which remained unregistered.

Species group “babinjensis”

40.3. Trechus babinjensis Jeannel, 1927

References. Trechus babinjensis: Jeannel, 1927: 267-268 (North Albanian Alps, loc. typ.); Casale

& Laneyrie, 1983: 141; Moravec et al., 2003: 327.

40.4. Trechus valbonensis Jeannel, 1927

References. Trechus valbonensis: Jeannel, 1927: 268 (North Albanian Alps, loc. typ.); Casale &

Laneyrie, 1983: 141; Moravec et al., 2003: 343.

Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m,

11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).

Species group “subnotatus”

40.5. Trechus subnotatus Dejean, 1831

40.5.1. Trechus subnotatus subnotatus Dejean, 1831

References. Trechus subnotatus subnotatus: Apfelbeck, 1904: 127 (Vlorë); Jeannel, 1927: 446 (Kruja;

Tomor Mt.: Kulmak); Pawlowski, 1973: 232; Moravec et al., 2003: 341. Trechus subnotatus: Jeannel,


1925: 23 (Tomor Mt.: Kulmak); Horion, 1941: 181; Schatzmayr, 1943b: 96 (Dukati; cave “Stere

e Madhe” near Dukati); Casale & Laneyrie, 1983: 147; Hieke & Wrase, 1988: 36; Nonveiller et

al., 1994: 9, 15; Guéorguiev & Guéorguiev, 1995: 75; Guéorguiev et al., 1997: 11; Sakalian &

Guéorguiev, 1997: 46.

Material studied. “Kruja, Alban., Mader”, 2 s. (HNHM).


m., D. Antonelli leg. (cAVT).

Notes. In MNHUB I have seen also additional 39 s., which remained not registered.

40.5.2. Trechus subnotatus ljubetensis Apfelbeck, 1908

References. Trechus subnotatus ljubetensis: Moravec et al., 2003: 341.

Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Albania” / “ljubetensis Apf.

R. Jeannel”, 1 m. (NMW).

Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908;

Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid. The identification

of the two specimens from Vlorë is doubtful since the ssp. ljubetnesis is normally treated as

local endemic of the Shar Planina Mts.

? species group “cardioderus”

? Trechus irenis Csiki, 1912 [= cardioderus balcanicus Jeannel, 1927]

References. Trechus cardioderus balcanicus: Tabaković-Tošić, 1988: 137.

Notes. Tabaković-Tošić (1988) noted the range of this taxon reaching Albania. Due to lack of

reliable records from the country the species is treated as questionable for Albania.

Species group “pulchellus”

40.6. Trechus merditanus Apfelbeck, 1906

References. Trechus merditanus: Apfelbeck, 1906: 1163 (Merdita: Mal i Shenjt Mt. and Zepë

Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Jeannel,

1927: 539-540 (Merdita: Mal i Shenjt Mt. and Zepë Mt.); Pawlowski, 1972: 312-314; Casale &


Material studied. “Merdita M. Scheit” / “merditanus n. sp. Apfelb.”, 2 s. probably syntypes (NMW).

Notes. According to Jeannel (1927), T. merditanus is a vicarious species of T. priapus.

40.7. Trechus priapus K. Daniel, 1902

40.7.1. Trechus priapus priapus K. Daniel, 1902

References. Trechus priapus: Meixner, 1939: 309; Pawlowski, 1973: 249; Hieke & Wrase, 1988:

38; Nonveiller et al., 1994: 9; Guéorguiev & Guéorguiev, 1995: 77; Guéorguiev et al., 1997: 10.

Trechus priapus priapus: Jeannel, 1927: 537 (North Albanian Alps; Gjalicë Mt.); Moravec et al.,

2003: 338.

Material studied. “Alban. Exped. Gjalica Ljums 17.-26.VI’18”, 2 s. (NMW).


Rubës, 2000 m, 8.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).

Notes. Except the two specimens above, in MNHUB I have seen additionally nearly 800 s.,



Species group “egreginus”

40.8. Trechus egregius Jeannel, 1927

References. Trechus egregius: Jeannel, 1927: 551-552 (North Albanian Alps, loc. typ.); Casale &


Species group “tristis”

40.9. Trechus nigrinus Putzeys, 1847 [= tristis (Duftschmid, 1812)]

References. Trechus nigrinus: Apfelbeck, 1904: 129 (Vlorë); Apfelbeck, 1907a: 498-499 (Mal i Shenjt

Mt.: top region, 1500 m); Jeannel, 1927: 555 (Vlorë); Moravec et al., 2003: 336. Trechus tristis: Csiki,

1940: 214 (Gjalicë Mt.); Pawlowski, 1973: 250; Hieke & Wrase, 1988: 38; Nonveiller et al., 1994: 9.


“Alban. Exp.’18. Pashtrik 4-14.7”, 1 s. (NMW); “Kruja, Alban. Mader”, 3 s. (HNHM); “Tomor”

/ “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff

1937”, 67 s. (MNHUB); “Mal i Gjer” / “Albania leg. Bischoff 1937”, 15 s. (MNHUB).

Notes. Except the material studied above, in MNHUB the present author has seen more 48 s.,


Species group “obtusiusculus”

40.10. Trechus albanicus Apfelbeck, 1905

References. Trechus albanicus: Apfelbeck, 1905: 241 (Northeastern Albania / Kosovo border

region: alpine belt of Koritnik Mt., loc. typ.); Apfelbeck, 1907b: 636-637 (alpine belt of Koritnik

Mt., loc. typ.); Jeannel, 1927: 563-564 (Koritnik Mt.; Gjalicë Mt.); Csiki, 1940: 214 (Gjalicë Mt.;

Korab Mt.); Casale & Laneyrie, 1982: 153; Drovenik & Peks, 1994: 39 (Koritnik Mt.); Drovenik

& Peks, 1999: 44 (Koritnik Mt.); Moravec et al., 2003: 326.

Material studied. “Latif Koritnik” / “Trechus albanicus Apfb. Typus! Coll. Apfelbeck”, 1 f. syntype

(HNHM); “Latif Koritnik”, 2 syntypes (DEI; NMW); “Latif Alban. spt.”, 1 s. (DEI); “Albania

Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Mts. Gyalica Lums” /

“Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki

1918”, 2 s. (HNHM); “Alban. Exped. Gjalica Ljums 17.-26.VI.’18”, 1 s. (NMW); “Gjalica” / “Albania

leg. Bischoff 1936”, 2 s. (NMW); “Gjallica” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB).

Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b)

and the latter has long time been accredited to be correct (see also Casale & Laneyrie, 1982;

Drovenik & Peks, 1994: 39; Drovenik & Peks, 1999: 44; Moravec et al., 2003: 326). However,

it is incorrect and the original description must be assigned to the older one.

40.11. Trechus kobingeri Apfelbeck, 1902

40.11.1. Trechus kobingeri kobingeri Apfelbeck, 1902 [= distinctissimus Roubal, 1920]

References. Trechus kobingeri: Jeannel, 1925: 23 (Çukall Mt.); Jeannel, 1927: 566 (Çukall Mt.);

Casale & Laneyrie, 1982: 153; Nonveiller et al., 1994: 8, 18 (near to Shkodër). Trechus distinctissimus:


Casale & Laneyrie, 1982: 153. Trechus kobingeri bulgaricus: Guéorguiev et al., 1997: 10. Trechus

kobingeri kobingeri: Moravec et al., 2003: 333.

Material studied. “Maja e Jezerce Albania sept.”, 1 m. (NMW); “Cucale” / “Albania leg.


Notes. Tråchus kobingeri bulgaricus Pawlowski, 1972 is a junior synonym of T. kobingeri pawlowskianus

P. Moravec & Lompe, 2003, and its record for Albania is erroneous. This race is known only

from a limited region in Western Bulgaria.

# Trechus obtusiusculus Ganglbauer, 1889

References. Trechus obtusiusculus: Nonveiller et al., 1994: 9.

Notes. The species is recorded with detailed records only from Bosnia Herzegovina and

Montenegro. The data for Albania is too doubtful, so that it is removed from the species list.

40.12. Trechus ravasinianus Lorenz, 1998 [= ravasinii Jeannel, 1925]

References. Trechus ravasinii: Jeannel, 1925: 24-25 (Tomor Mt.: Kulmak, Toik, Abbas Ali, loc.

typ.); Jeannel, 1927: 570-571 (Tomor Mt.: Abbas Ali, Kulmak, Toik); Casale & Laneyrie, 1982:

153. Trechus ravasinianus: Moravec et al., 2003: 339.

Material studied. “Tomor: Tomor Albania mer.” / “lg. Winkler Mai 1931”, 1 f. (HNHM);

“Tomor: Kulmak Albania mer.” / “lg. Winkler Mai 1931”, 12 s. (NMW).

Notes. Except the localities mentioned above, in MNHUB I have seen more than 100 s.,


! Trechini gen. indet.

! Trechini gen. & sp. indet.

Material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-Thatë Mt., cave

“Shpella e Haxise”, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT).

Notes. According to Vigna Taglianti (personal communication) the parts found in the cave

“Shpella e Haxise” may belong to an unknown genus close to Pheggomisetes Knirsch, 1923.

Subtribe TRECHODINA Jeannel, 1926

41. Genus Thalassophilus Wollaston, 1854

41.1. Thalassophilus longicornis (Sturm, 1825)

References. Thalassophilus longicornis: Apfelbeck, 1907a: 496 (Kiri River near Mesi).

Notes. Moravec et al. (2003: 346) have omitted the species for Albania. Actually, it needs


COLEOPTERA: CARABIDAE: HARPALINAE 89

Subfamily HARPALINAE Bonelli, 1810

Tribe CHLAENIINI Brullé, 1834

Subtribe CALLISTINA Laporte, 1834

42. Genus Callistus Bonelli, 1810

42.1. Callistus lunatus (Fabricius, 1775)

42.1.1. Callistus lunatus lunatus (Fabricius, 1775)

References. Callistus lunatus: Schatzmayr, 1943b: 99 (Lushnje); Murraj, 1968: 33 (Hibë; Sauk);

Hieke et al., 1986: 190 (Ibë near Krraba). Callistus lunatus lunatus: Kirschenhofer, 2003: 347.

Material studied. “Lomi Beratit Albanien”, 1 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr.

Grenze Penther’14”, 1 s. (NMW); “Hani Grabom Mai” / “Alban. montenegr. Grenze

Penther’14”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); Sauk,

9.VII.1961, 1 f. (MNST).

Notes. The records of Murraj (1968) are included here with minimal reservation since the

revision of a female from Sauk having the same label data as the published material has been

found to be correctly identified by him.

Subtribe CHLAENIINA Brullé, 1834

43. Genus Chlaenius Bonelli, 1810

Subgenus Chlaeniellus Reitter, 1908

43.1. Chlaenius flavipes Ménétries, 1832

References. Chlaenius flavipes: Apfelbeck, 1907a: 496, 498, 501 (Kiri River near Mesi; Fani i

Madh River near Oroshi); Schatzmayr, 1943b: 100 (Zezes River near Ura Zezes); Hieke et al.,

1986: 190 (Borshi near Vlorë); Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 349.

Material studied. “Oboti Bojana”, 1 s. (NMW); “Skutari Mesi”, 1 s. (NMW); “Merdita Oroschi”

/ “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Alban. Exped. Kula Ljums 18-28.V.’18”, 1 s.

(NMW); “Elbasan, Alb. 16.IX.18 Dr. H. Priesner”, 8 s. (NMW); “Elbasan Alb., Mader”, 8 s.

(NMW; MNHUB); “Tepelene juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); Tepelenë District, Vijosë

River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS).

43.2. Chlaenius nigricornis (Fabricius, 1787)

References. Chlaenius nigricornis: Apfelbeck, 1904: 158 (Durrës); Hieke & Wrase, 1988: 149;

Kirschenhofer, 2003: 350.

Material studied. “Scutari l. See-Ufer”, 1 s. (NMW); “Oboti Bojana”, 1 s. (HNHM); “Albania

Rittm. Matoha”, 1 s. (NMW); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 2 s. (MNHUB).


Notes. Murraj (1968: 33) cited this species from two localities (Tiranë; Tomor Mt.). Among

the material loaned from MNST, I found out 9 s. from the second place having the same label

data as the material published by Murraj as Chlaenius nigricornis. Actually, all the nine specimens

refer to Bembidion dalmatinum.

43.3. Chlaenius nitidulus (Schrank, 1781)

References. Chlaenius nitidulus: Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 350.

Material studied. “Maja ë Qori 2050 m, Albania”, 1 s. (NMW); Sauk, 5.III.1961, 1 m. (MNST).


43.4. Chlaenius terminatus Dejean, 1826

References. Chlaenius terminatus: Schatzmayr, 1943b: 100 (Stan-Karbunarë; Skrofotina; Vlorë).

Material studied. “Oboti Bojana”, 3 s. (HNHM).

Notes. Kirschenhofer (2003: 350) has omitted the species for Albania. Here it is confirmed

for the country.

43.5. Chlaenius tristis (Schaller, 1783)

43.5.1. Chlaenius tristis tristis (Schaller, 1783)

References. Chlaenius tristis: Hieke & Wrase, 1988: 149. Chlaenius tristis tristis: Kirschenhofer, 2003: 350.

Material studied. “Oboti Bojana”, 5 s. (HNHM); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s.

(MNHUB).


43.6. Chlaenius vestitus (Paykull, 1790)

References. Chlaenius vestitus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (Stan-

Karbunarë); Kirschenhofer, 2003: 351.

Material studied. “Bojana Oboti, Alb.”, 1 s. (NMW); “Lumi Dukatit Alban. mer.”, 4 s. (NMW);

“Maj’e Tartarit Albania mer.”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”,

1 s. (HNHM); “Alban. Exped. Kula Ljums 14.20.VII.’18”, 1 s. (NMW); “Elbasan, Alb. 15.IX.18 Dr.

H. Priesner”, 4 s. (NMW); “Alessio, Alb. Matzenauer”, 2 s. (NMW); “Miloti, Alb. Matzenauer”, 1 s.

(NMW); 40 s. from different localities (MNHUB); Librazhd, Prenjas, 750 m, 7.V.1995, 1 s., S.

Golovach, B. Petrov, P. Stoev leg. (NMNHS); Drina River, 3.VI.1995, 1 s., V. Sakalain leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë

and Gjirokastër), 350 m, 23.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Subgenus Chlaenites Motschulsky, 1860

43.7. Chlaenius spoliatus (P. Rossi, 1792)

43.7.1. Chlaenius spoliatus spoliatus (P. Rossi, 1792)

References. Chlaenius spoliatus: Schatzmayr, 1943b: 99 (Durrës); Hieke & Wrase, 1988: 147.

Chlaenius spoliatus spoliatus: Kirschenhofer, 2003: 351.


Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Oboti Bojana”, 2 s.

(HNHM); “Durazzo” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Himara juni” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 5 s.

(MNHUB); “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i

That” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s. (MNHUB).



Subgenus Chlaenius s.str.

43.8. Chlaenius festivus (Panzer, 1796)

43.8.1. Chlaenius festivus festivus (Panzer, 1796)

References. Chlaenius festivus: Apfelbeck, 1904: 157 (Vlorë; Velipoja near Lezhë); Apfelbeck,

1909: 298 (Vlorë); Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (Lushnje); Murraj,

1968: 33 (Radenjë); Hieke & Wrase, 1988: 148. Chlaenius festivus festivus: Gridelli, 1925: 120

(Shkodër; Berat); Kirschenhofer, 2003: 351.

Material studied. “Alban., Wgth. Valona”, 1 s. (NMW); “Oboti Bojana”, 1 s. (HNHM); “Skutari

l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk.

Exp. Csiki 1917”, 1 s. (HNHM); “Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner”, 1 s. (NMW); nearly

40 s. from different localities (MNHUB); Sauk, 18.II.1961, 1 f. (MNST); Rodenj, 26-28.VII.1962,

1 s. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS).


and Gjirokastër), 350 m, 23.IV.2003, 4 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Notes. The specimen collected at Liqen i Shkodrës was determined as C. velutinus (Duftschmid,

1812) by Kirschenhofer, but I refer this record to the nominotypical subspecies. The material

published by Murraj (1968) was found to be correctly determinated.

Subgenus Dinodes Bonelli, 1810

43.9. Chlaenius cruralis Fischer von Waldheim, 1829

References. Chlaenius cruralis: Kirschenhofer, 2003: 352.

Material studied. “Skutari” / “Strasser”, 3 s. (NMW); Ibë, 4.VII.1963, 1 m. (MNST).


43.10. Chlaenius decipiens (L. Dufour, 1820) [= azureus Duftschmid, 1812, nec Fabricius,

1775; laticollis Chaudoir, 1843; ambiguus Csiki, 1931]

References. Chlaenius decipiens: Apfelbeck, 1904: 154 (Vlorë); Hieke et al., 1986: 190 (Ibë near

Krraba); Hieke & Wrase, 1988: 148; Kirschenhofer, 2003: 352. Chlaenius azureus: Apfelbeck,

1909: 298 (Vlorë). Chlaenius decipiens var. ambiguus: Csiki, 1940: 219 (Gjalicë Mt.). Chlaenius

decipiens laticollis: Schatzmayr, 1943b: 100 (Lushnje).

Material studied. “Elbasan Albanien”, 1 s. (NMW); “Albania Mts. Gyalica Lums” / “Akad.

Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Tepelene Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”,

1 s. (NMW); “Tirane” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg.


Bischoff 1936”, 1 s. (MNHUB); “Ivan Geb Korce” / “Albania 1936 leg. Ramadan coll. Bischoff ”,

1 s. (MNHUB); Tiranë, 27.IX.1958, 1 m. (MNST); Tiranë, 10.V.1962, 1 m. (MNST).

Subgenus Epomis Bonelli, 1810

43.11. Chlaenius circumscriptus (Duftschmid, 1812)

References. Chlaenius circumscriptus: Kirschenhofer, 2003: 353.

Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Liqen i Butrintes”



43.12. Chlaenius dejeani (Dejean, 1831)

References. Chlaenius dejeani: Apfelbeck, 1904: 154 (Vlorë); Schatzmayr, 1943b: 99 (Lushnje);

Hieke & Wrase, 1988: 147. Epomis dejeani: Apfelbeck, 1909: 298 (Vlorë).

Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Murthi” / “Albania leg.

Bischoff 1936”, 2 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB).

Notes. Kirschenhofer (2003: 353) has omitted the species for Albania. Here it is confirmed

for the country.

Subgenus Trichochlaenius Seidlitz, 1887

43.13. Chlaenius aeneocephalus Dejean, 1826

43.13.1. Chlaenius aeneocephalus aeneocephalus Dejean, 1826

References. Chlaenius aeneocephalus: Apfelbeck, 1904: 156 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);


Notes. Kirschenhofer (2003: 355) has omitted the species for Albania, so that it needs of


Tribe CYCLOSOMINI Laporte, 1834

Subtribe MASOREINA Chaudoir, 1870

44. Genus Masoreus Dejean, 1821

Subgenus Masoreus s.str.

44.1. Masoreus wetterhallii (Gyllenhal, 1813)

44.1.1. Masoreus wetterhallii wetterhallii (Gyllenhal, 1813)

References. Masoreus wetterhallii: Hieke & Wrase, 1988: 152.

Material studied. “Tomor Tojk” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

Notes. Bousquet (2003b: 357) has omitted the species for Albania. Here it is confirmed for

the country with a first detailed record.


Tribe DRYPTINI Bonelli, 1810

45. Genus Drypta Latreille, 1796

Subgenus Drypta s.str.

45.1. Drypta dentata (P. Rossi, 1790)

References. Drypta dentata: Apfelbeck, 1904: 346 (Velipoja); Hieke & Wrase, 1988: 153; Baehr,

2003b: 358.

Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB).

Tribe HARPALINI Bonelli, 1810

Subtribe AMBLYSTOMINA Fauvel, 1889

46. Genus Amblystomus Erichson, 1837

46.1. Amblystomus levantinus Reitter, 1883

46.1.1. Amblystomus levantinus levantinus Reitter, 1883

References. Amblystomus levantinus: Apfelbeck, 1904 (Velipoja); Schatzmayr, 1943b: 101 (Stan-

Karbunarë; Lushnje); Hieke & Wrase, 1988: 165 (Velipoja). Amblystomus levantinus levantinus:

Jaeger & Wrase, 2003: 360.

Notes. According to D. Wrase (personal communication) the “metallescens-levantinus” group

needs revision. This notable specialist on the Palaearctic Harpalini wrote me: “I don’t know

really what levantinus is, as I haven’t checked yet the type. I have several species, clearly different

in genitalia belonging to that group. For the time being the best solution of that problem it to

continue the classification we have now.”.

46.2. Amblystomus metallescens (Dejean, 1829)

References. Amblystomus metallescens: Csiki, 1940: 219 (Kula Ljums; Gjalicë Mt.); Schatzmayr,

1943b: 101 (Lushnje; Mal i Krujes); Hieke & Wrase, 1988: 146; Jaeger & Wrase, 2003: 360.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Oboti Bojana”, 1 s. (HNHM);

“Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” /

“Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

46.3. Amblystomus niger (Heer, 1841)

References. Amblystomus niger: Hieke et al., 1986: 188 (Ibë near Krraba; Bizë near Shëngjergji);

Hieke & Wrase, 1988: 146; Jaeger & Wrase, 2003: 360.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Durres” / “Albania leg.

Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB);

“Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 8.II.1998 / 15.II.1998,



Subtribe ANISODACTYLINA Lacordaire, 1854

47. Genus Anisodactylus Dejean, 1829

Subgenus Anisodactylus s.str.

47.1. Anisodactylus binotatus (Fabricius, 1787) [= spurcaticornis Dejean, 1829]

References. Anisodactylus binotatus: Apfelbeck, 1909: 298 (Pashaliman); Hieke et al., 1986: 186

(North Albanian Alps near Theth); Hieke & Wrase, 1988: 112; Ito, 2003a: 360. Anisodactylus

binotatus ab. spurcaticornis: Apfelbeck, 1909: 298 (Pasha Liman).

Material studied. “Presba See” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Theth, 1-8.VIII.

1961, 1 m. (MNST); Theth, 800-1200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).

47.2. Anisodactylus nemorivagus (Duftschmid, 1812)

References. Anisodactylus nemorivagus: Apfelbeck, 1904: 212 (Velipoja); Horion, 1941: 247; Hieke

& Wrase, 1988: 112.

Notes. Ito (2003a: 361) has omitted the species for Albania. Actually, it needs further


47.3. Anisodactylus pueli Schauberger, 1933

47.3.1. Anisodactylus pueli pueli Schauberger, 1933

References. Anisodactylus pueli: Schauberger, 1933: 124 (Vlorë, loc. typ.); Sciaky, 1979: 15 (Vlorë); Turin,

1981: 150; Wrase, 1991: 12; Noonan, 1996: 34, 109 (Vlorë). Anisodactylus pueli pueli: Ito, 2003a: 361.

Material studied. “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).



Subgenus Pseudodichirius Lutshnik, 1921

47.4. Anisodactylus intermedius Dejean, 1829

References. Anisodactylus intermedius: Hieke & Wrase, 1988: 113; Ito, 2003a: 361.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Maja e Konispol Seraqin” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1936”, 1 s.

(MNHUB); Rodeni, 2-22.VII.1962, 1 s. (MNST); Gjirokastra, 6.V.1994, 1 m., P. Stoev leg. (NMNHS).


48. Genus Diachromus Erichson, 1837

48.1. Diachromus germanus (Linnaeus, 1758)

References. Diachromus germanus: Ito, 2003a: 362.

Material studied. “Albania Tirana” / “28.IX.1958 leg. Xh. Murraj”, 2 s. (HNHM).



49. Genus Gynandromorphus Dejean, 1829

49.1. Gynandromorphus etruscus (Quensel, 1806)

49.1.1. Gynandromorphus etruscus etruscus (Quensel, 1806)

References. Gynandromorphus etruscus: Schatzmayr, 1943b: 105 (Dukati); Hieke & Wrase, 1988:

113. Gynandromorphus etruscus etruscus: Ito, 2003a: 362.

Material studied. “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Radona, Shkiela,

21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).

Subtribe DITOMINA Bonelli, 1810

50. Genus Carterus Dejean, 1830

Subgenus Carterus s.str.

50.1. Carterus dama (P. Rossi, 1792)

References. Ditomus dama: Apfelbeck, 1909: 298 (Vlorë). Carterus dama: Hieke & Wrase, 1988:

145; Wrase, 1989: 39 (Mursi - Kara Ali Bey); Wrase, 2003b: 363.

Material studied. “Murrsi - Kara Ali Bey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);

Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).

Additional material included (studied D.W. Wrase). Oricum, south of Vlorë, X.1994, 2 f.,

K. Resl leg. (cWR).

50.2. Carterus gilvipes (Piochard de la Brûlerie, 1873)

References. Carterus gilvipes: Schatzmayr, 1943b: 101 (Dukati); Wrase, 2003b: 364.

50.3. Carterus rotundicollis Rambur, 1837

Material studied. Borsh, 15-28.V.1961, 1 f. (MNST).


51. Genus Ditomus Bonelli, 1810

51.1. Ditomus calydonius (P. Rossi, 1790)

51.1.1. Ditomus calydonius calydonius (P. Rossi, 1790)

References. Ditomus calydonius calydonius: Wrase, 2003b: 364.

51.2. Ditomus tricuspidatus (Fabricius, 1792)

References. Ditomus tricuspidatus: Ganglbauer, 1889: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);

Wrase, 2003b: 365. Carterus tricuspidatus: Apfelbeck, 1904: 173 (Vlorë).


Material studied. “Albania Avlona v. Oertzen”, 1 s. (MNHUB); “Brataj Ramica Albania mer.”

/ “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “M. e Tartarit juni 1932” / “Albania leg.

Bischoff 1932”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);

“Borshne” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); Borsh, 15-28.V.1961, 1 f. (MNST);

Ljukove, 100 m, 5.VI.1995, 1 m., G. Blagoev leg. (NMNHS).

Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 2 s., M.


52. Genus Dixus Billberg, 1820

52.1. Dixus clypeatus (P. Rossi, 1790)

References. Aristus clypeatus: Apfelbeck, 1909: 298 (Berat). Ditomus clypeatus: Hieke et al., 1986:

186 (Borshi near Vlorë; Mali me Gropë near Livadhet e Selitës; Mal i Dajtit Mt.: Shkall Priskë);

Hieke & Wrase, 1988: 146. Dixus clypeatus: Wrase, 2003b: 365.

Material studied. “Kruja, Alban., Mader”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff

1933”, 3 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Albanien

Dajti…6.6.55 Burmeister”, 1 s. (MNHUB); “N 111”, 1 s. (MNST), Sauk, 10-20.VI.1961, 1 s.

(MNST).



52.2. Dixus eremita (Dejean, 1825)

References. Ditomus eremita: Schatzmayr, 1943b: 102 (Skrofotina; Lushnje). Dixus eremita: Wrase,

2003b: 365; Wrase, 2005: 820.

52.3. Dixus obscurus (Dejean, 1825)

References. Aristus obscurus: Apfelbeck, 1909: 298 (Vlorë; Berat). Ditomus obscurus: Schatzmayr,

1943b: 101 (Lushnje); Hieke et al., 1986: 186 (Borshi near Vlorë; Poliçan near Tomor Mt.).

Dixus obscurus: Wrase, 2003b: 365.

Material studied. “Serandë” Piqeros” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Tirane”


53. Genus Graniger Motschulsky, 1864

53.1. Graniger femoralis (Coquerel, 1859)

References. Carterophonus femoralis: Sciaky & Franzini, 1976: 26 (Berat).


54. Genus Odontocarus Solier, 1835


54.1. Odontocarus robustus (Dejean, 1831)

Material studied. “Murrsi - Kara Alibey” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB);

“Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania

leg. Bischoff 1936”, 3 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

Butrinti, 4.VI.1995, 1 s., V. Sakalian leg. (NMNHS).


[genus Pachycarus Solier, 1835]

[subgenus Mystropterus Chaudoir, 1842]

[Pachycarus macedonicus V.B. Guéorguiev & B.V. Guéorguiev, 1997]

Notes. This species has been described from the Macedonian part of the Galičica Mt. very

close to the Albanian border, so that it may inhabit Albania too.

Subtribe HARPALINA Bonelli, 1810

55. Genus Acinopus Dejean, 1821

Subgenus Acinopus s.str.

55.1. Acinopus baudii A. Fiori, 1913

55.1.1. Acinopus baudii hellenicus Heinz, 1982

References. Acinopus subquadratus: Magistretti, 1965: 272; Guéorguiev & Guéorguiev, 1995:

205; Kataev & Wrase, 2003: 368. Acinopus baudii hellenicus: Heinz, 1982: 16 (Tepelenë; Maja e

Konispol, Seraqin); Kataev & Wrase, 2003: 368.

Material studied. “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Murthe bis Kara Alibey” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB).

55.2. Acinopus picipes (Olivier, 1795) [= emarginatus Chaudoir, 1842]

References. Acinopus emarginatus: Apfelbeck, 1904: 180 (Vlorë). Acinopus picipes: Apfelbeck,

1909: 297 (Vlorë); Schatzmayr, 1943b: 102 (Karbunara e Madhe; Tërpan; Lushnje); Hieke &

Wrase, 1988: 143; Kataev & Wrase, 2003: 368.

Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Mal i That”

/ “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Mal i Dajtit Mt.: Shkall Priskë, 26-30. VI.1961,

1 f. (MNST); Ibë, 4.VII.1962, 1 m. (MNST); Ibë, 8.VII.1966, 1 f. (MNST); “AL (Berat). Paß ca.

5 km S. Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 s. (MNHUB); Ljukove, 100 m, 5.VI.1995,

1 s., G. Blagoev leg. (NMNHS).

Additional material included (studied S. Hristovski and D.W. Wrase). Korcë District, Mai i

That Mt., Zwezde-Sattel, 1000-1100 m, 4.V.1993, 1 f., Heinz & Peks leg. (cWR); Hotel “Edva”

at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH); Dhermi, 200 m,

12.VII.2006, 2 s., M. Komnenov leg. (cSH).


# Acinopus subquadratus Brullé, 1832

References. Acinopus subquadratus: Magistretti, 1965: 272; Guéorguiev & Guéorguiev, 1995:

205; Kataev & Wrase, 2003: 368.

Notes. Guéorguiev & Guéorguiev (1995) mentioned this taxon for Albania, following an

older indication for South Albania (Magistretti, 1965: 272). Kataev & Wrase (2003: 368) have

most probably followed also the same sources. However, according to Heinz (1982) A.

subquadratus inhabits only the southern and central regions of Greece and the data on its

presence in Albania concerns A. baudii hellenicus; both species were often mixed in the past.

The species is removed from the list of the Albanian Carabidae.

Subgenus Oedematicus Bedel, 1897

55.3. Acinopus megacephalus (P. Rossi, 1794)

References. Acinopus megacephalus: Apfelbeck, 1904: 180 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);

Puel, 1934: 79; Schatzmayr, 1943b: 102 (Dukati; Skrofolina); Jeanne, 1984: 24; Hieke & Wrase,

1988: 143; Kataev & Wrase, 2003: 369.

Material studied. “Tirane” / “Albania leg. Bischoff 1933”, 3 s. (MNHUB); “Durrës” / “Albania

leg. Bischoff 1935”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB);

“Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 3 s. (MNHUB).

Additional material included (studied S. Hristovski and A. Vigna Taglianti). Lushnja-Divjaka,

21.VII.1993, 1 m., L. Latella leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of

Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva”

at Radhime (Vlorë District), 8-15.VII.2006, 3 s., leg. M. Komnenov (cSH).

Subgenus Osimus Fischer von Waldheim, 1829

55.4. Acinopus ammophilus Dejean, 1829

References. Acinopus ammophilus: Apfelbeck, 1907a: 498, 501 (Oroshi).

Notes. Kataev & Wrase (2003: 369) have omitted the species for Albania. Actually, it needs


56. Genus Harpalus Latreille, 1802

Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982

56.1. Harpalus tenebrosus Dejean, 1829 [= centralis Schauberger, 1929]

References. Harpalus tenebrosus var. centralis: Csiki, 1940: 220 (Kula Ljums). Harpalus tenebrosus:

Hieke & Wrase, 1988: 138; Kataev & Wrase, 2003: 370.


“Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).


Subgenus Harpalus s.str.

Species group “honestus”

56.2. Harpalus attenuatus Stephens, 1828

References. Harpalus attenuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Csiki, 1940: 220 (Shkëlzen Mt.; Kula Ljums); Schatzmayr, 1943b: 104 (“Poggio Boschetto”);

Hieke et al., 1986: 187 (Borshi near Vlorë; Bizë near Shëngjergji); Hieke & Wrase, 1988: 139;

Kataev & Wrase, 2003: 373.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Albania Skölsen” / “Akad.

Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”,

1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Burelle” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);

“Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja ae Konispol Seraqin” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Vlesh” / “30.VII.1958 leg. Xh. Murraj”,

1 s. (HNHM); “Albanien-Exp. DEI Borshi südl. Vlora 14.-27.V.1961”, 1 s. (MNHUB); Mal i

Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 2 f. (MNST); 2 km N from Durrës, 24.V.1993, 1 s., P.

Stoev & D. Zaprjanova leg. (NMNHS).

56.3. Harpalus honestus (Duftschmid, 1812)

56.3.1. Harpalus honestus honestus (Duftschmid, 1812)

References. Harpalus honestus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Csiki, 1940: 220 (Korab Mt.); Hieke & Wrase, 1988: 138. Harpalus honestus honestus: Kataev &

Wrase, 2003: 377.

Material studied. “Vermosa Penther”, 20 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr.

Grenze, Penther’14”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze,

Penther’14”, 1 s. (NMW); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

“Alban. Exp. Kula Ljums 18-28.V.’18”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”,

1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg.

Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 3 s.

(MNHUB); “Bashtrik” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Gjallica” / “Albania

leg. Bischoff 1936”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);

Ibë, 18.V.1960, 1 m. (MNST).

56.4. Harpalus rubripes (Duftschmid, 1812)

References. Harpalus rubripes: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Csiki, 1940: 220 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke & Wrase, 1988: 133; Kataev &

Wrase, 2003: 381.

Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania

Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Korab” / “Akad. Balk. Exp.

Csiki 1918”, 1 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1

s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Alban. Exp. 1918 Korab, 23-

31.VII.”, 1 s. (NMW); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal


i Shenjit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Okol Shkalla” / “Albania leg.


“AL (Tirana) 8 km N Psene. E Tirana 1450 m. 30.IV.1993 leg. H. + S. Peks”, 2 m. (MNHUB);

North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS);

Cafasan, 1230 m, 27.VII.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS).

56.5. Harpalus rufipalpis Sturm, 1818

56.5.1. Harpalus rufipalpis rufipalpis Sturm, 1818 [= rufitarsis (Duftschmid, 1812)]

References. Harpalus rufitarsis: Apfelbeck, 1904: 196 (Berat); Apfelbeck, 1907a: 499 (Mal i

Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Berat); Hieke et al., 1986: 188 (Bizë

near Shëngjergji); Hieke & Wrase, 1988: 138. Harpalus rufipalpis rufipalpis: Kataev & Wrase,

2003: 381.

Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Mal i Shenjit”

/ “leg. Bischoff Albania 1934”, 6 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 1 s.

(MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Maranaj” / “Albania leg.

Bischoff 1937”, 2 s. (MNHUB); “Albania Sauk” / “27.VII.1959 leg. Xh. Murraj”, 3 s. (HNHM).

Notes. In addition to the material studied above, in NMW I have seen more specimens, which


56.6. Harpalus sulphuripes Germar, 1824

56.6.1. Harpalus sulphuripes sulphuripes Germar, 1824

References. Harpalus sulphuripes: Apfelbeck, 1904: 196 (Vlorë); Anonymous, 1907: 284;

Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë);

Schauberger, 1926: 32 (Vlorë; Pasha Liman); Hieke et al., 1986: 188 (Tomor Mt.: Abbas

Ali, 2401 m); Hieke & Wrase, 1988: 139. Harpalus sulphuripes sulphuripes: Kataev & Wrase,

2003: 383.

Material studied. More than 50 s. from different localities (MNHUB); North Albanian Alps,

Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS); Saranda District,

Ljukove, 5.V.1995, 1 s., P. Stoev leg. (NMNHS); Gjirokastra, 6.V.1994, 1 s., P. Stoev leg.

(NMNHS); 12 km NE Erseka, 900 m, 12.V.1995, 1 s., L. Penev leg. (NMNHS).



Species group “atratus”

56.7. Harpalus atratus Latreille, 1804 [= subsinuatus (Duftschmid, 1812)]

References. Harpalus atratus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Hieke et al., 1986: 187 (Mal i Dajtit Mt.: Shkall Priskë); Hieke & Wrase, 1988: 137; Kataev &

Wrase, 2003: 373. Harpalus atratus var. subsinuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top

region, 1500 m).

Material studied. “Albania Zalle-Dardha” / “23.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); more

than 60 s. from different localities (MNHUB).


Additional material included (studied A. Vigna Taglianti). 5 km S of Tepelenë (crossroads

Permet), 350 m, 21-22.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Notes. In addition to the material studied above, in NMW I have seen more specimens, which


Species group “quadripunctatus”

56.8. Harpalus laevipes Zetterstedt, 1828 [= quadripunctatus Dejean, 1829]

References. Harpalus quadripunctatus: Hieke & Wrase, 1988: 133. Harpalus laevipes: Kataev &

Wrase, 2003: 378.

Material studied. “Gjallica” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).


Species group “serripes”

56.9. Harpalus flavicornis Dejean, 1829

56.9.1. Harpalus flavicornis flavicornis Dejean, 1829

References. Harpalus flavicornis: Apfelbeck, 1904: 198 (Vlorë); Apfelbeck, 1909: 298 (Vlorë);

Hieke & Wrase, 1988: 136. Harpalus flavicornis flavicornis: Kataev & Wrase, 2003: 376.

Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania

leg. Bischoff 1933”, 1 s. (MNHUB); “Lumi i Erzen” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);

“Mal i Dajtit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tirane” / “leg. Bischoff Albania

1934”, 3 s. (MNHUB); “Mal i That” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB);

“Tiranë” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff

1937”, 1 s. (MNHUB); “Albanien - Tirana Dajti-Vorberge 3.V.39…Müller J.”, 2 s. (MNHUB); Mal

i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 1 m. (MNST); Zooteknika, 14.V.1963, 1 s. (MNST); “AL

(Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 1 s. (MNHUB); “AL (Berat).

Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 f. (MNHUB); Cafasan, 1230 m,

27.VII.1993, 2 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 4 s., G.

Blagoev leg. (NMNHS); 5 km Leskovac, 650 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNH); Tiranë

Park, 11.I.1998 / 8.II.1998, 4 s., R. Ward leg. (CMNH).

56.10. Harpalus serripes (Quensel, 1806)

56.10.1. Harpalus serripes serripes (Quensel, 1806) [= subchalybaeus Reitter, 1900]

References. Harpalus serripes: Apfelbeck, 1904: 197 (Vlorë); Anonymous, 1907: 283; Apfelbeck,

1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Csiki, 1940:

220 (Korab Mt.); Hieke et al., 1986: 187 (Ibë near Krraba); Hieke & Wrase, 1988: 135. Harpalus

serripes ab. subchalybaeus: Schauberger, 1926: 25 (Dukati i Ri). Harpalus serripes serripes: Kataev &

Wrase, 2003: 382.

Material studied. L. Selitës, 3-9.VII.1961, 1 m. (MNST); Ibë, 4.VII.1963, 1 f. (MNST); “AL

(Vlore) Llogora-Paß 1025 m 29.IV.1993 leg. F. + S. Peks”, 10 s. (MNHUB); 2 km N from

Durrës, 24.V.1993, 2 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Cafasan, 1230 m, 27.VII.1993,

3 s., P. Stoev, D. Zaprjanova leg. (NMNHS); Lushnja, Divjaka National Park, 10.V.1995, 1 s., S.


Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m, 6.VI.1995, 3 s., G. Blagoev

leg. (NMNHS).

Additional material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë

District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).

Notes. In addition to the material studied above, in NMW and ZNHU I have seen more than

130 s. from different localities, which remained unregistered.

Species group “pumilus”

[Harpalus picipennis (Duftschmid, 1812)]

References. Harpalus picipennis: Apfelbeck, 1904: 200; Hieke & Wrase, 1988: 141; Kataev &

Wrase, 2003: 380.

Notes. Hieke & Wrase (1988) noted the species for Albania following Apfelbeck’s record

“Prisren” (Apfelbeck, 1904: 200). In fact this place is situated in the Kosovo Region of Serbia.

Most likely, Kataev & Wrase (2003) have used the same sources. For the time being the species

is treated here as probable but not yet proven for Albania (having in mind recent data from

the north parts of Republic of Macedonia, unpublished information).

56.11. Harpalus pumilus Sturm, 1818 [= vernalis Fabricius, 1801]

References. Harpalus vernalis: Müller, 1931: 69; Hieke & Wrase, 1988: 140; Kataev & Wrase,

2003: 380.

Material studied. “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).


Species group “anxius”

56.12. Harpalus anxius (Duftschmid, 1812)

References. Harpalus anxius: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);


Material studied. “Latif Cucali”, 1 s. (NMW); “Merdita Oroshi” / “Apfelb. Alban. spt.

IV.V.’05”, 2 s. (NMW); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B. Petrov, P.

Stoev leg. (NMNHS).

Notes. Hieke & Wrase (1988: 139) noted this species for Albania apparently following the

record “Jannina” (Apfelbeck, 1904: 199). In fact this locality is situated in Greece.

56.13. Harpalus subcylindricus Dejean, 1829

References. Harpalus subcylindricus: Hieke & Wrase, 1988: 140; Kataev & Wrase, 2003: 382.

Material studied. “Skutari” / “Strasser”, 1 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1

s. (NMW); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rapsh” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);

Mal i Dajtit Mt., 10.V.1965, 1 m. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë ~ 600 m 27.IV.1993

leg. Heinz u. Sonja Peks”, 3 s. (MNHUB); “AL (Berat). Paß ca. 5 km S Gllava 600 m. 27.IV.1993

leg. H. + S. Peks”, 1 m., 1 f. (MNHUB); North Albanian Alps, Maj’e Radohimes, 2400-2550


m, 29.V.1993, 3 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Radohimes,

1000-1100 m, 5.IX.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS).


Species group “flavescens”

56.14. Harpalus flavescens (Piller et Mitterpacher, 1783)

Material studied. “Elbasan, Alb. 11.IX.18 Dr. H. Priesner”, 1 s. (NMW).


Species group “tardus”

56.15. Harpalus albanicus Reitter, 1900

References. Harpalus albanicus: Reitter, 1900: 114 (? Shkodër, loc. typ.); Apfelbeck, 1904: 200

(Shkodër); Rambousek, 1912: 86; Horion, 1941: 230; Roudier, 1949: 176 (Shkodër); Magistretti,

1965: 308; Hieke & Wrase, 1988: 134; Kataev & Wrase, 2003: 372.

Material studied. “Skutari” / “coll. Reitter”, 1 f. syntype (HNHM).

Notes. Tschitschérine (1901: 154) expressed doubt about the correct interpretation of the

species type locality “Scutari”. It may as well represent the present city of Üsküdar located on

the Asian shore of the Bosporus right opposite the peninsula of Istanbul, or the present

Albanian City of Shkodër.

56.16. Harpalus tardus (Panzer, 1796)

References. Harpalus tardus: Apfelbeck, 1904: 198 (Vlorë); Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top

region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988:134; Kataev & Wrase, 2003: 383.

Material studied. “Latif Oroshi”, 1 s. (NMW); “Maranaj” / “Apfelb. Alban. spt. IV.V.’05”, 2

s. (NMW); “Alban. Exped. Kula Ljums 18-28.V.1918”, 1 s. (NMW).

Species group “latus”

56.17. Harpalus latus (Linnaeus, 1758)

References. Harpalus latus: Kataev & Wrase, 2003: 378.

Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); North

Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & P. Stoev leg. (NMNHS).


56.18. Harpalus solitaris Dejean, 1829

References. Harpalus solitaris: Kataev & Wrase, 2003: 382.

Material studied. North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 f., P. Beron, B.




Species group “smaragdinus”

56.19. Harpalus smaragdinus (Duftschmid, 1812)

References. Harpalus smaragdinus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500

m); Hieke & Wrase, 1988: 132; Kataev & Wrase, 2003: 382.

Material studied. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Mal i

Shenjit” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); Theth, 1-8.VIII.1961, 1 m. (MNST);

“AL (Shkodar). Paß SE Puke 850 m 2.V.1993 leg. H. & S. Peks”, 1 m. (MNHUB); North

Albanian Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & B. Petrov leg.

(NMNHS); Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS).

Additional material included (studied S. Hristovski and A. Vigna Taglianti). Rapsh, 15.V.2002,

3 m., 1 f., A. Paparistos leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006,

1 s., leg. M. Komnenov (cSH).

Species group “autumnalis”

# Harpalus anatolicus Tschitschérine, 1898

# Harpalus anatolicus anatolicus Tschitschérine, 1898 [= skutariensis Reitter, 1900]

References. Harpalus tristis: Apfelbeck, 1904: 199 (“Scutari”).

Notes. Reitter (1900: 107) indicated as type locality of H. skutariensis the site “Scutari” (see

also Kataev & Wrase, 1997). “Scutari” was the Greek name for the district of Istanbul now

known as Üsküdar, located on the Asian shore of the Bosporus right opposite the peninsula

of Istanbul. The same name was also used in the past for the present city of Shkodër.

Tschitschérine (1901: 146) has stated that H. skutariensis Reitter is identical with H. tristis. Apfelbeck

(1904: 199) accepted this opinion despite Daniel & Daniel (1902: 250-251) had already synonymized

H. tristis to H. danieli. Later on, Mlynář (1979: 83) considered H. skutariensis as synonym of H.

anatolicus. Kataev & Wrase (1997) confirmed this synonymy and correctly gave data only for the

Anatolian Peninsula. The species is excluded from the list of the Albanian Carabidae.

56.20. Harpalus autumnalis (Duftschmid, 1812)

References. Harpalus autumnalis: Hieke & Wrase, 1988: 136; Kataev & Wrase, 2003: 373.

Material studied. “Skutari Bojana”, 3 s. (NMW); “Lum i Tiranes” / “Albania leg. Bischoff

1933”, 1 s. (MNHUB).


# Harpalus danieli Reitter, 1900 [= tristis Tschitschérine, 1898]

References. Harpalus tristis: Apfelbeck, 1904: 199 (“Scutari”); Müller, 1931: 65 (“Skutari”).

Notes. Tschitschérine (1901: 146) stated H. skutariensis Reitter is synonym of H. tristis, but a

bit later Daniel & Daniel (1902: 250-251) synonymized H. tristis to H. danieli. Apfelbeck (1904:

199) accepted the opinion of Tschitschérine. Later on, Mlynář (1979) solved the problem (see

notes under H. anatolicus). Recently, Kataev & Wrase (2003: 375) have indicated H. danieli for

European Turkey and former Yugoslavia (Serbia and Montenegro). I consider, however, due

to lack of reliable records, the species should be excluded from the faunas of Albania and

adjacent territories in the north.


[Harpalus triseriatus A. Fleischer, 1897]

Notes. This Balkan endemic species lives in Bosnia Herzegovina, Macedonia, Bulgaria and

Greece (Kataev & Wrase, 2003: 383) and most probably inhabits Albania as well.

Species group “cupreus”

56.21. Harpalus cupreus Dejean, 1829

56.21.1. Harpalus cupreus fastuosus Faldermann, 1836

References. Harpalus cupreus fastuosus: Schatzmayr, 1943b: 104 (Lushnje).

Material studied. “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Murrsi-

Kara Alibey” / “leg. Bischoff Albania 1934” / “cotype” / “Harpalus bischoffi Schaub. det. Dr.

E. Schaub. 35” / “(H. bischoffi Schb. i. l.) = H. cupreus Z. Mlynar det. 1976”, 1 m. with extracted

genitalia (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).

Notes. Kataev & Wrase (2003: 374) have omitted the species for Albania. Here it is confirmed

for the country. Harpalus bischoffi Schauberger is nomen nudum.

56.22. Harpalus euchlorus Ménétriés, 1839

Material studied. “Skutari” / “Strasser” / “collectio Paganetti”, 2 s. (NMW).

Notes. New species for the fauna of Albania. B. Kataev (personal communication) expressed

the opinion that H. euchlorus may be a synonym of the preceding species.

Species group “dimidiatus”

56.23. Harpalus dimidiatus (P. Rossi, 1790)

References. Harpalus dimidiatus: Apfelbeck, 1904: 193 (Vlorë; Berat); Apfelbeck, 1907a: 495

(northeast of Shkodër); Apfelbeck, 1909: 298 (Vlorë; Berat); Csiki, 1940: 220 (Kula Ljums);

Schatzmayr, 1943b: 104 (“Poggio Boschetto”; Lushnje; Dukati); Hieke et al., 1986: 187 (North

Albanian Alps near Theth); Hieke & Wrase, 1988: 132; Kataev & Wrase, 2003: 375.

Material studied. “Merdita M. Scheit”, 2 s. (NMW); “Skutari Mesi”, 2 s. (NMW); “Albania

Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Alban. Exp. Kula Ljums 18-

28.V.’18”, 5 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 2 s. (NMW); more than 60 s. from

different localities (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 2 s. (HNHM);

“Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 12 s. (HNHM); “Albania Shkoder” /

“21.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); Ibë, 24.IV.1962, 2 f. (MNST); Ibë, 4.VII.1963, 2 f.

(MNST); Mal i Dajtit Mt., 10.V.1965, 3 m., 1 f. (MNST); Mal i Dajtit Mt., 24.V.1966, 1 m.

(MNST); Tiranë, 7.IV.1966, 1 m. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993

leg. Heinz u. S. Peks”, 10 s. (MNHUB); “AL (Vlore) Llogora-Paß 1025 m 29.IV.1993 leg. F. +

S. Peks”, 5 s. (MNHUB); “AL (Shkodar) Paß SE Puke 850 m 2.V.1993”, 1 s. (MNHUB);

Hotolisht - Librazhd, 300 m, 7.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS);

Librazhd - Prenjas, 750 m, 7.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS);

Tiranë, botanical garden, 8.V.1995, 3 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Dukati

Mt., 30 km S Vlorë, 450 m, 11.V.1995, 2 s., L. Penev leg. (NMNHS); Kiari Pass, 1160 m,

1.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Tiranë Park, 22.II.1998, 1 s., R. Ward leg. (CMNH).


Additional material included (studied S. Hristovski, A. Vigna Taglianti and D.W. Wrase).

Tiranë, XI.58, 1 s. (collector illegible) (cWR); Tiranë District, Kffaba-Paß, Südseite, 500 m,

29.IV.1993, 5 s., H. & S. Peks leg. (cWR); Korcë District, Mai i That Mt., Zwezde-Sattel, 1000-

1100 m, 4.V.1993, 6 s., Heinz & Peks leg. (cWR); Fier, 4.VII.1993, 1 f., L. Latella leg. (cAVT);

Tiranë, botanical garden, 4.VI.2002, 1 f., A. Paparistos leg. (cAVT); Tiranë, Mal i Dajtit Mt.,

7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelenë District), valley of

Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva”

at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).

Species group “dispar”

56.24. Harpalus punctatostriatus Dejean, 1829

References. Harpalus punctatostriatus: Apfelbeck, 1909: 298 (Vlorë; Sop); Schatzmayr, 1943b:

103 (Lushnje); Hieke & Wrase, 1988: 128; Kataev & Wrase, 2003: 380.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Albania 1918”, 1 s. (HNHM);

“Lum i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Murthi” / “Albania leg.


56.25. Harpalus pygmaeus Dejean, 1829

References. Harpalus pygmaeus: Apfelbeck, 1904: 195 (Vlorë; Berat); Apfelbeck, 1909: 298

(Berat); Hieke et al., 1986: 188 (Ibë near Krraba); Hieke & Wrase, 1988: 139; Kataev & Wrase,

2003: 380.

Material studied. “Albania 1918”, 1 s. (HNHM); “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 10 s.

(NMW); “Mal i Dajtit juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Ura Erzen” /

“Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1933”,

1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania

leg. Bischoff 1935”, 1 s. (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s.

(HNHM); “Albania Dajti” / “6.V.1959 leg. Xh. Murraj”, 1 s. (HNHM); Sauk, 9.VII.1961, 3 s.

(MNST); Tiranë, 10.V.1962, 1 s. (MNST); Tiranë Park, 22-31.X.1997, 2 s., R. Ward leg. (CMNH);

Tiranë Park, 16-21.II.1998, 2 s., R. Ward leg. (CMNH).

Notes. The specimens from the locality “Tiranë Park” are somewhat bigger (7.1-7.4 mm),

compared with specimens that inhabit Central Europe (cfr. Hůrka, 1996).

Species group “hospes”

56.26. Harpalus hospes Sturm, 1818

56.26.1. Harpalus hospes hospes Sturm, 1818

Material studied. “AL (Korcë) Qaf e Qarrit – 1100 m 4.V.1993, leg. Heinz u. S. Peks”, 9 s.

(MNHUB).



Species group “affinis”

56.27. Harpalus affinis (Schrank, 1781) [= aeneus (Fabricius, 1775); viridulus (Geoffroy,

1785); limbopunctatus Fuss, 1858]

References. Harpalus aeneus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Schatzmayr, 1943b: 103 (Lushnje). Harpalus affinis: Csiki, 1940: 219 (Gjalicë Mt.); Kataev &

Wrase, 2003: 371. Harpalus affinis ab. viridulus: Csiki, 1940: 219-220 (Gjalicë Mt.; Korab Mt.);

Harpalus affinis ab. limbopunctatus: Csiki, 1940: 220 (Gjalicë Mt.; Korab Mt.).

Material studied. Korcë, 10-20. VIII.1962, 1 m., 1 f. (MNST); North Albanian Alps, Theth,

800-1200 m, 28.V.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e

Radohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian

Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian

Alps, 5-9.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River,

900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 8-


Notes. In addition to the material studied above, in NMW and MNHUB I have seen more

specimens, which remained unregistered.

Species group “distinguendus”

56.28. Harpalus distinguendus (Duftschmid, 1812)

56.28.1. Harpalus distinguendus distinguendus (Duftschmid, 1812)

References. Harpalus distinguendus: Apfelbeck, 1904: 192 (Vlorë; Durrës); Apfelbeck, 1909:

298 (Vlorë; Berat); Schatzmayr, 1943b: 103 (Dukati; Lushnje); Hieke & Wrase, 1988: 131.

Harpalus distinguendus distinguendus: Kataev & Wrase, 2003: 376.

Material studied. Ibë, 18.V.1960, 1 f. (MNST); Sauk, 5.III.1961, 1 m. / 23.IV.1963, 1 f. (MNST);

Rojan, 6.III.1961, 1 f. (MNST); Rodeni, 26-28.VII.1962, 1 f. (MNST); Tiranë, V.1963, 1 m. /

7.IV.1966, 5 m., 6 f. (MNST); Mal i Dajtit Mt., 10.V.1965, 1 f. (MNST); Kurbneshi, VI.1993, 1

f., P. Beron & B. Petrov leg. (NMNHS); Saranda District, Himarë, 3-4.V.1994, 1 m., 2 f., P.

Stoev leg. (NMNHS); Saranda District, Qeparo, 4-5.V.1994, 2 s., P. Stoev leg. (NMNHS);

Saranda District, Ljukove, 3-4.VI.1994, 1 m., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994, 1

f., B. Guéorguiev leg. (NMNHS); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B.

Petrov, P. Stoev leg. (NMNHS); Librazhd - Prenjas, 750 m, 7.V.1995, 1 f., S. Golovach, B.

Petrov, P. Stoev leg. (NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 1 f., S. Golovach,

B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 22-31.X.1997 / 8.II.1998 / 23-28.II.1998, 41

s., R. Ward leg. (CMNH).


and Gjirokastër), 350 m, 23.IV.2003, 1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Notes. In addition to the material studied above, in NMW and MNHUB I have seen some

more specimens, which remained unregistered.


56.29. Harpalus saxicola Dejean, 1829 [= fugax Faldermann, 1836]

References. Harpalus fugax: Apfelbeck, 1907a: 498-499, 501 (Oroshi; Mal i Shenjt Mt.: top

region, 1500 m). Harpalus saxicola: Kataev & Wrase, 2003: 381.

Material studied. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 7 s. (NMW); “Rapsa 10-

18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Tartarit” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i That” / “Albania Ramadan coll. Bischoff ”, 1 s.

(MNHUB); “Albania Tirana” / “28.IX.1958 leg. Xh. Murraj”, 1 s. (HNHM).

Species group “oblitus”

56.30. Harpalus oblitus Dejean, 1829

56.30.1. Harpalus oblitus oblitus Dejean, 1829

References. Harpalus oblitus: Hieke & Wrase, 1988: 130. Harpalus oblitus oblitus: Kataev, 1993:

89, fig. 106; Kataev & Wrase, 2003: 379.

Notes. In NMW and MNHUB there are several specimens with labels indicating they come

from Albania. Additional labels with detailed localities revealed however that the material

originates from sites located in the valley of Sava River (Croatia and Serbia). Most probably,

Hieke & Wrase (1988: 130) have based their data for the species occurrence in Albania on the

wrongly labeled specimens in question. In spite of that Kataev (1993) displayed on map one

sure locality for the country.

Subgenus Pseudophonus Motschulsky, 1844

56.31. Harpalus griseus (Panzer, 1796)

References. Harpalus griseus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 103 (Dukati).

Material studied. “Elbasan, Alb., 28.VIII.18 Dr. H. Priesner”, 3 s. (NMW); Rodeni, 2-

22.VII.1962, 1 m. (MNST); Tiranë, Selit e Vogel, 17.IX.1993, 2 s., S. Beshkov leg. (NMNHS);

“Albania, Alban Alps, Okol, 900 m, swept singled, 16-19.VII.1996, leg. A. Orosz”, 1 m. (HNHM).


for the country.

56.32. Harpalus rufipes (DeGeer, 1774) [= pubescens (O.F. Müller, 1776)]

References. Harpalus pubescens: Apfelbeck, 1904: 187 (Vlorë; Berat); Schatzmayr, 1943b: 103

(Dukati). Ophonus pubescens: Apfelbeck, 1909: 298 (Berat). Harpalus rufipes: Csiki, 1940: 219

(Kula Ljums); Kataev & Wrase, 2003: 385. Pseudophonus rufipes: Hieke et al., 1986: 187 (Vlorë;

North Albanian Alps near Theth; Shkodër); Hieke & Wrase, 1988: 126.

Material studied. “Elbasan, Alb., 28.VIII.18 Dr. H. Priesner”, 1 s. (NMW); “Alban., Ricavac

Penther”, 1 s. (NMW); “Alban. Exped. Kula Ljums 14.20.VII’18”, 1 s. (NMW); Tiranë,

28.IX.1958, 1 f. (MNST); Ibë, 18.V.1960, 1 m., 1 f. (MNST); Theth, 1-8.VIII.1961, 3 f. (MNST);

Tiranë, 10.V.1961, 1 m., 2 f. (MNST); Rushkull, 6.IV.1962, 1 f. (MNST); Rodeni, 2-28.VII.1962,

2 m., 2 f. (MNST); Shkodër, 14.V.1963, 2 m. (MNST); Tiranë, 11.IX.1963, 2 f. (MNST); Shkodër,

12. VII.1965, 1 m. (MNST); Mal i Dajtit Mt., 24.V.1966, 1 m. (MNST); North Albanian Alps,


Bogë, 1000-1100 m, 5-9.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District,

Merkurth, 11.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).

Additional material included (studied S. Hristovski and A. Vigna Taglianti). Tropoia, valley

of Valbona River, 900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT); Durrës, Plepa, 15.VIII.1995,

1 m., 2 f., D. Antonelli leg. (cAVT); Tiranë, Mal i Dajtit Mt., 7.VI.2002, 1 f., A. Paparistos leg.

(cAVT); M. Polisit, 8.VII.2004, 1 f., A. Paparistos leg. (cAVT); Malj Cojepit 1300-1400,

17.VII.2004, 1 m., A. Paparistos leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-

15.VII.2006, 3 s., leg. M. Komnenov (cSH).

Notes. Besides the material studied above, in NMW and MNHUB I have seen more specimens,


57. Genus Ophonus Dejean, 1821

Subgenus Brachyophonus Sciaky, 1987

57.1 Ophonus krueperi Apfelbeck, 1902

References. Ophonus krueperi: Kataev & Wrase, 2003: 388.

Material included (studied D.W. Wrase). “AL (Kolonjë) Paß ca. 26 km n. Leskovik -1000 m

5.V.1993 Heinz leg.”, 1 m. (cWR; probably more specimens in cWH).

Notes. Apfelbeck published two descriptions for this taxon. First he described Ophonus krueperi

(Apfelbeck, 1902b: 543), and after than – Harpalus krueperi (Apfelbeck, 1904: 185). Incorrectly

the second description has long time been accredited to be the valid. Actually, this is incorrect

and the older description must be assigned as valid. Müller (1931: 48 sub Harpalus krueperi) and

Sciaky (1987: 69 sub Ophonus krueperi) followed the record “Chani Driskos” (Apfelbeck, 1904)

when they cited the species for Albania. However, this locality lies in Northwestern Greece.

According to D.W. Wrase (personal communication), Kataev & Wrase (2003: 388) used the

record near Leskovac, marking the species for Albania. First detailed record for the country.

Subgenus Hesperophonus Antoine, 1959

57.2. Ophonus azureus (Fabricius, 1775) [= oberthuri (Pater, 1938)]

References. Harpalus azureus: Apfelbeck, 1904: 184 (Vlorë); Csiki, 1940: 219 (Kula Ljums;

Ura i Lopez). Ophonus azureus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Apfelbeck, 1909: 298 (Vlorë); Hieke et al., 1986: 187 (North Albanian Alps near Theth; Bizë

near Shëngjergji); Hieke & Wrase, 1988: 125; Kataev & Wrase, 2003: 388. Harpalus azureus

oberthuri: Schatzmayr, 1943b: 102 (Lushnje).

Material studied. “Latif Orosi”, 1 s. (NMW); “Merdita Orosi” / “Apfelb. Alban. spt. IV.V.’05”,

2 s. (NMW); “Merdita M. Sheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Albania Kula

Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Ura e Loper” / “Akad. Balk.

Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Kula Ljums 18.-28.V.’18”, 12 s. (NMW);

“Alban. Exped. 1918 Bicaj 14.15.VI.”, 8 s. (NMW); “Alban. Exped.’18 Pashtrik, 4-14.7.”, 2 s.

(NMW); 53 s. from different localities (MNHUB); Ibë, 17-25.VI.1961, 1 f. (MNST); Rodens,

2-22. VII.1962, 1 s. (MNST); “AL (Berat). Paß ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. +

S. Peks”, 2 m., 1 f. (MNHUB); “AL (Korce) Qaf e Qarrit – 1100 m, 4.v.1993 leg. Heinz u.


Sonja Peks”, 1 m. (MNHUB); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993,

1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District, Merkurth, 11.VI.1993, 1 s., P. Beron

& B. Petrov leg. (NMNHS); Saranda District, Lyukove, 5.V.1994, 2 s., P. Stoev leg. (NMNHS);

Saranda District, Qeparo, 4-5.V.1995, 1 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove,

5.V.1995, 1 s., P. Stoev leg. (NMNHS); Dukati Mt., 30 km S Vlorë, 450 m, 11.V.1995, 1 s., L.

Penev leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 5 s., G. Blagoev leg. (NMNHS); Tiranë

Park, 15.II.1998, 1 s., from grass and soil around tree trunks, R. Ward leg. (CMNH); Tiranë

District, Petrela, 28.XI.2000, 1 s., A. Zhalov leg. (NMNHS).

Notes. In the region of Gllavë O. azureus was found to lives sympatrically with O. similis.

57.3. Ophonus cribricollis (Dejean, 1829)

References. Harpalus cribricollis: Apfelbeck, 1904: 184-185 (Vlorë; Berat); Csiki, 1940: 219

(Shkëlzen Mt.; Kula Ljums; Gostilë). Ophonus cribricollis: Apfelbeck, 1907a: 499 (Mal i Shenjt

Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë; Berat); Hieke & Wrase, 1988: 126;


Material studied. “Merdita Oroshi”, 3 s. (NMW); “Skutari Mesi” / “Apfelb. Alban. spt.

IV.V.’05”, 3 s. (NMW); “Pasha-Liman Albanien V.1908 H. Hopp”, 1 s. (MNHUB); “Alban.

Exped. Kula Ljums 18-28.V.1918”, 3 s. (NMW); “Devoll-Tal juni 1932” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tepelene

juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania

1934”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Dukati

Mt., 30 km S Vlorë, 450 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelenë District),

18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Notes. In addition to the material studied above, in HNHM the present author has seen more

specimens from the localities “Kula Lums”, “Delvine”, “Skölsen” and “Mont Peklen”. Sciaky

(1987: 66) cited the locality “Štimlje” for Albania, but actually this place is situated in the

Kosovo Region of Serbia.

57.4. Ophonus similis (Dejean, 1829)

Material studied. “Hani Grabom Mai” / “Alban. montenegr. Grenze “Alban. Exped. 1918

Bicaj 14.15.VI.”, 1 s. (NMW); “AL (Berat). Paß ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. +

S. Peks”, 2 f. (MNHUB).


57.5. Ophonus subquadratus (Dejean, 1829) [= meridionalis (Dejean, 1829)]

References. Ophonus meridionalis: Apfelbeck, 1909: 298 (Berat; Gramata).

Material examined. “Liqen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);

“Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).




for the country.


Subgenus Incisophonus Sciaky, 1987

57.6. Ophonus incisus (Dejean, 1829)

References. Ophonus incisus: Hieke & Wrase, 1988: 121; Kataev & Wrase, 2003: 389.

Material studied. “Murthi” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Albania Cerric”

/ “16.IV.1958 leg. Xh. Murraj”, 1 s. (HNHM); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS).


Subgenus Metophonus Bedel, 1897

57.7. Ophonus brevicollis (Audinet-Serville, 1821) [= albanensis (Schauberger, 1933)]

References. Harpalus parallelus albanensis: Schauberger, 1933: 65 (Vlorë, loc. typ.). Harpalus

albanensis: Turin, 1981: 156. Ophonus brevicollis: Sciaky, 1987: 92-93 (Vlorë); Kataev & Wrase,

2003: 389.

57.8. Ophonus cordatus (Duftschmid, 1812)

References. Ophonus cordatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);


Material examined. “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).

57.9. Ophonus laticollis Mannerheim, 1825 [= punctatulus (Duftschmid, 1812)]

References. Harpalus punctatulus: Csiki, 1940: 219 (Gjalicë Mt.). Ophonus punctulatus sic!: Hieke

et al., 1986: 187 (Mali me Gropë near Livadhet e Selitës). Ophonus laticollis: Kataev & Wrase,

2003: 390.


“Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

Notes. Hieke & Wrase (1988: 122 sub O. punctatulus) noted this species for Albania apparently

following the record “Prisren” (Apfelbeck, 1904: 183 sub Harpalus punctatulus). Actually this

locality is situated in the Kosovo Region of Serbia.

57.10. Ophonus melletii (Heer, 1837)

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB).


57.11 Ophonus parallelus (Dejean, 1829) [= albanicus Schauberger, 1933]

References. Ophonus parallelus albanicus: Schauberger, 1933a: 65 (Vlorë, loc. typ.). Ophonus

parallelus: Hieke & Wrase, 1988: 122; Kataev & Wrase, 2003: 390.

Material studied. “Alban. Exp.’18 Kruma 5.VI.”, 1 s. (NMW); “Maranaj” / “Albania leg.

Bischoff 1937”, 1 s. (MNHUB); “AL (Berat) Paß ca. 5 km s. Gllavë 600 m, 27.IV.1993 leg.

Heinz u. S. Peks”, 1 m. (MNHUB).


57.12. Ophonus puncticeps Stephens, 1828

References. Harpalus puncticeps: Horion, 1941: 212; Magistretti, 1965: 280; Geisthardt, 1975: 8. Ophonus

puncticeps: Hieke et al., 1986: 187 (North Albanian Alps near Theth); Kataev & Wrase, 2003: 390.

Material studied. “Mali i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “AL (Berat).

Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 f. (MNHUB).

57.13. Ophonus puncticollis (Paykull, 1798)

References. Ophonus puncticollis: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Apfelbeck, 1909: 298 (Berat); Kataev & Wrase, 2003: 390. Harpalus puncticollis: Müller, 1931:

50 (Tomor Mt.); Csiki, 1940: 219 (Kula Ljums); Hieke & Wrase, 1988: 122.


“Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).

57.14. Ophonus rufibarbis (Fabricius, 1792)

References. Ophonus rufibarbis: Kataev & Wrase, 2003: 391.

Material studied. “Berat…Alban.”, 1 s. (MNHUB); “AL (Vlorë) Llogora-Paß (N-Seite) 800

m, 29.IV.1993 leg. Heinz & Sonja Peks”, 1 f. (MNHUB).

Notes. Hieke & Wrase (1988: 124) noted the species for Albania apparently following the

record “Prisren” (Apfelbeck, 1904: 183 sub Harpalus rufibarbis). However, this place is situated

in the Kosovo Region of Serbia. First detailed records for Albania.

57.15. Ophonus rupicola (Sturm, 1818)

References. Harpalus rupicola: Apfelbeck, 1904: 183 (Velipoja); Hieke & Wrase, 1988: 124.

Ophonus rupicola: Kataev & Wrase, 2003: 391.

Material studied. Kjari Pass, 1160 m, 1.VI.1995, 1 s., G. Blagoev leg. (NMNHS).

57.16. Ophonus schaubergerianus (Puel, 1937)

References. Ophonus schaubergerianus: Kataev & Wrase, 2003: 391.

Material studied. “Mal i Shenit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).


Subgenus Ophonus s.str.

57.17. Ophonus ardosiacus (Lutshnik, 1922)

References. Ophonus ardosiacus: Kataev & Wrase, 2003: 391.

Material studied. Ibë, 4.VII.1962, 1 f. (MNST).

Additional material included (studied S. Hristovski and A. Vigna Taglianti). Durrës, Plepa,

15.VIII.1995, 2 m., D. Antonelli leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-

15.VII.2006, 3 s., leg. M. Komnenov (cSH).



57.18. Ophonus diffinis (Dejean, 1829)

References. Harpalus diffinis: Schatzmayr, 1943b: 102 (Mal i Krujes). Ophonus diffinis: Kataev &

Wrase, 2003: 391.

[Ophonus franziniorum Sciaky, 1987]

Notes. According to D.W. Wrase (personal communication), the occurrence of this winged

species in Albania is very probable. Except its proven presence in Bulgaria (Sciaky, 1987: 49;

Wrase, 1991: 14), there are reliable records for its occurrence in the Greek Ionic Islands and

European Turkey (D.W. Wrase, B. Guéorguiev, unpublished data).

57.19. Ophonus sabulicola (Panzer, 1796) [= columbinus (Germar, 1817)]

References. Ophonus sabulicola: Apfelbeck, 1909: 298 (Vlorë; Berat); Kataev & Wrase, 2003:

392. Harpalus sabulicola: Csiki, 1940: 219 (Kula Ljums). Harpalus sabulicola columbinus: Schatzmayr,

1943b: 102 (Tërpan; Dukati). Ophonus sabulicola columbinus: Hieke et al., 1986: 187 (Poliçan near

Tomor Mt.).

Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); 11 s.

from different localities (MNHUB); Ibë, 4.VII.1963, 1 f. (MNST); “AL (Berat) Paß ca. 5 km s.

Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 2 s. (MNHUB); “AL (Korcë) Qaf e Qarrit –

1100 m 4.V.1993, leg. Heinz u. S. Peks”, 18 s. (MNHUB).

Additional material included (studied S. Hristovski and D.W. Wrase). Korcë District, Mai i

That Mt., Zwezde-Sattel, 1000-1100 m, 4.V.1993, 1 f., W. Heinz & S. Peks leg. (cWR); “AL

(Kolonjë) Paß ca. 2 km ne. Leskovik -1000 m 5.V.1993 H. & S. Peks leg.”, 16 s. (cWR); 2 km N

from Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Hotel “Edva”, Radhime

(Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH).

57.20. Ophonus stictus Stephens, 1828

Material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 8-

15.VII.2006, 1 f., leg. M. Komnenov (cSH).


58. Genus Parophonus Ganglbauer, 1891

Subgenus Ophonomimus Schauberger, 1923

58.1. Parophonus hirsutulus (Dejean, 1829)

References. Ophonus hirsutulus: Apfelbeck, 1902a: 99 (Velipoja near Lezhë). Harpalus hirsutulus:

Apfelbeck, 1904: 186 (Velipoja). Parophonus hirsutulus: Apfelbeck, 1907a: 495 (Buenë River);

Sciaky, 1992: 50. Ophonomimus hirsutulus: Hieke & Wrase, 1988: 141.

Material studied. “Oboti Bojana”, 1 s. (HNHM); “Oboti Schirei”, 2 s. (HNHM).

Notes. Ito (2003b: 392) has omitted the species for Albania. Here it is confirmed for the

country.


Subgenus Parophonus s.str.

58.2. Parophonus dejeani (Csiki, 1932)

References. Parophonus dejeani: Ito, 2003b: 393.

58.3. Parophonus laeviceps (Ménétriés, 1832) [= fallax (Peyron, 1858); planicollis Apfel-

beck, 1904 non (Dejean, 1829)]

References. Ophonus planicollis: Apfelbeck, 1903: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë).

Harpalus planicollis: Apfelbeck, 1904: 186 (Vlorë). Tachyophonus fallax: Schatzmayr, 1943b: 104

(Stan-Karbunarë). Parophonus laeviceps: Hieke & Wrase, 1988: 142; Ito, 2003b: 393.

58.4. Parophonus maculicornis (Duftschmid, 1812)

References. Harpalus maculicornis: Apfelbeck, 1904: 186 (Velipoja; Vlorë; Berat). Ophonus

maculicornis: Apfelbeck, 1909: 298 (Vlorë; Berat). Trichotichnus maculicornis: Csiki, 1940: 220 (Kula

Ljums). Parophonus maculicornis: Hieke et al., 1986: 186 (Borshi near Vlorë); Hieke & Wrase,

1988: 142; Ito, 2003b: 393.

Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Pasha-Liman Albanien V.1908

H. Hopp”, 2 s. (MNHUB); “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Albania Kula

Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Elbasan, Alb. 2.IX.18 Dr. H. Priesner”,

1 s. (NMW); “Logara juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” /

“Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Burelle” / “leg. Bischoff Albania 1934”, 1 s.

(MNHUB); “Lum i Erzen Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murrsi-

Kara Alibey” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Cerrik”

/ “18.IV.1958 leg. Xh. Murraj”, 1 s. (HNHM); Lushnja, Divjaka National Park, 10.V.1995, 2 s., S.

Golovach, B. Petrov, P. Stoev leg. (cWR; NMNHS); Tiranë Park, 1.II.1998 / 5.II.1998 / 8.II.1998,


58.5. Parophonus mendax (P. Rossi, 1790)

References. Harpalus mendax: Apfelbeck, 1904: 187 (Vlorë). Ophonus mendax: Apfelbeck, 1909:

298 (Vlorë). Tachyophonus mendax: Schatzmayr, 1943b: 104 (Stan-Karbunarë). Parophonus mendax:

Sciaky, 1992: 55; Hieke & Wrase, 1988: 143; Ito, 2003b: 393.

Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Alessio, Alb. Matzenauer”,

1 s. (NMW); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 1-4.I.1998 / 11.I.1998

/ 1.II.1998, 10 s., R. Ward leg. (CMNH).

58.6. Parophonus planicollis (Dejean, 1829)

References. Ophonus planicollis: Apfelbeck, 1903: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë).

Harpalus planicollis: Apfelbeck, 1904: 186 (Vlorë). Parophonus planicollis: Ito, 2003b: 393.

Material studied. “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj” / “suturifer Rttr. det.

Jedlicka”, 1 f. (HNHM).


Notes. Hieke & Wrase (1988: 142) noted that the Apfelbeck’s figures for P. planicollis (Apfelbeck,

1904: 187) refer to P. laeviceps.

59. Genus Trichotichnus A. Morawitz, 1863

Subgenus Trichotichnus s.str.

59.1. Trichotichnus laevicollis (Duftschmid, 1812)

59.1.1. Trichotichnus laevicollis laevicollis (Duftschmid, 1812)

References. Trichotichnus laevicollis: Schatzmayr, 1926: 35 (Çukall Mt.); Horion, 1941: 233; Hieke

& Wrase, 1988: 141. Trichotichnus laevicollis laevicollis: Ito, 2003b: 396.

Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Cucale” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 4 s.

(MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., 3 f., P. Beron & B.


Subtribe STENOLOPHINA Kirby, 1837

60. Genus Acupalpus Latreille, 1829

Subgenus Acupalpus s.str.

60.1. Acupalpus brunnipes (Sturm, 1825)

Material studied. “Valona”, 1 s. (HNHM); “Oboti Bojana”, 1 s. (HNHM); “Albania Vlesh” /

“30.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM).


60.2. Acupalpus elegans (Dejean, 1829)

References. Acupalpus elegans: Hieke & Wrase, 1988: 116; Jaeger, 1992: 224; Jaeger, 1999: 957;

Jaeger & Kataev, 2003: 398.

Material studied. “Skutari Bojana”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB); “Liqen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS).


60.3. Acupalpus luteatus (Duftschmid, 1812)

References. Acupalpus luteatus: Hieke & Wrase, 1988: 118; Jaeger, 1992: 225; Jaeger, 1999: 959;



(HNHM); “Skutari Bojana”, 1 s. (HNHM); “Skutari Drin”, 1 s. (HNHM).

Additional material included (studied B. Jaeger). “Velipoja” / “Coll. Paganetti”, 1 s. (NMW).



60.4. Acupalpus maculatus (Schaum, 1860) [= immundus Reitter, 1900]

References. Acupalpus immundus: Apfelbeck, 1904: 206 (? Shkodër). Acupalpus maculatus: Csiki,

1940: 221 (Kula Ljums); Horion, 1941: 238; Focarile, 1959: 81; Hieke & Wrase, 1988: 118;

Jaeger, 1992: 225; Jaeger, 1999: 960; Jaeger & Kataev, 2003: 398.


“Busher Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg.

Bischoff 1935”, 1 s. (MNHUB); “Presba See” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);

Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS).

Additional material included (studied B. Jaeger). “Elbasan, VIII.1918, Priesner”, 2 s. (MTKD);

“Elbasan, Mader”, 1 s. (MTKD); Shkodër, VIII. 1941, 12 s., Capra leg. (MHNG; cBJ);

Rrogozhinë, 1 s. (cBJ).

Notes. The record “Scutari” (Apfelbeck, 1904) is questionable because the material of Strasser

could be collected from the city of Üsküdar - in the Asian seacoast of Bosporus.

60.5. Acupalpus meridianus (Linnaeus, 1761)

References. Acupalpus meridianus: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226; Jaeger, 1999:

960; Jaeger & Kataev, 2003: 398.

Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB).


60.6. Acupalpus notatus Mulsant & Rey, 1861

References. Acupalpus notatus: Horion, 1941: 238; Focarile, 1959: 82; Hieke et al., 1986: 188

(Uji Ftohte, south of Tepelenë; Ibë near Krraba); Hieke & Wrase, 1988: 118; Jaeger, 1988: 20

(Ljum, Shkumbin; Durrës: Rogozina; Elbasan; Mati: Burrelë; Tepelenë: Uji Ftohte; Ibë near

Krraba; Tiranë; Vlorë); Jaeger, 1992: 226; Jaeger, 1999: 961; Jaeger & Kataev, 2003: 398.

Material studied. “Elbasan Alb., Mader”, 3 s. (NMW; MNHUB); “Vorra, Alb. leg. Mader”, 2 s.

(NMW); “Ljum i Shkumbin, Juni 1932” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Burelle” /

“Albania leg. Bischoff 1934”, 16 s. (MNHUB); “Shullaz” / “Albania leg. Bischoff 1934”, 5 s. (MNHUB);

“Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Albanien-Exp. DEI Uji Ftohte, 200 m, südl. Tepelene lux 29.-31.V.1961”, 3

s. (MNHUB); Tiranë Park, 22.II.1998, 30 s., grasses along small sluggish stream, R. Ward leg. (CMNH).

Additional material included (studied B. Jaeger and A. Vigna Taglianti). “Elbasan, Mader”,

4 s. (MTKD, cBJ); “Elbasan, Priesner”, 4 s. (MTKD); Shkodër, VIII. 1941, 2 s., Capra leg.

(MHNG); Durrës, Plepa, 15.VIII.1995, 1 m., D. Antonelli leg. (cAVT).

60.7. Acupalpus paludicola Reitter, 1884 [= puncticollis Auct. nec (Coquerel, 1859)]

References. Acupalpus paludicola: Apfelbeck, 1904: 206 (Vlorë; Velipoja); Apfelbeck, 1909: 298

(Vlorë-Malket); Jaeger, 1989: 148 (Vlorë; Lum i Tiranes; Shkodër); Jaeger, 1992: 226; Jaeger,

1999: 961; Jaeger & Kataev, 2003: 399. Acupalpus puncticollis: Hieke et al., 1986: 188 (Uji Ftohte,

south of Tepelenë); Hieke & Wrase, 1988: 117.

Material studied. “Albania Avlona v. Oertzen”, 3 s. (MNHUB); “Skutari l. See-Ufer”, 1 s.

(NMW); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB; cBJ).


Notes. All older references and collection records in the museums about A. puncticollis (Coquerel,

1859) from Albania refer to A. paludicola.

60.8. Acupalpus parvulus (Sturm, 1825) [= dorsalis (Fabricius, 1787)]

References. Acupalpus dorsalis: Apfelbeck, 1904: 206 (Velipoja; Vlorë); Apfelbeck, 1909: 298

(Vlorë); Csiki, 1940: 221 (Kula Ljums); Hieke & Wrase, 1988: 117.


Tiranë Park, 8.II.1998 / 1.III.1998, 3 s., from grass and soil around tree trunks, R. Ward leg.

(CMNH).

Notes. Jaeger & Kataev (2003: 399) have omitted the species for Albania. Here it is confirmed

for the country. According to B. Jaeger and D.W. Wrase (personal communication) the records

from Albania may concern aberrant specimens of A. maculatus without darkened humeral

macula, but the only certainty for a correct identification is the study of male genitalia.

60.9. Acupalpus planicollis (Schaum, 1857)

References. Acupalpus planicollis: Apfelbeck, 1904: 205 (Velipoja); Apfelbeck, 1909: 298 (Vlorë-

Malket); Hieke & Wrase, 1988: 116; Guéorguiev & Guéorguiev, 1995: 177; Guéorguiev et al.,

1997: 20; Jaeger & Kataev, 2003: 399.

Material studied. “Liqen i Butrintes vivari” / “leg. Bischoff Albani 1934”, 1 s. (MNHUB).

Additional material included (studied B. Jaeger). “Velipoja, Mustjabeg”, 1 s. (ZSM).

60.10. Acupalpus suturalis Dejean, 1829

60.10.1. Acupalpus suturalis suturalis Dejean, 1829

References. Acupalpus suturalis: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226. Acupalpus suturalis

suturalis: Jaeger, 1999: 962; Jaeger & Kataev, 2003: 399.

Material studied. “Oboti Bojana”, 1 s. (HNHM); Tiranë Park, 22-31.X.1997, 1 s., R. Ward

leg. (CMNH); Tiranë Park, 22.II.1998, 3 s., R. Ward leg. (CMNH).

Additional material included (studied B. Jaeger). “Scutari, VIII. 1941, Capra”, 1 s. (MHNG).


61. Genus Anthracus Motschulsky, 1850

61.1. Anthracus longicornis (Schaum, 1857)

References. Anthracus longicornis: Apfelbeck, 1904: 207 (Velipoja); Hieke & Wrase, 1988: 118;



(HNHM); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB).

61.2. Anthracus quarnerensis (Reitter, 1884)

References. Anthracus quarnerensis: Apfelbeck, 1904: 208 (Velipoja); Hieke & Wrase, 1988:

119; Jaeger & Kataev, 2003: 400.


62. Genus Bradycellus Erichson, 1837

Subgenus Bradycellus s.str.

62.1. Bradycellus caucasicus (Chaudoir, 1846) [= collaris (Paykull, 1798)]

References. Bradycellus collaris: Csiki, 1940: 221 (Shkëlzen Mt.).

Material studied. “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM).

Notes. Jaeger & Kataev (2003: 400) have omitted the species for Albania. Here it is confirmed

for the country.

62.2. Bradycellus distinctus (Dejean, 1829)

References. Bradycellus distinctus: Jaeger, 1990: 10; Jaeger, 1999: 966; Jaeger & Kataev, 2003: 401.

Material studied. “Skutari Bojana”, 5 s. (NMW).

Additional material included (studied B. Jaeger). “Skutari Bojana”, 12 s. (MNHUB; DEI;

NMPC; cBJ).


62.3. Bradycellus ganglbaueri Apfelbeck, 1904

References. Bradycellus ganglbaueri: Jaeger, 1997: 352 (Mal i Krujes); Jaeger & Kataev, 2003: 401.

62.4. Bradycellus verbasci (Duftschmid, 1812)

References. Bradycellus verbasci: Hieke & Wrase, 1988: 120; Jaeger & Kataev, 2003: 401.

Material studied. Velipoja, 19.IX.1993, 8 s., S. Beshkov leg. (NMNHS; ZISP); Tiranë Park, 1-

9.XI.1997, 1 s., R. Ward leg. (CMNH).

Additional material included (studied B. Jaeger). “Velipoja Avola v. Oertzen”, 1 s. (MNHUB).


63. genus Dicheirotrichus Jaquelin du Val, 1857

Subgenus Dicheirotrichus s.str.

63.1. Dicheirotrichus obsoletus (Dejean, 1829)

References. Dicheirotrichus obsoletus: Apfelbeck, 1904: 211 (Velipoja); Hieke & Wrase, 1988:

120; Jaeger & Kataev, 2003: 402.


Notes. According to B. Jaeger (personal communication) E. Schauberger identified the material

from MNHUB as D. obsoletus, but a revision of that is needed.


64. Genus Loxoncus Schmidt-Göbel, 1846

64.1. Loxoncus procerus (Schaum, 1858)

References. Anoplogenius procerus: Apfelbeck, 1904: 202 (Velipoja near Lezhë); Apfelbeck, 1907a:

496 (Buenë River Outfall); Schatzmayr, 1943b: 105 (Lushnje). Loxoncus procerus: Wrase, 1991:

12; Kataev, 2002: 364 (Buenë River; Buenë River at Oboti); Jaeger & Kataev, 2003: 404.

Material studied. “Apfelb. Albanien”, 2 s. (MNHUB); “Apfelb. Albanien” / “Bojana

Mündung”, 3 s. (HNHM); “Oboti Bojana”, 2 s. (HNHM); “Skutari Bojana”, 3 s. (HNHM);

“Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

65. Genus Stenolophus Dejean, 1821

Subgenus Egadroma Motschulsky, 1855

65.1. Stenolophus marginatus Dejean, 1829

References. Egadroma marginata: Apfelbeck, 1904: 204 (Vlorë; Berat); Apfelbeck, 1907a: 496

(Buenë River); Hieke et al., 1986: 188 (Durrës; Uji Ftohte, south of Tepelenë); Hieke & Wrase,

1988: 116. Stenolophus marginatus: Apfelbeck, 1909: 298 (Vlorë); Jaeger & Kataev, 2003: 405.


(HNHM); “Skumbi Elbasan Alb. 10.8.1918 Dr. H. Priesner”, 3 s. (NMW); “Ibe Erzen” /

“Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 2 s.

(MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë” / “Albania leg.

Bischoff 1937”, 1 s. (MNHUB); “Myzeqe Ebene” / “Albania leg. Bischoff 1938”, 1 s.

(MNHUB); “Albanien-Exp. DEI Uji Ftohte, 200 m, südl. Tepelene lux 29.-31.V.1961”, 1 s.

(MNHUB).

Subgenus Stenolophus s.str.

65.2. Stenolophus abdominalis Gené, 1836

65.2.1. Stenolophus abdominalis persicus Mannerheim, 1844

Material studied. Tiranë Park, 22.II.1998, 1 s., shaded lake edge under leaves and wet, rotten

wood, R. Ward leg. (CMNH); Tiranë Park, 1.III.1998, 2 s., shaded lake edge under leaves and

wet, rotten wood, R. Ward leg. (CMNH).


65.3. Stenolophus discophorus (Fischer von Waldheim, 1823)

References. Stenolophus discophorus: Apfelbeck, 1904: 203 (Velipoja); Hieke & Wrase, 1988:

114; Jaeger & Kataev, 2003: 405. Acupalpus discophorus: Csiki, 1940: 221 (Kula Ljums).

Material studied. “Oboti Bojana”, 5 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp.

Csiki 1918”, 2 s. (HNHM); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB);

“Bojana Shkoder” / “Albania leg. Bischoff 1937”, 4 s. (MNHUB); Radona, Shkiela, 21.VI.1994,

1 s., B. Guéorguiev leg. (NMNHS).


65.4. Stenolophus mixtus (Herbst, 1784)

References. Stenolophus mixtus: Apfelbeck, 1904: 203 (Velipoja); Hieke & Wrase, 1988: 114;


65.5. Stenolophus proximus Dejean, 1829

References. Stenolophus proximus: Apfelbeck, 1904: 202 (Velipoja); Hieke & Wrase, 1988: 114;


65.6. Stenolophus skrimshiranus Stephens, 1828

References. Stenolophus skrimshiranus: Apfelbeck, 1904: 203 (Buenë River at Velipoja); Apfelbeck,

1909: 298 (Pasha Liman); Hieke & Wrase, 1988: 115; Jaeger & Kataev, 2003: 406.

Material studied. “Ljum i Schkumbin Juni 1932” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB).

65.7. Stenolophus teutonus (Schrank, 1781)

References. Stenolophus teutonus: Apfelbeck, 1904: 203 (Velipoja); Apfelbeck, 1909: 298 (Vlorë);

Murraj, 1968: 33 (Ibë); Hieke et al., 1986: 188 (Vlorë; Bizë near Shëngjergji); Hieke & Wrase,

1988: 115; Jaeger & Kataev, 2003: 406. Acupalpus teutonus: Csiki, 1940: 221 (Kula Ljums);

Schatzmayr, 1943b: 105 (Fieri).

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Alban. Exp.’18 Kula Ljums

26.VI.-3.VII.”, 2 s. (NMW); “Elbasan, Alb. 16.IX.18 Dr. H. Priesner”, 6 s. (NMW); “Berat,

Juli”, “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Busher, Juni” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara

Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Shkumbin, Juni” / “Albania leg.

Bischoff 1932”, 1 s. (MNHUB); “Maia e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Tepelene” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Burelle” / “Albania leg. Bischoff

1934”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1934”, 7 s. (MNHUB); “Shullaz” /

“Albania leg. Bischoff 1934”, 4 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1935”, 1 s.

(MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania

leg. Bischoff 1935”, 2 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);

“Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Tiranë, 24.VI.1961, 1 f. (MNST);

Rodeni, 2-22.VII.1962, 1 s. (MNST); Tiranë Park, 22.II.1998, 1 s., shaded lake edge under

leaves and wet, rotten wood, R. Ward leg. (CMNH).

Notes. The record of Murraj (1968) is included here with reservation because of the

misidentification found in his collection.


Tribe LEBIINI Bonelli, 1810

Subtribe CYMINDIDINA Laporte, 1834

66. Genus Cymindis Latreille, 1806

Subgenus Cymindis s.str.

66.1. Cymindis angularis Gyllenhal, 1810

Material studied. “Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania

leg. Bischoff 1935”, 15 s. (MNHUB); “Taja” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB);

“Gjallica” / “Albania leg. Bischoff 1936”, 20 s. (MNHUB); “Taja” / “Albania leg. Bischoff

1936”, 2 s. (MNHUB).

Notes. New species for the faunas of the Balkan Peninsula and Albania. Most likely, the

populations from the high mountains in North Albania are glacial relics.

66.2. Cymindis axillaris (Fabricius, 1794)

66.2.1. Cymindis axillaris axillaris (Fabricius, 1794)

References. Cymindis axillaris: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m);

Klaptocz, 1910: 399 (Oroshi); Hieke et al., 1986: 190 (North Albanian Alps near Theth).

Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Maja e

Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Mali i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Koritnik”

/ “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Maja e Policanit” / “Albania 1936 leg. Ramadan

coll. Bischoff ”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Boge” / “Albania Mueller leg. 1939”, 1 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania

Mueller leg. 1939”, 9 s. (MNHUB); “Iballe” / “Albania Mueller leg. 1939”, 1 s. (MNHUB).

Notes. Kabak (2003: 411) has omitted the species for Albania. Here it is confirmed for the

country.

66.2.2. Cymindis axillaris adriatica J. Müller, 1921

References. Cymindis axillaris adriatica: Müller, 1921a: 139 (Shkodër).

66.3. Cymindis coadunata Dejean, 1825

66.3.1. Cymindis coadunata coadunata Dejean, 1825

Material studied. “Çafa Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercoë”

/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 6

s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB).


66.4. Cymindis humeralis (Geoffroy, 1785)

References. Cymindis humeralis: Csiki, 1940: 222 (Shkëlzen Mt.); Hieke & Wrase, 1988: 161.


Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s.

(NMW); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Alban. Exp.’18

Pashtrik, 4-14.7.”, 1 s. (NMW); 68 s. from different localities (MNHUB).


country.

66.5. Cymindis lineata (Quensel, 1806)

References. Cymindis lineata: Apfelbeck, 1904: 343 (Vlorë); Apfelbeck, 1907a: 499 (Mal i Shenjt

Mt.: top region, 1500 m); Hieke & Wrase, 1988: 162; Kabak, 2003: 413.

References. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Rapsa 10-18.V.”

/ “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Alban. Exped. Kula Ljums 18.-

28.V.1918”, 2 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 3 s. (NMW); “Mal i Dajtit Tirane”

/ “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 1 s.

(MNHUB); “Mal i Scheinjit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tomor” / “leg.

Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll.

Bischoff ”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Borsne”

/ “Albania leg. Bischoff 1940”, 1 s. (MNHUB); “AL (Korcë) Vlacisht n. Korcë, 800 m 4.V.1993,

leg. Heinz u. S. Peks (ex. coll H. Peks)”, 1 s. (MNHUB).

66.6. Cymindis lineola L. Dufour, 1820

References. Cymindis axillaris lineola: Apfelbeck, 1907a: 495 (Buenë River).

Material studied. “Latif Skutari”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 2 s. (HNHM).


country.

? Cymindis pindicola Apfelbeck, 1901

Material studied. “Neres” / “Apfelb. 1906 Albania or.”, 1 s. (HNHM).

Notes. Apfelbeck described this species from the mountains of Peristeri, Veluchi and Oxya

situated in Northwestern and Central Greece. The specimen from “Neres” has been found to

be conspecific with a single syntype of C. pindicola labeled “Veluchi Gb. Stralcia” / “Coll.

Apfelbeck” (HNHM). However, I failed to find a place named “Neres” in Albania or in the

adjacent regions. Because of uncertain localization, this record remains questionable for Albania.

66.7. Cymindis scapularis Schaum, 1857

66.7.1. Cymindis scapularis scapularis Schaum, 1857

References. Cymindis scapularis var.: Apfelbeck, 1909: 298 (Vlorë-Fieri).

Notes. Kabak (2003: 411) has omitted the species for Albania. Actually, it needs further


Subgenus Menas Motschulsky, 1864



66.8. Cymindis miliaris (Fabricius, 1801)

Material studied. “Albania Rapsa Dr. Grabowski”, 3 s. (HNHM).


Subgenus Tarulus Bedel, 1906

66.9. Cymindis vaporariorum (Linnaeus, 1758)

References. Cymindis vaporariorum: Hieke & Wrase, 1988: 162.


(NMW); “Korab” / “Albania leg. Bischoff 1935”, 20 s. (MNHUB); “Maja e Jezercoë” / “Albania

leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB);

“Korab” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).



Subtribe DROMIINA Bonelli, 1810

67. Genus Dromius Bonelli, 1810


Subgenus Dromius s.str.


67.1. Dromius quadrimaculatus (Linnaeus, 1758)

Material studied. “Elbasan, Alb. 19.IX.18. Dr. H. Priesner”, 1 s. (NMW).


68. Genus Microlestes Schmidt-Göbel, 1846

68.1. Microlestes corticalis (L. Dufour, 1820) [= escorialensis (Brisout de Barneville, 1885)]

References. Microlestes plagiatus escorialensis: Holdhaus, 1904: 335 (Velipoja; Vlorë). Microlestes

corticalis escorialensis: Schatzmayr, 1943b: 112 (Lushnje). Microlestes corticalis: Hieke & Wrase, 1988:

160; Kabak, 2003: 422.

Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB).

68.2. Microlestes fissuralis (Reitter, 1901)

References. Microlestes fissuralis: Holdhaus, 1904: 332 (Velipoja; “Ardenica” near Vlorë); Csiki,

1940: 221 (Gjalicë Mt.); Horion, 1941: 343; Hieke et al., 1986: 190 (Kula e Lumës near Kukës);

Hieke & Wrase, 1988: 159; Kabak, 2003: 422.


Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica”

/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Kara Alibey” / “Albania leg. Bischoff 1936”,

4 s. (MNHUB); Librazhd Prenjas, 750 m, 7.V.1995, 1 s., L. Penev leg. (NMNHS).

68.3. Microlestes luctuosus Holdhaus, 1904

68.3.1. Microlestes luctuosus luctuosus Holdhaus, 1904

References. Microlestes exilis luctuosus: Holdhaus, 1904: 331 (Berat). Microlestes luctuosus: Hieke &

Wrase, 1988: 159. Microlestes apterus: Wrase, 1991: 18.

Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Kjori” / “Albania leg.

Bischoff 1932”, 2 s. (MNHUB); “Murrsi - Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s.

(MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

Notes. According to Wrase (1991) the material cited as M. exilis Schmidt-Göbel, 1846 and M.

exilis luctuosus (Apfelbeck, 1904: 330-331) may concern either M. apterus or M. luctuosus. I refer

it provisionally to the latter species, which was proved to inhabit the country. Kabak (2003:

423) has omitted or M. luctuosus for Albania. Here, the species is confirmed for the country.

68.4. Microlestes maurus (Sturm, 1827)

68.4.1. Microlestes maurus maurus (Sturm, 1827)

References. Microlestes maurus: Holdhaus, 1904: 332 (Velipoja; Vlorë); Hieke & Wrase, 1988:

159; Kabak, 2003: 423.

68.5. Microlestes minutulus (Goeze, 1777)

References. Blechrus minutulus: Apfelbeck, 1909: 298 (Vlorë; Berat).

Material studied. “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Mal i Krujes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg.



country.

68.6. Microlestes negrita (Wollaston, 1854)

68.6.1. Microlestes negrita apfelbecki Holdhaus, 1904

References. Microlestes minutulus apfelbecki: Holdhaus, 1904: 330 (Velipoja). Microlestes negrita:


Material studied. “Cafa Liqenit Iballe” / “Albania leg. Mueller 1939”, 1 s. (MNHUB).


country.

68.7. Microlestes plagiatus (Duftschmid, 1812)

References. Microlestes plagiatus: Apfelbeck, 1907a: 495 (Buenë River); Csiki, 1940: 221 (Korab Mt.).

Notes. Kabak (2003: 423) has omitted the species for Albania. Actually it needs further



69. Genus Paradromius Fowler, 1887

Subgenus Manodromius Reitter, 1905

69.1. Paradromius linearis (Olivier, 1795)

69.1.1. Paradromius linearis linearis (Olivier, 1795)

References. Dromius linearis: Hieke & Wrase, 1988: 155.

Material studied. “Vorra, Alb. 25.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Bushek juni” /

“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Durres juni” / “Albania leg. Bischoff 1932”, 2

s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranës” /

“leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Lum i Tiranës” / “Albania leg. Bischoff 1936”,

3 s. (MNHUB); Tiranë Park, 8.II.1998, 2 s., R. Ward leg. (CMNH).

Notes. Kabak (2003: 424) has omitted the species for Albania. Here P. linearis is confirmed for


70. Genus Philorhizus Hope, 1838

subgenus Philorhizus s.str.

70.1. Philorhizus notatus (Stephens, 1827)

References. Dromius notatus: Hieke & Wrase, 1988: 156.

Material studied. “Oboti Bojana” / “Coll. Apfelbeck”, 1 s. (HNHM); “Tomor” / “Albania




70.2. Philorhizus sigma (P. Rossi, 1790)

References. Dromius sigma: Apfelbeck, 1904: 337 (Velipoja); Hieke & Wrase, 1988: 157; Kabak,

2003: 426.

Subtribe LEBIINA Bonelli, 1810

71. Genus Lebia Latreille, 1802

Subgenus Lamprias Bonelli, 1810


71.1. Lebia cyanocephala (Linnaeus, 1758)

71.1.1. Lebia cyanocephala cyanocephala (Linnaeus, 1758)

Material studied. “Skutari Drin”, 1 s. (HNHM); “Tirane” / “leg. Bischoff Albania 1934”, 1 s.

(MNHUB).


Notes. The mention of Hieke & Wrase (1988: 153) for Albania is apparently result of a

following the record “Prisren” (Apfelbeck, 1904: 323). However, this locality is situated in the

Kosovo Region of Serbia. Kabak (2003: 427) has most likely followed one of these sources,

too. Since there is no previous reliable record from Albania, L. cyanocephala is treated here as

new species for the fauna of Albania.

Subgenus Lebia s.str.

71.2. Lebia cruxminor (Linnaeus, 1758)

71.2.1. Lebia cruxminor cruxminor (Linnaeus, 1758)

References. Lebia cruxminor: Hieke et al., 1986: 190 (North Albanian Alps near Theth; Kula e

Lumës near Kukës); Hieke & Wrase, 1988: 153.

Material studied. “Skutari Drin”, 1 s. (HNHM); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”,

1 s. (NMW); “Cucale Geb.” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Kjori” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albanien-Exp.

DEI Nordalban. Thethi, 1.-4.VIII.61”, 1 s. (MNHUB).

Notes. Kabak (2003: 428) has omitted the species for Albania. Here L. cruxminor is confirmed

for the country.

71.3. Lebia humeralis Dejean, 1825

References. Lebia humeralis: Hieke & Wrase, 1988: 154.

Material studied. “Albania Djakova” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Alban.

Exped. Gjalica Ljums 17-26.VI.’18”, 1 s. (NMW); “Lum i Tiranës” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).



71.4. Lebia scapularis (Geoffroy, 1785)

71.4.1. Lebia scapularis scapularis (Geoffroy, 1785)

References. Lebia scapularis: Hieke & Wrase, 1988: 154.

Material studied. “Serandë Piqeros” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

Notes. Kabak (2003: 429) has omitted the species for Albania. Here L. scapularis is confirmed

for the country with a first detailed record.

71.5. Lebia trimaculata (Villers, 1789)

Material studied. 12 km NE Erseka, 900 m, 12.V.1995, 2 s., L. Penev leg. (NMNHS).



Subtribe LIONYCHINA Jeannel, 1948

72. Genus Apristus Chaudoir, 1846

72.1. Apristus reticulatus Schaum, 1857

Material studied. “Mesi Kiri-Ufer” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).


72.2. Apristus subaeneus Chaudoir, 1846

References. Apristus subaeneus: Apfelbeck, 1907a: 495 (Liqen i Shkodrës); Müller, 1921a: 139;

Müller, 1924: 139; Hieke & Wrase, 1988: 161.

Material studied. “Skutari Drin”, 1 s. (NMW); “Skutari Vraka”, 6 s. (HNHM); “Skutari Vraka” /

“Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW).

Notes. Kabak (2003: 431) has omitted the species for Albania. Here it is confirmed for the country.

73. Lionychus Wissmann, 1846


Subgenus Lionychus s.str.


73.1. Lionychus maritimus Fairmaire, 1862

Material studied. “Himara juni” / “Albania leg. Bischoff 1932”, 17 s. (MNHUB).

Notes. New species for the faunas of the Balkan Peninsula and Albania.

73.2. Lionychus quadrillum (Duftschmid, 1812)

Material studied. “Medua, Alb. 10.5.914”, 1 s. (HNHM).


74. Genus Syntomus Hope, 1838

74.1. Syntomus impressus (Dejean, 1825)

74.1.1. Syntomus impressus impressus (Dejean, 1825)

References. Metabletus impressus: Apfelbeck, 1907a: 495 (Buenë River). Syntomus impressum sic!:


Material studied. “Albanien Avlona v. Oertzen.”, 2 s. (MNHUB); Saranda District, Ljukove,

3-4.VI.1994, 5 s., G. Blagoev leg. (NMNHS).


country.


74.2. Syntomus obscuroguttatus (Duftschmid, 1812)

References. Metabletus obscuroguttatus: Apfelbeck, 1904: 327 (Vlorë; Velipoja); Apfelbeck, 1909:

298 (Vlorë). Syntomus obscuroguttatus: Hieke & Wrase, 1988: 157; Kabak, 2003: 433.

Material studied. “Oboti Bojana”, 4 s. (HNHM); “Skutari Bojana”, 1 s. (HNHM); “Logara”

/ “leg. Bischoff Albania 1934”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg.

(NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Radona,

Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS); Tiranë Park, 15.II.1998, 320 s., from

grass and soil around tree trunks, R. Ward leg. (CMNH).


m., D. Antonelli leg. (cAVT).

74.3. Syntomus pallipes (Dejean, 1825)

References. Metabletus pallipes: Apfelbeck, 1909: 298 (Vlorë). Syntomus pallipes: Hieke & Wrase,

1988: 157.

Material studied. “Okol Shkalla” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).


74.4. Syntomus truncatellus (Linnaeus, 1761)

References. Metabletus truncatellus: Csiki, 1940: 221 (Korab Mt.). Syntomus truncatellus: Hieke &

Wrase, 1988: 158.

Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 3 s. (HNHM); “Albania

Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Korab” / “Albania leg. Bischoff

1935”, 3 s. (MNHUB).


Tribe LICININI Bonelli, 1810

Subtribe LICININA Bonelli, 1810

75. Genus Badister Clairville, 1806

Subgenus Badister s.str.

75.1. Badister bullatus (Schrank, 1798)

Material studied. “Scutari Bojana”, 1 s. (HNHM); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”,

1 s. (NMW).


75.2. Badister unipustulatus Bonelli, 1813

References. Badister unipustulatus: Apfelbeck, 1904: 163 (Velipoja); Murraj, 1968: 33 (Tiranë);

Hieke & Wrase, 1988: 151; Baehr, 2003c: 440.


Material studied. “Oboti Bojana”, 2 s. (HNHM).

Notes. The record of Murraj (1968) is included here with reservation due to misidentifications

found in his collection.

Subgenus Baudia Ragusa, 1884

75.3. Badister dilatatus Chaudoir, 1837

References. Badister dilatatus: Apfelbeck, 1904: 164 (Velipoja); Hieke & Wrase, 1988: 151;

Baehr, 2003c: 441.

75.4. Badister peltatus (Panzer, 1796)

75.4.1. Badister peltatus peltatus (Panzer, 1796)

References. Badister peltatus peltatus: Baehr, 2003c: 441.

76. Genus Licinus Latreille, 1802

Subgenus Licinus s.str.

76.1. Licinus cassideus (Fabricius, 1792)

76.1.1. Licinus cassideus cassideus (Fabricius, 1792) [= latus Dejean, 1831]

References. Licinus cassideus latus: Magistretti, 1965: 263; Hieke & Wrase, 1988: 150. Licinus cassideus

cassideus: Baehr, 2003c: 442.

Material studied. “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” /

“Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Tomor-Abas Ali juli 1932” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Tomor-Tojk juli 1932” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB);

“Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 4

s. (MNHUB); “Boge Aufstieg z. Rab” / “Albania leg. Mueller 1939”, 1 s. (MNHUB).

Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelenë District),

18.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Notes. Csiki (1931: 911-912) treated dohrni, graecus and submarginatus as synonyms of cassideus.

Here, I follow the arrangement of Baehr (2003c: 442). In spite of that, a revision of the

Balkan taxa close to L. cassideus is needed. First detailed records for Albania.

76.1.2. Licinus cassideus dohrni Fairmaire, 1863

References. Licinus cassideus dohrhi: Fairmaire, 1863: 131 (Albania, loc. typ.); Reitter, 1900: 150;

Apfelbeck, 1904: 162; Baehr, 2003c: 442.

Material studied. “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB).

76.2. Licinus graecus Apfelbeck, 1901 [= submarginatus J.R. Sahlberg, 1903]

References. Licinus graecus: Apfelbeck, 1901a: 431 (Berat); Apfelbeck, 1909: 298 (Berat). Licinus

cassideus graecus: Apfelbeck, 1904: 162 (Berat). Licinus submarginatus: Schatzmayr, 1943b: 100

(Tërpan); Baehr, 2003c: 442.


Notes. Apfelbeck published two descriptions of L. graecus. First he described L. graecus

(Apfelbeck, 1901a: 431), and after that L. cassideus graecus (Apfelbeck, 1904: 162). Incorrectly

the second taxon has been accredited to be the valid, which create a future complication.

In 1904, Apfelbeck (1904: 399, 402 note 1) synonymized L. submarginatus J.R. Sahlberg,

1903 with L. cassideus graecus Apfelbeck, 1904, not with L. graecus Apfelbeck, 1901. The

only explanation of this fact is that this outstanding specialist seemed to have forgotten

his earlier description. Later, the subsequent authors (Schatzmayr, 1943b; Baehr, 2003c)

followed this view, but treated L. submarginatus as valid species, and L. cassideus graecus

Apfelbeck, 1904 – for its synonym. I believe, however, that both Licinus - species described

by Apfelbeck are identical based on identical type localities. According to the rules of

ISZN (1999) an original description of a taxon is the oldest one. Therefore, the following

nomenclatorial act is proposed: Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn.

nov. of Licinus (Licinus) graecus Apfelbeck, 1901.

76.3. Licinus silphoides (P. Rossi, 1790)

References. Licinus silphoides: Apfelbeck, 1909: 298 (Vlorë-Fieri); Schatzmayr, 1943b: 100 (Karbunara

e Madhe; Stan-Karbunarë); Focarile, 1959: 76; Hieke & Wrase, 1988: 150; Baehr, 2003c: 442.

Subgenus Neorescius Bedel, 1906

76.4. Licinus oertzeni Reitter, 1889

76.4.1. Licinus oertzeni oertzeni Reitter, 1889

References. Licinus oertzeni: Apfelbeck, 1907a: 499, 501,503 (Mal i Shenjt Mt.: top region, 1500 m).

76.4.2. Licinus oertzeni albanicus J. Müller, 1932

References. Licinus oertzeni albanicus: Müller, 1932: 209 (Tomor Mt. near Berat, loc. typ.;

Northwestern Greece, “Buč-Gebirge”, loc. typ.); Baehr, 2003c: 443.

Material studied. “Merdita Orochi”, 1 s. (HNHM); “Maja e Tartarit” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor” /

“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Çafa Kjores” / “Albania leg. Bischoff 1935” /

“Licin. Oertzeni albanicus m det. J. Müller”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shen

Ili” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shkalla Harushe” / “Korab” / “Albania


Notes. The distinction of the two subspecies of L. oertzeni s.l. is very easy due to their clear

external differences. Probably both taxa deserve species status.

Tribe ODACANTHINI Laporte, 1834

Notes. New tribe for the fauna of Albania.

77. Genus Odacantha Paykull, 1798



Subgenus Odacanta s.str.


77.1. Odacantha melanura (Linnaeus, 1767)

Material studied. “Skutari Bojana” / “Coll. Apfelbeck”, 1 s. (HNHM).


Tribe OODINI LaFerté-Sénectère, 1851

78. Genus Oodes Bonelli, 1810

78.1. Oodes gracilis A. Villa & G.B. Villa, 1833

References. Oodes gracilis: Apfelbeck, 1904: 159 (Velipoja); Focarile, 1959: 75; Hieke & Wrase,

1988: 150; Bousquet, 2003c: 445.

Material studied. Shkodër District, Mali Tarabosh Mt., 100 m, 24.XI.2000, 3 m., limestone

terrain, A. Zhalov leg. (NMNHS).

78.2. Oodes helopioides (Fabricius, 1792)

78.2.1. Oodes helopioides helopioides (Fabricius, 1792)

References. Oodes helopioides: Apfelbeck, 1904: 159 (Velipoja near Lezhë); Hieke & Wrase,

1988: 150; Bousquet, 2003c: 445.

Tribe OMPHREINI Ganglbauer, 1891

79. Genus Omphreus Dejean, 1828

Notes. This genus needs revision. The author’s examinations revealed only a small number of

specimens of the most species and these in bad condition (excluding O. morio s.l.) in the largest

Central European collections (DEI; HNHM; NMW). Contrary, in MNHUB the collection of

Omphreus is better represented.

Subgenus Omphreus s.str.

79.1. Omphreus lonai Winkler, 1933

References. Omphreus lonai: Winkler, 1933b: 118 (Llogora, 1200 m, southeast of Vlorë, loc.

typ.); Bousquet, 2003d: 446.

Material studied. “Llogora” / “Albania leg. Bischoff 1935”, 1 m. (MNHUB); “Llogora” /

“Albania leg. Bischoff 1936”, 3 f. (MNHUB).


79.2. Omphreus morio Dejean, 1828

79.2.1. Omphreus morio albanicus Apfelbeck, 1906

References. Omphreus albanicus: Apfelbeck, 1906: 1666-1667 (Merdita: Mal i Shenjt Mt., loc.

typ.; Zepë Mt., loc. typ.); Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck,

1918c: 160, 165 (Mal i Shenjt Mt.; Zepë Mt.; North Albanian Alps, 1300-2000 m); Hieke,

1981: 82. Omphreus morio: Anonymous, 1907: 286. Omphreus morio albanicus: Winkler, 1933b: 117

(Mal i Dajtit Mt.; Mal i Shenjt Mt.; Zepë Mt.; Gjalicë Mt.; Peshtrik Mt.); Drovenik & Peks,

1994: 75 (Peshtrik Mt.); Drovenik & Peks, 1999: 87 (Peshtrik Mt.).

Material studied. “Merdita M. Scheit” / “Coll. Apfelbeck”, 4 s. (HNHM); “Merdita M. Scheit”,

3 s. (NMW); “Merdita Zebia”, 6 s. (HNHM; NMW; MNHUB); “Merdita Munela”, 1 s. (HNHM);

“Oroši (Lat.) Alban. spt.” / “Zebja”, 2 s. (DEI; MNHUB); “Merdita Munela” / “Wgth., 1906

Merdita”, 1 s. (NMW); “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Alban.

Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Alban. Exped. Gjalica Ljums 17.-26.VI’18”,

2 s. (NMW); “Mal i Dajtit juni” / “Albania leg. Bischoff 1932”, 2 m., 1 f. (MNHUB); “Mal i

Krujes” / “leg. Bischoff Albania 1934”, 1 m. (MNHUB); “Gjallica” / “Albania leg. Bischoff

1936”, 1 m. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 m. (MNHUB); “Mal

i Krujes” / “Albania leg. Bischoff 1939”, 1 f. (MNHUB); Reshen District, near Merkurth,

920-980 m, 11.VI.1993, 1 s., grassland, P. Beron & B. Petrov leg. (NMNHS); Mal i Dajtit Mt.,

20 km east of Tiranë, 1000 m, 9.V.1995, 3 s., L. Penev leg. (NMNHS).

Notes. Bousquet (2003d: 446) has omitted O. morio for Albania. Here the species is confirmed

for the country.

79.2.2. Omphreus morio malissorum Winkler, 1933

References. Omphreus morio malissorum: Winkler, 1933b: 117 (North Albanian Alps: Maja e

Jezercë, loc. typ.; Starca; Çukall Mt.).

Material studied. “Maja e Jezerce Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 1

syntype (NMW); “Maja e Jezercoë” / “Albania leg. Bischoff 1931”, 1 syntype (MNHUB);

“Cukale Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 1 syntype (NMW); “Cucale

Geb.” / “Albania leg. Bischoff 1931”, 3 syntypes (MNHUB); “Maja e Jezercoë” / “Albania


“Cucale” / “Albania leg. Bischoff 1936”, 9 s. (MNHUB); “M. Mulecifit” / “Albania leg. Bischoff

1936”, 3 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maranaj”

/ “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Maja e Dragomirit” / “Albania Mueller leg.

1939”, 3 s. (MNHUB); “M. Raba Massiv” / “Albania Mueller leg. 1939”, 6 s. (MNHUB);

North Albanian Alps, 1400-1800 m, 1.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS).

79.3. Omphreus ravasinii J. Müller, 1923

References. Omphreus ravasinii: Müller, 1923: 111-112 (Tomor Mt., alpine zone, loc. typ.);

Winkler, 1933b: 118 (Tomor Mt.); Bousquet, 2003d: 446.

Material studied. “Tomor-Abas Ali juli 1932” / “Albania leg. Bischoff 1932”, 1 f. (MNHUB).

! Omphreus ravasinii ssp.

Material studied. “Korab” / “Albania leg. Bischoff 1936”, 1 f. (MNHUB).

Notes. It seems this specimen belong to a vicarious form close to O. ravasinii. Unfortunately

its abdomen is destroyed and further material is needed to specify its right status.


Tribe PANAGAEINI Bonelli, 1810

80. Genus Panagaeus Latreille, 1802

Subgenus Panagaeus s.str.

80.1. Panagaeus cruxmajor (Linnaeus, 1758) [= schaumi Ganglbauer, 1892]

References. Panagaeus cruxmajor ab. schaumi: Apfelbeck, 1904: 153 (Velipoja near Lezhë). Panagaeus

cruxmajor: Hieke & Wrase, 1988: 152; Baehr, 2003d: 447.

Material studied. “Scutari Drin”, 3 s. (HNHM); “Oboti Bojana”, 2 s. (HNHM).

Tribe PLATYNINI Bonelli, 1810

81. Genus Agonum Bonelli, 1810

Subgenus Agonum s.str.

81.1. Agonum afrum (Duftschmid, 1812)

Material studied. Tiranë Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 7 s., shaded lake edge

under leaves and wet, rotten wood, R. Ward leg. (CMNH).


81.2. Agonum carbonarium Dejean, 1828

81.2.1. Agonum carbonarium carbonarium Dejean, 1828 [= alpestre (Heer, 1838)]

References. Agonum alpestre: Schmidt, 1995: 168 (Korab Mt.).

Notes. Bousquet (2003e: 448 sub A. alpestre) has omitted A. carbonarium for Albania. The

species occurrence in the central area of the Balkans is interesting from the biogeographic

point of view since the Albanian locality is distant from the main Western European range of

the species, as well as from its Asian shelters.

81.3. Agonum duftschmidi J. Schmidt, 1994 [= moestum (Duftschmid, 1812)]

References. Platynus viduus var. moestus: Apfelbeck, 1904: 292 (Velipoja). Platynus viduus moestus:

Apfelbeck, 1907a: 495 (Liqen i Shkodrës). Platynus moestus: Apfelbeck, 1909: 297 (Vlorë). Agonum

moestum: Hieke & Wrase, 1988: 86. Agonum duftschmidi: Bousquet, 2003e: 450.

81.4. Agonum extensum Ménétriés, 1849 [= binaghii Burlini, 1939]

References. Agonum monachum var. binaghii: Burlini, 1939: 60 (Vlorë). Agonum extensum: Schatzmayr,

1943b: 111 (Skrofotina).

Material studied. “Apfelb. Avlona”, 1 s. (HNHM); “Myzeqe Ebene” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB).

Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for the country.


81.5. Agonum gisellae Csiki, 1931 [= angustatum Dejean, 1828]

References. Agonum angustatum: Hieke & Wrase, 1988: 86; Schmidt, 1994: 27 (Buenë River

near Shkodër). Agonum gisellae: Bousquet, 2003e: 450.

81.6. Agonum hypocrita (Apfelbeck, 1904)

References. Agonum hypocrita: Csiki, 1940: 218 (Korab Mt.).

Material studied. “Scutari l. See-Ufer”, 3 s. (HNHM); “Murrsi-Kara Alibey” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Bushnesh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);

“Mamurash” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB).

Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for

the country.

81.7. Agonum longicorne Chaudoir, 1846 [= holdhausi (Apfelbeck, 1904)]

References. Platynus holdhausi: Apfelbeck, 1904: 294 (Velipoja); Apfelbeck, 1907a: 495 (Buenë

River). Agonum holdhausi: Hieke & Wrase, 1988: 87. Agonum longicorne: Schmidt, 1994: 32 (Buenë

River near Oboti); Bousquet, 2003e: 450.

Material studied. “Latif Pulaj”, 2 s. (HNHM); “Oboti Wald” / “Apfelb. Alban. spt. IV.V.’05”,

1 paralectotype of Platynus holdhausi (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1938”,

1 s. (MNHUB).

Notes. In NMW I found a paralectotype not listed by Schmidt (1994: 31) among the type

material of A. longicorne, though the author included the record of this material (Oboti) in his

list of species distribution.

81.8. Agonum marginatum (Linnaeus, 1758)

References. Agonum marginatum: Hieke & Wrase, 1988: 85.

Material studied. “Skutari Vraka” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Maja e Konispol

Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Valona” / “Albania leg. Bischoff

1936”, 2 s. (MNHUB); Tiranë Park, 8.II.1998, shaded lake edge under leaves and wet, rotten

wood and 22.II.1998, from grasses along small sluggish stream, 22 s., R. Ward leg. (CMNH).


and Gjirokastër), 350 m, 23.IV.2003, 1 m., P. Crucitti & C. Cavalieri leg. (cAVT).



81.9. Agonum muelleri (Herbst, 1784)

Material studied. “Skutari l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Alban.

Vermosa Penther”, 3 s. (NMW).

Notes. The mention of Hieke & Wrase (1988: 85) for Albania is apparently a result of following

the record “Prisren” (Apfelbeck, 1904: 290 sub Platynus muelleri). However, currently this locality


is situated in the Kosovo Region of Serbia. Bousquet (2003e: 451) has most likely followed

one of these sources, too. Since there is no previous reliable record from Albania, A. muelleri

is treated here as new species for the fauna of Albania.

81.10. Agonum permoestum Puel, 1938

References. Agonum permoestum: Puel, 1938: 194 (Velipoja); Schmidt, 1994: 25 (Buenë River

near Shkodër; Bushnesh; Mamurash; Murrsi-Kara Alibey; Buenë River near Oboti; Rushkull;

Liqen i Shkodrës; Sukthi near Gurse); Bousquet, 2003e: 451.

Material studied. “Scutari l. See-Ufer”, 1 s. (HNHM); “Skutari l. See-Ufer” / “Apfelb. Alban.

spt. IV.V.’05”, 8 s. (NMW); “Oboti Bojana”, 1 s. (HNHM).

81.11. Agonum sexpunctatum (Linnaeus, 1758)

References. Agonum sexpunctatum: Csiki, 1940: 218 (Korab Mt.); Hieke et al., 1986: 189 (Lurë

e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 84.

Material studied. “Alban. Vermosa Penther”, 1 s. (NMW); “Ricavac 1300 m, 25.VI.-2.VII.” /

“Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Albania Tirana” / “3.VI.1958 leg. Xh.

Murraj”, 3 s. (HNHM).


the country.

81.12. Agonum sordidum Dejean, 1828

81.12.1. Agonum sordidum sordidum Dejean, 1828

References. Platynus sordidus: Apfelbeck, 1904: 290 (Velipoja; Vlorë); Apfelbeck, 1909: 297

(Vlorë). Agonum sordidum: Schatzmayr, 1943b: 111 (Stan-Karbunarë); Hieke et al., 1986: 189

(Poliçan near Tomor Mt.); Hieke & Wrase, 1988: 85.

Material studied. “Elbasan, Alb. IX… Dr. H. Priesner”, 1 s. (NMW); “Liquen i Butrintes

Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff

1935”, 2 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis

Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB).


81.13. Agonum viridicupreum (Goeze, 1777)

80.13.1. Agonum viridicupreum viridicupreum (Goeze, 1777)

References. Platynus viridicupreus: Apfelbeck, 1904: 290 (Velipoja). Agonum viridicupreum: Hieke

et al., 1986: 189 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 85.

Material studied. “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);

“Tiranë” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Liquen i Butrintes Vivari” / “Albania

leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s.

(MNHUB); “Han-Hot” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Sukthi bis Gurse” /

“Albania leg. Bischoff 1938”, 1 s. (MNHUB); “Cafa Liqenit Iballe” / “Albania Mueller leg.

1939”, 5 s. (MNHUB); Tiranë Park, 11.I.1998 / 1.III.1998 / 8.III.1998, 4 s., shaded lake edge




the country.

Subgenus Europhilus Chaudoir, 1859

81.14. Agonum antennarium (Duftschmid, 1812)

References. Platynus antennarius: Apfelbeck, 1904: 295 (Velipoja). Agonum antennarium: Hieke

& Wrase, 1988: 87.

Material studied. “Mustajbeg Velipoja Albania”, 2 s. (MNHUB); “Maja Rogamit” / “Albania

leg. Bischoff 1935”, 11 s. (MNHUB); Tiranë Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 8 s.,

shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH).


81.15. Agonum thoreyi Dejean, 1828

81.15.1. Agonum thoreyi thoreyi Dejean, 1828

References. Platynus thoreyi: Apfelbeck, 1904: 296 (Velipoja). Agonum thoreyi: Hieke & Wrase,

1988: 87; Bousquet, 2003e: 453.

82. Genus Anchomenus Bonelli, 1810

Subgenus Anchomenus s.str.

82.1. Anchomenus dorsalis (Pontoppidan, 1763)

References. Platynus dorsalis: Apfelbeck, 1904 (Vlorë; Berat; Velipoja); Apfelbeck, 1909: 297

(Vlorë; Berat); Hieke & Wrase, 1988: 83. Platynus dorsale: Schatzmayr, 1943b: 111 (Lushnje);

Hieke et al., 1986: 189 (North Albanian Alps near Theth).

Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Alban. Vermos Penther”,

3 s. (NMW); “Skutari Kolcekai” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW);

“Bushek bei Elbasan” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Bushek juni” / “Albania

leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Lum i Tiranes” / “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Liquen i Butrintes Vivari” /

“leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 12 s.

(MNHUB); “Murrsi-Kara Ali Bey” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB); “Piqeros”

/ “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”,

1 s. (MNHUB); “Am Wege von Murthi nach Konispol” / “Albania leg. Bischoff 1936”, 1 s.

(MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Cafa e Ceromis Iballe”

/ “Albania Mueller leg. 1939”, 2 s. (MNHUB); “Albanien Tirana Dajti-Vorberge

3.V.39…Mueller”, 1 s. (MNHUB); “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 5 s.

(HNHM); Sauk, 14.III.1961, 1 f. (MNST); Rhesen District, Merkurth, 11.VI.1993, 1 s., P.

Beron & B. Petrov leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 1 s., S. Golovach, B.

Petrov, P. Stoev leg. (NMNHS); Ljukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS).


Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., 1 f., P.


Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006,



the country.

83. Genus Olisthopus Dejean, 1828

83.1. Olisthopus fuscatus Dejean, 1828

References. Olisthopus fuscatus: Schatzmayr, 1943b: 111 (Lushnje); Hieke & Wrase, 1988: 88.

Material studied. “Alban. Vermosa Penther”, 2 s. (NMW); “Ibe Erzen” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); Saranda District, Lyukove, 5.V.1994, 1 s., P.

Stoev leg. (NMNHS); Saranda District, Lyukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS);

Tiranë, botanical garden, 8.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).


the country.

83.2. Olisthopus glabricollis (Germar, 1817)

References. Olisthopus glabricollis: Schatzmayr, 1943b: 111 (Lushnje); Hieke & Wrase, 1988: 88.

Material studied. “Alban. Exp. Kula Ljums 18-28.V.’18”, 4 s. (NMW); “Alban. Exp.’18. Kruma

5.VI.”, 4 s. (NMW); “Burelle” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene”



the country.

? Olisthopus rotundatus (Paykull, 1790)

? Olisthopus rotundatus rotundatus (Paykull, 1790)

References. Olisthopus rotundatus: Horion, 1941: 310; Hieke & Wrase, 1988: 88. Olisthopus

rotundatus rotundatus: Bousquet, 2003e: 461.

Notes. Horion (1941: 310) and Hieke & Wrase (1988) noted this species for Albania apparently

following the record “Prisren” (Apfelbeck, 1904: 287). Actually this locality is situated in the

Kosovo Region of Serbia. Most probably, Bousquet (2003e: 461) has followed the above

sources. For the time being P. rotundatus is regarded as questionable (possible but yet not

proven) for Albania.

84. Genus Paranchus Lindroth, 1974


84.1. Paranchus albipes (Fabricius, 1796)

Material studied. “Alban. Exp.’18. Kruma 5.VI.”, 1 s. (NMW).



85. Genus Platynus Bonelli, 1810

Subgenus Batenus Motschulsky, 1865

85.1. Platynus scrobiculatus (Fabricius, 1801)

85.1.1. Platynus scrobiculatus turcicus Apfelbeck, 1904

References. Platynus scrobiculatus turcicus: Schmidt, 1996: 347.

Material studied. “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze Penther’14”,

5 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 26 s. (MNHUB).

Notes. Apfelbeck (1904: 288) described P. scrobiculatus turcicus from Bosnia Herzegovina. In the

same year, Csiki (1904) described P. serbicus Csiki, 1904 from Southeast Serbia. Recently, Bousquet

(2003e: 463) treated both taxa as different species. However, judging by the description of these

taxa and based on research on different populations from the Eastern Balkans (Guéorguiev &

Muilwijk, 2001), I consider the diagnostic characters indicated by Apfelbeck and Csiki reflect

rather geographical variations than being of specific importance. On account of that, P. turcicus

is treated here as conspecific with P. scrobiculatus (Fabricius, 1801): Platynus (Batenus) scrobiculatus

turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus Apfelbeck, 1904. Recently, we

also treated Csiki’s taxon as subspecies of P. scrobiculatus, omitting formal change in rank

(Guéorguiev & Muilwijk, 2001). Here this act is introduced: Platynus (Batenus) scrobiculatus serbicus

Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904. The Albanian populations are

provisionally assigned to the ssp. turcicus. Bousquet (2003e: 463 sub Platynus turcicus) has omitted

the species for Albania. Here it is confirmed for the country with first detailed records.

Subgenus Platynus s.str.

85.2. Platynus assimilis (Paykull, 1790)

References. Platynus assimilis: Apfelbeck, 1904: 296 (Vlorë; Berat; Velipoja); Hieke & Wrase, 1988: 84.

Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 19 s.

(NMW); “Bushnesh” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Mamurash” / “Albania leg.

Bischoff 1935”, 5 s. (MNHUB); “Pylla Thumana” / “Albania leg. Bischoff 1935”, 10 s. (MNHUB).


Tribe PTEROSTICHINI Bonelli, 1810

86. Genus Abax Bonelli, 1810

Subgenus Abacopercus Ganglbauer, 1891

86.1. Abax carinatus (Duftschmid, 1812)

86.1.1. Abax carinatus carinatus (Duftschmid, 1812) [= latus (Dejean, 1828)]

References. Abax carinatus var. latus: Apfelbeck, 1904: 216 (Velipoja). Abax carinatus: Hieke &

Wrase, 1988: 79. Abax carinatus carinatus: Bousquet, 2003f: 469.

Material studied. “Velipoja Albania”, 1 s. (HNHM); 18 s. from different localities (MNHUB).


Subgenus Abax s.str.

86.2. Abax ovalis (Duftschmid, 1812)

References. Abax ovalis: Hieke & Wrase, 1988: 79; Bousquet, 2003f: 470.

Material studied. “Koritnik” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Shkëlzën” /



87. Genus Molops Bonelli, 1810

Notes. The species recognition in this group stay difficult due to the great external resemblance

of the adults that often resulted in impossibility to distinguish the females of closely related

species. Besides, a great similarity and variations in the shape of the aedeagus in close species

occur. The current systematics of the genus must not be treated as definitive (Mlynář, 1977).

Except the “spartanus” group, all the other groups of species of Molops s.l. display North

Aegean origin. The type material used by Müller (1917; 1918a) to describe five Albanian taxa

of Molops s.str. is still preserved in NMW; the same concerns the type material published by

Apfelbeck (1918c).

Subgenus Molops s.str.

Species group “robustus”

87.1. Molops malshentianus Apfelbeck, 1917 [= maderi J. Müller, 1933]

References. Molops malshetianus: Apfelbeck, 1917: 224 (Merdita: Mal i Shenjt Mt. near Oroshi,

loc. typ.); Apfelbeck, 1918b: 88-89 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Apfelbeck,

1930: 167-168 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Mlynář, 1977: 144 (Mal i

Shenjt Mt. near Oroshi). Molops maderi: Müller, 1933: 217 (Kruja Mt., 600 m, loc. typ.); Mařan,

1940c: 192 (Kruja Mt.); Mařan, 1945: 60 (Kruja Mt.); Mlynář, 1977: 39-40 (Kruja Mt.); Turin,

1981: 130; Bousquet, 2003f: 473.

Material studied. “Kruja, Alban., Mader”, 1 s. (NMW); “Ibe-Erzen” / “Albania leg. Bischoff

1932”, 1 f. (MNHUB); “Maja e Tartarit” / “leg. Bischoff Albania 1934”, 1 f. (MNHUB); “Mal

i Krujes” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); “Mal i Krujes” / “Albania leg.

Bischoff 1936”, 2 s. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 f., L.

Penev leg. (NMNHS).

Notes. Apfelbeck published three consecutive descriptions for M. malshentianus (Apfelbeck,

1917; Apfelbeck, 1918b; Apfelbeck, 1930), the second one has been accredited to be the valid.

Actually this is incorrect and the original description must be assigned to the oldest one.

According to Bousquet (2003f: 473) M. malshentianus is synonym of M. maderi, but since the

former has been described earlier, M. malshentianus must be treated as senior synonym: Molops

(Molops) maderi syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917. Notwithstanding,

revision of the types is needed to elaborate this association and to designate lectotypes. The

record from “Maja e Tartarit” should be treated with caution since it is based on a single

female specimen.


# Molops weiratheri J. Müller, 1930 [= alinae J. Müller, 1932]

References. Molops parnassicola alinae: Müller, 1932: 220 ((“Südalbanien (Epiros)”); Mařan,

1940c: 190, 192.

Notes. Mařan (1940c) specified that the Müller’s record for Albania (Müller, 1932) refers to

Greece. M. weiratheri inhabits the mountains in South Macedonia and North Greece reaching

in the south the massif of Olympus Mt. (Mlynář, 1977: 41-42).

Species group “spartanus”

87.2. Molops spartanus (Schaum, 1862)

? Molops spartanus parnassicola Kraatz, 1875

References. Molops cf. parnassicola: Müller, 1936: 105 (Gjirokastër).

Notes. As Müller (1936) has determinated his material only to species level, the subspecific

treatment of the population near Gjirokastër needs further specification.

87.2.1. Molops spartanus valonensis J. Müller, 1936

References. Molops spec.? parnassicola: Ganglbauer, 1889: 54 (Vlorë). Molops parnassicola:

Apfelbeck, 1904: 217, 220 (Vlorë); Apfelbeck, 1909: 297 (Kanina near Vlorë; Hagion Elias =

Keraun); Müller, 1918a: 7 (Vlorë). Molops parnassicola valonensis: Müller, 1936: 105 (Reza e Kanalit

Mt. near Llogora, loc. typ.; Maja e Tartarit Mt.; Shëndelli Mt.); Mařan, 1940c: 190, 192 (Llogora);

Mařan, 1945: 59 (Vlorë). Molops spartanus valonensis: Mlynář, 1977: 47-48 (Reza e Kanalit Mt.

near Llogora; Llogora; Maja ë Qori; Cafa ë Qori); Bousquet, 2003f: 475.

Material studied. “Albania coll. Lichtn.”, 2 s. (HNHM); “Llogora, Albania mer.”, 4 s.

(MNHUB); “Maj’e Tartarit Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);

“Reza e Kanalit Logara, Albania m.” / “Winkler, Lona, Bischoff, V.1931”, 5 s. (NMW); “Logara”

/ “Albania leg. Bischoff 1932”, 158 s. (MNHUB); “Maja e Qori juni 1932” / “Albania leg.

Bischoff 1932”, 10 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 36 s.

(MNHUB); “Mato-Hasanaj juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen-Ili

juni” / “Albania leg. Bischoff 1932”, 35 s. (MNHUB); “Logara” / “leg. Bischoff Albania

1934”, 48 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 30 s. (MNHUB); Ibë,

24.IV.1962, 1 m. (MNST).

Notes. Before the description of this subspecies, all records from South Albania had been

attributed to M. spartanus parnassicola Kraatz, 1875. Bousquet (2003f: 475) indicated an incorrect

page of the original description of ssp. valonensis.

Species group “simplex”

87.3. Molops simplex Chaudoir, 1868

# Molops simplex simplex Chaudoir, 1868

References. Molops simplex simplex: Bousquet, 2003f: 475.

Notes. Mlynář (1977: 54) noted that the older data for this subspecies from Albania concerns

M. simplex meridionalis, because the nominotypical subspecies inhabits regions removed from

the borders of the country.


87.3.1. Molops simplex meridionalis Mlynář, 1977

References. Molops simplex: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m);

Müller, 1918a: 7 (Maranajt Mt.; Rapshë). Molops simplex meridionalis: Mlynář, 1977: 55; Bousquet,

2003f: 474.

Notes: The subspecies occurs only in the marginal northwestern part of the country.

Species group “alpestris”

87.4. Molops alpestris (Dejean, 1828)

87.4.1. Molops alpestris imitator J. Müller, 1917

References. Molops alpestris: Apfelbeck, 1907a: 497-498 (Maranajt Mt.: top region, 1500-1576 m);

Hieke & Wrase, 1988: 81. Molops alpestris imitator: Müller, 1917: 347-348 (Rapshë, loc. typ.); Müller,

1918a: 11 (North Albanian Alps: Rapshë); Mlynář, 1977: 80-81 (Rapshë); Hieke, 1981: 81.

Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 3 syntypes

(NMW); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 3 s., P. Beron & B. Petrov leg.

(NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 8 s., P. Beron & B. Petrov

leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron & B.


Notes. Bousquet (2003f: 472) has omitted the species for Albania. Here, it is confirmed for

the country.

Species group “elatus”

87.5. Molops parreyssi Kraatz, 1875

87.5.1. Molops parreyssi lonae J. Müller, 1921

References. Molops parreyssi: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m);

Klaptocz, 1910: 399 (Theth); Müller, 1918a: 7 (Maranajt Mt.). Molops parreyssi lonae: Müller,

1921b: 28-30 (Çukall Mt., loc. typ.); Mlynář, 1977: 96 (Çukall Mt.: cave “Kunor Vukai”);

Bousquet, 2003f: 473.

Material studied. “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Cucale

Albania sept.” / “leg. Bischoff 1931”, 4 s. (NMW); nearly 100 s. from locality “Cucale”

(MNHUB); 4 s. from locality “Maranay” (MNHUB).

Notes. The species occurs only in the marginal northwestern parts of Albania.


87.6. Molops merditanus Apfelbeck, 1906

87.6.1. Molops merditanus merditanus Apfelbeck, 1906

References. Molops merditanus: Apfelbeck, 1906: 1664 (Merdita: Mal i Shenjt Mt. near Oroshi,

loc. typ.; Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1907a: 499, 503 (Mal i Shenjt Mt.:

top region, 1500 m); Müller, 1918a: 10 (Merdita: Mal i Shenjt Mt. near Oroshi; Merdita: Zepë

Mt. near Fandi); Müller, 1930 (Merdita). Molops merditanus merditanus: Mlynář, 1977: 106-107

(Mal i Shenjt Mt. near Oroshi; Zepë Mt.); Bousquet, 2003f: 473.


Material studied. “Merdita M. Scheit”, 1 paralectotype (DEI); “Merdita M. Scheit” / “Apfelb.

Alban. spt. IV.V.’05” / “Molops merditanus typ. † Apfelb.”, 1 f. (NMW); “Merdita Zebia”, 2

paralectotypes (DEI); “Merdita Zebia” / “Molops merditanus typ. ‡ Apfelb.”, 1 f. (NMW).

Notes. Mlynář (1977: 106) noted that the lectotype of M. merditanus is in the collection of

HNHM, but I could not find it there.

87.6.2. Molops merditanus intermedius J. Müller, 1917

References. Molops obtusangulus intermedius: Müller, 1917: 348-349 (Montenegro: Podgorica,

loc. typ.; Albania / Montenegro border: Ricavac, loc. typ; Albania: Vermosh, loc. typ). Molops

merditanus intermedius: Mlynář, 1977: 107-108 (Ricavac; Vermosh); Drovenik & Peks, 1994: 71

(Ricavac); Drovenik & Peks, 1999: 83 (Ricavac); Bousquet, 2003f: 473.

Material studied. “Ricavac juni” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype (NMW);

“Vermosa 10-21.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype (NMW).

Notes. Each of the three localities mentioned in the original description (Müller, 1917) could

be a potential “locus typicus”, however a lectotype designation still has to be done.

Species group “curtulus”

Notes. This taxonomically difficult species complex is treated here differently than previously

(Mlynář, 1977):

Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri

reiseri Apfelbeck, 1904

Molops (Molops) apfelbecki corallipes J. Müller, 1936, new assignment of Molops (Molops) reiseri

corallipes J. Müller, 1936

Molops (Molops) apfelbecki pseudoalpestris Mlynář, 1977, new assignment of Molops (Molops) reiseri

pseudoalpestris Mlynář, 1977

Molops (Molops) apfelbecki victoris J. Müller, 1918, new assignment of Molops (Molops) reiseri

victoris J. Müller, 1918

Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J.

Müller, 1917

87.7. Molops albanicus Apfelbeck, 1904

References. Molops albanicus: Apfelbeck, 1906: 1665 (Mal i Shenjt Mt.; Zepë Mt.); Apfelbeck,

1907a: 499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 8 (Merdita: Mal i Shenjt

Mt.; Merdita: Zepë Mt.); Müller, 1930: 124 (Merdita: Zepë Mt.); Mlynář, 1977: 118, 120 (Mal

i Shenjt Mt.; Zepë Mt.; Oroshi); Bousquet, 2003f: 472.

Material studied. “Merdita Zebia”, 9 s. (DEI; HNHM; NMW); “Merdita M. Scheit” / “Coll.

Apfelbeck”, 1 m. (HNHM); “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 21 s. (DEI;

NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 3 s. (NMW); “Alban. Exped. Pashtrik’18

4-17.7”, 1 s. (NMW); 9 s. from locality “Merdita” (MNHUB); 33 s. from locality “Llogora”

(MNHUB); 11 s, from locality “Mal i Shenjit” (MNHUB); 3 s. from locality “Iballe” (MNHUB);

1 s. from locality “Mal i Gjer” (MNHUB); 3 s. from locality “Shen-Ili” (MNHUB).

Additional material included (studied A. Vigna Taglianti). Valbona, Hellit Kuq Mt., south

slope, 930 m, 6.VII.1993, 1 m., L. Latella leg. (cAVT).


Notes. Apfelbeck (1904: 222) described this species basing on a single male from the type

locality “Prizren”. This place is situated in the Kosovo Region of Serbia. The binomen Molops

transfuga Apfelbeck (Müller, 1930: 124; Mlynář, 1977: 120) is nomen nudum.

87.8. Molops apfelbecki Ganglbauer, 1891

Notes. In the past, M. apfelbecki and M. reiseri have been treated as separate species. Probable

cause for this state is that Apfelbeck (1904: 223), while describing M. reiseri, missed to compare

it with M. apfelbecki. Later, Mlynář (1977) similarly did not compare the aedeagus of both species,

but noticed the resemblance in their female genital structures. The present author has studied

the aedeagus of M. reiseri holdhausi (syntype, NMW), M. reiseri (lectotype, HNHM), M. reiseri

victoris (syntype, NMW; specimens from Koprivnik Mt. and Shkëlzen Mt., HNHM) and M. apfelbecki

(lectotype, HNHM) and found that M. reiseri holdhausi is different in the shape of the median

lobe from the other three taxa. In fact, the last taxa have identical aedeagus. Therefore, M. reiseri

reiseri and M. apfelbecki apfelbecki are considered as geographical races of single species, and M.

holdhausi – as valid species which occurs together with M. apfelbecki s.l.

87.8.1. Molops apfelbecki victoris J. Müller, 1918

References. Molops victoris: Müller, 1918a: 1-3, 6 (Albania / Montenegro border: North Albanian

Alps, 1300-2000 m, loc. typ.). Molops reiseri victoris: Mlynář, 1977: 114-115 (North Albanian

Alps; Shkëlzen Mt.).

Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14” /

“victoris n. det. J. Müller”, 2 m. syntypes of Molops victoris (NMW); “Albania Skölsen” / “Akad.

Balk. Exp. Csiki 1917”, 1 m. with extracted penis (HNHM); “Maja Peis” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 5 s. (MNHUB); “Shkëlzën”

/ “Albania leg. Bischoff 1936”, 22 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Bischoff,

Mueller 1939” / “Type” / “Apfelbecki sbsp. Bischoffi m. det. Müller”, 1 s. (MNHUB); “M. Raba

Massiv” / “Albania leg. Bischoff, Mueller 1939” / “Type”, 5 s. (MNHUB); “Maja e Dragomirit”

/ “Albania leg. Bischoff, Mueller 1939” / “M. Apfelbecki sbsp. Bischoffi m. det. G. Müller 1940”,

1 s. (MNHUB); “Maja e Dragomirit” / “Albania leg. Bischoff, Mueller 1939”, 5 s. (MNHUB).

Additional material studied. Bosnia Herzegovina / Montenegro. “Apfelbeck Volujak” /

“Coll. Apfelbeck”, 1 m. lectotype and 3 paralectotypes of Molops apfelbecki (HNHM).

Montenegro. “Vunšaj, Cafa glava, Alban. Penther” / “reiseri Apf.”, 1 m. of Molops apfelbecki

victoris (NMW). Serbia (Kosovo Region). “Montes Koprivnik” / “Akad. Balk. Exp. Csiki 1917”,

1 m. of Molops apfelbecki victoris (HNHM). Serbia. “Reiser 1898 Kopaonik” / “Coll. Apfelbeck”,

1 m. lectotypus and 2 paralectotypes of Molops reiseri (HNHM); “Reiser 1899 Kopaonik” /

“reiseri Apf. B& typus” / “type”, 1 m. of Molops apfelbecki reiseri (NMW); “Reiser 1899 Kopaonik”

/ “reiseri Apf. Serb. turk. Grence” / “type”, 1 m., 1 f. of Molops apfelbecki reiseri (NMW); “Reisser

1899 Kopaonik”, 1 m., 1 f. of Molops apfelbecki reiseri (MNHUB).


Rubës, 2000 m, 8.VII.1993, 1 f., L. Latella leg. (cAVT).

Notes. Müller (1930: 126) revised the material of Apfelbeck (1918c: 166 sub M. reiseri from

Vunšaj, Cafa glava) and determinated it as M. victoris. However, Bousquet (2003f: 474) has omitted

M. reiseri victoris (currently M. apfelbecki victoris, see above) for Albania in spite of the fact that

Mlynář (1977: 115) gave a detailed record. Here the subspecies is confirmed for the country. The

trinomen M. apfelbecki bischoffi G. Müller 1940, found on labels of two specimens, is nomen nudum.


87.9. Molops curtulus Ganglbauer, 1891

87.9.1. Molops curtulus cakorensis Mlynář, 1977

References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps). Molops curtulus:

Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Müller, 1918a: 10 (Maranajt

Mt.; North Albanian Alps: Rapshë, Grebeni, Vermosh, Ricavac); Csiki, 1940: 217 (Shkëlzen

Mt.). Molops curtulus cakorensis: Mlynář, 1977: 111.


(NMW); “Vermosa Penther”, 1 s. (NMW); “13.15.VI.14 Vermosa ca 1000 m”, 1 s. (NMW);

“Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Rapsa

10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 8 s. (NMW); “Grebeni 16.VI.” / “Alban.

montenegr. Grenze, Penther’14”, 5 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban.

montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Rogamit” / “Albania leg. Bischoff

1935”, 3 f. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 9 s. (MNHUB); “Boge

Aufstieg z. Rab” / “Albania leg. Bischoff, Mueller 1939”, 1 s. (MNHUB); North Albanian

Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS).

Notes. Bousquet (2003f: 472) has omitted the species for Albania. Here it is confirmed for

the country.

87.9.2. Molops curtulus winklerianus J. Müller, 1936

References. Molops winklerianus: Müller, 1936: 103 (Tomor Mt.: Kulmak, loc. typ.; Shëndelli

Mt. near Tepelenë; Llogora); Molops curtulus winklerianus: Mlynář, 1977: 111 (Tomor Mt.: Kulmak;

Llogora); Bousquet, 2003f: 472.

Material studied. “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 4 s. (NMW).

Notes. Mlynář (1977: 111) stated in a footnote that the populations of M. curtulus from Central

Albania may belong to a separate race, so further investigations of material from this region

of the country will reveal more details for elucidation that statement.

87.10. Molops holdhausi J. Müller, 1917

References. Molops holdhausi: Müller, 1917: 352-353 (Albania / Montenegro border: North

Albanian Alps, 1300-2000 m, loc. typ.); Müller, 1918a: 6 (North Albanian Alps); Müller, 1930:

126 (North Albanian Alps, Grebeni near Vermosh). Molops reiseri: Apfelbeck, 1918c: 166 (North

Albanian Alps, Grebeni near Vermosh). Molops reiseri holdhausi: Mlynář, 1977: 112, 114 (North

Albanian Alps).

Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14” /

“holdhausi n. det. J. Müller”, 1 m. syntype of Molops holdhausi (NMW); “Grebeni, Vermoša

Alban., Penther 16 Juni, am Schnee” / “reiseri Apf.”, 2 m. (NMW).

Additional material studied. Montenegro. “Durmitor”, 4 s. (HNHM); “Montenegro

Durmitor, Penther’04”, 1 m., 1 f. (NMW); “Jugoslavia mer. Prokletija Gusinje Matìjka VI.1937”,

1 m., 1 f. (MNHUB); “Jugoslavia mer. Prokletja Gusinje Kodym 20.VI.1937”, 2 s. (HNHM).

Notes. Müller described M. holdhausi and M. victoris as separate species (Müller 1917; Müller

1918a). Mlynář (1977: 112-113) neglected the fact that M. reiseri holdhausi and M. reiseri victoris

occur sympatrically and expressed doubt about the status of both taxa due to the lack of

enough specimens at hand (Mlynář (1977: 115). The present author compared the penis

structures of several males, previously determined as ssp. holdhausi or ssp. victoris, and ascertained


the first is good species - Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops)

reiseri holdhausi J. Müller, 1917. Bousquet (2003f: 474) has omitted M. holdhausi for Albania

though Müller (1930) gave a detailed record. Here the species is confirmed for the country.

87.11. Molops plurisetosus J. Müller, 1918 [= malissorum Apfelbeck, 1918]

References. Molops apfelbecki plurisetosus: Müller, 1918a: 3-4, 11 (Albania / Montenegro border:

North Albanian Alps, 1200-2000 m, loc. typ.; Albania / Montenegro border: Ricavac, loc. typ.;

Albania: Vermosh, loc. typ.). Molops malissorum: Apfelbeck, 1918c: 160, 167-168 (Montenegro:

North Albanian Alps, Vusanje, loc. typ.; Albania: pastures over Theth). Molops plurisetosus: Mlynář,1977: 120-121 (North Albanian Alps; Ricavac; Vermosh).

Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 4

syntypes of Molops plurisetosus (NMW); “Ricavac Juni” / “Alban. montenegr. Grenze, Penther’14”,

1 syntype of Molops plurisetosus (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze,

Penther’14”, 2 syntypes of Molops plurisetosus (NMW); North Albanian Alps, Theth, 800-1200 m,

28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Redohimes,

2400-2550 m, 29.V.1993, 3 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps,

Bogë, 1800-1900 m, 20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian

Alps, Bogë, cave “No 25”, 23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Shkodër

District, Kozuja, 1800 m, 27.VIII.1996, 1 m., 1 f., T. Ivanova leg. (NMNHS).

Additional material studied. Montenegro. “Vunšaj, Prokletija-Geb., Albania Dr. Penther” /

“Molops malissorum n. sp. Apf.”, 2 syntypes of Molops malissorum (NMW).

Notes. Bousquet (2003f: 474) has omitted the species for Albania. Here it is confirmed for the

country. Currently, the exact publishing dates of the descriptions of M. plurisetosus and M. malissorum

were not found, so the problem with the priority of one of the two names is pending. Each of

the different localities mentioned in the original descriptions of M. plurisetosus (Müller, 1918a)

could be a potential “locus typicus”, however lectotype designation still has to be done.

Species group “rufipes”

87.12. Molops osmanilis Apfelbeck, 1904

[Molops osmanilis osmanilis Apfelbeck, 1904]

Notes. The record from “Prisren” (Apfelbeck, 1904: 229) refers to the Kosovo Region of Serbia,

so perhaps Bousquet (2003f: 473) has followed this source when listing M. osmanilis osmanilis for

Albania. The nominotypical subspecies is treated here as likely to be found in the country.

87.12.1. Molops osmanilis tomoricensis J. Müller, 1923

References. Molops tomoricensis: Müller, 1923: 109-110 (Tomor Mt., alpine zone, loc. typ.). Molops

osmanilis tomoricensis: Mlynář, 1977: 123 (Tomor Mt.: Tomor, Abbas Ali); Bousquet, 2003f: 473.

Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 5 s. (NMW;

MNHUB); “Tomor: Kulmak Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);

“Tomor: Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor:

Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 3 (NMW); “Tomor: Tomor Alban.

mer.” / “Winkler, Lona, Bischoff, V.1931”, 7 s. (NMW); more than 90 s. from Tomor Mt.

(MNHUB); “Mal i Dajtit Tirane” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara


juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit Tirane” / “leg. Bischoff

Albania 1934”, 1 s. (MNHUB); “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff ”,

13 s. (MNHUB); “Maja e Dhembelit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s.

(MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 2 s. (MNHUB); “AL (Tirana) 8 km

N psene E Tirana 1450 m. 30.IV.1993 leg. H. & S. Peks”, 2 s. (MNHUB).

Notes. All the specimens examined from Albania are put to this race more or less provisionally.

However, for instance specimens from Southwest Albania (localities “Logara” and “Maja e

Dhembelit”) have a shape of the pronotum like spp. osmanilis, whereas these from Central

Albania (localities “Mal i Dajtit” and “Mal i Krujes”) – like ssp. tomoricensis.

87.13. Molops rufipes Chaudoir, 1843

87.13.1. Molops rufipes rufipes Chaudoir, 1843

References. Molops rufipes: Rambousek, 1912: 93; Mlynář, 1977: 124; Hieke, 1981: 81; Hieke &

Wrase, 1988: 82. Molops rufipes rufipes: Mlynář, 1977: 124 (Mal i Dajtit Mt.).

Material studied. “Mal i Daitit b. Tirana, Alb. C.” / “leg. Bischoff 1931 coll. A. Winkler”, 2

s. (NMW); “Tomor: Tomor Alban. mer.”, 1 m. (MNHUB); “Tomor-Buranj juli 1932” / “Albania

leg. Bischoff 1932”, 1 m. (MNHUB); more than 50 s. from locality “Iballe” (MNHUB); more

than 50 s. from locality “Mal i Dajtit” (MNHUB); Tiranë, 26.VII.1965, 1 f. (MNST); Mal i

Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 4 m., 3 f., L. Penev leg. (NMNHS).

Notes. Bousquet (2003f: 474) has omitted the subspecies for Albania. Here it is confirmed

for the country. The material of Rambousek (1912) was not revised and therefore it is placed

in the nominotypical race provisionally.

# Molops rufipes klisuranus Apfelbeck, 1902

References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps).

Notes. Two wrong records (Müller, 1930: 125-126, revised as M. curtulus s.l.) for this taxon

have been recorded from the Western Balkan (South Serbia, Sandzak: Raška near Novi Pazar,

Apfelbeck, 1904: 222; North Albanian Alps, Apfelbeck, 1918c: 166). Therefore the taxon is

removed from the list of the Albanian Carabidae.

87.13.2. Molops rufipes munelensis Apfelbeck, 1906

References. Molops sturanyi munelensis: Apfelbeck, 1906: 1665-1666 (Munella Mt., loc. typ.);

Müller, 1918a: 9 (Merdita: Munella Mt.). Molops rufipes munelensis: Mlynář, 1977: 128, 130 (Munella

Mt.; Munella Mt.: Oroshi; Berishe); Bousquet, 2003f: 474.

Material studied. “Orosi”, 5 s. (HNHM); “Oroši (Ptr.) Alban. spt.” / “Munela” / “Coll. Apfelbeck”,

1 lectotype (HNHM); “Merdita Munela”, 3 paralectotypes (HNHM); “Merdita Munela”, 9 s. (DEI;

MNHUB); “Merdita Munela Apfb.”, 4 s. (MNHUB); “Merdita” / “Apfb.”, 2 s. (DEI); “Merdita

Munela” / “Wgth. 1906 Merdita”, 21 s. (NMW); “Merdita Zebia”, 9 s. (NMW); “Alban. Exp.’18

Pashtrik, 29.V.-4.VI.”, 9 s. (NMW); “Alban. Exp.’18 Pashtrik, 4-17.7”, 5 s. (NMW); “M. Mulecifit”

/ “Albania leg. Bischoff 1936”, 6 s. (MNHUB); “Berishe” / “Albania leg. Mueller 1939”, 6 s.

(MNHUB); “AL (Shkodar). Paß SE Puke 850 m 2.V.1993” / “leg. H. & S. Peks”, 1 m. (MNHUB).

[Molops rufipes pseudoosmanilis Mlynář, 1977]

Notes. Since this subspecies was hitherto recorded only from the Macedonian section of the

Korabi Mt., it is expected to live in Albania, too.


87.13.3. Molops rufipes steindachneri Apfelbeck, 1908

References. Molops rufipes steindachneri: Mlynář, 1977: 128 (Korab Mt.).

Notes. Bousquet (2003f: 474) has omitted this subspecies for Albania. Actually its occurrence in

the country needs confirmation. Apfelbeck published three consecutive descriptions of this taxon

(Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid.

87.13.4. Molops rufipes sturanyi Apfelbeck, 1906

References. Molops sturanyi: Apfelbeck, 1906: 1665 (Merdita: Mal i Shenjt Mt., loc. typ.; Merdita:

Munella, loc. typ.; Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 497, 499, 503 (Maranajt Mt.:

top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 9 (Merdita:

Zepë Mt.; Merdita: Mal i Shenjt Mt.; Maranajt Mt.); Müller, 1930: 123 (Gjalicë Mt.). Molops

steindachneri: Csiki, 1940: 217 (Gjalicë Mt.; Korab Mt.). Molops rufipes sturanyi: Mlynář, 1977: 128

(Mal i Shenjt Mt.; Zepë Mt.; Gjalicë Mt.; Korab Mt.); Bousquet, 2003f: 474.

Material studied. “Albania” / “Merdita”, 2 s. (DEI); “Merdita M. Schèit” / “Apfelb. Alban.

spt. IV.V.’05”, 1 lectotype (HNHM); “Merdita Zebia”, 4 paralectotypes (HNHM); “Merdita

Zebia”, 6 s. (DEI; MNHUB); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s.

(HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM);

“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 11 s. (HNHM); “Alban. Exp. 1918

Gjalica Ljums 17-26.VI.”, 49 s. (NMW); 15 s. from locality “Bjeshka Matroxh” (MNHUB); 20

s. from localaity “Derzë” (MNHUB); more than 300 s. from locality “Gjallica” (MNHUB);

more than 500 s. from locality “Korab” (MNHUB); 2 s. from locality “Taja” (MNHUB).

Notes. The record from the Maranajt Mt. (Müller, 1918a: 9) has to be treated cautiously

because it is the only record of this species north of the Drin River.

Species group “piceus”

87.14. Molops piceus (Panzer, 1793)

87.14.1. Molops piceus balcanicus Mlynář, 1977

References. Molops austriacus: Müller, 1918a: 9 (Rapshë; North Albanian Alps); Horion, 1941: 302.

Molops piceus balcanicus: Mlynář, 1977: 136; Hieke, 1981: 81. Molops piceus: Hieke & Wrase, 1988: 82.


(NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s.

(NMW); “Maja Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania


Notes. Bousquet (2003f: 473) has omitted the species for Albania. Here it is confirmed for

the country.

Subgenus Stenochoromus L. Miller, 1866

87.15. Molops montenegrinus (L. Miller, 1866)

87.15.1. Molops montenegrinus cavernicola J. Müller, 1916

References. Molops montenegrinus cavernicola: Müller, 1916: 81 (Çukall Mt.: cave “Sckamit”, loc.

typ.); Müller, 1918a: 11-12 (Çukall Mt.: cave “Sckamit”; North Albanian Alps: Ricavac, Vermosh);



Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, more

than 20 s. (NMW); “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Starka

Alb. sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 6 s. (NMW); “Maja e Jezerce Albania sept.”

/ “leg. Bischoff 1931 coll. A. Winkler”, 2 s. (NMNHS); “Maja e Jezercoë” / “Albania leg.

Bischoff 1933”, 9 s. (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 27 s.

(MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cucale” / “Albania


“Maranaj” / “Albania leg. Bischoff 1937”, 3 s. (MNHUB); “Boge Aufstieg Z. Rab” / “Albania

leg. Mueller 1939”, 2 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Mueller 1939”, 7 s.

(MNHUB); “Maja e Dragomirit” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); North Albanian

Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 2 m., P. Beron & B. Petrov leg. (NMNHS); North

Albanian Alps, Maj’e Redohimes, 1800-2200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg.

(NMNHS); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron &

B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 3 s., P. Beron



Rubës, 2000 m, 8.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Mali-i-Thatë Mt., cave “Shpella e

Haxise, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2250 m,


Notes. All the data from Albania are placed provisionally to the ssp. cavernicola, since no

intraspecific revision of M. montenegrinus s.l. is made. In addition to the material studied above,

in NMW I have seen more specimens, which remained unregistered.

88. Genus Myas Sturm, 1826

Subgenus Myas s.str.

88.1. Myas chalybaeus (Palliardi, 1825) [= rugosicollis Brullé, 1832]

References. Myas chalybaeus: Tschitschérine, 1899: 70; Anonymous, 1907: 284; Hieke & Wrase,

1988: 69; Bousquet, 2003f: 475. Myas rugosicollis: Tschitschérine, 1899: 70.

Material studied. “Alban. Exp.’18. Kruma 5.VI.”, 1 s. (NMW); “Mal i Dajtit Tiranë” / “Albania

leg. Bischoff 1931”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB);

“Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania

leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit Tiranë” / “Albania 1936 leg. Ramadan coll.

Bischoff ”, 7 s. (MNHUB); “Valbona Çerem” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Berishe” / “Albania Mueller leg. 1939”, 3 s. (MNHUB); “Boge Ober D. Quelle” / “Albania

Mueller leg. 1939”, 3 s. (MNHUB); “Çafa e Ceromis Iballe” / “Albania Mueller leg. 1939”, 1 s.

(MNHUB); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov

leg. (NMNHS); 5 km Leskoec, 2.VII.1994, 1 s., P. Tenchev leg. (NMNHS); Stebleva, cave “Shpella

e Hadzhi i Malit”, 5-8.V.2006, 1 s., A. Zhalov, D. Raychev & J. Tsvetanov leg. (NMNHS).



89. Genus Pedius Motschulsky, 1850

89.1. Pedius inquinatus (Sturm, 1824)

References. Pterostichus inquinatus: Apfelbeck, 1904: 257 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);

Schatzmayr, 1930: 198; Horion, 1941: 279; Schatzmayr, 1942: 65; Schatzmayr, 1943b: 110

(Mal i Krujes); Hieke & Wrase, 1988: 71. Pedius inquinatus: Bousquet, 2003f: 478.

Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Lum i Tiranes” / “leg.

Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s.

(MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); Tiranë Park, 22-

31.X.1997, 3 s., R. Ward leg. (CMNH).

89.2. Pedius longicollis (Duftschmid, 1812)

References. Pedius longicollis: Bousquet, 2003f: 478.

90. Genus Poecilus Bonelli, 1810

Subgenus Ancholeus Dejean, 1828

90.1. Poecilus puncticollis (Dejean, 1828)

References. Pterostichus puncticollis: Schatzmayr, 1943b: 110 (Kruja; Skrofotina; Stan-Karbunarë).

Poecilus puncticollis: Hieke & Wrase, 1988: 71; Bousquet, 2003f: 480.

Material studied. “Liquen i Butrintes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB);

“Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).

Subgenus Poecilus s.str.

90.2. Poecilus cupreus (Linnaeus, 1758)

90.2.1. Poecilus cupreus cupreus (Linnaeus, 1758)

References. Pterostichus cupreus: Apfelbeck, 1904: 253 (Velipoja); Apfelbeck, 1907a: 495 (Buenë

River); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1930: 184 (Tomor Mt.); Schatzmayr, 1943b:

109 (Kruja; Stan-Karbunarë; Lushnje). Poecilus cupreus: Hieke & Wrase, 1988: 70. Poecilus cupreus

cupreus: Bousquet, 2003f: 482.

Material studied. “Oboti Bojana”, 1 s. (NMW); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”,

1 s. (NMW); “Alban. Exp.’18 Pashtrik 4-14.7.”, 7 s. (NMW); “Albanien Ura Gjolés Umg. Tirana

Marz 1942 Muller Ausbeute”, 1 s. (MNHUB); “Albania Ostrovice” / “7.VII.1959 leg. Xh.

Murraj”, 2 s. (HNHM); Sauk, 14.V.1961, 1 m. (MNST); Sauk, 3.IV.1963, 1 m. (MNST); Saranda

District, Butrinti, 16.IV.1994, 3 s., S. Beshkov leg. (NMNHS); Tiranë Park, 8.II.1998, 2 m.,

shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH).

Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A.

Paparistos leg. (cAVT); Rapsh, Reç, 15.V.2002, 1 f., A. Paparistos leg. (cAVT).

Notes. In addition to the material studied, in MNHUB I have seen other more 24 s. from

different localities, which remained unregistered.


90.3. Poecilus cursorius (Dejean, 1828)

90.3.1. Poecilus cursorius cursorius (Dejean, 1828)

References. Pterostichus cursorius: Schatzmayr, 1930: 187; Schatzmayr, 1943b: 110 (Lushnje).

Poecilus cursorius: Hieke & Wrase, 1988: 71. Poecilus cursorius cursorius: Bousquet, 2003f: 482.

Material studied. “Tirane” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Rushkull” / “Albania

leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bei Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

90.4. Poecilus koyi Germar, 1824

90.4.1. Poecilus koyi koyi Germar, 1824

References. Pterostichus koyi dinaricus: Schatzmayr, 1930: 176 (Tomor Mt.); Schatzmayr, 1942:

55 (Tomor Mt.). Pterostichus koyi: Schatzmayr, 1943b: 109 (Tërpan). Poecilus koyi: Csiki, 1940:

216 (Korab Mt.); Hieke et al., 1986: 188 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988:

69. Poecilus koyi koyi: Bousquet, 2003f: 482.

Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 18 s. (NMW);

“Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Albania Mts. Korab” /

“Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Korab 1918 24.31.VII.”, 1 s.

(NMW); “Albania Rapsa Dr. Grabowski”, 1 s. (HNHM); “Albania Zalle-Dardha” / “23.V.1958

leg. Xh. Murraj”, 3 s. (HNHM); Bizë, 9-16.VII.1961, 2 f. (MNST); Mal i Dajtit Mt., 10-11.V.1965,

5 m., 1 f.; Mal i Dajtit Mt., 24.V.1966, 2 f. (MNST); Tiranë, 30.V. 1965, 2 m. (MNST); 2 km N

Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS).

90.5. Poecilus lepidus (Leske, 1785)

90.5.1. Poecilus lepidus lepidus (Leske, 1785)

References. Pterostichus lepidus: Csiki, 1940: 216 (Korab Mt.); Schatzmayr, 1942: 56. Poecilus

lepidus: Hieke & Wrase, 1988: 70. Poecilus lepidus lepidus: Bousquet, 2003f: 483.

Material studied. “Alban. Vermosa Penther”, 4 s. (NMW); “Vermosa’14 Penther”, 1 s. (NMW);

“Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania

Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Shkëlzën” / “Albania leg. Bischoff

1936”, 2 s. (MNHUB).

90.6. Poecilus rebeli (Apfelbeck, 1904)

References. Pterostichus rebeli: Apfelbeck, 1904: 254 (Velipoja; Vlorë); Apfelbeck, 1907a: 495

(Buenë River); Scatzmayr, 1930: 184; Schatzmayr, 1942: 59; Bousquet, 2003f: 483.

Material studied. “Velipoja Albania” / “Coll. Apfelbeck”, 1 syntype (HNHM); “Mustajbeg

Velipoja Albania”, 2 syntypes (NMW); “Oboti Albania”, 1 s. (NMW); “Scutari Bojana” /

“Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW).

90.7. Poecilus striatopunctatus (Duftschmid, 1812)

References. Poecilus striatopunctatus: Apfelbeck, 1907a: 495 (Buenë River); Hieke & Wrase, 1988:

71; Bousquet, 2003f: 484.

Material studied. “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).


90.8. Poecilus versicolor (Sturm, 1824) [= coerulescens (Linnaeus, 1758)]

Material studied. “Alban. Ricavac Penther”, 2 s. (NMW); “Alban. Vermosa Penther”, 4 s.

(NMW); “Vermosa’14 Penther”, 2 s. (NMW); “Albania Korab” / “Akad. Balk. Exp. Csiki

1918”, 1 s. (HNHM); “Albania Mali-Thate” / “17.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM);

“Albania Sauk” / “27.VII.1959 leg. Xh. Murraj”, 1 s. (HNHM); Golem, 10.VII. 1965, 1 s.

(MNST); Tiranë, 26.VIII.1965, 1 m. (MNST).

Notes. The mention of Hieke & Wrase (1988: 70) for Albania bases apparently on the record

“Prisren” (Apfelbeck, 1904: 255 sub Pterostichus coerulescens). However, currently this locality is

situated in the Kosovo Region of Serbia. Bousquet (2003f: 484) has most likely followed one

of these sources too. Since there is no previous reliable record from Albania, P. versicolor is

treated here as new species for the fauna of Albania.

91. Genus Pterostichus Bonelli, 1810

Subgenus Argutor Dejean, 1821

91.1. Pterostichus cursor (Dejean, 1828)

References. Pterostichus vernalis cursor: Apfelbeck, 1904: 257 (Velipoja). Pterostichus cursor: Hieke

& Wrase, 1988: 73; Bousquet, 2003f: 486.

Material studied. “Skutari Bojana”, 1 s. (NMW); “Skutari l. See-Ufer”, 1 s. (NMW); “Skutari

Mesi”, 1 s. (NMW); “Skutari Vraka”, 1 s. (NMW); “Vermosa’14 Penther”, 1 s. (NMW); “Liquen

i Butrintes vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg.


Subgenus Bothriopterus Chaudoir, 1835

91.2. Pterostichus oblongopunctatus (Fabricius, 1787)

91.2.1. Pterostichus oblongopunctatus oblongopunctatus (Fabricius, 1787)

References. Pterostichus oblongopunctatus: Schatzmayr, 1943a: 85; Hieke & Wrase, 1988: 74.

Pterostichus oblongopunctatus oblongopunctatus: Bousquet, 2003f: 487.

Material studied. “Vermosa’14 Penther”, 2 s. (NMW); “Maja Rogamit” / “Albania leg. Bischoff

1935”, 2 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).


Subgenus Cheporus Latreille, 1829

91.3. Pterostichus burmeisteri Heer, 1838

91.3.1. Pterostichus burmeisteri burmeisteri Heer, 1838 [= metallicus (Fabricius, 1792)]

References. Pterostichus metallicus: Klaptocz, 1910: 399 (Theth).

Material studied. “Vermosa’14 Penther” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW);

“Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja

e Jezercë Çafa Ostars” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Shkëlzën” / “Albania


leg. Bischoff 1935”, 18 s. (MNHUB); “Maja Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB);

“Maja Rogamit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Boge Aufstieg z. Rab” /

“Albania Mueller leg. 1939”, 1 s. (MNHUB); “Boge Çafa Shtegot” / “Albania Mueller leg. 1939”,

1 s. (MNHUB); “Maja e Dragomirit” / “Albania Mueller leg. 1939”, 1 s. (MNHUB).


900 m, 15.VII.1993, 1 f., L. Latella leg. (cAVT).

Notes. Bousquet (2003f: 488) has omitted the species for Albania. Here it is confirmed for the country.

Subgenus Feronidius Jeannel, 1942

91.4. Pterostichus melas (Creutzer, 1799)

91.4.1. Pterostichus melas depressus (Dejean, 1828)

References. Pterostichus melas: Apfelbeck, 1904: 265 (Vlorë; Velipoja); Apfelbeck, 1909: 297

(Vlorë; Berat); Schatzmayr, 1943b: 109 (Kruja; Dukati; Tërpan); Hieke et al., 1986: 189 (Tiranë;

Borshi near Vlorë; Poliçan near Tomor Mt.). Pterostichus melas depressus: Hieke & Wrase, 1988:

76; Bousquet, 2003f: 495.

Material studied. “Mustajbeg Velipoja Albania”, 4 s. (HNHM); “Sudalbanien Kopstein”, 1 s.

(NMW); “Skutari Penther”, 4 s. (NMW); “Hani Grabom mai” / “Alban. montenegr. Grenze,

Penther’14”, 1 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW);

“Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 1 s.

(HNHM); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 3 s. (HNHM); “Albania Sauk” /

“16.VI.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Terbac” / “6.VI.1960 leg. Xh. Murraj”, 1 s.

(HNHM); Sauk, 9.III.1961, 1 m. (MNST); Tepelenë, 1-3.VI.1961, 1 m. (MNST); Rodenj, 26-

28.VII.1962, 2 m. (MNST); Kurbneshi, VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Tiranë,

Selit e Vogel, 17.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Saranda District, Ljukove, 5.V.1994, 6 s.,

P. Stoev leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg.

(NMNHS); Tiranë Park, 22-31.X.1997 / 1-9.XI.1997, 5 s., R. Ward leg. (CMNH).

Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 f., A. Paparistos

leg. (cAVT); Rapsh, Reç, 15.V.2002, 1 f., A. Paparistos leg. (cAVT).

Notes. Besides the material studied above, in MNHUB I have seen other more 63 s. from

Albania, which remained unregistered.

Subgenus Melanius Bonelli, 1810

91.5. Pterostichus atterimus (Herbst, 1784)

91.5.1. Pterostichus atterimus atterimus (Herbst, 1784)

References. Pterostichus atterimus: Apfelbeck, 1904: 258 (Velipoja); Schatzmayr, 1943a: 87.

Pterostichus atterimus atterimus: Bousquet, 2003f: 499.

Material studied. “Gjormi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

91.6. Pterostichus elongatus (Duftschmid, 1812)

References. Pterostichus elongatus: Schatzmayr, 1930: 210 (Shkodër); Hieke & Wrase, 1988: 76;



Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Mamurash” / “Albania leg.


Subgenus Morphnosoma Lutshnik, 1915

91.7. Pterostichus melanarius (Illiger, 1798)

91.7.1. Pterostichus melanarius melanarius (Illiger, 1798)

References. Pterostichus melanarius: Hieke & Wrase, 1988: 75. Pterostichus melanarius melanarius:


Material studied. “Alban. Exp. 1918 Gjallica Ljums 17-26.VI.”, 1 s. (NMW); “Lesh” / “Albania



900 m, 8-16.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT).


# subgenus Oreoplatysma Jakobson, 1907

# Pterostichus lodosi Heinz, 1977

References. Pterostichus lodosi: Bousquet, 2003f: 505.

Notes. Most probably, Bousquet (2003f: 505) has listed this Caucasian endemic species for

the country by mistake. Therefore, it is removed from the list of the Albanian ground beetles.

Subgenus Parahaptoderus Jeanne, 1969


91.8. Pterostichus brevis (Duftschmid, 1812)

91.8.1. Pterostichus brevis brevis (Duftschmid, 1812)

Material studied. “Vermosa’14 Penther”, 1 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” /

“Alban. montenegr. Grenze Penther’14”, 7 s. (NMW).


Subgenus Phonias Gosis, 1886

91.9. Pterostichus apfelbecki Csiki, 1908 [= convexiusculus Apfelbeck, 1904]

References. Pterostichus convexiusculus: Apfelbeck, 1904: 261-262 (Velipoja, loc. typ.); Magistretti,

1965: 380; Jeanne, 1988: 74. Pterostichus apfelbecki: Schatzmayr, 1943a: 82 (Velipoja); Bousquet,

2003f: 507. Argutor convexiusculus: Jeanne, 1988: 74.

91.10. Pterostichus diligens (Sturm, 1824)

References. Pterostichus diligens: Hieke & Wrase, 1988: 72; Bousquet, 2003f: 507.

Material studied. “Vermosa’14 Penther”, 2 s. (NMW); “Çafa Markofçes” / “Albania leg.




91.11. Pterostichus strenuus (Panzer, 1796)

References. Pterostichus strenuus: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 234; Hieke

& Wrase, 1988: 72; Bousquet, 2003f: 508. Argutor strenuus: Jeanne, 1988: 74.

Material studied. “Skutari Albania”, 3 s. (NMW); “Mustajbeg Velipoja Albania”, 5 s. (HNHM;

NMW); “Buschnesh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Pyla Thumana” /


Subgenus Platypterinus Reitter, 1908

91.12. Pterostichus ottomanus Apfelbeck, 1908

91.12.1. Pterostichus ottomanus ottomanus Apfelbeck, 1908

References. Pterostichus ottomanus: Schatzmayr, 1943a: 106. Pterostichus ottomanus ottomanus:


Material studies. “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkalle

Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB).

Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908;

Apfelbeck, 1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. First detailed

records for Albania.

Subgenus Platysma Bonelli, 1810

91.13. Pterostichus niger (Schaller, 1783)

91.13.1. Pterostichus niger niger (Schaller, 1783)

References. Pterostichus niger: Schatzmayr, 1930: 221 (Tomor Mt.); Csiki, 1940: 216 (Kula Ljums).

Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Mamurash” /

“Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Pyla Shullazit” / “Albania leg. Bischoff 1935”,

1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Bizë, 27.VII.1966, 1

m. (MNST).

Additional material included (studied A. Vigna Taglianti). Fier, 4.VII.1993, 1 f., L. Latella

leg. (cAVT).

Notes. Bousquet (2003f: 509) has omitted this species for Albania. Here it is confirmed for the

country.

Subgenus Pseudomaseus Chaudoir, 1838

91.14. Pterostichus anthracinus (Illiger, 1798)

91.14.1. Pterostichus anthracinus anthracinus (Illiger, 1798)

References. Pterostichus anthracinus: Apfelbeck, 1909: 297 (Pasha Liman); Hieke & Wrase, 1988:

74. Pterostichus anthracinus anthracinus: Bousquet, 2003f: 510.

Material studied. 36 s. from different localities (MNHUB); “Albania Shkoder” / “25.V.1958

leg. Xh. Murraj”, 1 s. (HNHM); Tiranë Park, 22.II.1998 / 8.III.1998, 6 s., shaded lake edge



91.14.2. Pterostichus anthracinus biimpressus (Küster, 1853)

References. Pterostichus anthracinus biimpressus: Apfelbeck, 1904: 260 (Velipoja; Vlorë); Bucciareli

& Sopracordevole, 1958: 198 (Mamurash); Bousquet, 2003f: 510.

Notes. Magistretti (1965: 378) noted by mistake that the type locality of ssp. biimpressus is

“Almissa (Albania)”. In fact this Italian name concerns the town of Omiš at the Dalmatian

coast of Croatia.

91.15. Pterostichus gracilis (Dejean, 1828)

91.15.1. Pterostichus gracilis gracilis (Dejean, 1828)

References. Pterostichus gracilis: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 219;

Schatzmayr, 1943a: 84; Hieke & Wrase, 1988: 74. Pterostichus gracilis gracilis: Bousquet,

2003f: 511.

91.16. Pterostichus minor (Gyllenhal, 1827)

91.16.1. Pterostichus minor minor (Gyllenhal, 1827)

References. Pterostichus minor: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 220; Hieke

& Wrase, 1988: 74. Pterostichus minor minor: Bousquet, 2003f: 511.

91.17. Pterostichus nigrita (Paykull, 1790)

Material studied. Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten

wood, R. Ward leg. (CMNH).


Subgenus Pterostichus s.str. [= Aphaon Heyden, 1906, syn. nov.]

Notes. The principal character used to separate Pterostichus s.str. and Aphaon (replacement

name for Phaon Heyden, 1883) is the number of lateral setiferous pores on pronotum

(Heyden, 1883). However, this character is inapplicable to divide the species of the two

subgenera. For instance, Pterostichus fasciatopunctatus (Creutzer, 1799), type species of the

genus and nominotypical subgenus has sometimes 1-3 marginal seta on the lateral border

of pronotum. On the other hand, P. rhilensis Rottenberg, 1874, the type species of Aphaon

also has 1-3 (sometimes 4) such setae. In fact, no other species from Aphaon shares this

character. The remaining four diagnostic features of Aphaon, e.g. transverse keel on the

last visible sternum in the males, number of the marginal pores on the same sternum in

both sexes, reduction of the scutellar stria and sharp basal border of elytra, cannot be of

subgeneric importance since these present also in other species of Pterostichus s.str. For

this reason Aphaon is treated as junior synonym of the latter subgenus and the following

nomenclatorial acts are introduced:

Pterostichus (Pterostichus) bischoffianus Jedlička, 1936, new assignment from Pterostichus (Aphaon)

bischoffianus Jedlička, 1936

Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment from Pterostichus (Aphaon) brucki

Schaum, 1859


Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment from Pterostichus

(Aphaon) latifianus latifianus Apfelbeck, 1906

Pterostichus (Pterostichus) latifianus strupii J. Müller, 1935, new assignment from Pterostichus (Aphaon)

latifianus strupii J. Müller, 1935

Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment from Pterostichus

(Aphaon) lumensis lumensis Apfelbeck, 1905

Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment from Pterostichus (Aphaon)

lumensis litae Csiki, 1940

Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment from Pterostichus

(Aphaon) lumensis ljubetensis Apfelbeck, 1908

Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment from Pterostichus (Aphaon)

malissorum Apfelbeck, 1905

Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment from Pterostichus (Aphaon) meisteri

Reitter, 1885

Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment from Pterostichus (Aphaon)

merklii (J. Frivaldszky, 1879)

Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment from Pterostichus (Aphaon)

pentheri Apfelbeck, 1918

Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment from Pterostichus (Aphaon)

reiseri Ganglbauer, 1889

Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment from Pterostichus

(Aphaon) rhilensis rhilensis Rottenberg, 1874

Pterostichus (Pterostichus) rhilensis kourili Mařan, 1933, new assignment from Pterostichus (Aphaon)

rhilensis kourili Mařan, 1933

Pterostichus (Pterostichus) rhilensis vitosensis Mařan, 1933, new assignment from Pterostichus (Aphaon)

rhilensis vitosensis Mařan, 1933

Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment from Pterostichus (Aphaon) walteri

Reitter, 1883

91.18. Pterostichus bischoffianus Jedlička, 1936

References. Pterostichus bischoffianus: Jedlička, 1936: 167 (Gjalicë Mt., loc. typ.); Schatzmayr,

1943a: 109 (Gjalicë Mt.); Bousquet, 2003f: 515.

Material studied. “Gjallica” / “Albania leg. Bischoff 1936” / “Typus” / “bischoffianus sp.n.

type det. Ing. Jedlička”, 1 m. syntype (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 2

m. (MNHUB).

91.19. Pterostichus brucki Schaum, 1859

References. Pterostichus brucki: Apfelbeck, 1918c: 166 (North Albanian Alps, Vermosh, 1100 m);

Jedlička, 1936: 167 (Koritnik Mt.; Gjalicë Mt.); Csiki, 1940: 216 (Gjalicë Mt.); Schatzmayr, 1943a:

107 (Koritnik Mt.; Gjalicë Mt.); Hieke, 1981: 80; Hieke & Wrase, 1988: 78; Bousquet, 2003f: 511.

Material studied. “Vermoša 1100 m” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW);

“Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp.’18

Gjalica Ljums 17-26.VI.”, 1 s. (NMW); “Gjalica” / “Albania leg. Bischoff 1935”, 1 s. (NMW);

“Koritnik” / “Albania leg. Bischoff 1935”, 8 s. (NMW; MNHUB); “Gjallica” / “Albania leg.


Bischoff 1935”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1936”, 1 s.

(MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 32 s. (MNHUB); “Shkalla Harushe” /

“Korab” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB).

91.20. Pterostichus latifianus Apfelbeck, 1906

91.20.1. Pterostichus latifianus latifianus Apfelbeck, 1906 [= munella Jedlička, 1936]

References. Pterostichus latifianus: Apfelbeck, 1906: 1663 (North Albania, Merdita: Mal Shenjt

Mt. and Munella Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 502 (Mal i Shenjt Mt.: top region,

1500 m); Schatzmayr, 1930: 256; Jedlička, 1936: 168 (Merdita; Dërze); Jedlička, 1939: 52;

Schatzmayr, 1943a: 109 (Merdita; Dërze). Pterostichus malissorum ab. munella: Jedlička, 1936: 169

(Munella Mt., loc. typ.); Jedlička, 1939: 52; Schatzmayr, 1943a: 108 (Munella Mt.). Pterostichus

latifianus latifianus: Bousquet, 2003f: 485.

Material studied. “Apfelb. Albanien”, 2 syntypes (MNHUB); “Latif Alban. sept.”, 4 s.

(DEI; HNHM); “Latif Munela” / “Latif Alban. spt.”, 1 s. (HNHM); “Latif Orosi” / “latifianus

typus! Apfelb.” / “typus”, 1 f. syntype (NMW); “Latif Orosi”, 2 syntypes (NMW); “Merdita

Mal Shêit” / “latifianus Apf. cotype det. Apflb.”, 1 m. syntype (MNHUB); “Merdita M.

Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (HNHM); “Merdita M. Scheit” / “Coll.

Apfelbeck”, 12 syntypes (HNHM); “Merdita M. Scheit”, 4 s. probably syntypes (NMW);

“Merdita Munela”, 8 syntypes (NMW; MNHUB); “Merdita Munela” / “Apfelb. Albanien”,

1 syntype (MNHUB); “Merdita Zebia”, 4 syntypes (MNHUB); “Merdita Zebia” / “Wgth.

1906 Merdita”, 10 s. (NMW); “Merdita Oroschi”, 10 s. (HNHM; NMW); “Derzë” / “Albania


Notes. Jedlička (1939: 52) stated that his P. malissorum ab. munella is identical with P. latifianus,

based on revision of type material, but subsequent authors (Schatzmayr, 1943a; Lorenz, 1998:

261) overlooked his opinion.

91.20.2. Pterostichus latifianus strupii J. Müller, 1935

References. Pterostichus latifianus strupii: Müller, 1935: 156 (Qaf e Stames, northeast of Kruja,

loc. typ.); Schatzmayr, 1943a: 109 (Qaf e Stames); Bousquet, 2003f: 485.

Notes. The ssp. struppi may be a distinct species since its type locality (Qaf e Stames Pass) is

situated in the geographically isolated massif of Skënderbe Mt.

91.21. Pterostichus lumensis Apfelbeck, 1905

91.21.1. Pterostichus lumensis lumensis Apfelbeck, 1905

References. Pterostichus lumensis: Apfelbeck, 1905: 242 (Albania / Kosovo border, Koritnik

Mt., loc. typ.); Apfelbeck, 1907: 637 (Koritnik Mt., loc. typ.); Apfelbeck, 1917: 225, note 1

(Koritnik Mt.); Apfelbeck, 1930: 169, note 1 (Koritnik Mt.); Schatzmayr, 1930: 255; Jedlička,

1936: 167 (Koritnik Mt.; Korab Mt.); Schatzmayr, 1943a: 108 (Koritnik Mt.). Pterostichus latifianus

ssp.?: Schatzmayr, 1930: 256 (Korab Mt.). Pterostichus lumensis lumensis: Drovenik & Peks, 1994:

67 (Koritnik Mt.); Drovenik & Peks, 1999: 78 (Koritnik Mt.); Bousquet, 2003f: 515.

Material studied. “Latif Koritnik”, 2 syntypes (NMW); “Latif Alban. spt.”, 1 s. probably

syntype (HNHM); “Latif Alban. spt.” / “P. lumensis Apfelb. Typen”, 1 s. probably syntype

(HNHM); “Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 3 s. (NMW); “Alban. Exp. Korab 1918

24.-31.VII.”, 16 s. (NMW); “Korab Albania Bischoff ”, 1 s. (NMW); “Bjeshka Matroxh”, 11 s.


(MNHUB); “Korab” / “Albania leg. Bischoff 1935”, more than 120 s. (MNHUB); “Gjallica”

/ “Albania leg. Bischoff 1936”, 14 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”,

more than 120 s. (MNHUB).

Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck,

1907b), and the older one is correctly treated as valid. Two specimens (1 m., 1 f.) labelled

“Latif Alban. sept.” and identified as Pterostichus djumanus Apfelbeck were found in DEI.

They are put provisionally here, since both specimens resemble P. lumensis. Pterostichus djumanus

Apfelbeck is nomen nudum.

91.21.2. Pterostichus lumensis litae Csiki, 1940

References. Pterostichus lumensis litae: Csiki, 1940: 216 (Gjalicë Mt., loc. typ.; Korab Mt., loc.

typ.); Schatzmayr, 1943a: 108 (Gjalicë Mt.; Korab Mt.); Bousquet, 2003f: 515.

Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 m. syntype

(HNHM); “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 4 syntypes (HNHM); “Albania

Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 21 syntypes (HNHM).

? Pterostichus lumensis ljubetensis Apfelbeck, 1908 [= chalybaeus Apfelbeck, 1908]

References. Pterostichus lumensis ljubetensis: Schatzmayr, 1930: 255; Bousquet, 2003f: 515.

Notes. Apfelbeck published three descriptions of this taxon (Apfelbeck, 1908; Apfelbeck,

1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. All data in the literature

and collections about ssp. ljubetensis come from the subalpine and alpine regions of the Shar

Planina Mts. that do not extend into Albania. Here, P. lumensis ljubetensis is treated as doubtful

for the fauna of Albania.

91.22. Pterostichus malissorum Apfelbeck, 1905 [= nigripes Jedlièka, 1936]

References. Pterostichus malissorum: Apfelbeck, 1905: 241-242 (Çukall Mt., loc. typ.); Apfelbeck,

1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1907b: 637 (Çukall Mt., loc.

typ.); Schatzmayr, 1930: 255; Jedlička, 1936: 169 (Malissora; Çukall Mt.; Cafa Valbones; Cafa

Pejs; Maja Pejs); Schatzmayr, 1943a: 108 (Çukall Mt.; Malissora; Cafa Valbones; Cafa Pejs;

Maja Pejs); Bousquet, 2003f: 515. Pterostichus latifianus ab. nigripes: Jedlička, 1936: 168 (North

Albanian Alps: Quf e Valbone, loc. typ.); Jedlička, 1939: 53. Pterostichus latifianus strupii (=

nigripes): Schatzmayr, 1943a: 109 (Quf e Valbone).

Material studied. “Latif Alban. sept.”, 4 s. (DEI; HNHM); “Latif Cukali” / “Coll.

Apfelbeck”, 1 m. syntype (HNHM); “Latif Cukali”, 2 syntypes (NMW); “Maranai Apfelb.”,

1 s. (HNHM); “Maranai Apf.”, 1 s. (HNHM); “Apfelb. Albanien”, 1 m. syntype (MNHUB);

“Apfelb. Albanien” / “Alban. spt. Maranaj”, 1 f. probably syntype (MNHUB); “Apfelb. Alban

spt” / “Maranaj Geb.”, 2 s. (HNHM); “Apfelb. Alban. spt. IV.V.’05” / “Maranaj”, 6 s. (NMW);

“Kunor Vukai Zukali Albanien Lona Giugno 1914”, 6 s. (NMW; MNHUB); “Cukale Albania

Bischoff ”, 12 s. (MNHUB); “Cukale-Geb.” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB);

“Çafa Stamës” / “leg. Strupi Albania 1934” / “Cotype” / “latifianus a. nigripes n. ab. det. Ing.

Jedlička”, 1 f. designated as syntype of Pterostichus latifianus ab. nigripes Jedlička, 1936

(MNHUB); “Cukale” / “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Çafa Valbonës” /

“Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja Pejs” / “Albania leg. Bischoff 1935”, 3

s. (MNHUB); “M. Mulecifit” / “Albania leg. Bischoff 1936”, 14 s. (MNHUB); “Maranaj” /

“Albania leg. Bischoff 1937”, 23 s. (MNHUB); “Boge ober D. Quelle” / “Albania leg. Mueller


1939”, 1 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Mueller 1939”, 10 s. (MNHUB);

“Maja e Dragomirit” / “Albania leg. Mueller 1939”, 4 s. (MNHUB); North Albanian Alps,

Maj’e Redohimes, 1900-2200 m, 28.V.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); North

Albanian Alps, Maj’e Redohimes, 2200-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg.

(NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov

leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron &

B. Petrov leg. (NMNHS).

Notes. Apfelbeck published two descriptions of this species (Apfelbeck, 1905; Apfelbeck,

1907b), and the older one is correctly treated as valid. Jedlička (1939: 52) stated that his P.

latifianus ab. nigripes Jedlička, 1936 is identical with P. malissorum Apfelbeck, 1905. Later,

Schatzmayr (1943a), however, synonymized P. latifianus ab. nigripes Jedlička, 1936 with P. latifianus

strupii J. Müller, 1935. I support the view of Jedlička because the type locality of the former is

quite close to the known findings of P. malisstorum, whereas P. latifianus strupii lives further in

the south: Pterostichus (Pterostichus) latifianus ab. nigripes Jedlička, 1936, syn. rest. of Pterostichus

(Pterostichus) malissorum Apfelbeck, 1905.

91.23. Pterostichus pentheri Apfelbeck, 1918

References. Pterostichus pentheri: Apfelbeck, 1918c: 161, 166 (North Albanian Alps, loc. typ.);

Schatzmayr, 1930: 255; Jedlička, 1936: 168 (North Albanian Alps; Shkëlzen Mt.; Maja e Jezercë

Mt.); Schatzmayr, 1943a: 108 (North Albanian Alps; Shkëlzen Mt.; Maja e Jezercë Mt.); Bousquet,

2003f: 515.

Material studied. “Albania coll. Lichtn.”, 2 s. (HNHM); “Prokletija Albanien Dr. Penther”, 2

syntypes (NMW); “Prokletija” / “Alban. montenegr. Grenze Penther’14”, 5 syntypes (DEI;

NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 5 syntypes

(NMW); “Maja e Jezercë” / “Albania leg. Bischoff 1931”, 7 s. (MNHUB); “Maja e Jezercë

Çafa Pejs-Çafa Gstars” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Çafa e Markofçes” /

“Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Maja e Jezercë” / “Albania leg. Bischoff 1935”,

12 s. (MNHUB); more than 90 s. from locality “Shkëlzën” (MNHUB); North Albanian Alps,

Maj’e Redohimes, 2400-2550 m, 29.V.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS).


Rubës, 2000 m, 8.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2250 m,

11.VII.1993, 2 m., 2 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2000 m, 14.VII.1993, 2

m., A. Gobetti leg. (cAVT).

91.24. Pterostichus reiseri Ganglbauer, 1889

References. Pterostichus reiseri: Apfelbeck, 1918c: 160, 166 (North Albanian Alps, Vermosh,

Grebeni, 1100-1700 m); Jedlička, 1939: 52; Schatzmayr, 1943a: 108 (Grebeni).

Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 2 s.

(NMW); “Vermoša Albanien Dr. Penther”, 2 s. (NMW); “Vermosa 20-21.VII.” / “Alban.

montenegr. Grenze, Penther’14”, 1 s. (NMW).

Notes. Bousquet (2003f: 516) has omitted P. reiseri for Albania. Here the species is confirmed

for the country. The record “Albanien: Žljeb plan.” (Jedlička, 1936: 168) refers to the massif

shared among Montenegro and the Kosovo Region of Serbia.



? Pterostichus epiroticus Csiki, 1930 [= corax Ganglbauer, 1889]

References. Pterostichus epiroticus: Schatzmayr, 1943a: 107.

Notes. The record “Peristerigebirge bei Jannina (Albanien)” (Schatzmayr, 1943a) is based on an older

record of Apfelbeck (1904: 267 sub P. corax). Currently this record concerns the territory in

Northwestern Greece. In NMW, besides specimens from Central Greece (Oeta Mt.; Korax Mt.), I

found five recently collected specimens from the region near to Joannina and Metsovon, which is very

close to the Albanian territory. Therefore, P. epiroticus is treated here as likely to be found in Albania.

92. Genus Speluncarius Reitter, 1886

Subgenus Hypogium Tschitschérine, 1900 [= Hypogeobium Tschitschérine, 1903]

92.1. Speluncarius albanicus (Tschitschérine, 1900)

References. Platysma albanicum: Tschitschérine, 1900: 49-51 (“Albanie”, loc. typ.). Pterostichus

albanicus: Apfelbeck, 1904: 236-237; Schatzmayr, 1930: 329. Tapinopterus albanicus: Reitter, 1914:

262; Müller, 1937: 134; Schatzmayr, 1943a: 131; Jeannel, 1950: 157, 159-160. Hypogeobium

albanicum: Jeannel, 1950: 160, 162; Jeannel, 1953: 11. Speluncarius albanicus: Sciaky, 1982: 28;

Casale & Giachino, 1991: 215-216; 219; Bousquet, 2003f: 516.

Material studied. “Albania” / G.C.Champion Coll. B.M.1927-409” / “Platysma (Hypogium)

albanicum m. Typ. Tschitscherin det” / “syntype” (round blue coloured label), 1 f. (BMNH).

Notes. The type locality of S. albanicus remains not yet specified.

Subgenus Speluncarius s.str.

92.2. Speluncarius minutulus J. Müller, 1937

References. Tapinopterus minutulus: Müller, 1937: 132-123 (Shkëlzen Mt., loc. typ.); Schatzmayr, 1943a:

128 (Shkëlzen Mt.); Turin, 1981: 127. Speluncarius minutulus: Jeanne, 1982: 41-42 (Shkëlzen Mt.); Sciaky,

1982: 20; Drovenik & Peks, 1994: 68; Drovenik & Peks, 1999: 79; Bousquet, 2003f: 516.

Material studied. “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 f. designated as paralectotype

by P.M. Giachino (MNHUB).

Notes. Drovenik & Peks (1994) and Drovenik & Peks (1999) noted S. minutulus for Montenegro

by mistake because it is a local endemic species for Northeast Albania.

93. Genus Stomis Clairville, 1806

Subgenus Stomis s.str.

93.1. Stomis pumicatus (Panzer, 1796)

93.1.1. Stomis pumicatus pumicatus (Panzer, 1796)

References. Stomis pumicatus: Apfelbeck, 1904: 213 (Velipoja); Hieke & Wrase, 1988: 68;



Material studied. “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Bojana

Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB).

93.2. Stomis rostratus (Duftschmid, 1812)

93.2.1. Stomis rostratus rostratus (Duftschmid, 1812)

References. Stomis rostratus: Apfelbeck, 1918c: 166 (North Albanian Alps, Grebeni near

Vermosh, 1100-1800 m); Magistretti, 1965: 362; Bousquet, 2003f: 518.

Material studied. “Grebeni 16.VI.” / “Alban. montenegr. Grenze, Penther’14”, 3 s. (NMW);

“Alban. Vermosa Penther”, 2 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr.

Grenze, Penther’14”, 4 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze,

Penther’14”, 4 s. (NMW); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).

94. Genus Tapinopterus Schaum, 1858

Subgenus Tapinopterus s.str.

Notes. The species from this subgenus are placed here in three groups of species in agreement

with a forthcoming arrangement proposed by P. M. Giachino and L. Picciau (personal

communication).

I. South Aegean groups.

Species group “extensus”

# Tapinopterus extensus (Dejean, 1831)

# Tapinopterus extensus abreshensis (Lutshnik, 1916) [= convexiusculus Apfelbeck, 1904]

References. Pterostichus extensus convexiusculus: Apfelbeck, 1904: 251 (“Turcia – Abresch”, loc.

typ.); Schatzmayr, 1943a: 127.

Notes. According to Schatzmayr (1943a) the type locality of this race (“Abresch”) must be

somewhere between Albania and Palestine. Bousquet (2003f: 519) has noted the subspecies

for Greece. Based on Apfelbeck’ short description (Apfelbeck, 1904: 251 sub Pterostichus extensus

convexiusculus), I suppose that ssp. abreshensis is not subordinate to T. extensus s.l., taking into

consideration also the fact that Apfelbeck (1904) compared the former with T. kaufmanni

Ganglbauer, 1897. Dealing with the taxa of Tapinopteris extensus s.l., Mařan (1939b: 298, 303)

without hesitation separated T. extensus abreshensis from the remaining races of the species.

Only a revision of Apfelbeck’s type (probably in Museum Sarajevo) could establish the

systematic position of the taxon in question. Presently, both the species and subspecies are

removed from the list of the Albanian Carabidae.

94.1. Tapinopterus extensoides Straneo, 1936 [= extensoides Jedlièka, 1936]

References. Tapinopterus extensoides Straneo: Straneo, 1936: 155-156 (Llogora, typ. loc.); Müller,

1937: 132, note 1 (Llogora; cave “Paftali” near Berat; Shëndelli Mt.); Schatzmayr, 1943a: 127

(Llogora; cave “Paftali” near Berat; Shëndelli Mt.). Tapinopterus extensoides Jedlička: Jedlička,

1936: 169 (Llogora, loc. typ.); Turin, 1981: 127. Tapinopterus extensus extensoides “Straneo et


Jedl.”: Mařan, 1939b: 295-299, 302-303 (Llogora). Tapinopterus extensus extensoides Jedlička:


Material studied. “Logara Juni” / “Albania leg. Bischoff 1932”, 1 m. syntype (NMW); “Maja

e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Reza e Kanalit Logara, Alban. m.”

/ “Albania leg. Bischoff 1932”, 1 f. syntype (NMW); “Logara juni” / “Albania leg. Bischoff

1932”, 3 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 7 s. (MNHUB); “Llogora,

Alb. Mer.” / “Albania leg. Bischoff 1935”, 1 f. syntype of extensoides Jedlička, 1936 (MNHUB);

“Llogora” / “leg. Bischoff Albania 1936”, 17 s. (MNHUB); “Shen Ili” / “leg. Bischoff Albania

1936”, 1 s. (MNHUB); “Borshe” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); “Shpella Buz

i Lugit Mal i Piqs” / “Borshe” / “Albania leg. Bischoff 1940”, 2 s. (MNHUB); “Albania,

Periferi Tepelenë, a gorge 1.5 km W of Progonat,” / “785 m, 9.X.2004, Z. Fehér, J. Kontschán

& D. Murányi (No. 133)”, 2 f. (HNHM).

Notes. In 1936 Jedlička (1936) and Straneo (1936) described T. extensoides having at disposal

identical specimens collected by Bischoff from the region of Llogora. Later, Müller (1937:

132, note 1) stated that the Straneo’s description has priority over that of Jedlička. This opinion

is correct in spite of Mařan’s statement that both are authors of this taxon (Mařan, 1939b:

302). It is Mařan who first regarded extensoises as subspecies of T. extensus. However, in my

view T. extensoides Straneo is clearly distinct from all the other subspecies of T. extensus by an

evident apomorphic character – 2 (instead 1) posterior supraorbital setiferous pores (see also

Straneo, 1936). The check of all specimens at hand demonstrated that this character is constant

(as only a single female from HNHM has a single left and two right posterior supraorbital

pores) so that it may define a separate species: Tapinopterus (Tapinopterus) extensoides (Straneo,

1936), stat. rest. from Tapinopterus (Tapinopterus) extensus extensoides (Straneo, 1936).

Species group “protensus”

94.2. Tapinopterus protensus (Schaum, 1857)

Notes. Bousquet (2003f: 520) noted that T. protensus aetolicus Ganglbauer, 1889 inhabits

Montenegro, and not Albania. In spite of that Schatzmayr (1943: 122) cited it for the studied

country (see below). Here, the species records and reference data from Albania are referred to

two defined and two undefined subspecies. The occurrence of T. protensus in Montenegro

needs serious confirmations.

94.2.1. Tapinopterus protensus peristericus Apfelbeck, 1901

References. Tapinopterus aetolicus peristericus: Müller, 1935: 153 (Shëndelli Mt. near Tepelenë).

Material studied. “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s.

(MNHUB); “Maja e Policanit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 6 s. (MNHUB);

“Shen Ili” / “leg. Bischoff Albania 1936”, 3 s. (MNHUB); “Maja e Policanit” / “Albania leg.


Notes. This subspecies is confined to the area between the valleys of Drina River and Vijosë

River and lives also in Northwestern Greece. Müller (1935: 153-154) noted the identity of

syntypes of T. peristericus Apfelbeck, 1901 with specimens from the Shëndelli Mt. In MNHUB,

I found specimens from Shëndelli Mt. labelled “aetolicus sbsp. peristericus det. J. Müller”, and

others - “aetolicus subsp. nova m. det. J. Müller”.


94.2.2. Tapinopterus protensus tomoricensis J. Müller, 1935

References. Tapinopterus aetolicus tomoricensis: Müller, 1935: 153-154 (Tomor Mt., loc. typ.).

Tapinopterus thessalicus tomoricensis: Mařan, 1940b: 59 (Tomor Mt.). Tapinopterus protensus tomoricensis:

Schatzmayr, 1943a: 122 (Tomor Mt.); Bousquet, 2003f: 520.

Material studied. “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 2 syntypes

(NMW); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 8 syntypes (NMW);

40 s. from Tomor Mt. (MNHUB).

! Tapinopterus protensus ssp.1

Material studied. “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 1 m. (MNHUB).

Notes. Josef Müller has noted on the label that the specimen from “Mal i Dajtit” belongs to

an undescribed taxon. This specimen has gigantic body dimensions (21.5 mm) in comparison

with the other races from Albania, but more material is needed for the correct classification

of this local population.

! Tapinopterus protensus ssp.2

References. Tapinopterus aetolicus: Müller, 1935: 154 (Reza e Kanalit Mt. near Llogora).

Tapinopterus protensus aetolicus: Schatzmayr, 1943a: 122 (Reza e Kanalit Mt. near Llogora).

Material studied. “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Logara” / “leg.

Bischoff Albania 1934”, 2 s. (MNHUB); “Maja e Tartarit” / “leg. Bischoff Albania 1934”, 1 s.

(MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 7 s. (MNHUB).

Notes. This still not defined race inhabits the mountains west of the valleys of Vijosë River

and Drina River. Müller has identified the specimens from “Llogara” as ssp. aetolicus, and that

from “Maja e Tartarit” as “aetolicus sbsp. valonensis m. det. J. Müller”. Both populations from

Albania may not belong to the ssp. aetolicus (as Müller stated, 1935), if considering the

geographical factor, since the type locality of ssp. aetolicus is the Korax Mt. (Central Greece),

situated nearly 270 km south of the area of Llogora.

II. North Aegean groups.

Species group “balcanicus”

94.3. Tapinopterus bischoffi J. Müller, 1936

References. Tapinopterus bischoffi: Müller, 1936: 106-107 (Mal i Dajtit Mt., loc. typ.); Schatzmayr,

1943a: 127 (Mal i Dajtit Mt.); Turin, 1981: 127; Bousquet, 2003f: 519.

Material studied. “Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Mal

i Dajtit” / “leg. Bischoff Albania 1934”, 6 s. (MNHUB); “Mal i Drini” / “leg. Bischoff Albania

1934” / “cotype” / “Tapinopterus bischoffi m. det. J. Müller”, 1 m. (NMW); “Rushkull” / “Albania

leg. Bischoff 1935”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll. Bischoff ”,

5 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Krujes”

/ “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1939”,

7 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); “AL (Kruje)

Paß ca. 5 km ne. Kruje 700-900 m, 1.V.1993 leg. Heinz & Sonja Peks”, 1 f. (MNHUB); Mal i

Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 m., 2 f., L. Penev leg. (NMNHS).


# Tapinopterus comita Jedlička, 1935

References. Tapinopterus eomita sic!: Turin, 1981: 127. Tapinopterus comita: Bousquet, 2003f: 519.

Notes. I do not find any exact records in the literature or material in the collection of this

species from Albania, thus the indications of Turin (1981) and Bousquet (2003f) for the country

may be treated as incorrect. T. comita inhabits only the Pelister Mt. (Jedlička, 1935; Drovenik &

Peks, 1994: 68), a massif shared between Macedonia and Greece. The species is removed

from the list of the Albanian Carabidae.

94.4. Tapinopterus detonii Schatzmayr, 1943

References. Tapinopterus detonii: Schatzmayr, 1943b: 110 (Mal i Krujes Mt., 1000 m, loc. typ.);

Turin, 1981: 127; Bousquet, 2003f: 519.

Material studied. “Albania”, 1 s. (DEI).

Notes. Further taxonomic work is needed to demonstrate the differentiation of T. detonii

from T. bischoffi J. Müller, 1936, since the former seems not well defined and may be a subspecies

or synonym of the latter.

94.5. Tapinopterus dochii (Apfelbeck, 1906)

References. Pterostichus dochii: Apfelbeck, 1906: 1663-1664 (Merdita: Mal i Shenjt Mt., loc. typ.;

Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 499, 502 (Mal i Shenjt Mt.: top region, 1500 m).

Tapinopterus dochii: Jedlička, 1935: 186 (Korab Mt.); Müller, 1936: 107 (Korab Mt.); Schatzmayr,

1943a: 126 (Mal i Shenjt Mt.; Zepë Mt.; Munella Mt.); Hieke, 1981: 81; Bousquet, 2003f: 519.

Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05” / “Tapinopterus dochii

Apf.”, 1 f. syntype (NMW); more than 100 s. from localities “Berishe”, “Cucale”, “Maja e

Shpatit”, “Cafa e Sharit”, “Maja Shales Vogel Iballe”, “Korab”, “Munela”, “Derzë”, “Taja”,

“M. Mulecifit”, “Merdita M. Sheit” (MNHUB).

[Tapinopterus miridita (Apfelbeck, 1904)]

Notes. The only records for T. miridita s.l. from Albania are those of Hieke (1981: 81) and

Bousquet (2003f: 520). However, the present author cannot find any exact data in the literature

or collections to prove the species occurrence in Albania. The type locality “Sveti Troice”

(Apfelbeck, 1904: 245), or more correct “Sveta Trojica”, concerns a monastery near the village

of Mušutište, a place situated in the northern slopes of the Shar Planina Mt. (in the Kosovo

Region of Serbia) (see also Jedlička, 1935: 186). Oddly enough, the “loc. typ.” and known

records of the species are quite removed from the region of Mirdita (situated in the south part

of North Albania, south of the Drin River), after which the species was named. T. miridita is

treated here as likely to be found in the country.

[Tapinopterus miridita miridita (Apfelbeck, 1904)]

Notes. The occurrence of this race in the marginal northeastern regions of Albania (the

Koritnik Mt.; the Gjalicë Mt.) is possible.

[Tapinopterus miridita jablanicensis Mařan, 1935]

Notes. This race was described from the Macedonian part of the Jablaniča Mt. (shared between

Albania and Macedonia), but so far the subspecies is not properly cited for Albania.


# Tapinopterus miridita jacupicensis Jedlička, 1935

Notes. Bousquet (2003f: 520) cited this subspecies for Albania. However, this taxon is a local

endemic for the massif of the Jakupiča Mt. situated in the central part of Republic of

Macedonia. Probably the author in question has had in mind the spp. jablanicensis when he

mentioned ssp. jacupicensis for Albania. The latter subspecies is here excluded from the list of

the Albanian Carabidae.

94.6. Tapinopterus monastirensis Reitter, 1913

94.6.1. Tapinopterus monastirensis gestroi J. Müller, 1923

References. Tapinopterus gestroi: Müller, 1923: 110-111 (Tomor Mt.: Toik, loc. typ.). Tapinopterus

monastirensis gestroi: Müller, 1936: 107; Mařan, 1939e: 82, 84 (Tomor Mt.: Toik); Schatzmayr,

1943a: 126 (Tomor Mt.); Bousquet, 2003f: 520.

Material studied. “Albania”, 1 s. (DEI); “Tomorica Kulmak” / “Albania 1922 Ravasini,

Lona”, 1 s. (NMW); “Tomor: Kulmak Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1

s. (NMW); “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW);

“Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 4 s. (NMW); “Tomor-

Tojk juli 1932” / “Albania leg. Bischoff 1932”, 32 s. (MNHUB); “Tomor” / “Albania leg.


Tribe SPHODRINI Laporte, 1834

Subtribe ATRANOPSINA Baehr, 1982

95. Genus Platyderus Stephens, 1827

Notes. The present knowledge about the Balkan representatives of this genus poorly

corresponds to the actual status of the species.

Subgenus Platyderus s.str.

95.1. Platyderus dalmatinus L. Miller, 1881

95.1.1. Platyderus dalmatinus dalmatinus L. Miller, 1881

References. Platyderus dalmatinus: Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme,

300 m, and Vorfai Siperme, 700 m).

Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 m. (NMW).

Notes. Hovorka & Sciaky (2003a: 522) have omitted this species for Albania. Here, it is

confirmed for the fauna of Albania.

95.2. Platyderus graecus (Reiche & Saulcy, 1855)

References. Platyderus graecus: Apfelbeck, 1909: 297 (Vlorë).

Notes. Hovorka & Sciaky (2003a: 522) have omitted this species for Albania. Further study is

needed to confirm its occurrence in the country.


95.3. Platyderus minutus (Reiche & Saulcy, 1855)

95.3.1. Platyderus minutus obtusangulus J. Müller, 1923

References. Platyderus minutus: Apfelbeck, 1907a: 498, 501 (Mal i Shenjt Mt.: top region, 1500

m). Platyderus minutus obtusangulus: Müller, 1923: 112 (Tomor Mt.: Kulmak, loc. typ.); Hovorka

& Sciaky, 2003a: 523.

Material studied. “Albania”, 1 f. (BMNH).

95.4. Platyderus rufus (Duftschmid, 1812)

95.4.1. Platyderus rufus rufus (Duftschmid, 1812)

References. Platyderus rufus: Apfelbeck, 1904: 279 (Velipoja near Lezhë); Hieke & Wrase, 1988:

89. Platyderus rufus rufus: Hovorka & Sciaky, 2003a: 523.

Material studied. Nearly 50 s. from different localities (MNHUB); Rojan, 6.III.1961, 1 f. (MNST).

Subtribe CALATHINA Laporte, 1834

96. Genus Calathus Bonelli, 1810

Subgenus Bedelinus Ragusa, 1885

96.1. Calathus circumseptus Germar, 1824

References. Calathus circumseptus: Hovorka & Sciaky, 2003b: 524.

Material studied. “Albania”, 1 s. (HNHM); “Durazzo”, 1 s. (HNHM); “Apfelb. 1893 Durazzo”,

1 s. (HNHM); “Berat” / “Albania leg. Bischoff 1931”, 4 s. (MNHUB); “Tepelene juni 1932” /

“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Lum i Erzen” / “leg. Bischoff Albania 1934”,

1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania

leg. Bischoff 1935”, 3 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB);

“Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); Saranda District, Qeparo,

4-5.V.1994, 2 s., P. Stoev leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg.

(NMNHS); Kruja Castle, 16.XI.1997, 2 s., R. Ward leg. (CMNH).


Subgenus Calathus s.str.

96.2. Calathus bosnicus Ganglbauer, 1891

References. Calathus bosnicus: Hovorka & Sciaky, 2003b: 524.

Material studied. “Prokletija 1300-2000 m” / “Alban.-montenegr. Grence, Penther’14”, 2 s.

(NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Gjalica Ljums 17-26.VI.”, 1

s. (NMW); “Llogora…”, 1 s. (NMW); 60 s. from localities “Shkëlzën”, “Çafa Markofçes”,

“Maja Pejs”, “Korab”, “Maja e Jezercoë”, “Valbona Çerem” (MNHUB); North Albanian Alps,

Bogë, 1000-1100 m, 5-9.VI.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS).

Notes. First detailed records for Albania. The record “Llogora…” either concerns another congener

or is “patria falsa”, since the locality is quite removed in the south from the range of the species.


[Calathus distinguendus Chaudoir, 1846]

Notes. According to Battoni & Vereschagina (1984: 141) this species may live in Albania, but

so far this statement is not proved.

96.3. Calathus ellipticus Reitter, 1889

96.3.1. Calathus ellipticus ellipticus Reitter, 1889

Material studied. “Çafa e Dhembellit”, 2 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff

1932”, 26 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB);

“Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Maja e Policanit” / “Albania leg.


Notes. New species for the fauna of Albania. After a comparison of type material with

specimens from Albania, D.W. Wrase (personal communication) concluded that they are

conspecific, being in agreement with specimens from Western Greece. A revision of all related

taxa involved (including also C. giganteus and C. korax) is needed to clarify the status of the

Albanian population.

96.4. Calathus fuscipes (Goeze, 1777)

96.4.1. Calathus fuscipes graecus Dejean, 1831 [= latus (Audinet-Serville, 1821) nec Linnaeus, 1758]

References. Calathus fuscipes: Apfelbeck, 1904: 284 (Vlorë; Berat); Apfelbeck, 1909: 297 (Vlorë;

Berat); Csiki, 1940: 217 (Shkëlzen Mt.; Kula Ljums); Csiki, 1943: 214; Schatzmayr, 1943b: 110

(“Poggio Boschetto”; Kruja; Dukati); Battoni & Vereschagina, 1984: 152; Hieke et al., 1986:

189 (Tiranë; Tomor Mt.: Abbas Ali Monastery; Bizë near Shëngjergji; Lurë e Vjetër near Lura

Mt.); Hieke & Wrase, 1988: 89. Calathus fuscipes var. graecus: Wohlberedt, 1909: 127 (around

Liqen i Shkodrës). Calathus fuscipes graecus: Schatzmayr, 1937: 27 (Gjirokastër); Hovorka &

Sciaky, 2003b: 525. Calathus fuscipes var. latus: Csiki, 1940: 217 (Gjalicë Mt.; Korab Mt.). Calathus

fuscipes fuscipes: Hovorka & Sciaky, 2003b: 525.

Material studied. More than 180 s. from different localities (MNHUB); L. Selitës, 8-9.VII.1961,

5 s. (MNST); Lurë, 18-25.VII.1961, 2 s. (MNST); Theth, 1-8.VIII.1961, 2 f. (MNST); Ibë,

14.VII.1962, 1 f. (MNST); Shkodër, 15.V.1963, 1 m.; Mal i Dajtit Mt., 10-11.V.1965, 1 m., 1 f.

(MNST); 2 km N Durrës, 24.V.1993, 1 m., 1 f., P. Stoev & D. Zaprjanova leg. (NMNHS);

North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 f., P. Beron & B. Petrov

leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 f., P. Beron & B.

Petrov leg. (NMNHS); Reshen, 11.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Cafasan,

1230 m, 27.VII.1993, 3 m., P. Stoev & D. Zaprjanova leg. (NMNHS); Tiranë, botanical garden,

8.V.1995, 1 m, 1 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Vlorë District, Dukati Mt.,

450 m, 11.V.1995, 2 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m,

11.V.1995, 2 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 /

8.II.1998, 5 s., R. Ward leg. (CMNH).

Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., 3 f., A.

Paparistos leg. (cAVT).

Notes. According to A. Vigna Taglianti (personal communication) the populations from Albania

belong to ssp. graecus. Besides the material studied above, in NMW I have seen more specimens,



96.5. Calathus giganteus Dejean, 1828

References. Calathus giganteus: Apfelbeck, 1904: 282 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);

Schatzmayr, 1937: 2, 6; 21 (Vlorë; Sopoti near Gjirokastër); Hovorka & Sciaky, 2003b: 525.

Material studied. “Llogora juni” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Tepelene

juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania

1934”, 1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Maja e Konispol

Seraqin” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Llogora

Pass, 1025 m, 11.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS).

Additional material included (studied S. Hristovski and D.W. Wrase). “Logara” / “Albania,

leg Bischoff 1932”, 1 s. (MNHUB); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006,


? Calathus glabricollis Dejean, 1828

Material studied. “coll. L.W. Schaufuß” / “Calathus glabricollis Albanien verus”, 1 m. (MNHUB).

Notes. It is not sure if the record above concerns the present territory of Albania, so the

species is treated here as doubtful for the country.

96.6. Calathus korax Reitter, 1889

References. Calathus corax: Schatzmayr, 1937: 2, 7; 21 (Gjirokastër).

Notes. Hovorka & Sciaky (2003b: 525) have omitted C. korax for Albania. Further collections

are needed to confirm its occurrence in the country.

96.7. Calathus ravasinii J. Müller, 1935

96.7.1. Calathus ravasinii ravasinii J. Müller, 1935

References. Calathus ravasinii: Müller, 1935: 152-153 (Tomor Mt.: Kulmak, loc. typ.); Schatzmayr,

1937: 2, 7, 22 (Tomor Mt.); Turin, 1981: 136; Hovorka & Sciaky, 2003b: 526.

Material studied. “M. Toik Alban. mer.”, 1 s. probably syntype (NMW); “Tomorica Kulmak”

/ “Albania 1922 Ravasini, Lona”, 3 syntypes (NMW); “Tomor” / “Albania leg. Bischoff 1932”,

22 s. (MNHUB).

Notes. Schatzmayr (1937: 22) treated C. ravasinii as separate species, but hinted that it resembles

C. macedonicus. Based on the material studied, the present author considers the two taxa are only

well-defined subspecies of one species: Calathus (Calathus) ravasinii macedonicus Mařan, 1935, stat.

nov. for Calathus (Calathus) macedonicus Mařan, 1935. Müller (1935) published his description in

Bollettino della Società Entomologica Italiana, volume 67; this issue was printed on 26.II.1935.

On the other hand, Mařan (1935) described C. macedonicus in Sborník Entomologického Odděleni

Národního Musea v Praze, volume 13; this issue was printed on 1.XII.1935. The nominotypical

taxon is easily recognized from ssp. macedonicus by the anteriorly narrowed sides of pronotum.

96.7.2. Calathus ravasinii macedonicus Mařan, 1935

References. Calathus macedonicus: Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 525.


“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); “Bjeshka Matroxh” /


“Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 23 s.

(MNHUB); “Teja” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkalla Harushe” / “Korab”



Subgenus Neocalathus Ball & Nègre, 1972

96.8. Calathus albanicus Apfelbeck, 1906

References. Calathus albanicus: Apfelbeck, 1906: 1666 (Merdita: Zepë Mt., loc. typ.); Apfelbeck,

1907a: 500, 503; Schatzmayr, 1937: 2, 8; 36 (Zepë Mt.); Csiki, 1940: 218 (Gjalicë Mt.; Korab

Mt.); Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 526.

Material studied. More than 160 s. with three kinds of labeling: “Albania Galica Lums” /

“Akad. Balk. Exp. Csiki 1917” (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp.

Csiki 1918” (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918” (HNHM); more

than 150 s. from the massifs of Shar Planina Mt., Merdita, Munella Mt., Korab Mt., Gjalica

Ljums and others (NMW); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 9 s. (MNHUB);

“Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff

1935”, 22 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Mal i

Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Fusha Korabit” / “Albania leg. Bischoff

1936”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 12 s. (MNHUB); “Berishe”

/ “Albania leg. Mueller 1939”, 2 s. (MNHUB); “Maja Shalles i Vogel-Ibale” / “Albania leg.

Mueller 1939”, 1 s. (MNHUB); “Kunora Drunit Iballe” / “Albania leg. Bischoff 1939”, 1 s.

(MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB).

Notes. This species inhabits the mountain chain Shar Planina Mt. - Korabi Mt. and massifs in

the area of Mirdita, but not the North Albanian Alps, where C. albanicus most probably is

replaced by C. metallicus.

96.9. Calathus ambiguus (Paykull, 1790)

96.9.1. Calathus ambiguus ambiguus (Paykull, 1790)

Material studied. “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”,

3 s. (MNHUB).

Notes. The mention of Hieke & Wrase (1988: 90) for Albania is apparently result of following

the record “Prisren” (Apfelbeck, 1904: 285). Currently this locality is situated in the Kosovo

Region of Serbia. Hovorka & Sciaky (2003b: 526) have most likely followed one of these

sources too. Since there is no previous reliable record from Albania, C. ambiguus is treated here

as new species for the fauna of the country.

96.10. Calathus cinctus Motschulsky, 1850 [= erythroderus Gemminger & Harold, 1868]

References. Calathus mollis f. erythroderus: Schatzmayr, 1943b: 111 (Lushnje). Calathus cinctus:

Aukema, 1990: 138.

Material studied. “Mal i Daitit” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Mal i Krujes”

/ “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”,

7 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Mal i Daitit”


/ “Albania 1936 leg. Ramadan coll. Bischoff ”, 4 s. (MNHUB); Kruja Castle, 16.XI.1997, 1 s.,

R. Ward leg. (CMNH).

Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A.

Paparistos leg. (cAVT).

Notes. Hovorka & Sciaky (2003b: 527) have omitted C. cinctus for Albania. Here the species is


96.11. Calathus erratus (C.R. Sahlberg, 1827)

96.11.1. Calathus erratus erratus (C.R. Sahlberg, 1827)

References. Calathus erratus: Hieke & Wrase, 1988: 90. Calathus erratus erratus: Hovorka &

Sciaky, 2003b: 527.

Material studied. “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Shkëlzën”



900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT).

Notes. First detailed records for Albania. In addition to the material studied above, the present

author has seen several further specimens in NMW, which remained unregistered.

96.12. Calathus melanocephalus (Linnaeus, 1758)

96.12.1. Calathus melanocephalus melanocephalus (Linnaeus, 1758)

References. Calathus melanocephalus: Csiki, 1940: 218 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.);

Hieke et al., 1986: 189 (Bizë near Shëngjergji; Lurë e Vjetër near Lura Mt.); Aukema, 1990:

138. Calathus melanocephalus melanocephalus: Hovorka & Sciaky, 2003b: 527.

Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); 30 s.

from localities “Shkëlzën”, “Bjeshka Matroxh”, “Tomor”, “Korab”, “Koritnik”, “Gjallica”,

“Mal i Shenjit”, “Boge”, “Cucale”, “Taja”, “Llogora”, “Cafa e Sherit Iballe”, “Çafa Markofçes”

(MNHUB); Tomor Mt., 21.IX.1961, 1 m., 1 f. (MNST).



Notes. The mention of Hieke & Wrase (1988: 91) for Albania refers apparently to the record

“Prisren” (Apfelbeck, 1904: 286). Actually this locality is presently situated in the Kosovo

Region of Serbia. Besides the specimens above, I have seen several further specimens in

NMW, which remained unregistered.

96.13. Calathus metallicus Dejean, 1828

96.13.1. Calathus metallicus prokletijensis Mařan, 1938

References. Calathus metallicus: Hieke & Wrase, 1988: 90. Calathus metallicus metallicus: Hovorka

& Sciaky, 2003b: 527.


(NMW); nearly 100 s. from localities “Shkëlzën”, “Çafa Markofçes”, “Maja e Jezerze”

(MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., 1 f., P. Beron & B.

Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., P. Beron




2250 m, 11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT).

Notes. First detailed records for Albania. The absence of C. metallicus in the massifs of Shar

Planina and Korab Mt. may be a result of its replacement by C. albanicus there.

96.14. Calathus mollis (Marsham, 1802)

96.14.1. Calathus mollis mollis (Marsham, 1802)

References.Calathus mollis: Apfelbeck, 1904: 285 (Vlorë; Berat); Apfelbeck, 1909: 297 (Vlorë;

Berat); Hieke & Wrase, 1988: 91.

Material studied. “Albania Avlona v. Oertzen”, 1 s. (MNHUB).

Notes. Hovorka & Sciaky (2003b: 528) have omitted C. mollis for Albania. Here the species is

confirmed for the country. C. mollis is exclusively a littoral species and most probably the record

from Berat (Apfelbeck, 1904; 1909) concerns C. cinctus (A. Vigna Taglianti, personal communication).

97. Genus Synuchidius Apfelbeck, 1908

97.1. Synuchidius ganglbaueri Apfelbeck, 1908

References. Synuchidius ganglbaueri: Csiki, 1940: 218 (Gjalicë Mt.; Korab Mt.); Hieke, 1981: 82;

Hovorka & Sciaky, 2003b: 529.

Material studied. 66 s. from three kinds of labeling: “Albania Korab” / “Akad. Balk. Exp. Csiki

1917” (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918” (HNHM);

“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918” (HNHM); “Alban. Exped. Gjalica Ljums

17-26.VI.”, 12 s. (NMW); “Alban. Exped. Korab 1918 24-31..VII.”, 19 s. (NMW); “Jablanica coll.

Lichtneckert”, 2 s. (HNHM); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB);

“Shkalla Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB).

Notes. Apfelbeck published three consecutive descriptions for the subgenus and species (Apfelbeck,

1908; Apfelbeck, 1917; Apfelbeck, 1930), and the oldest ones are correctly treated as valid.

Subtribe SPHODRINA Laporte, 1834

98. Genus Laemostenus Bonelli, 1810

Subgenus Antisphodrus Schaufuss, 1865

98.1. Laemostenus cavicola (Schaum, 1858)

98.1.1. Laemostenus cavicola albanicus J. Müller, 1914

References. Laemostenus cavicola albanicus: Müller, 1914: 1005-1006 (Çukall Mt.: cave “Scpedha n’

kunor Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha Nre Gegs n’ gur t’barzodhs”, loc. typ.); Pretner,

1978: 135, 148 (caves in Çukall Mt.; near Shkodër); Genest & Juberthie, 1994: 593 (Çukall Mt.:

cave “n’Kunor Vukai”, cave “Nre Gegs n’gur t’barzodhs”); Casale, 1988: 708, 716, 720 (Çukall

Mt.); Casale, 2003: 534. Laemostenus cavicola var. modestus: Apfelbeck, 1918c: 168 (Rapshë, 700-

1400 m). Laemosthenes cavicola: Jeannel, 1937b: 89; Magistretti, 1965: 439. Laemostenus cavicola modestus:

Pretner, 1978: 135, 148, 168 (Rapshë); Casale, 1988: 719 (Rapshë).


Material studied. “Albanie”, 1 s. (MNHUB); “Albanien”, 1 s. (MNHUB); “Scpedha n’Kunor

Zukali Albanien Lona Giuno 1914”, 1 syntype (DEI); “Prokletija 1300-2000 m” / “Alban.

montenegr. Grenze Penther’14”, 12 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze

Penther’14”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze

Penther’14”, 3 s. (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze Penther’14”, 2 s.

(NMW); “Albania Rapsa Dr. Grabowski”, 6 s. (HNHM); “Shpella Kodra Çarit” / “Cukale”, 3 s.

(MNHUB); “Brigie Hot” / “leg. Bischoff Albania 1934” / “Shpella e Lugstit”, 1 s. (MNHUB).

# Laemostenus cavicola modestus (Schaufuss, 1862)

References. Laemostenus cavicola var. modestus: Apfelbeck, 1918c: 168 (Rapshë, 700-1400 m).

Laemostenus cavicola modestus: Pretner, 1978: 135, 148, 168 (Rapshë); Casale, 1988: 719 (Rapshë).

Notes. I follow Casale (2003: 534) who noted ssp. modestus only for Croatia and Bosnia Herzegovina.

98.2. Laemostenus elongatus (Dejean, 1828)

98.2.1. Laemostenus elongatus pentheri J. Müller, 1924

References. Laemostenus elongatus: Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme,

300 m, and Vorfai Siperme, 700 m). Laemostenus elongatus pentheri: Müller, 1924: 70-71 (North

Albanian Alps: state attribution of loc. typ. in original description not specified; Rikavac;

Vermosh; Rapshë); Pretner, 1978: 136, 148, 168 (Rikavac; Vermosh; Rapshë); Casale, 1988:

700, 704 (Okol near Theth: designated as loc. typ.; Maranaj; Vermosh, Rapshë).

Material studied. 49 s. from localities “Maranaj”, “Shkëlzën”, “Maja e Jezercoë”, “Çafa Peis”,

“Cucale”, “Boge”, “Maja e Dragomirit”, “Maja Rogamit” (MNHUB); North Albanian Alps,

Bogë, 1400-1800 m, 1.VI.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District,

Merkuth, 920-980 m, 11.VI.1933, 1 m., grassland, P. Beron & B. Petrov leg. (NMNHS).


Mali i Thatë Mt., 18.VII.1993, 1 f., L. Latella leg. (cAVT).

Notes. Casale (2003: 534) has omitted the species for Albania. Here it is confirmed for the country.

Subgenus Laemostenus s.str.

98.3. Laemostenus dalmatinus (Dejean, 1828)

Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW).


98.4. Laemiostenus janthinus (Duftschmid, 1812)

98.4.1. Laemiostenus janthinus janthinus (Duftschmid, 1812)

References. Laemiostenus janthinus: Csiki, 1940: 218 (Gjalicë Mt.).

Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).

Notes. Casale (2003: 536) has omitted the species for Albania. Here it is confirms for the country.

98.5. Laemostenus venustus (Dejean, 1828)

References. Laemostenus venustus: Apfelbeck, 1904: 275 (Velipoja); Casale, 1988: 476 (Velipoja);

Hieke & Wrase, 1988: 93; Casale, 2003: 536.


Material studied. “Mamurash” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Mamurash”

/ “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Gjormi” / “Albania leg. Bischoff 1935”, 2 s.

(MNHUB); “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Sukthi bis Gurse”


Subgenus Pristonychus Dejean, 1828

98.6. Laemostenus cimmerius (Fischer von Waldheim, 1823)

98.6.1. Laemostenus cimmerius cimmerius (Fischer von Waldheim, 1823)

References. Sphodrus cimmerius: Anonymous, 1907: 285. Prustonychus cimmerius: Magistretti, 1965:

433. Laemostenus cimmerius: Wrase, 1991: 11. Laemostenus cimmerius cimmerius: Casale, 1988: 809

(Durrës); Casale, 2003: 537.

Material studied. Durrës, amphitheater, 12.VI.1993, 1 f., B. Petrov leg. (NMNHS); Tiranë

Vore, 27.VI.1995, 1 m., S. Abadjiev leg. (NMNHS).

98.6.2. Laemostenus cimmerius weiratheri J. Müller, 1932

References. Laemostenus cimmerius weiratheri: Casale, 1988: 809 (M. Sopot); Sakalian & Guéorguiev,

1997: 48; Casale, 2003: 537.

98.7. Laemostenus terricola (Herbst, 1784)

98.7.1. Laemostenus terricola punctatus (Dejean, 1828)

References. Pristonychus terricola: Hieke et al., 1986: 189 (Mali me Gropë). Laemostenus punctatus:

Hieke & Wrase, 1988: 93. Laemostenus terricola punctatus: Casale, 1988: 787 (Mali me Grope,

Dolinen-Gez); Casale, 2003: 539.

Material studied. “Vermosa’14 Penther” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW);

“Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Tomor-Tojk juli

1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”,

3 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 8 s. (MNHUB); “Vukaj” / “Cucale” /

“Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”,

2 s. (MNHUB); “Mal i That” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albanien – Exp.

DEI Mali me Gropë, Dolinengebiet 1350 m, 6.VII.1961”, 1 s. (DEI).

Subtribe SYNUCHINA Lindroth, 1956

99. Genus Synuchus Gyllenhal, 1810

Subgenus Synuchus s.str.

99.1. Synuchus vivalis (Illiger, 1798)

99.1.1. Synuchus vivalis vivalis (Illiger, 1798) [= nivalis (Panzer, 1796)]

References. Synuchus nivalis: Hieke & Wrase, 1988: 88. Synuchus vivalis vivalis: Hovorka & Sciaky,

2003c: 546.


Material studied. “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Theth”

/ “23.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM).


Tribe ZABRINI Bonelli, 1810

Subtribe AMARINA C. Zimmermann, 1831

100. Genus Amara Bonelli, 1810

Subgenus Amara s.str.

100.1. Amara aenea (DeGeer, 1774)

References. Amara aenea: Apfelbeck, 1904: 300 (Vlorë; Berat; Velipoja); Apfelbeck, 1907a:

497, 499 (Maranajt Mt.: top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m);

Apfelbeck, 1909: 297 (Vlorë); Csiki, 1940: 215 (Shkëlzen Mt.; Kula Ljums; Ura i Lopez; “Stičen”

near Gjalicë Mt.; Nangë; Gostilë); Schatzmayr, 1943b: 105 (Dukati; Tërpan); Hieke et al.,

1986: 189 (Tomor Mt.: Ali Abbas Monastery; North Albanian Alps near Theth); Hieke &

Wrase, 1988: 100; Hieke, 2003: 547.

Material studied. Tiranë, 7.IV.1960, 1 s. (MNST); Ibë, 18.V.1960, 1 s. (MNST); Rodeni, 2-

22.VII.1962, 4 s. (MNST); Zooteknica, 16.V.1963, 1 s. (MNST); Theth, 800-1200 m, 28.V.1993,

1 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, cave No 25, 23.VI.1993,

1 s., P. Beron, B. Petrov leg. (NMNHS); Saranda Dhermi, 2.V.1994, 1 s., P. Stoev leg. (NMNHS);

Saranda District, Ljukove, 3-4.VI.1994, 2 s., G. Blagoev leg. (NMNHS); 5 km Leskovic,

2.VII.1994, 1 s., P. Tenchev leg. (NMNHS); Petrela, 15 km NE Tiranë, 300 m, 9.V.1995, 2 s., S.

Golovach, B. Petrov, P. Stoev leg. (NMNHS); 5 km Saranda, 3.VI.1995, 1 s., G. Blagoev leg.

(NMNHS); Butrinti, 4.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Lyukove, 5.VI.1995, 1 s., V.

Sakalian leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 12-14.II.1998 / 16-21.II.1998 / 23-

28.II.1998, 56 s., R. Ward leg. (CMNH).

Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona, 900

m, 5.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Durrës, Plepa, 15.VIII.1995, 1 f., D. Antonelli

leg. (cAVT); Rapsh, 15.V.2002, 2 m., 1 f., A. Paparistos leg. (cAVT).

Notes. In addition to the material studied above, the present author has seen numerous further

specimens in HNHM, NMW and MNHUB, which remained unregistered.

100.2. Amara anthobia A. Villa & G.B. Villa, 1833

References. Amara anthobia: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke

& Wrase, 1988: 102; Hieke, 2003: 547.

Material studied. “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Logara” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Cafa

e Dhembellit” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Vlorë District, Llogora Pass,

1050 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS); Dhermi, 100 m, 11.V.1995, 1 s., S. Golovach,

B. Petrov, P. Stoev leg. (NMNHS).


100.3. Amara convexior Stephens, 1828

References. Amara convexior: Hieke & Wrase, 1988: 97; Hieke, 2003: 548.

Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB).


100.4. Amara curta Dejean, 1828

References. Amara curta: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576 m;

Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 215 (Ura i Lopez; Gjalicë Mt.; Korab Mt.);

Hieke & Wrase, 1988: 98; Hieke, 2003: 548.

Material studied. “Apfelb. Alban. spt.. IV.V.’05”, 3 s. (NMW); “Prokletjia 1300-2000” / “Alban.

Montenegr. Grenze, Penther’14”, 2 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban.

Montenegr. Grenze, Penther’14”, 1 s. (NMW); “Vermosa’14 Penther”, 2 s. (NMW); “Albania,

Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Maja e Tartarit” /

“Albania leg. Bischoff 1932”, 5 s. (MNHUB); “Berat-Bargulash” / “leg. Bischoff Albania

1934”, 1 s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Shkëlzën”

/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 5 s.

(MNHUB); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron, B. Petrov

leg. (NMNHS); Shkoder, 12.VI.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS).

100.5. Amara eurynota (Panzer, 1796)

References. Amara eurynota: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke

& Wrase, 1988: 101; Hieke, 2003: 549.

Material studied. “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Bashtrik” /

“Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff

1936”, 3 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Saranda

District, Qeparo, 4-5.V.1994, 1 s., P. Stoev leg. (NMNHS).

? Amara famelica C. Zimmermann, 1832

Material studied. “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”, 7 s. (NMW); “Alban. Exp.

1918 Korab, 23-31.VII.”, 1 s. (NMW).

Notes. F. Hieke (personal communication) expressed doubt about the correct identification

of the specimens in NMW.

100.6. Amara familiaris (Duftschmid, 1812)

References. Amara familiaris: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë);


Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Logara”

/ “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor” / “Albania leg. Bischoff 1932”, 1 s.

(MNHUB); “Logara” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).


100.7. Amara lucida (Duftschmid, 1812)

References. Amara lucida: Apfelbeck, 1904: 301 (Berat); Apfelbeck, 1909: 297 (Berat); Csiki,

1940: 215 (Ura i Lopez; Korab Mt.); Hieke & Wrase, 1988: 102; Hieke, 2003: 549.

Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW);

“Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Ura i Lopez” /

“Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Krabe” / “Albania leg. Bischoff 1932”, 1

s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Shen Ili” /

“Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor-Burany juli 1932”, 3 s. (MNHUB);

“Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “leg.

Bischoff Albania 1934”, 4 s. (MNHUB); “Mal i Shejit” / “leg. Bischoff Albania 1934”, 1

s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Logara” / “Albania

leg. Bischoff 1936”, 3 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1

s. (MNHUB).

100.8. Amara morio Ménétriés, 1832

100.8.1. Amara morio nivium Tschitschérine, 1900

References. Amara morio nivium: Hieke, 1995: 288 (Gjalicë Ljums); Hieke, 2003: 550.

Material studied. “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM;

MNHUB).

100.9. Amara ovata (Fabricius, 1792)

References. Amara ovata: Hieke, 2003: 550.

Material studied. “Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Llogora” / “leg.

Bischoff Albania 1936”, 2 s. (MNHUB).

Additional material included (studied F. Hieke). “Tomor, Kulmak, Alban. mer.”, 1 s. (MCG);

“Lona, Albania”, 1 s. (MSNM).

Notes. The mention of Hieke & Wrase (1988: 96) for Albania refers apparently to the record

“Prisren” (Apfelbeck, 1904: 299). However, this locality is situated in the Kosovo Region of

Serbia. Besides the material studied above, I have seen more specimens in NMW, which remained

unregistered. First detailed records for Albania.

100.10. Amara proxima Putzeys, 1866

References. Amara proxima: Hieke & Wrase, 1988: 98; Hieke, 2003: 551.

Material studied. “Shendeli Alban. Mer.” / “Winkler. Lona, Bischoff, V.1931”, 6 s. (NMW;

MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania

leg. Bischoff 1936”, 3 s. (MNHUB); “Maja e Tartarit juni 1937” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB).

Additional material included (studied F. Hieke). “Argiro Kastron, S.Alban”, 2 s. (MCG;

MTKD); “Ismen Ili, Albania”, 1 s. (NMPC).



100.11. Amara saphyrea Dejean, 1828

References. Amara saphyrea: Hieke & Wrase, 1988: 96; Hieke, 2003: 551.

Material included (studied F. Hieke). “Tomor, Kulmak, Alban. mer.”, 1 s. (NMW).


100.12. Amara similata (Gyllenhal, 1810)

Material studied. Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten

wood, R. Ward leg. (CMNH).


# Amara spreta Dejean, 1831

References. Amara spreta: Csiki, 1940: 215 (Gjalicë Mt.).

Notes. Basing on the most recent information about the species distribution (Hieke, 2003:

551), the Csiki’s identification is most probably incorrect and that the material in question

concerns another congener. F. Hieke (personal communication) expressed similar opinion, so

A. spreta is removed from the list of the Albanian Carabidae.

100.13. Amara tibialis (Paykull, 1798)

References. Amara tibialis: Csiki, 1940: 215 (Korab Mt.); Hieke & Wrase, 1988: 103; Hieke,

2003: 551.

Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM).

Subgenus Amarocelia Motschulsky, 1862

100.14. Amara erratica (Duftschmid, 1812)

References. Amara erratica: Csiki, 1940: 216 (Gjalicë Mt.); Hieke & Wrase, 1988: 103; Hieke,

2003: 556.

Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 1 s. (HNHM); “Prokletija 1300-2000

m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Skëlzën” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB).

Subgenus Bradytus Stephens, 1827

100.15. Amara apricaria (Paykull, 1790)

References. Amara apricaria: Hieke et al., 1986: 190 (Borshi near Vlorë; Bizë near Shëngjergji);


Material studied. “Vermosa’14 Penther”, 1 s. (NMW); “Albania Mts. Korab” / “Akad. Balk.

Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp.’18 Pashtrik, 4-14.7.”, 3 s. (NMW); 28 s. from

localities “Shkalle Harushe-Korab”, “Korab”, “Shkëlzën”, “Mal i Sheniit”, “Mal i Tartarit”,

“Taja”, “Giallica” (MNHUB); “Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 2 s. (HNHM).


100.16. Amara consularis (Duftschmid, 1812)

Material studied. Mal i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 1 s. (MNST).


Subgenus Celia C. Zimmermann, 1832

100.17. Amara bifrons (Gyllenhal, 1810)

References. Amara bifrons: Csiki, 1940: 216 (Korab Mt.); Hieke & Wrase, 1988: 105; Hieke, 2003: 555.

Material studied. “Ricavac 1300 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1

s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW);

“Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp. 1918

Korab 23-31.VII.”, 4 s. (NMW); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB);

“Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercë” / “Albania leg.


“Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM).



100.18. Amara hannemanni Hieke, 1991

References. Amara hannemanni: Hieke, 1991: 1-3 (Shkëlzen Mt., loc. typ.); Hieke, 2003: 556.

Material studied. “Skëlzën” / “Albania leg. Bischoff 1935”, 1 paratype (MNHUB); “Skëlzën”

/ “Albania leg. Bischoff 1936”, 6 paratypes (MNHUB).

100.19. Amara montana Dejean, 1828

References. Amara montana: Hieke, 2003: 557.

Material studied. “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Shkoder” /

“25.V.1958 leg. Xh. Murraj”, 2 s. (HNHM); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS);

Tiranë Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH).


? Amara praetermissa (C.R. Sahlberg, 1827)

References. Amara praetermissa: Hieke & Wrase, 1988: 105.

Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 2 s. (HNHM); “Latif Koritnik”, 3 s. (NMW).

Notes. Hieke (2003: 557) has omitted the species for Albania in spite of his previous data

(Hieke & Wrase, 1988: 105). He stated me (personal communication) that the specimens above

might belong to A. hannemanni, so that A. praetermissa is treated here as doubtful for Albania.

Subenus Curtonotus Stephens, 1827

100.20. Amara aulica (Panzer, 1796)

References. Amara aulica: Hieke & Wrase, 1988: 109; Hieke, 2003: 559.


Material studied. “Tomor Albanien”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff

1932”, 1 s. (MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).


Subgenus Paracelia Bedel, 1899

100.21. Amara dalmatina Dejean, 1828

References. Amara dalmatina: Hieke (in press).

Material included (studied F. Hieke). “SE of Medua, Albania”, 1 s. (MNHUB); “Shendeli,

Albania mer.”, 1 s. (MNHUB).


100.22. Amara quenseli (Schönherr, 1806)

100.22.1. Amara quenseli quenseli (Schönherr, 1806)

References. Amara quenseli: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576

m; Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 216 (Korab Mt.); Holdhaus & Lindroth,

1940: 146-147, 250 (Koritnik Mt.; Korab Mt.); Hieke & Wrase, 1988: 106. Amara quenseli quenseli:

Hieke, 2003: 564.

Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 2 s. (HNHM); “Latif Koritnik”, 8 s.

(NMW); “Merdita Munela” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Prokletija 1300-

2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Prokletija 1300-2000 m” /

“Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania Galica Lums” / “Akad. Balk.

Exp. Csiki 1917”, 2 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki

1918”, 3 s. (HNHM); “Korab” / “Albania leg. Bischoff 1935”, 16 s. (MNHUB); “Korab” /

“Albania leg. Bischoff 1936”, 19 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 23

s. (MNHUB).

Subgenus Parapercosia Tschitschérine, 1899

100.23. Amara taurica (Motschulsky, 1844) [= fallaciosa (J. Müller, 1946)]

References. Zabrus fallaciosus: Müller, 1946: 107-110 (South Albania: Shëndelli Mt. near Tepelenë,

loc. typ.); Turin, 1981: 148. Amara taurica: Hieke, 2003: 565.

Notes. Hieke (1995: 35-36) synonymized Zabrus fallaciosus Müller, 1946 with Amara taurica,

noting the type locality of the former as “Patria falsa!”, but later the same has cited A. taurica

for Albania (Hieke, 2003: 565). According to F. Hieke (personal communication) A. taurica is

known from different parts of Turkey and from two localities in Greece, so that the type

locality of Zabrus fallaciosus in Albania should not be considered as doubtful (Hieke, 2005).

Subgenus Percosia C. Zimmermann, 1832

100.24. Amara equestris (Duftschmid, 1812)

100.24.1. Amara equestris equestris (Duftschmid, 1812) [= tomoricensis J. Müller, 1923]


References. Amara tomoricensis: Müller, 1923: 108-109 (Tomor Mt.: Kulmak, loc. typ.). Amara

equestris: Csiki, 1940: 216 (Korab Mt.); Hieke, 1978: 229, 257 (Tomor Mt.: Kulmak; Koritnik

Mt.; Korab Mt.); Hieke & Wrase, 1988: 108. Amara equestris equestris: Hieke, 2003: 565.

Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM);

“Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Korab” / “Albania leg. Bischoff

1935”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Albania

Zalle-Dardha” / “24.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM).

Subgenus Xenocelia Hieke, 2001

100.25. Amara bischoffi Jedlička, 1946

References. Amara municipalis: Csiki, 1940: 215 (Gjalicë Mt.; Korab Mt.); Jedlička, 1946: 7

Peshtrik Mt. (loc. typ.); Hieke & Wrase, 1988: 104. Amara bischoffi: Turin, 1981: 145; Hieke,

1983: 328-329, 334 (Peshtrik Mt.; Taja); Hieke, 2001: 33 (two localities). Amara municipalis

bischoffi: Hieke, 2003: 557.

Material studied. “Taja” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Bashtrik” / “Albania

leg. Bischoff 1936”, 1 syntype (MNHUB).

Notes. According to F. Hieke (personal communication) the records of Csiki (1940) are

problematic since they may concern A. bischoffi, as well as A. messae.

100.26. Amara cursitans C. Zimmermann, 1832 [= properans C. Zimmermann, 1832]

References. Amara properans: Csiki, 1940: 215 (Gjalicë Mt.; Korab Mt.). Amara cursitans: Hieke

& Wrase, 1988: 104; Hieke, 2001: 50 (five localities); Hieke, 2003: 556.


(NMW); “Skëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg.

Bischoff 1936”, 5 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB);

“Maja e Dragomit” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB).

100.27. Amara fusca Dejean, 1828 [= complanata Dejean, 1828]

References. Amara complanata: Csiki, 1940: 215 (Gjalicë Mt.). Amara fusca: Hieke & Wrase,

1988: 104; Hieke, 2001: 42 (“Koritza-Ebene”, 1 s. in MNHN); Hieke, 2003: 556.

100.28. Amara messae Baliani, 1924

References. Amara messae: Hieke & Wrase, 1988: 104; Hieke, 2001: 65; Hieke, 2003: 557.

Material studied. “Giallica” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercoë”

/ “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cotype” / “Bashtrik” / “Albania leg. Bischoff

1936”, 1 s. (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron, B.




Subgenus Zezea Csiki, 1929

100.29. Amara chaudoiri Schaum, 1858

100.29.1. Amara chaudoiri incognita Fassati, 1946

References. Amara chaudoiri incognita: Hieke, 1970: 138 (Skrofotino near Vlorë); Hieke, 2003:

567. Amara chaudoiri: Hieke & Wrase, 1988: 94.

# Amara erythrocnema Dejean, 1828

References. Amara erythrocnema: Schatzmayr, 1943b: 105 (Skrofotina).

Notes. At present the species is treated as local endemic for the island of Corfu (Hieke, 2003:

567). F. Hieke (personal communication) expressed the opinion that the record for A.

erythrocnema from Albania most probably concerns A. reflexicollis. Therefore, the former species

is removed from the list of Albanian Carabidae.

100.30. Amara fulvipes (Audinet-Serville, 1821)

References. Amara fulvipes: Apfelbeck, 1904: 298 (Velipoja); Hieke, 1970: 157 (Velipoja; Lum;

Tiranës; Lum i Tiranës; Mal i Dajtit Mt.); Hieke & Wrase, 1988: 94; Hieke, 2003: 567.

Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane”

/ “leg. Bischoff Albania 1934”, 6 s. (MNHUB).

100.31. Amara reflexicollis Motschulsky, 1844

100.31.1. Amara reflexicollis reflexicollis Motschulsky, 1844

References. Amara erythrocnema: Schatzmayr, 1943b: 105 (Skrofotina). Amara reflexicollis

reflexicollis: Hieke, 2003: 568.

100.32. Amara tricuspidata Dejean, 1831

References. Amara tricuspidata: Hieke & Wrase, 1988: 94; Hieke, 2003: 568.

Material studied. “Museum Paris Albanie Starova (env. de Koritza) Caporal Vuillaume 1918”

/ “Albanie Armée d’Orient 1916-1918” / “Avril” / “Mai”, 1 s. (MNHUB).

Additional material included (studied F. Hieke). 1 s. labelled as that up above (MNHP).


Subtribe ZABRINA Bonelli, 1810

101. Genus Zabrus Clairville, 1806

Subgenus Pelor Bonelli, 1810

Notes. The species from this subgenus are arranged here in three groups of species with the

aim to delineate the related lineages.


Species group “aetolus”

101.1. Zabrus aetolus Schaum, 1864

[Zabrus aetolus kodymi Mařan, 1940]

Notes. This race has been described from the Montenegrin part of the North Albanian Alps

(Mařan, 1940a: 33), basing on the label data of the material collected by O. Kodym and C.

Purkynì in 1937. The ssp. kodymi may inhabit the Albanian part of the mountain, too.

101.1.1. Zabrus aetolus winkleri J. Müller, 1946

References. Zabrus winkleri: Müller, 1946: 110-111 (Maja e Tartarit Mt., south of Vlorë, loc.

typ.; Maja e Djembellit Mt., east of Gjirokastër; Maja e Policanit Mt., east of Gjirokastër);

Turin, 1981: 148. Zabrus aetolus winkleri: Freude, 1989: 83, 95 (Maja e Tartarit Mt.; Maja e

Dhembellit Mt.; Maja e Policanit Mt.); Serrano & Andújar, 2003: 570.

Material studied. “Argirokastron Sud-Albania”, 1 s. (MNHUB); “Maja e Tartarit” / “Albania


101.2. Zabrus albanicus Apfelbeck, 1904

101.2.1. Zabrus albanicus albanicus Apfelbeck, 1904

References. Zabrus albanicus latifianus: Ganglbauer, 1931: 39 (Koritnik Mt.); Mařan, 1939d:

145-146 (Koritnik Mt.); Schatzmayr, 1943b: 108 (Koritnik Mt.); Freude, 1989: 83, 88, 97

(Koritnik Mt.); Drovenik & Peks, 1994: 96 (Koritnik Mt.); Drovenik & Peks, 1999: 111 (Koritnik

Mt.). Zabrus albanicus: Csiki, 1940: 221 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Schatzmayr,

1943b: 108 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke, 1981: 83. Zabrus albanicus albanicus:

Serrano & Andújar, 2003: 570.

Material studied. “Latif Koritnik” / “albanicus typ. Apfelb.”, 12 s. (NMW); “Latif Alban. spt.”,

3 s. (DEI); more than 140 s. from localities “Albania Galica Lums” (or “Gyalica Lums”), “Skölsen”

and “Korab” (HNHM); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 4

s. (NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 2 s. (NMW); “Alban. Exped. 1918 Gjalica

Ljums 17-26.VI.”, 20 s. (NMW); “Alban. Exped. Korab 1918 24-31.VII.”, 71 s. (NMW).

Notes. The populations from the Koritnik Mt., as well as the records for these (Ganglbauer,

1931; Mařan, 1939d; Schatzmayr, 1943b; Freude, 1989; Drovenik & Peks, 1994; 1999), are

assigned provisionally to the nominotypical subspecies rather than to the ssp. latifianus, since

the Zepë Mt. (an unmarked, but potential “loc. typ.” of the latter taxon) has an more southern

and well-isolated position from the other massifs in North Albania. Besides the material studied

above, in MNHUB the present author has seen other more 140 s. of Z. albanicus s.l. from

different localities, which remained unregistered.

[Zabrus albanicus jablanicensis Mařan, 1939]

References. Zabrus albanicus jablanicensis: Freude, 1989: 88 (“Mittel-Ost-Albanien an der jugo-

slavischen Grenze”).

Notes. Serrano & Andújar (2003: 570) indicated Bosnia Herzegovina as region of the

distribution of ssp. jablanicensis by mistake. In fact, this race was described from the Jablanitsa

Mt. shared between Albania and Macedonia and so far is cited only from the Macedonian part

of the mountain. Most likely the record of Freude (1989) is not based on material coming


from collections, and may bases of supposition. The subspecies most probably inhabits the

Albanian part of the Jablanitsa Mt., too.

101.2.2. Zabrus albanicus latifianus Ganglbauer, 1915

References. Zabrus albanicus: Apfelbeck, 1907a: 503 (Zepë Mt.). Zabrus albanicus latifianus:

Ganglbauer, 1931: 39 (Zepë Mt.); Mařan, 1939d: 145-146 (Zepë Mt.); Schatzmayr, 1943b: 108

(Zepë Mt.); Freude, 1989: 83, 88, 97 (Zepë Mt.); Serrano & Andújar, 2003: 570.

Material studied. “Merdita Munela”, 1 s. (NMW); “Merdita Zebia”, 2 s. (NMW).

Notes. In my opinion, only the populations from the region of Merdita have to be considered

as belonging to spp. latifianus.

! Zabrus albanicus ssp.

Zabrus albanicus: Schatzmayr, 1943b: 108 (Tomor Mt.: Abbas Ali and Toik).

Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 4 s. (NMW;

MNHUB); “Maja e Dhembelit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 8 s. (MNHUB);

“Albania 1936 leg. Ramadan coll. Bischoff ” / “Maja e Papingut”, 21 s. (MNHUB).

Notes. The specimens from the massifs of Tomor Mt., Maja e Dhembelit Mt. and Maja e

Papingut Mt. (all in South Albania) may concern another subspecies, since the latter are far

removed in the south from the massifs of the Zepë Mt. and Koritnik Mt.

# Zabrus apfelbecki Ganglbauer, 1915

References. Zabrus apfelbecki: Magistretti, 1965: 360.

Notes. Magistretti (1965) referred older records of Z. ignavus Csiki, 1907 to Z. apfelbecki,

but I failed to find any records in the literature. The latter species lives only in Bosnia

Herzegovina (Serrano & Andújar, 2003: 570), so that it is removed from the list of the

Albanian ground beetles.

101.3. Zabrus ganglbaueri Apfelbeck, 1906

101.3.1. Zabrus ganglbaueri ganglbaueri Apfelbeck, 1906

References. Zabrus ganglbaueri: Apfelbeck, 1906: 1667 (Merdita: Mal i Shenjt Mt. near Oroshi,

loc. typ.); Apfelbeck, 1907a: 499-500, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1936:

108 (Mal i Shenjt Mt. near Oroshi); Schatzmayr, 1943b: 108 (Mal i Shenjt Mt. near Oroshi);

Hieke, 1981: 75. Zabrus ganglbaueri ganglbaueri: Freude, 1989: 82, 110 (Mal i Shenjt Mt. near

Oroshi; Mal i That Mt. near Gjirokastër); Serrano & Andújar, 2003: 570.

Material studied. “Merdita M. Scheit”, 1 s. (NMW); “Merdita 18.19. n. 20/5 1934”, 2 s. (NMW).

Notes. Freude (1989) stated that the populations from the Mal i Shenjt Mt. (Merdita District)

and the Mal i That Mt. (Gjirokastër District) belong to the nominotypical race. In fact both

populations inhabit opposite outskirts of the country, so the specimens from the latter massif

may belong either to the following subspecies or to a separate geographical race.

101.3.2. Zabrus ganglbaueri lonae J. Müller, 1923

References. Zabrus lonae: Müller, 1923: 107-108 (Tomor Mt.: Toik, loc. typ.); Müller, 1935:

154 (Tomor Mt.: Toik); Mařan, 1940a: 30-31, 33, 35. Zabrus lonai: Schatzmayr, 1943b: 108

(Tomor Mt.: Toik). Zabrus ganglbaueri lonae: Freude, 1989: 83, 111 (Tomor Mt.: Toik); Serrano

& Andújar, 2003: 570.


Material studied. “Albania 1922 Ravasini, Lona” / “Tomor Monte Toik”, 1 s. (DEI); “Tomor:

Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Tojk” / “Albania


Species group “incrassatus”

101.4. Zabrus bischoffi J. Müller, 1936

References. Zabrus bischoffi: Müller, 1936: 107-108 (Mal i Dajtit Mt. near Tiranë, loc. typ.;

Llogora); Schatzmayr, 1943b: 106, 109 (Kruja; Mal i Dajtit Mt.; Llogora); Turin, 1981: 148.

Zabrus incrassatus bischoffi: Freude, 1989: 86, 120 (Mal i Dajtit Mt. near Tiranë; Llogara, south of

Vlorë); Serrano & Andújar, 2003: 571.

Material studied. 19 s. from “Mal i Dajtit” (MNHUB); 5 s. from “Mal i Krues” (MNHUB);

3 s. from “Mal i Shenjit” (MNHUB); 37 s. from “Llogora” (MNHUB); Llogora Pass, 1025 m,

6.6.1995, 1 m., G. Blagoev leg. (NMNHS).

Notes. This species has a quite different apex of the median lobe and sides of pronotum

narrower anteriorly, which is in contrast with the respective characteristics in Z. incrassatus s.l.

In my opinion, the former is separate species not subspecies of the latter: Zabrus (Pelor) bischoffi

J. Müller, 1936, stat. rest. of Zabrus (Pelor) incrassatus bischoffi J. Müller, 1936.

101.5. Zabrus boldorii Schatzmayr, 1943

References. Zabrus boldori: Schatzmayr, 1943: 106, 108 (Dukati, south of Vlorë, loc. typ.);

Serrano & Andújar, 2003: 570. Zabrus baldori sic!: Turin, 1981: 148. Zabrus incrassatus boldorii:

Freude, 1989: 86, 119 (Dukati near Vlorë).

Material studied. “Südalbanien Kopstein” / “coll. Prof. F. Werner” / “Pašaliman”, 1 m.

(NMW); “Südalbanien Kopstein” / “Valona”, 2 f. (NMW).

101.6. Zabrus incrassatus (Ahrens, 1814)

101.6.1. Zabrus incrassatus incrassatus (Ahrens, 1814)

References. Zabrus incrassatus: Apfelbeck, 1904: 310 (Berat; Vlorë); Anonymous, 1907: 285;

Apfelbeck, 1907a: 495-496 (northeast of Shkodër; Maranajt Mt.: between Vorfai Postme, 300

m, and Vorfai Siperme, 700 m); Apfelbeck, 1909: 297 (Vlorë; Berat); Müller, 1936: 108 (Shëndelli

Mt.; Tepelenë; Maja e Tartarit Mt.; Tiranë; Gjirokastër; Berat; Vlorë); Schatzmayr, 1943b: 106,

108 (“Poggio Boschetto”; Lushnje; Berat; Vlorë; Shen Ili; Tepelenë; Maja e Tartarit Mt.; Tiranë;

Shkodër); Hieke & Wrase, 1988: 110.

Material studied. “Südalbanien Kopstein”, 2 s. (NMW); “Albanien Ochrida”, 1 s. (NMW);

“Hani Grabom Mai” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Rapsa 10-

18.V.” / “Alban. montenegr. Grenze, Penther’14”, 4 s. (NMW); “Alban. Exp. 1918 Bicaj

14.15.VI.”, 3 s. (NMW); Ibë, 2-11.VII. 1962, 1 m. (MNST); Shkodër, 15.V.1963, 3 m., 1 f.

(MNST); 2 km N Durrës, 24.V.1993, 1 m., P. Stoev leg. (NMNHS); Vlorë, 1.V.1994, 2 m., 3 f.,

P. Stoev leg. (NMNHS); Saranda District, Himarë, 3-4.V.1994, 1 f., P. Stoev leg. (NMNHS);

Saranda District, Borsh, 5.V.1994, 1 f., P. Stoev leg. (NMNHS); Saranda District, Ljukova, 3-

4.VI.1995, 1 f., G. Blagoev leg. (NMNHS); Vlorë District, Llogora Mt., 1027 m, 4.VI.1995, 1

f., G. Blagoev leg. (NMNHS).


Notes. Serrano & Andújar (2003: 571) have omitted the species for Albania, so it is here

confirmed for the country. Besides the material studied above, in MNHUB I have seen other

more than 100 s. from different localities (“Shen Ili”, “Tirane”, “Murthi”, “Tepelene”, “Munela”,

“Burelle”, “Vlorë”, “Mal i Gjer”, “Myzege”, “Bose”, “Maja e Dhembellit”, “Tarabosh”, “Mal

i Tartarit”, etc.).

Species group “spinipes”

101.7. Zabrus spinipes (Fabricius, 1798)

101.7.1. Zabrus spinipes insignis J. Müller, 1932

References. Zabrus spinipes: Burmeister, 1939: 127; Hieke et al., 1986: 190 (Tomor Mt.: Abbas

Ali Monastery); Hieke & Wrase, 1988: 110; Freude, 1989: 147.

Material studied. “Shen-Ili Juni” / “Albania leg. Bischoff 1932”, 11 s. (MNHUB); “Mal i

That” / “Albania leg. Bischoff 1937”, 22 s. (MNHUB).

Notes. Serrano & Andújar (2003: 572) have omitted the species for Albania, so it is here


Subgenus Zabrus s.str.

101.8. Zabrus tenebrioides (Goeze, 1777)

101.8.1. Zabrus tenebrioides tenebrioides (Goeze, 1777)

References. Zabrus tenebrioides: Schatzmayr, 1943b: 105-106 (Dukati; Lushnje). Zabrus tenebrioides

tenebrioides: Serrano & Andújar, 2003: 573.

Material studied. “Myzeqe” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); 11 km west of

Tiranë, 20.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 5 s., B.

Guéorguiev leg. (NMNHS).

Additional material included (studied A. Vigna Taglianti). Durrës, Currila, 8.V.2002, 1 f., A.

Paparistos leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of Vijosë River, 300-

360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Tribe ZUPHIINI Bonelli, 1810

Subtribe ZUPHIINA Bonelli, 1810

102. Genus Polystichus Bonelli, 1810

102.1. Polystichus connexus (Geoffroy, 1785)

References. Polystichus connexus: Hieke & Wrase, 1988: 163.

Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).

Notes. Baehr (2003e: 574) has omitted the species for Albania. Here it is confirmed for the

country as first detailed record is given.


103. Genus Zuphium Latreille, 1806

Subgenus Zuphium s.str.

103.1. Zuphium olens (P. Rossi, 1790)

103.1.1. Zuphium olens olens (P. Rossi, 1790)

References. Zuphium olens: Schatzmayr, 1943b: 112 (Stan-Karbunarë); Hieke & Wrase, 1988:

162. Zuphium olens olens: Baehr, 2003e: 574.


Results

When omitting doubtful species and taxa still not well defined, the Albanian Carabidae

listed in this catalogue represent 543 species. They are distributed among 11 subfamilies

(Nebriinae – 25, Cicindelinae – 10, Carabinae – 20, Elaphrinae – 1, Omophroninae –

1, Brachininae – 17, Siagoninae – 1, Scaritinae – 28, Apotominae – 2, Trechinae – 115,

and Harpalinae – 323), 31 tribes and 103 genera (Table 1).

One tribe, 10 genera, 11 subgenera and 55 species are new for the fauna of Albania,

and 1 subgenus and 3 species are new for the fauna of the Balkan Peninsula (Appendix

A.1-2). From these, 1 tribe, 1 subtribe, 7 genera, 20 subgenera and 72 species are

reported firstly with detailed record/s for the country (Appendix A.3). Altogether

119 species, omitted in the recent Palaearctic Catalogue (Löbl & Smetana, 2003), are

here confirmed for Albania (Appendix A.4).

Twelve taxa from the species group remain still not well defined, as likely most of

them represent new for the science species or subspecies (Appendix A.5). Two

subgenera and 22 species and subspecies are doubtful for the country (Appendix

A.6); generally the published records about them concern adjacent regions in

Montenegro, the Kosovo Region of Serbia, Macedonia or Northwestern Greece. Most

of the species and subspecies from the last category may inhabit Albania, but until

now we do not have reliable data to confirm this. Two subgenera and 29 species and

subspecies are deleted from the list (Appendix A.7). The systematic list includes also

some endemics described from massifs shared between Albania and neighbour

countries (but not recorded yet from the former), as well as several more widespread

species found from localities very close to Albania.

A moderate taxonomic contribution is also achieved (Appendix B). Principally, it is

based on study of types, topotypes or more extensive series of previously problematic

species and subspecies from Albania and neighbouring countries. In result of that 38

various taxonomic innovations (new synonyms, new status or revalidations) are

introduced. Further, nine taxa from the species group (5 valid species and 4 synonyms),

all described by V. Apfelbeck, have been found to have assigned an incorrect year of

description. Accordingly, the relevant earlier years of their descriptions are fixed. Finally,

eight nomina nuda were also traced out, as one of them may be a result of mislabeling,

and one is supplied with year of description in the literature.

RESULTS 187


Conclusions

Undoubtedly the Balkan Peninsula contains the richest ground beetle diversity

compared to other diverse European regions like the Iberian Peninsula, Italy or the

Caucasus Major (Turin, 1981; Kryzhanovskij et al., 1995; Serrano, 2003; Vigna Taglianti,

1993; 2004; 2005a; 2005b). Currently an impressive total of at least 1487 species of

Carabidae, following the classification used in Löbl & Smetana, 2003, is reported

from this territory (unpublished data). This figure is provisional while the systematics,

diversity and distribution of the Carabidae in the Balkan Peninsula are better known.

Until now, the carabid fauna of Bulgaria is probably the best investigated one in the

region (Hieke & Wrase, 1988; Guéorguiev & Guéorguiev, 1995). The species

composition of no other Balkan regions has been examined in details; only that of

ex-Yugoslavia was subject of uncritical registration (Drovenik & Peks, 1994; Drovenik

& Peks, 1999). Therefore, the Carabidae of Slovenia, Croatia, Bosnia Herzegovina,

Montenegro, Serbia, Macedonia, Romania, Greece and European Turkey need scrutiny

and writing of up-to-date checklists.

Bearing in mind the surface of Albania, the number of the carabid species and

subspecies there seems relatively high (543), if compared it with that of other Balkan

or Mediterranean regions. In Bulgaria there are known more than 720 species

(Guéorguiev & Guéorguiev, 1995), from Macedonia there are reported about 420

species (Guéorguiev, 1998; Hristovski, personal communication), and for ex-Yugoslavia

more than 1500 species and subspecies have been cited (Drovenik & Peks, 1999).

Both the Iberian Peninsula and Italy, having territories from 10 to 20 times bigger

than Albania, have a number of species only 2.13-2.42 times higher than is that in the

last country (Serrano, 2003; Vigna Taglianti, 2005a; 2005b).

Subfamily Tribe Subtribe Genus Nr. Subgenus Nr.

species species

Nebriinae Nebriini Eurynebria 1

Leistus 7 Leistus 2

Pogonophorus 5

Nebria 9 Alpaeus 5

Boreonebria 1

Eunebria 1

Nebria 2

Notiophilini Notiophilus 8

Cicindelinae Cicindelini Calomera 2

Cicindela 4 Cicindela 4

Cylindera 3 Cylindera 1

Eugrapha 2

Myriochila 1 Myriochila 1

Carabinae Carabini Calosoma 4 Calosoma 2

Microcallisthenes 2

Carabus 15 Archicarabus 1

Carabus 1

Chaetocarabus 1

Megodontus 3

Oreocarabus 3

Pachystus 2

Procerus 1

Procrustes 1

Tachypus 1

Tomocarabus 1

Cychrini Cychrus 1

Elaphrinae Elaphrini Elaphrus 1 Neoelaphrus 1

Omophroninae Omophronini Omophron 1 Omophron 1

Brachininae Brachinini Aptinus 3 Aptinus 3

Brachinus 14 Brachinus 5

Brachynidius 4

Brachynolomus 1

Cnecostolus 3

insertae sedis 1

Siagoninae Siagonini Siagona 1

Scaritinae Clivinini Clivinina Clivina 3

Reicheiina Reicheadella 1 Reicheadella 1

Dyschiriini Dyschirius 21 Chiridysus 1

Dyschiriodes 14

Dyschirius 1

Eudyschirius 3

Paradyschirius 2

Scaritini Scaritina Scarites 3 Parallelomorphus 2

Scallophorites 1

Apotominae Apotomini Apotomus 2

Trechinae Bembidiini Bembidiina Asaphidion 4

Bembidion 63 Bembidion 1

Taxonomic structure of Albanian Carabidae (Table 1)

TAXONOMIC STRUCTURE OF ALBANIAN CARABIDAE 189


Bembidionetolitzkya 9

Chlorodium 1

Diplocampa 2

Emphanes 5

Euperyphus 3

Metallina 2

Microserrullula 1

Nepha 3

Notaphemphanes 1

Notaphus 1

Ocydromus 3

Ocyturanes 6

Odontium 2

Omoperyphus 1

Peryphanes 4

Peryphus 4

Philochthus 6

Phyla 1

Princidium 1

Talanes 1

Terminophanes 1

Testedium 1

Trepanes 3

Ocys 1

Sinechostictus 3 Pseudolimnaeum 1

Sinechostictus 2

Tachyina Lymnastis 1

Paratachys 3

Polyderis 1

Sphaerotachys 1

Tachys 1

Tachyta 1 Tachyta 1

Tachyura 2 Tachyura 2

Patrobini Deltomerus 2 Deltomerus 1

Paradeltomerus 1

Pogonini Pogonistes 2

Pogonus 2 Pogonus 2

Trechini Perileptina Perileptus 1 Perileptus 1

Trechina Albanotrechus 1

Duvalius 10 Duvalius 9

Euduvalius 1

Neotrechus 3

Trechus 12 Trechus 12

Trechodina Thalassophilus 1

Harpalinae Chlaeniini Callistina Callistus 1

Chlaeniina Chlaenius 13 Chlaeniellus 6

Chlaenites 1

Chlaenius 1

Dinodes 2

Epomis 2

Trichochlaenius 1

Cyclosomini Masoreina Masoreus 1 Masoreus 1

Dryptini Drypta 1 Drypta 1


species species

Harpalini Amblystomina Amblystomus 3

Anisodactylina Anisodactylus 4 Anisodactylus 3

Pseudodichirius 1

Diachromus 1

Gynandromorphus 1

Ditomina Carterus 3 Carterus 3

Ditomus 2

Dixus 3

Graniger 1

Odontocarus 1

Harpalina Acinopus 4 Acinopus 2

Oedematicus 1

Osimus 1

Harpalus 32 Cryptophonus 1

Harpalus 29

Pseudophonus 2

Ophonus 20 Brachyophonus 1

Hesperophonus 4

Incisophonus 1

Metophonus 10

Ophonus 4

Parophonus 6 Ophonomimus 1

Parophonus 5

Trichotichnus 1 Trichotichnus 1

Stenolophina Acupalpus 10 Acupalpus 10

Anthracus 2

Bradycellus 4 Bradycellus 4

Dicheirotrichus 1 Dicheirotrichus 1

Loxoncus 1

Stenolophus 7 Egadroma 1

Stenolophus 6

Lebiini Cymindidina Cymindis 9 Cymindis 7

Menas 1

Tarulus 1

Dromiina Dromius 1 Dromius 1

Microlestes 7

Paradromius 1 Manodromius 1

Philorhizus 2 Philorhizus 2

Lebiina Lebia 5 Lamprias 1

Lebia 4

Lionychina Apristus 2

Lionychus 2 Lionychus 2

Syntomus 4

Licinini Licinina Badister 4 Badister 2

Baudia 2

Licinus 4 Licinus 3

Neorescius 1

Odacanthini Odacantha 1 Odacantha 1

Oodini Oodes 2 Oodes 2

Omphreini Omphreus 3 Omphreus 3

Panagaeini Panagaeus 1 Panadaeus 1

Platynini Agonum 15 Agonum 13

Europhilus 2


species species

TAXONOMIC STRUCTURE OF ALBANIAN CARABIDAE 191


Anchomenus 1 Anchomenus 1

Olisthopus 2

Paranchus 1

Platynus 2 Batenus 1

Platynus 1

Pterostichini Abax 2 Abacopercus 1

Abax 1

Molops 15 Molops 14

Stenochoromus 1

Myas 1 Myas 1

Pedius 2

Poecilus 8 Angoleus 1

Poecilus 7

Pterostichus 24 Argutor 1

Bothriopterus 1

Cheporus 1

Feronidius 1

Melanius 2

Morphnosoma 1

Parahaptoderus 1

Phonias 3

Platypterinus 1

Platysma 1

Pseudomaseus 4

Pterostichus 7

Speluncarius 2 Hypogium 1

Speluncarius 1

Stomis 2 Stomis 2

Tapinopterus 6 Tapinopterus 6

Sphodrini Atranopsina Platyderus 4 Platyderus 4

Calathina Calathus 14 Bedelinus 1

Calathus 6

Neocalathus 7

Synuchidius 1

Sphodrina Laemostenus 7 Antisphodrus 2

Laemostenus 3

Pristonychus 2

Synuchina Synuchus 1 Synuchus 1

Zabrini Amarina Amara 32 Amara 13

Amarocelia 1

Bradytus 2

Celia 3

Curtonotus 1

Paracelia 2

Parapercosia 1

Percosia 1

Xenocelia 4

Zezea 4

Zabrina Zabrus 8 Pelor 7

Zabrus 1

Zuphiini Polystichus 1

Zuphium 1 Zuphium 1

11 subfamilies 31 tribes 103 genera 543 species


species species

APPENDIX 193

Appendix

A. Faunistic contributions

A.1. New taxa for the fauna of the Balkan Peninsula

Subgenera

1. Terminophanes Müller-Motzfeld, 1998

Species

1. Bembidion (Terminophanes) terminale Heer, 1841

2. Cymindis (Cymindis) angularis Gyllenhal, 1810

3. Lionychus (Lionychus) maritimus Fairmaire, 1862

A.2. Taxa new for the fauna of Albania

Tribes

1. ODACANTHINI Laporte, 1834

Genera

1. Eurynebria Ganglbauer, 1891

2. Myriochila Motschulsky, 1858

3. Lymnastis Motschulsky, 1862

4. Polyderis Motschulsky, 1862

5. Pogonistes Chaudoir, 1871

6. Odontocarus Solier, 1835

7. Dromius Bonelli, 1810

8. Lionychus Wissmann, 1846

9. Odacantha Paykull, 1798

10. Paranchus Lindroth, 1974

Subgenera

1. Myriochila Motschulsky, 1858 s.str.

2. Dyschirius Bonelli, 1810 s.str.

3. Talanes Motschulsky, 1864

4. Terminophanes Müller-Motzfld, 1998

5. Pseudolimnaeum Kraatz, 1888

6. Menas Motschulsky, 1864

7. Dromius Bonelli, 1810 s.str.

8. Lamprias Bonelli, 1810

9. Lionychus Wissmann, 1846 s.str.

10. Odacanta Paykull, 1798 s.str.

11. Parahaptoderus Jeanne, 1969

Species (subspecies not considered)

1. Eurynebria complanata (Linnaeus, 1767)

2. Leistus (Pogonophorus) apfelbecki Ganglbauer, 1891


3. Nebria (Alpaeus) germarii Heer, 1837

4. Myriochila (Myriochila) melancholica (Fabricius, 1798)

5. Aptinus (Aptinus) acutangulus Chaudoir, 1876

6. Dyschirius (Dyschiriodes) luticola Chaudoir, 1850

7. Dyschirius (Dyschirius) latipennis Seidlitz, 1867

8. Scarites (Parallelomorphus) terricola Bonelli, 1813

9. Asaphidion stierlini (Heyden, 1880)

10. Bembidion (Bembidion) quadripustulatum Audinet-Serville, 1821

11. Bembidion (Euperyphus) testaceum (Duftschmid, 1812)

12. Bembidion (Peryphanes) deletum Audinet-Serville, 1821

13. Bembidion (Peryphus) femoratum Sturm, 1825

14. Bembidion (Philochthus) mannerhaimii C.R. Sahlberg, 1827

15. Bembidion (Talanes) subfasciatum Chaudoir, 1850

16. Bembidion (Terminophanes) terminale Heer, 1841

17. Bembidion (Trepanes) maculatum Dejean, 1831

18. Sinechostictus (Pseudolimnaeum) doderoi (Ganglbauer, 1891)

19. Sinechostictus (Sinechostictus) millerianus (Heyden, 1883)

20. Lymnastis galilaeus Piochard de la Brûlerie, 1876

21. Polyderis cardioderus (Chaudoir, 1850)

22. Pogonistes convexicollis Chaudoir, 1871

23. Pogonistes gracilis (Dejean, 1828)

24. Carterus (Carterus) rotundicollis Rambur, 1837

25. Odontocarus robustus (Dejean, 1831)

26. Harpalus (Harpalus) euchlorus Ménétriés, 1839

27. Harpalus (Harpalus) flavescens (Piller et Mitterpacher, 1783)

28. Harpalus (Harpalus) hospes Sturm, 1818

29. Ophonus (Hesperophonus) similis (Dejean, 1829)

30. Ophonus (Metophonus) melletii (Heer, 1837)

31. Ophonus (Ophonus) stictus Stephens, 1828

32. Acupalpus (Acupalpus) brunnipes (Sturm, 1825)

33. Stenolophus (Stenolophus) abdominalis Gené, 1836

34. Cymindis (Cymindis) angularis Gyllenhal, 1810

35. Cymindis (Cymindis) coadunata Dejean, 1825

36. Cymindis (Menas) miliaris (Fabricius, 1801)

37. Dromius (Dromius) quadrimaculatus (Linnaeus, 1758)

38. Lebia (Lamprias) cyanocephala (Linnaeus, 1758)

39. Lebia (Lebia) trimaculata (Villers, 1789)

40. Apristus reticulatus Schaum, 1857

41. Lionychus (Lionychus) maritimus Fairmaire, 1862

42. Lionychus (Lionychus) quadrillum (Duftschmid, 1812)

43. Badister (Badister) bullatus (Schrank, 1798)

44. Odacantha (Odacantha) melanura (Linnaeus, 1767)

45. Agonum (Agonum) afrum (Duftschmid, 1812)

46. Agonum (Agonum) muelleri (Herbst, 1784)

47. Paranchus albipes (Fabricius, 1796)

48. Poecilus (Poecilus) versicolor (Sturm, 1824)

APPENDIX 195

49. Pterostichus (Parahaptoderus) brevis (Duftschmid, 1812)

50. Pterostichus (Pseudomaseus) nigrita (Paykull, 1790)

51. Calathus (Calathus) ellipticus Reitter, 1889

52. Calathus (Neocalathus) ambiguus (Paykull, 1790)

53. Laemostenus (Laemostenus) dalmatinus (Dejean, 1828)

54. Amara (Amara) similata (Gyllenhal, 1810)

55. Amara (Bradytus) consularis (Duftschmid, 1812)

A.3. Taxa firstly reported with detailed record/s for Albania

Tribes

1. CYCLOSOMINI Laporte, 1834

Subtribes

1. MASOREINA Chaudoir, 1870

Genera

1. Tachys Dejean, 1821

2. Masoreus Dejean, 1821

3. Diachromus Erichson, 1837

4. Paradromius Fowler, 1887

5. Myas Sturm, 1826

6. Synuchus Gyllenhal, 1810

7. Polystichus Bonelli, 1810

Subgenera

1. Boreonebria Jeannel, 1937

2. Eunebria Jeannel, 1937

3. Procerus Dejean, 1821

4. Notaphemphanes Netolitzky, 1920

5. Omoperyphus Netolitzky, 1931

6. Phyla Motschulsky, 1844

7. Masoreus Dejean, 1821 s.str.

8. Pseudodichirius Lutshnik, 1921

9. Brachyophonus Sciaky, 1987

10. Incisophonus Sciaky, 1987

11. Tarulus Bedel, 1906

12. Manodromius Reitter, 1905

13. Batenus Motschulsky, 1865

14. Myas Sturm, 1826 s.str.

15. Bothriopterus Chaudoir, 1835

16. Morphnosoma Lutshnik, 1915

17. Platypterinus Reitter, 1908

18. Bedelinus Ragusa, 1885

19. Synuchus Gyllenhal, 1810 s.str.

20. Curtonotus Stephens, 1827

Species (subspecies not considered)

1. Leistus (Leistus) nitidus (Duftschmid, 1812)

2. Leistus (Pogonophorus) parvicollis Chaudoir, 1869


3. Nebria (Boreonebria) rufescens (Strøm, 1768)

4. Nebria (Eunebria) jockischi Sturm, 1815

5. Notiophilus aquaticus (Linnaeus, 1758)

6. Notiophilus germinyi Fauvel, 1863

7. Calosoma (Calosoma) inquisitor (Linnaeus, 1758)

8. Carabus (Procerus) gigas Creutzer, 1799

9. Clivina collaris (Herbst, 1784)

10. Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837

11. Bembidion (Emphanes) axillare (Motschulsky, 1844)

12. Bembidion (Notaphemphanes) ephippium (Marsham, 1802)

13. Bembidion (Odontium) foraminosum Sturm, 1825

14. Bembidion (Omoperyphus) hypocrita Dejean, 1831

15. Bembidion (Philochthus) guttula (Fabricius, 1792)

16. Bembidion (Phyla) tethys Netolitzky, 1926

17. Paratachys micros (Fischer von Waldheim, 1828)

18. Tachys scutellaris Stephens, 1828

19. Tachyura (Tachyura) parvula (Dejean, 1831)

20. Chlaenius (Chlaeniellus) nitidulus (Schrank, 1781)

21. Chlaenius (Chlaeniellus) tristis (Schaller, 1783)

22. Chlaenius (Dinodes) cruralis Fischer von Waldheim, 1829

23. Chlaenius (Epomis) circumscriptus (Duftschmid, 1812)

24. Masoreus (Masoreus) wetterhallii (Gyllenhal, 1813)

25. Anisodactylus (Pseudodichirius) intermedius Dejean, 1829

26. Diachromus germanus (Linnaeus, 1758)

27. Harpalus (Harpalus) laevipes Zetterstedt, 1828

28. Harpalus (Harplaus) pumilus Sturm, 1818

29. Harpalus (Harpalus) subcylindricus Dejean, 1829

30. Harpalus (Harpalus) latus (Linnaeus, 1758)

31. Harpalus (Harpalus) solitaris Dejean, 1829

32. Harpalus (Harpalus) autumnalis (Duftschmid, 1812)

33. Ophonus (Brachyophonus) krueperi Apfelbeck, 1902

34. Ophonus (Incisophonus) incisus (Dejean, 1829)

35. Ophonus (Metophonus) rufibarbis (Fabricius, 1792)

36. Ophonus (Metophonus) schaubergerianus (Puel, 1937)

37. Ophonus (Ophonus) ardosiacus (Lutshnik, 1922)

38. Acupalpus (Acupalpus) elegans (Dejean, 1829)

39. Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)

40. Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)

41. Acupalpus (Acupalpus) suturalis Dejean, 1829

42. Bradycellus (Bradycellus) distinctus (Dejean, 1829)

43. Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)

44. Cymindis (Tarulus) vaporariorum (Linnaeus, 1758)

45. Paradromius (Manodromius) linearis (Olivier, 1795)

46. Philorhizus (Philorhizus) notatus (Stephens, 1827)

47. Lebia (Lebia) humeralis Dejean, 1825

48. Lebia (Lebia) scapularis (Geoffroy, 1785)

APPENDIX 197

49. Licinus (Licinus) cassideus (Fabricius, 1792)

50. Agonum (Agonum) marginatum (Linnaeus, 1758)

51. Platynus (Batenus) scrobiculatus (Fabricius, 1801)

52. Abax (Abax) ovalis (Duftschmid, 1812)

53. Myas (Myas) chalybaeus (Palliardi, 1825)

54. Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)

55. Pterostichus (Morphnosoma) melanarius (Illiger, 1798)

56. Pterostichus (Phonias) diligens (Sturm, 1824)

57. Pterostichus (Platypterinus) ottomanus Apfelbeck, 1908

58. Calathus (Bedelinus) circumseptus Germar, 1824

59. Calathus (Calathus) bosnicus Ganglbauer, 1891

60. Calathus (Neocalathus) erratus (C.R. Sahlberg, 1827)

61. Calathus (Neocalathus) metallicus Dejean, 1828

62. Synuchus (Synuchus) vivalis (Illiger, 1798)

63. Amara (Amara) convexior Stephens, 1828

64. Amara (Amara) ovata (Fabricius, 1792)

65. Amara (Amara) proxima Putzeys, 1866

66. Amara (Amara) saphyrea Dejean, 1828

67. Amara (Celia) montana Dejean, 1828

68. Amara (Curtonotus) aulica (Panzer, 1796)

69. Amara (Paracelia) dalmatina Dejean, 1828

70. Amara (Xenocelia) messae Baliani, 1924

71. Amara (Zezea) tricuspidata Dejean, 1831

72. Polystichus connexus (Geoffroy, 1785)

A.4. Species omitted for Albania in Löbl & Smetana (2003); here their presence in the

country is confirmed

1. Leistus (Leistus) fulvibarbis Dejean, 1826

2. Leistus (Pogonophorus) magnicollis Motschulsky, 1866

3. Nebria (Eunebria) jockischi Sturm, 1815

4. Notiophilus aestuans Dejean, 1826

5. Notiophilus aquaticus (Linnaeus, 1758)

6. Notiophilus biguttatus (Fabricius, 1779)

7. Notiophilus danieli Reitter, 1897

8. Notiophilus germinyi Fauvel, 1863

9. Notiophilus palustris (Duftschmid, 1812)

10. Notiophilus rufipes Curtis, 1829

11. Notiophilus substriatus C.R. Waterhouse, 1833

12. Calomera fischeri (M.F. Adams, 1817)

13. Carabus (Oreocarabus) preslii Dejean, 1830

14. Elaphrus (Neoelaphrus) uliginosus Fabricius, 1792

15. Brachinus (Brachinus) elegans Chaudoir, 1842

16. Brachinus (Brachinus) plagiatus Reiche, 1868

17. Brachinus (Cnecostolus) bayardi Dejean, 1831

18. Dyschirius (Dyschiriodes) nitidus (Dejean, 1825)


19. Dyschirius (Dyschiriodes) cylindricus (Dejean, 1825)

20. Dyschirius (Dyschiriodes) laeviusculus Putzeys, 1846

21. Dyschirius (Eudyschirius) gracilis (Heer, 1837)

22. Scarites (Scallophorites) buparius (Forster, 1771)

23. Asaphidion flavipes (Linnaeus, 1761)

24. Bembidion (Bembidionetolitzkya) conforme (Dejean, 1831)

25. Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837

26. Bembidion (Bembidionetolitzkya) rhodopense Apfelbeck, 1902

27. Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)

28. Bembidion (Bembidionetolitzkya) varicolor (Fabricius, 1803)

29. Bembidion (Chlorodium) splendidum Sturm, 1825

30. Bembidion (Diplocampa) assimile Gyllenhal, 1810

31. Bembidion (Emphanes) axillare (Motschulsky, 1844)

32. Bembidion (Emphanes) latiplaga Chaudoir, 1850

33. Bembidion (Emphanes) tenellum Erichson, 1837

34. Bembidion (Euperyphus) scapulare Dejean, 1831

35. Bembidion (Metallina) properans (Stephens, 1828)

36. Bembidion (Microserrullula) quadricolle (Motschulsky, 1844)

37. Bembidion (Nepha) callosum Küster, 1847

38. Bembidion (Nepha) caucasicum (Motschulsky, 1844)

39. Bembidion (Notaphemphanes) ephippium (Marsham, 1802)

40. Bembidion (Ocyturanes) pindicum Apfelbeck, 1901

41. Bembidion (Odontium) foraminosum Sturm, 1825

42. Bembidion (Odontium) striatum (Fabricius, 1792)

43. Bembidion (Omoperyphus) hypocrita Dejean, 1831

44. Bembidion (Peryphanes) brunnicorne Dejean, 1831

45. Bembidion (Peryphanes) grandipenne Schaum, 1862

46. Bembidion (Peryphanes) tetracolum Say, 1823

47. Bembidion (Philochthus) guttula (Fabricius, 1792)

48. Bembidion (Philochthus) inoptatum Schaum, 1857

49. Bembidion (Philochthus) lunulatum (Geoffroy, 1785)

50. Bembidion (Phyla) tethys Netolitzky, 1926

51. Bembidion (Princidium) punctulatum Drapiez, 1820

52. Bembidion (Testedium) bipunctatum (Linnaeus, 1761)

53. Bembidion (Trepanes) octomaculatum (Goeze, 1777)

54. Ocys harpaloides (Audinet-Serville, 1821)

55. Sinechostictus (Sinechostictus) tarsicus (Peyron, 1858)

56. Paratachys bistriatus (Duftschmid, 1812)

57. Paratachys fulvicollis (Dejean, 1831)

58. Paratachys micros (Fischer von Waldheim, 1828)

59. Sphaerotachys haemorrhoidalis (Ponza, 1805)

60. Tachys scutellaris Stephens, 1828

61. Tachyta (Tachyta) nana (Gyllenhal, 1810)

62. Tachyura (Tachyura) diabrachys (Kolenati, 1845)

63. Tachyura (Tachyura) parvula (Dejean, 1831)

64. Pogonus (Pogonus) littoralis (Duftschmid, 1812)

APPENDIX 199

65. Chlaenius (Chlaeniellus) terminatus Dejean, 1826

66. Chlaenius (Epomis) dejeani (Dejean, 1831)

67. Masoreus (Masoreus) wetterhallii (Gyllenhal, 1813)

68. Harpalus (Harpalus) cupreus Dejean, 1829

69. Harpalus (Pseudophonus) griseus (Panzer, 1796)

70. Ophonus (Hesperophonus) subquadratus (Dejean, 1829)

71. Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)

72. Acupalpus (Acupalpus) parvulus (Sturm, 1825)

73. Bradycellus (Bradycellus) caucasicus (Chaudoir, 1846)

74. Cymindis (Cymindis) axillaris (Fabricius, 1794)

75. Cymindis (Cymindis) humeralis (Geoffroy, 1785)

76. Cymindis (Cymindis) lineola L. Dufour, 1820

77. Cymindis (Tarulus) vaporariorum (Linnaeus, 1758)

78. Microlestes luctuosus Holdhaus, 1904

79. Microlestes minutulus (Goeze, 1777)

80. Microlestes negrita (Wollaston, 1854)

81. Paradromius (Manodromius) linearis (Olivier, 1795)

82. Philorhizus (Philorhizus) notatus (Stephens, 1827)

83. Lebia (Lebia) cruxminor (Linnaeus, 1758)

84. Lebia (Lebia) humeralis Dejean, 1825

85. Lebia (Lebia) scapularis (Geoffroy, 1785)

86. Apristus subaeneus Chaudoir, 1846

87. Syntomus impressus (Dejean, 1825)

88. Syntomus pallipes (Dejean, 1825)

89. Syntomus truncatellus (Linnaeus, 1761)

90. Omphreus (Omphreus) morio Dejean, 1828

91. Agonum (Agonum) extensum Ménétriés, 1849

92. Agonum (Agonum) hypocrita (Apfelbeck, 1904)

93. Agonum (Agonum) marginatum (Linnaeus, 1758)

94. Agonum (Agonum) sexpunctatum (Linnaeus, 1758)

95. Agonum (Agonum) sordidum Dejean, 1828

96. Agonum (Agonum) viridicupreum (Goeze, 1777)

97. Agonum (Europhilus) antennarium (Duftschmid, 1812)

98. Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)

99. Olisthopus fuscatus Dejean, 1828

100. Olisthopus glabricollis (Germar, 1817)

101. Platynus (Batenus) scrobiculatus (Fabricius, 1801)

102. Platynus (Platynus) assimilis (Paykull, 1790)

103. Molops (Molops) alpestris (Dejean, 1828)

104. Molops (Molops) apfelbecki Ganglbauer, 1891 (considering former Molops reseri Apfelbeck, 1904)

105. Molops (Molops) curtulus Ganglbauer, 1891

106. Molops (Molops) holdhausi J. Müller, 1917

107. Molops (Molops) plurisetosus J. Müller, 1918

108. Molops (Molops) piceus (Panzer, 1793)

109. Pterostichus (Cheporus) burmeisteri Heer, 1838

110. Pterostichus (Platysma) niger (Schaller, 1783)


111. Pterostichus (Pterostichus) reiseri Ganglbauer, 1889

112. Platyderus (Platyderus) dalmatinus L. Miller, 1881

113. Calathus (Neocalathus) cinctus Motschulsky, 1850

114. Calathus (Neocalathus) mollis (Marsham, 1802)

115. Laemostenus (Antisphodrus) elongatus (Dejean, 1828)

116. Laemiostenus (Laemostenus) janthinus (Duftschmid, 1812)

117. Zabrus (Pelor) incrassatus (Ahrens, 1814)

118. Zabrus (Pelor) spinipes (Fabricius, 1798)

119. Polystichus connexus (Geoffroy, 1785)

A.5. Taxa still not well defined

1. Nebria (Alpaeus) germarii ssp.

2. Dalmatoreicheia sp.

3. Reicheadella (Reicheadella) sp.

4. Bembidion (Bembidionetolitzkya) sp. (aff. complanatum Heer, 1937)

5. Bembidion (Bembidionetolitzkya) sp. (ex. gruppo “fasciolatum”)

6. Bembidion (Terminophanes) terminale ssp.

7. Adriaphaenops sp.

8. Trechini gen. & sp. indet. (? aff. Pheggomisetes Knirsch, 1923)

9. Omphreus (Omphreus) ravasinii ssp.

10. Tapinopterus (Tapinopterus) protensus ssp.1

11. Tapinopterus (Tapinopterus) protensus ssp.2

12. Zabrus (Pelor) albanicus ssp.

A.6. Taxa (cited, mapped or found in collections) with doubtful presence in Albania

Subgenera

1. Platycarabus A. Morawitz, 1886

2. Trachycarabus Géhin, 1876

Species and subspecies

1. Nebria (Alpaeus) bosnica tresnjevikensis Horvatovich, 1974

2. Nebria (Alpaeus) dahli (Duftschmid, 1812) (considering ssp. montenegrina Apfelbeck, 1904)

3. Carabus (Platycarabus) irregularis Fabricius, 1792 (considering ssp. ramanus Sokolář, 1909)

4. Carabus (Trachycarabus) scabriusculus Olivier, 1795 (considering ssp. bulgarus Lapouge, 1908)

5. Siagona europaea oberleitneri Dejean, 1830

6. Dyschirius (Eudyschirius) gracilis (Heer, 1837) (considering nominotypical subspecies)

7. Asaphidion cyanicorne (Pandellé, 1867) (considering nominotypical subspecies)

8. Bembidion (Bembidion) humerale Sturm, 1825

9. Bembidion (Bembidion) quadrimaculatum (Linnaeus, 1761) (considering nominotypical subspecies)

10. Bembidion (Bembidionetolitzkya) relictum Apfelbeck, 1904

11. Deltomerus (Deltomerus) malissorum Apfelbeck, 1918

12. Deltomerus (Paradeltomerus) paradoxus paradoxus Apfelbeck, 1908

13. Pogonus (Pogonus) luridipennis (Germar, 1823)

14. Trechus (Trechus) irenis Csiki, 1912

15. Cymindis (Cymindis) pindicola Apfelbeck, 1901

APPENDIX 201

16. Olisthopus rotundatus (Paykull, 1790) (considering nominotypical subspecies)

17. Molops (Molops) spartanus parnassicola Kraatz, 1875

18. Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908

19. Pterostichus (?) epiroticus Csiki, 1930

20. Calathus (Calathus) glabricollis Dejean, 1828

21. Amara (Amara) famelica C. Zimmermann, 1832

22. Amara (Celia) praetermissa (C.R. Sahlberg, 1827)

A.7. Taxa deleted from the list of Albanian Carabidae

Subgenera

1. Eucarabus Géhin, 1876

2. Oreoplatysma Jakobson, 1907

Species and subspecies

1. Nebria (Nebria) salina Fairmaire & Laboulbène, 1854

2. Cicindela (Cicindela) hybrida Linnaeus, 1758

3. Carabus (Eucarabus) catenulatus Scopoli, 1763

4. Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812

5. Cychrus semigranosus semigranosus Palliardi, 1825

6. Cychrus semigranosus balcanicus Hopffgarten, 1881

7. Cychrus semigranosus montanus Winkler, 1939

8. Brachinus (Brachinus) alexandri F. Battoni, 1984

9. Dyschirius (Dyschiriodes) punctatus (Dejean, 1825)

10. Bembidion (Bembidionetolitzkya) ascendens K. Daniel, 1902

11. Bembidion (Bembidionetolitzkya) fasciolatum (Duftschmid, 1812)

12. Bembidion (Peryphanes) milleri Jacquelin du Val, 1852

13. Tachyura (Tachyura) sexstriata (Duftschmid, 1812)

14. Duvalius (Bicharotrechus) maglicensis Winkler, 1933

15. Trechus (Trechus) obtusiusculus Ganglbauer, 1889

16. Acinopus (Acinopus) subquadratus Brullé, 1832

17. Harpalus (Harpalus) anatolicus Tschitschérine, 1898

18. Harpalus (Harpalus) danieli Reitter, 1900

19. Molops (Molops) weiratheri J. Müller, 1930

20. Molops (Molops) simplex simplex Chaudoir, 1868

21. Molops (Molops) rufipes klisuranus Apfelbeck, 1902

22. Pterostichus (Oreoplatysma) lodosi Heinz, 1977

23. Tapinopterus (Tapinopterus) comita Jedlička, 1935

24. Tapinopterus (Tapinopterus) extensus (Dejean, 1831)

25. Tapinopterus (Tapinopterus) miridita jacupicensis Jedlička, 1935

26. Laemostenus (Antisphodrus) cavicola modestus (Schaufuss, 1862)

27. Amara (Amara) spreta Dejean, 1831

28. Amara (Zezea) erythrocnema Dejean, 1828

29. Zabrus (Pelor) apfelbecki Ganglbauer, 1915


B. Taxonomic contributions

B.1. Nomenclatorial acts

Nebriini

Nebria (Alpaeus) macedonica Mařan, 1938, stat. rest. for Nebria (Alpaeus) aetolica macedonica Mařan, 1938

Nebria (Alpaeus) macedonica galicica Horvatovich, 1974a, new assignment of Nebria (Alpaeus)

aetolica galicica Horvatovich, 1974a

Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974a, new assignment of Nebria (Alpaeus)

aetolica papingutica Horvatovich, 1974a

Nebria (Alpaeus) macedonica rambouseki Mařan, 1938, new assignment of Nebria (Alpaeus) aetolica

rambouseki Mařan, 1938

Carabini

Carabus (Oreocarabus) neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri

Schaum, 1856

Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii Dejean, 1830

Cychrini

Cychrus semigranosus albanicus Hopp, 1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck,

1904

Cychrus semigranosus dinaricus Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck,

1904

Licinini

Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn. nov. of Licinus (Licinus) graecus Apfelbeck, 1901

Platynini

Platynus (Batenus) scrobiculatus serbicus Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904

Platynus (Batenus) scrobiculatus turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus

Apfelbeck, 1904

Pterostichini

Molops (Molops) maderi J. Müller, 1933, syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917

Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri

reiseri Apfelbeck, 1904

Molops (Molops) apfelbecki corallipes J. Müller, 1936, new assignment of Molops (Molops) reiseri

corallipes J. Müller, 1936

Molops (Molops) apfelbecki pseudoalpestris Mlynář, 1977, new assignment of Molops (Molops) reiseri

pseudoalpestris Mlynář, 1977

Molops (Molops) apfelbecki victoris J. Müller, 1918, new assignment of Molops (Molops) reiseri

victoris J. Müller, 1918

Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J. Müller, 1917

Aphaon Heyden, 1906, syn. nov. of Pterostichus s. str. Bonelli, 1810

Pterostichus (Pterostichus) bischoffianus Jedlička, 1936, new assignment of Pterostichus (Aphaon)

bischoffianus Jedlička, 1936

APPENDIX 203

Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment of Pterostichus (Aphaon) brucki

Schaum, 1859

Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment of Pterostichus

(Aphaon) latifianus latifianus Apfelbeck, 1906

Pterostichus (Pterostichus) latifianus strupii J. Müller, 1935, new assignment of Pterostichus (Aphaon)

latifianus strupii J. Müller, 1935

Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment of Pterostichus

(Aphaon) lumensis lumensis Apfelbeck, 1905

Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment of Pterostichus (Aphaon) lumensis

litae Csiki, 1940

Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment of Pterostichus

(Aphaon) lumensis ljubetensis Apfelbeck, 1908

Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment of Pterostichus (Aphaon)


Pterostichus (Pterostichus) latifianus ab. nigripes Jedlička, 1936, syn. rest. for Pterostichus (Pterostichus)


Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment of Pterostichus (Aphaon) meisteri

Reitter, 1885

Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment of Pterostichus (Aphaon)

merklii (J. Frivaldszky, 1879)

Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment of Pterostichus (Aphaon) pentheri

Apfelbeck, 1918

Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment of Pterostichus (Aphaon) reiseri

Ganglbauer, 1889

Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment of Pterostichus

(Aphaon) rhilensis rhilensis Rottenberg, 1874

Pterostichus (Pterostichus) rhilensis kourili Mařan, 1933, new assignment of Pterostichus (Aphaon)

rhilensis kourili Mařan, 1933

Pterostichus (Pterostichus) rhilensis vitosensis Mařan, 1933, new assignment of Pterostichus (Aphaon)

rhilensis vitosensis Mařan, 1933

Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment of Pterostichus (Aphaon) walteri

Reitter, 1883

Tapinopterus (Tapinopterus) extensoides (Straneo, 1936), stat. rest. for Tapinopterus (Tapinopterus)

extensus extensoides (Straneo, 1936).

Sphodrini

Calathus (Calathus) ravasinii macedonicus Mařan, 1935, stat. nov. for Calathus (Calathus) macedonicus

Mařan, 1935

Zabrini

Zabrus (Pelor) bischoffi J. Müller, 1936, stat. rest. for Zabrus (Pelor) incrassatus bischoffi J. Müller, 1936.

B.2. Year of description

The following taxa (valid or synonyms) have long time been accredited to have a correct year

of description. However, it was found out that this is incorrect. Below, the relevant earlier


years of publication dates of the original descriptions are given. All the descriptions considered

fulfil the conditions of the ICZN (1999).

Carabus croaticus ljubetensis Apfelbeck, 1917 (synonym of Carabus croaticus durmitorensis Apfelbeck,

1904) instead Carabus croaticus ljubetensis Apfelbeck, 1918a

Carabus violaceus merditanus Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857)

instead Carabus violaceus merditanus Apfelbeck, 1918a

Carabus violaceus shardagensis Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857)

instead Carabus violaceus shardagensis Apfelbeck, 1918a

Aptinus merditanus Apfelbeck, 1917 (valid species) instead Aptinus merditanus Apfelbeck, 1918b

Aptinus merditanus var. granulipennis Apfelbeck, 1917 (synonym of Aptinus merditanus Apfelbeck,

1917) instead Aptinus merditanus var. granulipennis Apfelbeck, 1918b

Trechus albanicus Apfelbeck, 1905 (valid species) instead Trechus albanicus Apfelbeck, 1907b

Ophonus krueperi Apfelbeck, 1902 (valid species) instead Ophonus krueperi (Apfelbeck, 1904)

Licinus graecus Apfelbeck, 1901 (valid species) instead Licinus cassideus graecus Apfelbeck, 1904

Molops malshentianus Apfelbeck, 1917 (valid species) instead Molops malshentianus Apfelbeck, 1918b

B.3. Nomina nuda found in both collections and literature

Eurynebria complanata orientalis Apfelbeck

Eurynebria complanata albanica Apfelbeck

Eurynebria complanata var. albanica Apfelbeck

? Nebria ganglbaueri ditana Apfelbeck (or incorrect spelling of Nebria ganglbaueri merditana

Apfelbeck, 1906)

Aptinus lugubris rufipes Apfelbeck (according to Apfelbeck, 1907a: 499, 501)

Bembidion merditanum Apfelbeck

Harpalus bischoffi Schauberger

Molops transfuga Apfelbeck (according to Müller, 1930: 124; Mlynář, 1977: 120)

Molops apfelbecki bischoffi J. Müller

Pterostichus djumanus Apfelbeck

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Index

(valid taxa are given in bold)

Abacopercus 138

Abax 138

Abax s.str. 139

abdominalis (Stenolophus) 119

abdominalis persicus (Stenolophus) 119

Acinopus 97

Acinopus s.str. 97

Acupalpus 12, 115

Acupalpus s.str. 115

acutangulus (Aptinus) 39

Adriaphaenops 79

aenea (Amara) 174

aeneocephalus (Chlaenius) 92

aeneocephalus aeneocephalus (Chlaenius) 92

aeneus (Dyschirius) 46

aeneus (Harpalus) 107

aeneus aeneus (Dyschirius) 46

aestuans (Notiophilus) 23

aetolica (Nebria) 18

aetolica galicica (Nebria) 18, 21

aetolica korabica (Nebria) 21

aetolica macedonica (Nebria) 18

aetolica papingutica (Nebria) 18, 21, 22

aetolica rambouseki (Nebria) 18

aetolica tomoricensis (Nebria) 21, 22

aetolicum (Bembidion) 63

aetolicum aetolicum (Bembidion) 63

aetolicus (Carabus) 30

aetolicus (Tapinopterus) 163

aetolicus peristericus (Tapinopterus) 162

aetolicus tomoricensis (Tapinopterus) 163

aetolus (Zabrus) 182

aetolus kodymi (Zabrus) 182

aetolus winkleri (Zabrus) 182

affinis (Harpalus) 107

affinis ab. limbopunctatus (Harpalus) 107

affinis ab. viridulus (Harpalus) 107

afrum (Agonum) 133

agnatus (Dyschirius) 46

Agonum 133

Agonum s.str. 133

albanensis (Carabus) 31

albanensis (Harpalus) 111

albanensis (Ophonus) 111

albanicola (Carabus) 34

albanicum (Bembidion) 68

albanicum (Hypogeobium) 160

albanicum (Platysma) 160

albanicus (Calathus) 169, 171

albanicus (Carabus) 35

albanicus (Cychrus) 37

albanicus (Harpalus) 103

albanicus (Molops) 142

albanicus (Omphreus) 132

albanicus (Ophonus) 111

albanicus (Pterostichus) 160

albanicus (Speluncarius) 160

albanicus (Tapinopterus) 160

albanicus (Trechus) 87

albanicus (Zabrus) 182, 183

albanicus albanicus (Zabrus) 182

albanicus latifianus (Zabrus) 182, 183

albanicus jablanicensis (Zabrus) 182

Albanotrechus 79

albipes (Paranchus) 137

alexandri (Brachinus) 40

alinae (Molops) 140

Alpaeus 18

alpestre (Agonum) 133

alpestris (Molops) 141

alpestris imitator (Molops) 141

Amara 12, 174

Amara s.str. 174

AMARINA 174

Amarocelia 177

ambiguus (Calathus) 169

ambiguus (Chlaenius) 91

ambiguus ambiguus (Calathus) 169

INDEX 225

AMBLYSTOMINA 93

Amblystomus 93

ammophilus (Acinopus) 98

anatolicus (Harpalus) 104

anatolicus anatolicus (Harpalus) 104

Ancholeus 149

Anchomenus 136

Anchomenus s.str. 136

andreae (Bembidion) 68

andreae albanicum (Bembidion) 68

andreae bualei (Bembidion) 68

angularis (Cymindis) 121

angustatum (Agonum) 134

ANISODACTYLINA 94

Anisodactylus 12, 94

Anisodactylus s.str. 94

antennarium (Agonum) 136

antennarius (Platynus) 136

anthobia (Amara) 174

anthracinus (Pterostichus) 154

anthracinus anthracinus (Pterostichus) 154

anthracinus biimpressus (Pterostichus) 155

Anthracus 117

Antisphodrus 171

anxius (Harpalus) 102

apfelbecki (Leistus) 17

apfelbecki (Molops) 143

apfelbecki (Nebria) 19

apfelbecki (Pterostichus) 153

apfelbecki (Zabrus) 183

apfelbecki apfelbecki (Molops) 143

apfelbecki bischoffi (Molops) 143

apfelbecki corallipes (Molops) 142

apfelbecki kulti (Nebria) 19

apfelbecki plurisetosus (Molops) 145

apfelbecki pseudoalpestris (Molops) 142

apfelbecki reiseri (Molops) 142, 143

apfelbecki victoris (Molops) 142, 143

Aphaenopsis 79

Aphaon 155

apicalis (Dyschirius) 47

APOTOMINAE 51, 187

APOTOMINI 51

Apotomus 51

apricaria (Amara) 177

Apristus 127

apterus (Microlestes) 124

Aptinus 39

Aptinus s.str. 39

aquaticus (Notiophilus) 24

Archicarabus 30

ardosiacus (Ophonus) 112

arenaria (Cicindela) 28

arenaria (Cylindera) 28

arenaria viennensis (Cylindera) 28

areolatus (Perileptus) 79

areolatus areolatus (Perileptus) 79

Argutor 151

articulatum (Bembidion) 72

Asaphidion 52

ascendens (Bembidion) 56, 57

assimile (Bembidion) 58

assimilis (Platynus) 138

ATRANOPSINA 165

atratus (Harpalus) 100

atratus var. subsinuatus (Harpalus) 100

atrocaeruleum (Bembidion) 56

attemsi (Nebria) 19

attenuatus (Harpalus) 99

atterimus (Pterostichus) 152

atterimus atterimus (Pterostichus) 152

aulica (Amara) 178

auriculatus (Dyschirius) 47

austriacus (Molops) 147

autumnalis (Harpalus) 104

axillare (Bembidion) 58

axillare occiduum (Bembidion) 58

axillaris (Cymindis) 121

axillaris axillaris (Cymindis) 121

axillaris adriatica (Cymindis) 121

axillaris lineola (Cymindis) 122

azurescens (Bembidion) 59

azurescens azurescens (Bembidion) 59

azureus (Chlaenius) 91

azureus (Harpalus) 109

azureus (Ophonus) 109, 110

azureus oberthuri (Harpalus) 109

babinjensis (Carabus) 32

babinjensis (Trechus) 85

bacillus (Dyschirius) 47

bacillus arbensis (Dyschiriodes) 47

bacillus arbensis (Dyschirius) 47


bacillus bacillus (Dyschirius) 47

bacillus lucanus (Dyschirius) 47

Badister 128

Badister s.str. 128

balcanicum (Bembidion) 64

balcanicum balcanicum (Bembidion) 64, 65, 66

baldori sic! (Zabrus) 184

Batenus 138

Baudia 129

baudii (Acinopus) 97

baudii hellenicus (Acinopus) 97, 98

bayardi (Brachinus) 43

Bedelinus 166

BEMBIDIINA 52

BEMBIDIINI 12, 52

Bembidion 12, 53

Bembidion s.str. 53

Bembidionetolitzkya 54

beroni (Albanotrechus) 12, 79

Bicharotrechus 79

bifrons (Amara) 178

biguttatus (Notiophilus) 24

binaghii (Agonum) 133

binotatus (Anisodactylus) 94

binotatus ab. spurcaticornis (Anisodactylus) 94

bipunctatum (Bembidion) 71

bipunctatum nivale (Bembidion) 71

bipunctatum var. pyritosum (Bembidion) 71

bipustulatus (Brachinus) 43

bisbimaculatus (Tachys) 76

bischoffi (Amara) 180

bischoffi (Duvalius) 80

bischoffi (Harpalus) 105

bischoffi (Neotrechus) 83

bischoffi (Reicheadella) 45

bischoffi (Reicheia) 45

bischoffi (Tapinopterus) 163, 164

bischoffi (Zabrus) 184

bischoffianus (Pterostichus) 155, 156

bistriatus (Paratachys) 74

bistriatus (Tachys) 74

bodemeyeri (Brachinus) 41

boldorii (Zabrus) 184

bombarda (Aptinus) 39

Boreonebria 22

bosnica (Nebria) 19

bosnica apfelbecki (Nebria) 19

bosnica jezercensis (Nebria) 19

bosnica kulti (Nebria) 19

bosnica prokletiensis (Nebria) 19

bosnica sturanyi (Nebria) 19

bosnica tresnjevikensis (Nebria) 20

bosnicus (Calathus) 166

Bothriopterus 151

Brachynidius 41

BRACHININAE 7, 39, 187

BRACHININI 39

Brachynolomus 42

Brachinus 40

Brachinus s.str. 40

Brachyophonus 109

Bracteon 57

Bradycellus 12, 118

Bradycellus s.str. 118

Bradytus 177

brevicollis (Brachinus) 42

brevicollis (Nebria) 22, 23

brevicollis (Ophonus) 111

brevicollis var. salina (Nebria) 22, 23

brevis (Pterostichus) 153

brevis brevis (Pterostichus) 153

brucki (Pterostichus) 155, 156

brunnicorne (Bembidion) 66

brunnicorne brunnicorne (Bembidion) 66

brunnipes (Acupalpus) 115

brunoi (Bembidion) 56, 57

bualei (Bembidion) 68

bualei bualei (Bembidion) 68

bucephalus (Carabus) 35

bullatus (Badister) 128

buparius (Scarites) 51

burmeisteri (Pterostichus) 151

burmeisteri burmeisteri (Pterostichus) 151

caelatus (Carabus) 31

caelatus caelatus (Carabus) 31

caelatus caelatus m. malissorum (Carabus) 31

caelatus caelatus n. metalkanus (Carabus) 31

caelatus dalmatinus (Carabus) 32

caelatus dalmatinus n. procerus (Carabus) 32

caelatus malissorum (Carabus) 31

caelatus metalkanus (Carabus) 31

caelatus metalkanus v. malissorum (Carabus) 31

INDEX 227

caelatus sarajevensis sic! (Carabus) 31

caelatus sarajevoensis (Carabus) 31

caelatus var. procerus (Carabus) 32

CALATHINA 166

Calathus 166

Calathus s.str. 166

CALLISTINA 89

Callistus 89

callosum (Bembidion) 61

callosum callosum (Bembidion) 61

Calomera 25

Calosoma 29

Calosoma s.str. 29

calydonius (Ditomus) 95

calydonius calydonius (Ditomus) 95

campestris (Cicindela) 26, 27

campestris campestris (Cicindela) 26

campestris olivieria (Cicindela) 26, 27

campestris pontica (Cicindela) 26

cancellatus (Carabus) 36

cancellatus alessiensis (Carabus) 36

cancellatus emarginatus (Carabus) 36

cancellatus subgraniger var. alessiensis (Carabus) 36

cancellatus var. alessiensis (Carabus) 36

CARABINAE 7, 29, 187

CARABINI 29

caraboides (Asaphidion) 52, 53

caraboides balcanicus (Asaphidion) 52

caraboides var. balcanicus (Asaphidion) 52

Carabus 29

Carabus s.str. 30

carbonarium (Agonum) 133

carbonarium carbonarium (Agonum) 133

cardioderus (Polyderis) 74

cardioderus balcanicus (Trechus) 86

carinatus (Abax) 138

carinatus carinatus (Abax) 138

carinatus var. latus (Abax) 138

Carterus 95

Carterus s.str. 95

cassideus (Licinus) 129

cassideus cassideus (Licinus) 129

cassideus dohrni (Licinus) 129

cassideus graecus (Licinus) 129, 130

cassideus latus (Licinus) 129

catenatus (Carabus) 31

catenatus var. albanensis (Carabus) 31

catenulatus (Carabus) 31

catenulatus catenulatus (Carabus) 31

caucasicum (Bembidion) 62

caucasicus (Bradycellus) 118

cavernosus (Carabus) 34

cavernosus cavernosus (Carabus) 34

cavernosus n. cavernosus (Carabus) 34

cavernosus var. sterbai (Carabus) 34

cavicola (Laemostenus) 171

cavicola albanicus (Laemostenus) 171

cavicola modestus (Laemostenus) 171, 172

cavicola var. modestus (Laemostenus) 171, 172

Celia 178

centralis (Harpalus) 98

Cephalota 26

Chaetocarabus 30

chalybaeus (Myas) 148

chalybaeus (Pterostichus) 158

chalybeus (Dyschirius) 49

chalybeus gibbifrons (Dyschirius) 49

chaudoiri (Amara) 181

chaudoiri incognita (Amara) 181

Cheporus 151

chionobatus (Carabus) 30

chionophilus (Carabus) 36

Chiridysus 46

Chlaeniellus 89

CHLAENIINA 89

CHLAENIINI 89

Chlaenites 90

Chlaenius 89

Chlaenius s.str. 91

Chlorodium 58

Cicindela 26

Cicindela s.str. 26

CICINDELINAE 7, 12, 25, 187

CICINDELINI 25

cimmerius (Laemostenus) 173

cimmerius (Pristonychus) 173

cimmerius (Sphodrus) 173

cimmerius cimmerius (Laemostenus) 173

cimmerius weiratheri (Laemostenus) 173

cinctus (Calathus) 169, 170, 171

circumdata (Cephalota) 26

circumdata circumdata (Cephalota) 26


circumscriptus (Chlaenius) 92

circumseptus (Calathus) 166

clathratus (Carabus) 31

clathratus auraniensis (Carabus) 31

Clivina 44

CLIVININA 44

CLIVININI 44

clypeatus (Aristus) 96

clypeatus (Ditomus) 96

clypeatus (Dixus) 96

Cnecostolus 43

coadunata (Cymindis) 121

coadunata coadunata (Cymindis) 121

coerulescens (Poecilus) 151

coerulescens (Pterostichus) 151

coeruleum (Bembidion) 57

coeruleum concoeruleum (Bembidion) 57

collaris (Bradycellus) 118

collaris (Clivina) 44

columbinus (Ophonus) 113

combustum (Bembidion) 60, 78

combustum combustum (Bembidion) 60

comita (Tapinopterus) 164

complanata (Amara) 180

complanata (Eurynebria) 16

complanata albanica (Eurynebria) 16

complanata orientalis (Eurynebria) 16

complanata var. albanica (Eurynebria) 16

complanatum (Bembidion) 55

concoeruleum (Bembidion) 57

concoeruleus (Ocydromus) 57

conforme (Bembidion) 55

connexus (Polystichus) 185

consularis (Amara) 178

convexicollis (Pogonistes) 78

convexior (Amara) 175

convexiusculus (Argutor) 153

convexiusculus (Pterostichus) 153

convexiusculus (Tapinopterus) 161

convexus (Carabus) 36

convexus dilatatus (Carabus) 36

convexus dilatatus n. miliusi (Carabus) 36

convexus dilatatus var. miliusi (Carabus) 36

convexus miliusi (Carabus) 36

convexus var. chionophilus (Carabus) 36

corax (Pterostichus) 160

corax sic! (Calathus) 168

cordatus (Ophonus) 111

coriaceus (Carabus) 35

coriaceus cerisyi (Carabus) 35

coriaceus cerisyi m. weiratheriana (Carabus) 35

coriaceus excavatus (Carabus) 35

coriaceus excavatus dalmaticus (Carabus) 36

coriaceus excavatus m. dalmatica (Carabus) 35

coriaceus excavatus n. joniensis (Carabus) 35

coriaceus joniensis (Carabus) 36

coriaceus joniensis var. albanicus (Carabus) 36

coriaceus mediterraneus (Carabus) 36

coriaceus rugosus (Carabus) 35

coriaceus rugosus n. joniensis m. albanicus (Carabus) 35

coriaceus subrugosus (Carabus) 35

coriaceus subrugosus v. weiratherianus (Carabus) 35

coriaceus var. hopffgarteni (Carabus) 35

corticalis (Microlestes) 123

corticalis escorialensis (Microlestes) 123

crepitans (Brachinus) 40

crepitans ab. fallax (Brachinus) 40

cribricollis (Harpalus) 110

cribricollis (Ophonus) 110

croaticus (Carabus) 32

croaticus babinjensis (Carabus) 32

croaticus durmitorensis (Carabus) 32

croaticus durmitorensis var. babinjensis (Carabus) 32

croaticus durmitorensis var. ljubetensis (Carabus) 32

croaticus ljubetensis (Carabus) 32

croaticus ljubetensis m. durmitorensis (Carabus) 32

croaticus ljubetensis n. ljubetensis (Carabus) 32

cruciatum (Bembidion) 68

cruciatum albanicum (Bembidion) 68

cruralis (Chlaenius) 91

cruxmajor (Panagaeus) 133

cruxmajor ab. schaumi (Panagaeus) 133

cruxminor (Lebia) 126

cruxminor cruxminor (Lebia) 126

Cryptophonus 98

cupreus (Harpalus) 105

cupreus (Poecilus) 149

cupreus (Pterostichus) 149

cupreus cupreus (Poecilus) 149

cupreus fastuosus (Harpalus) 105

cursitans (Amara) 180

cursor (Pterostichus) 151

INDEX 229

cursorius (Poecilus) 150

cursorius (Pterostichus) 150

cursorius cursorius (Poecilus) 150

curta (Amara) 175

Curtonotus 178

curtulus (Molops) 144

curtulus cakorensis (Molops) 144

curtulus winklerianus (Molops) 144

cyaneomicans (Bembidion) 61

cyanicorne (Asaphidion) 52

cyanicorne cyanicorne (Asaphidion) 52

cyanocephala (Lebia) 125, 126

cyanocephala cyanocephala (Lebia) 125

CYCHRINI 37

Cychrus 37

CYCLOSOMINI 92

Cylindera 28

Cylindera s.str. 28

cylindricus (Dyschirius) 47

cylindricus hauseri (Dyschirius) 47

CYMINDIDINA 121

Cymindis 121

Cymindis s.str. 121

dahli (Nebria) 20

dahli montenegrina (Nebria) 20

dahli velebitica (Nebria) 20

dalmaticum sic! (Bembidion) 67

dalmaticus (Carabus) 35

dalmatina (Amara) 179

dalmatinum (Bembidion) 67, 90

dalmatinum dalmatinum (Bembidion) 67

dalmatinus (Laemostenus) 172

dalmatinus (Platyderus) 165

dalmatinus dalmatinus (Platyderus) 165

dalmatinus malissorum (Neotrechus) 84

dalmatinus pentheri (Neotrechus) 84

dalmatinus procerus (Carabus) 32

dalmatinus suturalis var. pentheri (Neotrechus) 84

Dalmatoreicheia 45

dama (Carterus) 95

dama (Ditomus) 95

danieli (Harpalus) 104

danieli (Notiophilus) 24

decipiens (Chlaenius) 91

decipiens laticollis (Chlaenius) 91

decipiens var. ambiguus (Chlaenius) 91

decolor (Bembidion) 69

decorum (Bembidion) 63

decorum decorum (Bembidion) 63

dejeani (Chlaenius) 92

dejeani (Epomis) 92

dejeani (Parophonus) 114

deletum (Bembidion) 67

deletum deletum (Bembidion) 67

Deltomerus 76

Deltomerus s.str. 76

dentata (Drypta) 93

depressa (Siagona) 44

detonii (Tapinopterus) 164

diabrachys (Tachys) 76

diabrachys (Tachyura) 76

Diachromus 94

Dicheirotrichus 118

Dicheirotrichus s.str. 118

didonnai (Duvalius) 80

diffinis (Harpalus) 113

diffinis (Ophonus) 113

dilatatus (Badister) 129

diligens (Pterostichus) 153

dimidiatus (Harpalus) 105

dinaricus (Cychrus) 37

Dinodes 91

Diplocampa 58

discophorus (Acupalpus) 119

discophorus (Stenolophus) 119

distinctissimus (Trechus) 87

distinctus (Bradycellus) 118

distinguendus (Calathus) 167

distinguendus (Harpalus) 107

distinguendus distinguendus (Harpalus) 107

DITOMINA 95

Ditomus 95

Dixus 96

djumanus (Pterostichus) 158

dochii (Pterostichus) 164

dochii (Tapinopterus) 164

doderoi (Sinechostictus) 73

dorsale (Platynus) 136

dorsalis (Acupalpus) 117

dorsalis (Anchomenus) 136

dorsalis (Platynus) 136


DROMIINA 123

Dromius 123

Dromius s.str. 123

droveniki (Duvalius) 79

Drypta 93

Drypta s.str. 93

DRYPTINI 93

duftschmidi (Agonum) 133

Duvalius 79

Duvalius s.str. 80

dyrrhachii (Bembidion) 63

DYSCHIRIINI 46

Dyschiriodes 46

Dyschirius 12, 46

Dyschirius s.str. 49

effluviorus (Sinechostictus) 73

Egadroma 119

egregius (Trechus) 87

ejaculans (Brachinus) 41

ELAPHRINAE 38, 187

ELAPHRINI 38

Elaphrus 38

elegans (Acupalpus) 115

elegans (Bembidion) 52, 53, 57, 58

elegans (Brachinus) 41

ellipticus (Calathus) 167

ellipticus ellipticus (Calathus) 167

elongatum (Bembidion) 73

elongatum tarsicum (Bembidion) 73

elongatum var. effluviorum (Bembidion) 73

elongatus (Laemostenus) 172

elongatus (Pterostichus) 152

elongatus (Sinechostictus) 73

elongatus pentheri (Laemostenus) 172

emarginatus (Acinopus) 97

Emphanes 58

eomita sic! (Tapinopterus) 164

ephippium (Bembidion) 62

epirensis (Carabus) 31

epiroticus (Pterostichus) 160

Epomis 92

equestris (Amara) 179, 180

equestris equestris (Amara) 179, 180

eremita (Ditomus) 96

eremita (Dixus) 96

erratica (Amara) 177

erratus (Calathus) 170

erratus erratus (Calathus) 170

erythrocnema (Amara) 181

erythroderus (Calathus) 169

escorialensis (Microlestes) 123

etruscus (Gynandromorphus) 95

etruscus etruscus (Gynandromorphus) 95

Eucarabus 31

euchlorus (Harpalus) 105

Euduvalius 82

Eudyschirius 49

Eugrapha 28

Eunebria 22

Euperyphus 60

europaea (Siagona) 44

europaea europaea (Siagona) 44

europaea oberleitneri (Siagona) 44

europaea var. oberleitneri (Siagona) 44

Europhilus 136

Eurynebria 16

eurynota (Amara) 175

exhalans (Brachinus) 43

exilis (Microlestes) 124

exilis luctuosus (Microlestes) 124

explodens (Brachinus) 42

explodens ab. glabratus (Brachinus) 42

explodens ab. strepens (Brachinus) 42

explodens sichemita (Brachinus) 42

extensoides Jedlička, 1936 (Tapinopterus) 161, 162

extensoides Straneo, 1936 (Tapinopterus) 161, 162

extensum (Agonum) 133

extensus (Tapinopterus) 161, 162

extensus abreshensis (Tapinopterus) 161

extensus convexiusculus (Pterostichus) 161

extensus extensoides (Tapinopterus) 161, 162

fallaciosa (Amara) 179

fallaciosus (Zabrus) 179

fallax (Brachinus) 40

fallax (Parophonus) 114

fallax (Tachyophonus) 114

famelica (Amara) 175

familiaris (Amara) 175

fasciatopunctatus (Pterostichus) 155

fasciolatum (Bembidion) 56, 57

femoralis (Carterophonus) 96

femoralis (Graniger) 96

INDEX 231

femoratum (Bembidion) 68

femoratum femoratum (Bembidion) 68

Feronidius 152

festivus (Chlaenius) 91

festivus festivus (Chlaenius) 91

fischeri (Calomera) 25

fischeri (Cicindela) 25

fischeri fischeri (Calomera) 25

fissuralis (Microlestes) 123

flavescens (Harpalus) 103

flavicornis (Harpalus) 101

flavicornis flavicornis (Harpalus) 101

flavipes (Asaphidion) 52, 53

flavipes (Chlaenius) 89

fodori (Duvalius) 82

foraminosum (Bembidion) 66

fossor (Clivina) 45

fossor fossor (Clivina) 45

franziniorum (Ophonus) 113

fuchsi (Duvalius) 81

fugax (Harpalus) 108

fulvibarbis (Leistus) 16

fulvibarbis fulvibarbis (Leistus) 16

fulvicollis (Paratachys) 74

fulvicollis (Tachys) 74

fulvipes (Amara) 181

fumigatum (Bembidion) 58

fusca (Amara) 180

fuscatus (Olisthopus) 137

fuscipes (Calathus) 167

fuscipes fuscipes (Calathus) 167

fuscipes graecus (Calathus) 167

fuscipes var. graecus (Calathus) 167

fuscipes var. latus (Calathus) 167

galilaeus (Lymnastis) 73

ganglbaueri (Brachinus) 41

ganglbaueri (Bradycellus) 118

ganglbaueri (Nebria) 20

ganglbaueri (Synuchidius) 171

ganglbaueri (Zabrus) 183

ganglbaueri ditana (Nebria) 21

ganglbaueri ganglbaueri (Nebria) 20

ganglbaueri ganglbaueri (Zabrus) 183

ganglbaueri gyalicaensis (Nebria) 20

ganglbaueri korabensis (Nebria) 20

ganglbaueri lonae (Zabrus) 183

ganglbaueri merditana (Nebria) 20, 21

ganglbaueri zebiaensis (Nebria) 21

genei illigeri (Bembidion) 62

geniculatum (Bembidion) 55

geniculatum geniculatum (Bembidion) 55

georgii (Duvalius) 80

germanica (Cicindela) 28

germanica (Cylindera) 28

germanica germanica (Cylindera) 28

germanica muelleri (Cylindera) 28

germanus (Diachromus) 94

germarii (Nebria) 21

germarii durmitorensis (Nebria) 21

germinyi (Notiophilus) 24

gestroi (Tapinopterus) 165

gibbifrons (Dyschirius) 49

giganteus (Calathus) 167, 168

gigas (Carabus) 35

gigas gigas (Carabus) 35

gigas parnassicus (Carabus) 35

gilvipes (Carterus) 95

gisellae (Agonum) 134

glabratus (Brachinus) 42

glabricollis (Calathus) 168

glabricollis (Olisthopus) 137

globosus (Dyschirius) 49

gracilis (Dyschirius) 49, 50

gracilis (Oodes) 131

gracilis (Pogonistes) 78

gracilis (Pterostichus) 155

gracilis gracilis (Dyschirius) 49

gracilis gracilis (Pterostichus) 155

graecus (Carabus) 35

graecus (Licinus) 129, 130

graecus (Platyderus) 165

graecus graecus (Carabus) 35

graecus morio (Carabus) 35

graecus morio var. neowinkleri (Carabus) 35

graecus morio var. winkleri (Carabus) 35

graecus n. winkleri (Carabus) 35

graecus winkleri (Carabus) 35

grandipenne (Bembidion) 67

grandipenne grandipenne (Bembidion) 67

Graniger 96

granulatus (Carabus) 30

granulatus aetolicus (Carabus) 30


granulatus granulatus m. merdita sic! (Carabus) 30

granulatus granulatus n. interstitialis (Carabus) 30

granulatus interstitialis (Carabus) 30

granulatus interstitialis v. miridita (Carabus) 30

granulatus miridita (Carabus) 30

granulatus m. miridita (Carabus) 30

granulatus var. miridita (Carabus) 30

granulipennis (Aptinus) 39

griseus (Harpalus) 108

gudenzii (Bembidion) 64

guttula (Bembidion) 69

guttula guttula (Bembidion) 69

gyllenhali (Nebria) 22

Gynandromorphus 95

haemorrhoidalis (Sphaerotachys) 75

haemorrhoidalis (Tachys) 75

hannemanni (Amara) 178

HARPALINA 97

HARPALINAE 7, 89, 187

HARPALINI 12, 93

harpaloides (Ocys) 72

Harpalus 12, 98

Harpalus s.str. 99

helopioides (Oodes) 131

helopioides helopioides (Oodes) 131

herzegowinensis (Carabus) 34

Hesperophonus 109

hilleri (Carabus) 34

hirsutulus (Harpalus) 113

hirsutulus (Ophonomimus) 113

hirsutulus (Ophonus) 113

hirsutulus (Parophonus) 113

holdhausi (Agonum) 134

holdhausi (Molops) 142, 143, 144, 145

holdhausi (Platynus) 134

honestus (Harpalus) 99

honestus honestus (Harpalus) 99

hopffgarteni (Carabus) 35

hortensis (Carabus) 33, 34

hortensis hortensis (Carabus) 34

hortensis neumeyeri (Carabus) 33, 34

hortensis neumeyeri n. herzegowinensis (Carabus) 34

hortensis neumeyeri n. neumeyeri (Carabus) 34

hortensis neumeyeri var. albanicola (Carabus) 34

hortensis neumeyeri var. hilleri (Carabus) 34

hortensis neumeyeri var. pindicus (Carabus) 34

hortensis preslii (Carabus) 33

hortensis preslii m. jonica (Carabus) 34

hortensis rhodopensis (Carabus) 34

hospes (Harpalus) 106

hospes hospes (Harpalus) 106

humerale (Bembidion) 53

humeralis (Cymindis) 121

humeralis (Lebia) 126

hybrida (Cicindela) 27

hybrida albanica (Cicindela) 27

hybrida riparia (Cicindela) 27

Hygrocarabus 31

hypocrita (Agonum) 134

hypocrita (Bembidion) 66

hypocrita (Notiophilus) 24

hypocrita illyricum (Bembidion) 66

Hypogeobium 160

Hypogium 160

hypsobius (Carabus) 36

ignavus (Zabrus) 183

illigeri (Bembidion) 62

immaculicornis (Brachinus) 42

immaculicornis immaculicornis (Brachinus) 42

immundus (Acupalpus) 116

importunus (Dyschirius) 50

importunus importunus (Dyschirius) 50

impressum sic! (Syntomus) 127

impressus (Metabletus) 127

impressus (Syntomus) 127

impressus impressus (Syntomus) 127

inaequalis (Tachys) 76

incertus (Brachinus) 43

Incisophonus 111

incisus (Ophonus) 111

incrassatus (Zabrus) 184

incrassatus bischoffi (Zabrus) 184

incrassatus boldorii (Zabrus) 184

incrassatus incrassatus (Zabrus) 184

inoptatum (Bembidion) 70

inquinatus (Pedius) 149

inquinatus (Pterostichus) 149

inquisitor (Calosoma) 29

inquisitor inquisitor (Calosoma) 29

inserticeps (Bembidion) 61

intermedius (Anisodactylus) 94

intermedius (Dyschirius) 46

INDEX 233

intricatus (Carabus) 30

intricatus chinobatus (Carabus) 30

intricatus intricatus (Carabus) 30

intricatus intricatus m. montenegrinus (Carabus) 30

intricatus intricatus n. weiratheri (Carabus) 30

intricatus intricatus var. weiratheri (Carabus) 30

intricatus montenegrinus (Carabus) 30

intricatus var. montenegrinus (Carabus) 30

irenis (Trechus) 86

iricolor (Bembidion) 70

irregularis (Carabus) 35

irregularis bucephalus (Carabus) 35

irregularis ramanus (Carabus) 35

janthinus (Laemostenus) 172

janthinus janthinus (Laemostenus) 172

javurkovae (Bembidion) 68

jockischi (Nebria) 22

jockischi jockischi (Nebria) 22

jonicus (Carabus) 34

joniensis (Carabus) 35

justinae (Bembidion) 55

kaufmanni (Tapinopterus) 161

klisuranus (Molops) 144, 146

kobingeri (Trechus) 87

kobingeri bulgaricus (Trechus) 88

kobingeri kobingeri (Trechus) 87, 88

kobingeri pawlowskianus (Trechus) 88

korabensis (Carabus) 33

korabica (Nebria) 21

korax (Calathus) 167, 168

koyi (Poecilus) 150

koyi (Pterostichus) 150

koyi dinaricus (Pterostichus) 150

koyi koyi (Poecilus) 150

kratteri (Nebria) 23

kratteri kratteri (Nebria) 23

kratteri valonensis (Nebria) 23

krueperi (Harpalus) 109

krueperi (Ophonus) 109

Laemostenus 171

Laemostenus s.str. 172

laeviceps (Parophonus) 114, 115

laevicollis (Trichotichnus) 115

laevicollis laevicollis (Trichotichnus) 115

laevifrons (Clivina) 45

laevigatus (Scarites) 50

laevipes (Harpalus) 101

laeviusculus (Dyschirius) 47

lafertei (Dyschirius) 49

Lamprias 125

lampros (Bembidion) 60

lampros var. properans (Bembidion) 61

laterale (Bembidion) 61

laterale pseudolaterale (Bembidion) 61

laticollis (Chlaenius) 91

laticollis (Ophonus) 111

latifianus (Pterostichus) 157

latifianus ab. nigripes (Pterostichus) 158, 159

latifianus latifianus (Pterostichus) 156, 157

latifianus strupii (Pterostichus) 156, 157,

158, 159

latipennis (Dyschirius) 49

latiplaga (Bembidion) 59

latiplaga latiplaga (Bembidion) 59

latus (Abax) 138

latus (Calathus) 167

latus (Harpalus) 103

latus (Licinus) 129

Lebia 125

Lebia s.str. 126

LEBIINA 125

LEBIINI 121

Leistus 16

Leistus s.str. 16

leonhardi stenocephalus (Duvalius) 81

leonhardi sturanyi (Duvalius) 81, 82

lepidus (Poecilus) 150

lepidus (Pterostichus) 150

lepidus lepidus (Poecilus) 150

levantinus (Amblystomus) 93

levantinus levantinus (Amblystomus) 93

LICININA 128

LICININI 128

Licinus 129

Licinus s.str. 129

limbatum (Omophron) 38

limbopunctatus (Harpalus) 107

Limnocarabus 31

linearis (Dromius) 125

linearis (Paradromius) 125

linearis linearis (Paradromius) 125

lineata (Cymindis) 122


lineola (Cymindis) 122

LIONYCHINA 127

Lionychus 127

Lionychus s.str. 127

literata sic! (Cicindela) 28

litorale (Bembidion) 57, 58

litoralis sic! (Pogonus) 78

litterata (Cicindela) 28

littoralis (Calomera) 26

littoralis (Pogonus) 78

littoralis nemoralis (Calomera) 26

ljubetensis (Carabus) 32

lodosi (Pterostichus) 153

lonae (Neotrechus) 82

lonae (Trechus) 82

lonae atavus (Neotrechus) 83

lonae lonae (Neotrechus) 82, 83

lonae shkodrensis (Neotrechus) 83

lonai (Omphreus) 131

lonai sic! (Neotrechus) 82

lonai sic! atavus (Neotrechus) 83

lonai sic! lonai sic! (Neotrechus) 82

lonai sic! shkodrensis (Neotrechus) 83

longicollis (Pedius) 149

longicorne (Agonum) 134

longicornis (Anthracus) 117

longicornis (Thalassophilus) 88

Loxoncus 119

lucida (Amara) 176

lucidus (Dyschirius) 46

luctuosus (Microlestes) 124

luctuosus luctuosus (Microlestes) 124

lugubris rufipes (Aptinus) 39, 40

lumensis (Pterostichus) 157, 158

lumensis litae (Pterostichus) 156, 158

lumensis ljubetensis (Pterostichus) 156, 158

lumensis lumensis (Pterostichus) 156, 157

lunatus (Callistus) 89

lunatus lunatus (Callistus) 89

lunulata (Cicindela) 26

lunulata var. nemoralis (Cicindela) 26

lunulatum (Bembidion) 70

luridipennis (Pogonus) 78

luteatus (Acupalpus) 115

luticola (Dyschirius) 47

luticola luticola (Dyschirius) 47

Lymnastis 73

macedonica (Nebria) 18, 21

macedonica galicica (Nebria) 18, 21

macedonica papingutica (Nebria) 18, 21

macedonica rambouseki (Nebria) 18, 22

macedonicus (Calathus) 168

macedonicus (Pachycarus) 97

macroderus (Dyschirius) 48

macroderus macroderus (Dyschirius) 48

maculatum (Bembidion) 72

maculatum maculatum (Bembidion) 72

maculatus (Acupalpus) 116, 117

maculicornis (Harpalus) 114

maculicornis (Ophonus) 114

maculicornis (Parophonus) 114

maculicornis (Trichotichnus) 114

maderi (Molops) 139

maglicensis (Duvalius) 79

magnicollis (Leistus) 17

malissorum (Carabus) 31, 32

malissorum (Deltomerus) 76, 77

malissorum (Neotrechus) 83, 84

malissorum (Molops) 145

malissorum (Pterostichus) 156, 158, 159

malissorum (Trechus) 84

malissorum ab. munella (Pterostichus) 157

malshentianus (Molops) 139

mannerheimii (Bembidion) 70

Manodromius 125

marginata (Egadroma) 119

marginatum (Agonum) 134

marginatus (Stenolophus) 119

maritimus (Lionychus) 127

MASOREINA 92

Masoreus 92

Masoreus s.str. 92

maurus (Microlestes) 124

maurus maurus (Microlestes) 124

megacephalus (Acinopus) 98

Megodontus 29, 31

meisteri (Pterostichus) 156

melanarius (Pterostichus) 153

melanarius melanarius (Pterostichus) 153

melancholica (Myriochila) 28

melancholica melancholica (Myriochila) 28

Melanius 152

INDEX 235

melanocephalus (Calathus) 170

melanocephalus melanocephalus (Calathus) 170

melanura (Odacantha) 131

melas (Pterostichus) 152

melas depressus (Pterostichus) 152

melletii (Ophonus) 111

Menas 122

mendax (Harpalus) 114

mendax (Ophonus) 114

mendax (Parophonus) 114

mendax (Tachyophonus) 114

merditana (Nebria) 20, 21

merditanum (Bembidion) 57

merditanus (Aptinus) 39

merditanus (Carabus) 33

merditanus (Molops) 141, 142

merditanus (Trechus) 86

merditanus intermedius (Molops) 142

merditanus merditanus (Aptinus) 39, 40

merditanus merditanus (Molops) 141

merditanus var. granulipennis (Aptinus) 40

meridianus (Acupalpus) 116

meridionale (Bembidion) 59

meridionalis (Ophonus) 110

merklii (Pterostichus) 156

meschniggi (Bembidion) 56

messae (Amara) 180

metalkanus (Carabus) 31

metallescens (Amblystomus) 93

metallicus (Calathus) 169, 170, 171

metallicus (Pterostichus) 151

metallicus metallicus (Calathus) 170

metallicus prokletijensis (Calathus) 170

Metallina 60

Metophonus 111

Microcallisthenes 29

Microlestes 12, 123

micros (Paratachys) 74

micros (Tachys) 74

Microserrullula 61

miliaris (Cymindis) 123

miliusi (Carabus) 36

milleri (Bembidion) 67, 68

milleri milleri (Bembidion) 68

millerianus (Sinechostictus) 73

minor (Pterostichus) 155

minor minor (Pterostichus) 155

minutulus (Blechrus) 124

minutulus (Microlestes) 124

minutulus (Speluncarius) 160

minutulus (Tapinopterus) 160

minutulus apfelbecki (Microlestes) 124

minutus (Dyschirius) 48

minutus (Platyderus) 166

minutus albanicus (Dyschirius) 48

minutus obtusangulus (Platyderus) 166

miridita (Tapinopterus) 164

miridita jablanicensis (Tapinopterus) 164, 165

miridita jacupicensis (Tapinopterus) 165

miridita miridita (Tapinopterus) 164

miriditus (Carabus) 30

mixtus (Stenolophus) 120

moestum (Agonum) 133

moestus (Platynus) 133

mollis (Calathus) 171

mollis f. erythroderus (Calathus) 169

mollis mollis (Calathus) 171

Molops 139

Molops s.str. 139

monachum var. binaghii (Agonum) 133

monastirensis (Tapinopterus) 165

monastirensis gestroi (Tapinopterus) 165

montana (Amara) 178

montenegrinus (Carabus) 30

montenegrinus (Molops) 147, 148

montenegrinus cavernicola (Molops) 147

monticola (Cicindela) 27

monticola albanica (Cicindela) 27

montivagus (Carabus) 30

montivagus m. sutomorensis (Carabus) 30

montivagus vellepiticus (Carabus) 30

morio (Amara) 176

morio (Dyschirius) 48

morio (Omphreus) 132

morio albanicus (Omphreus) 132

morio malissorum (Omphreus) 132

morio nivium (Amara) 176

Morphnosoma 153

muelleri (Agonum) 134, 135

muelleri (Duvalius) 80

muelleri (Platynus) 134

munella (Pterostichus) 157


municipalis (Amara) 180

municipalis bischoffi (Amara) 180

Myas 148

Myas s.str. 148

Myriochila 28

Myriochila s.str. 28

Mystropterus 97

nana (Tachyta) 75

Nebria 12, 18

Nebria s.str. 22

NEBRIINAE 7, 16, 187

NEBRIINI 16

nebulosum (Asaphidion) 52, 53

nebulosum balcanicum (Asaphidion) 52, 58

negrita (Microlestes) 124

negrita apfelbecki (Microlestes) 124

nemoralis nemoralis (Lophyridia) 26

nemorivagus (Anisodactylus) 94

Neocalathus 169

Neoelaphrus 38

Neorescius 130

Neotrechus 82

neowinkleri (Carabus) 35

Nepha 61

neresheimeri (Bembidion) 70

neumeyeri (Carabus) 33, 34

niger (Amblystomus) 93

niger (Pterostichus) 154

niger niger (Pterostichus) 154

nigricornis (Brachinus) 43

nigricornis (Chlaenius) 67, 89, 90

nigrinus (Trechus) 87

nigripes (Pterostichus) 158

nigrita (Pterostichus) 155

nitidulus (Chlaenius) 90

nitidus (Dyschirius) 46

nitidus (Leistus) 17

nitidus nitidus (Dyschirius) 46

nivalis (Synuchus) 173

nodulosus (Carabus) 31

normannum (Bembidion) 59

normannum apfelbecki (Bembidion) 59

normannum mediterraneum (Bembidion) 59

normannum meridionale (Bembidion) 59

Notaphemphanes 62

Notaphus 63

notatus (Acupalpus) 116

notatus (Dromius) 125

notatus (Philorhizus) 125

NOTIOPHILINI 23

Notiophilus 12, 23

oberthuri (Ophonus) 109

oblitus (Harpalus) 108

oblitus oblitus (Harpalus) 108

oblongopunctatus (Pterostichus) 151

oblongopunctatus oblongopunctatus

(Pterostichus) 151

oblongum (Bembidion) 60

obscuroguttatus (Metabletus) 128

obscuroguttatus (Syntomus) 128

obscurus (Aristus) 96

obscurus (Ditomus) 96

obscurus (Dixus) 96

obsoletus (Dicheirotrichus) 118

obtusangulus intermedius (Molops) 142

obtusiusculus (Trechus) 88

obtusus (Trechus) 85

obtusus obtusus (Trechus) 85

octomaculatum (Bembidion) 72

Ocydromus 63

Ocys 72

Ocyturanes 64

Odacantha 130

Odacanta s.str. 131

ODACANTHINI 130

Odontium 66

Odontocarus 97

Oedematicus 98

oertzeni (Licinus) 130

oertzeni albanicus (Licinus) 130

oertzeni oertzeni (Licinus) 130

olens (Zuphium) 186

olens olens (Zuphium) 186

Olisthopus 12, 137

Omoperyphus 66

Omophron 38

Omophron s.str. 38

OMOPHRONINAE 7, 38, 187

OMOPHRONINI 38

OMPHREINI 131

Omphreus 131

Omphreus s.str. 131

INDEX 237

Oodes 131

OODINI 131

Ophonomimus 113

Ophonus 12, 109

Ophonus s.str. 112

Oreocarabus 33

Oreoplatysma 153

Osimus 98

osmanilis (Molops) 145

osmanilis osmanilis (Molops) 145, 146

osmanilis tomoricensis (Molops) 145, 146

ottomanus (Pterostichus) 154

ottomanus ottomanus (Pterostichus) 154

ovalis (Abax) 139

ovata (Amara) 176

Pachycarus 97

Pachystus 34

pallipes (Metabletus) 128

pallipes (Syntomus) 128

paludicola (Acupalpus) 116, 117

palustris (Notiophilus) 24

PANAGAEINI 133

Panagaeus 133

Panagaeus s.str. 133

Paracelia 179

Paradeltomerus 77

paradoxus (Deltomerus) 77

paradoxus korabensis (Deltomerus) 77

paradoxus paradoxus (Deltomerus) 77

Paradromius 125

Paradyschirius 50

Parahaptoderus 153

Parallelomorphus 50

parallelus (Dyschirius) 50

parallelus (Ophonus) 111

parallelus albanensis (Harpalus) 111

parallelus albanicus (Ophonus) 111

parallelus ruficornis (Dyschiriodes) 50

parallelus ruficornis (Dyschirius) 50

Paranchus 137

Parapercosia 179

Paratachys 74

parnassicola (Molops) 140

parnassicola alinae (Molops) 140

parnassicola valonensis (Molops) 140

parnassicus (Carabus) 35

Parophonus 12, 113

Parophonus s.str. 114

parreyssi (Molops) 141

parreyssi lonae (Molops) 141

parumpunctatus (Deltomerus) 77

parvicollis (Leistus) 17

parvula (Tachyura) 76

parvulus (Acupalpus) 117

parvulus (Tachys) 76

PATROBINI 76

Pedius 149

Pelor 181

peltatus (Badister) 129

peltatus peltatus (Badister) 129

pentheri (Calosoma) 29

pentheri (Pterostichus) 156, 159

pentheroides (Calosoma) 29

Percosia 179

peregrinus (Brachinus) 42

PERILEPTINA 79

Perileptus 79

Perileptus s.str. 79

peristericus (Tapinopterus) 162

permoestum (Agonum) 135

Peryphanes 66

Peryphus 68

Phaon 155

Pheggomisetes 88

Philochthus 69

Philorhizus 125

Philorhizus s.str. 125

Phonias 153

Phyla 70

picenus peristericus (Carabus) 33

piceus (Molops) 147

piceus balcanicus (Molops) 147

picipennis (Harpalus) 102

picipes (Acinopus) 97

pindicola (Cymindis) 122

pindicum (Bembidion) 64, 65

pindicus (Carabus) 34

plagiatus (Brachinus) 40, 41

plagiatus (Microlestes) 124

plagiatus escorialensis (Microlestes) 123

planicollis (Acupalpus) 117

planicollis Apfelbeck (Harpalus) 114


planicollis Apfelbeck (Ophonus) 114

planicollis Apfelbeck (Parophonus) 114, 115

planicollis Dejean (Harpalus) 114

planicollis Dejean (Ophonus) 114

planicollis Dejean (Parophonus) 114

Platycarabus 35

Platyderus 165

Platyderus s.str. 165

PLATYNINI 133

Platynus 138

Platynus s.str. 138

Platypterinus 154

Platysma 154

plurisetosus (Molops) 145

Poecilus 149

Poecilus s.str. 149

POGONINI 78

Pogonistes 78

Pogonophorus 17

Pogonus 78

Pogonus s.str. 78

Polyderis 74

Polystichus 185

pominii (Duvalius) 81

praetermissa (Amara) 178

praeustum (Bembidion) 65, 74

praeustum praeustum (Bembidion) 65

praeustum viridifluum (Bembidion) 65

preslii (Carabus) 33, 34

preslii neumeyeri (Carabus) 34

priapus (Trechus) 86

priapus priapus (Trechus) 86

Princidium 71

Pristonychus 173

Procerus 29, 35

procerus (Anoplogenius) 119

procerus (Carabus) 32

procerus (Loxoncus) 119

Procrustes 29, 35

properans (Amara) 180

properans (Bembidion) 61

protensus (Tapinopterus) 162

protensus aetolicus (Tapinopterus) 162, 163

protensus peristericus (Tapinopterus) 162

protensus tomoricensis (Tapinopterus) 163

proxima (Amara) 176

proximus (Stenolophus) 120

Pseudodichirius 94

pseudolaterale (Bembidion) 61

Pseudolimnaeum 73

Pseudomaseus 154

Pseudophonus 108

psophia (Brachinus) 41

PTEROSTICHINI 138

Pterostichus 151

Pterostichus s.str. 155

pubescens (Harpalus) 108

pubescens (Ophonus) 108

pueli (Anisodactylus) 94

pueli pueli (Anisodactylus) 94

pumicatus (Stomis) 160

pumicatus pumicatus (Stomis) 160

pumilus (Harpalus) 102

punctatostriatus (Harpalus) 106

punctatulus (Harpalus) 111

punctatulus (Ophonus) 111

punctatus (Dyschirius) 48

punctatus (Laemostenus) 173

punctatus (Leistus) 18

punctatus albanicus (Dyschirius) 48

puncticeps (Harpalus) 112

puncticeps (Ophonus) 112

puncticollis Coquerel (Acupalpus) 117

puncticollis Auct. nec Coquerel (Acupalpus) 116

puncticollis (Harpalus) 112

puncticollis (Ophonus) 112

puncticollis (Poecilus) 149

puncticollis (Pterostichus) 149

punctulatum (Bembidion) 71

punctulatum punctulatum (Bembidion) 71

punctulatus sic! (Ophonus) 111

pusillus (Dyschirius) 48

pusillus (Notiophilus) 23

pygmaeus (Harpalus) 106

quadricolle (Bembidion) 61

quadriguttatum (Fabricius) (Bembidion) 53

quadriguttatum (Olivier) (Bembidion) 54

quadrillum (Lionychus) 127

quadrimaculatum (Bembidion) 53, 54

quadrimaculatum quadrimaculatum

(Bembidion) 53

quadrimaculatus (Dromius) 123

INDEX 239

quadripunctatus (Harpalus) 101

quadripustulatum (Bembidion) 54

quadripustulatum quadripustulatum

(Bembidion) 54

quadristriatus (Trechus) 85

quarnerensis (Anthracus) 117

quenseli (Amara) 179

quenseli quenseli (Amara) 179

ravasinianus (Trechus) 88

ravasinii (Calathus) 168

ravasinii (Omphreus) 132

ravasinii (Trechus) 88

ravasinii macedonicus (Calathus) 168

ravasinii ravasinii (Calathus) 168

rebeli (Poecilus) 150

rebeli (Pterostichus) 150

reflexicollis (Amara) 181

reflexicollis reflexicollis (Amara) 181

regiszogui (Duvalius) 81

Reicheadella 45

Reicheadella s.str. 45

REICHEINA 45

reiseri (Bembidion) 66

reiseri (Molops) 143, 144

reiseri (Pterostichus) 156, 159

reiseri corallipes (Molops) 142

reiseri holdhausi (Molops) 142, 143, 144, 145

reiseri pseudoalpestris (Molops) 142

reiseri reiseri (Bembidion) 65

reiseri reiseri (Molops) 142, 143

reiseri victoris (Molops) 142, 143, 144

reiseri vranense (Bembidion) 65

relictum (Bembidion) 55

relictum (Calosoma) 29

relictum var. pentheroides (Calosoma) 29

reticulatus (Apristus) 127

rhilensis (Pterostichus) 155

rhilensis kourili (Pterostichus) 156

rhilensis rhilensis (Pterostichus) 156

rhilensis vitosensis (Pterostichus) 156

rhodopense (Bembidion) 54

rhodopensis (Carabus) 34

rilvensis (Carabus) 32

riparia (Cicindela) 27

riparius (Pogonus) 78

rivulare (Bembidion) 58

robustus (Odontocarus) 97

rossii (Asaphidion) 53

rostratus (Stomis) 161

rostratus rostratus (Stomis) 161

rotundatus (Olisthopus) 137

rotundatus rotundatus (Olisthopus) 137

rotundicollis (Carterus) 95

rubripes (Harpalus) 99

rufescens (Nebria) 22

rufescens rufescens (Nebria) 22

rufibarbis (Harpalus) 112

rufibarbis (Ophonus) 112

ruficornis (Dyschirius) 50

rufipalpis (Harpalus) 100

rufipalpis rufipalpis (Harpalus) 100

rufipes (Harpalus) 108

rufipes (Molops) 146

rufipes (Notiophilus) 25

rufipes (Pseudophonus) 108

rufipes klisuranus (Molops) 146

rufipes munelensis (Molops) 146

rufipes pseudoosmanilis (Molops) 146

rufipes rufipes (Molops) 146

rufipes steindachneri (Molops) 147

rufipes sturanyi (Molops) 147

rufitarsis (Harpalus) 100

rufithorax (Apotomus) 51

rufomarginatus (Leistus) 18

rufus (Apotomus) 51

rufus (Platyderus) 166

rufus rufus (Platyderus) 166

rugosicollis (Myas) 148

rugosus (Carabus) 35

rumelicum (Bembidion) 63

rupicola (Harpalus) 112

rupicola (Ophonus) 112

sabulicola (Harpalus) 113

sabulicola (Ophonus) 113

sabulicola columbinus (Harpalus) 113

sabulicola columbinus (Ophonus) 113

sahlbergi albanica (Cicindela) 27

salina (Nebria) 23

salinus (Dyschirius) 49

salinus striatopunctatus (Dyschirius) 49

saphyrea (Amara) 177

sarajevoensis (Carabus) 31


saxicola (Harpalus) 108

scabriusculus (Carabus) 37

scabriusculus bulgarus (Carabus) 37

Scallophorites 51

scapulare (Bembidion) 60

scapulare oblongum (Bembidion) 60

scapularis (Cymindis) 122

scapularis (Lebia) 126

scapularis scapularis (Cymindis) 122

scapularis scapularis (Lebia) 126

Scarites 50

SCARITINA 50

SCARITINAE 7, 12, 44, 187

SCARITINI 50

schaubergerianus (Ophonus) 112

schaumi (Panagaeus) 133

scrobiculatus (Platynus) 138

scrobiculatus serbicus (Platynus) 138

scrobiculatus turcicus (Platynus) 138

scutellaris (Tachys) 75

semigranosus (Cychrus) 37

semigranosus albanicus (Cychrus) 37, 38

semigranosus balcanicus (Cychrus) 37

semigranosus dinaricus (Cychrus) 37, 38

semigranosus jakupicensis (Cychrus) 38

semigranosus montanus (Cychrus) 37

semigranosus montenegrinus (Cychrus) 37, 38

semigranosus n. balcanicus (Cychrus) 37

semigranosus n. montenegrinus (Cychrus) 37

semigranosus semigranosus (Cychrus) 37

semigranosus travnikanus (Cychrus) 38

serbicus (Platynus) 138

serripes (Harpalus) 101

serripes ab. subchalybaeus (Harpalus) 101

serripes serripes (Harpalus) 101

sexpunctatum (Agonum) 135

sexstriata (Tachyura) 76

sexstriatus (Tachys) 76

sexstriatus var. bisbimaculatus (Tachys) 76

shardagensis (Carabus) 33

shljebensis (Carabus) 32

Siagona 44

SIAGONINAE 7, 44, 187

SIAGONINI 44

sichemita (Brachinus) 42

siculum (Bembidion) 64

siculum smyrnense (Bembidion) 64

sigma (Dromius) 125

sigma (Philorhizus) 125

signatipenne (Bembidion) 65

silphoides (Licinus) 130

similata (Amara) 177

similis (Ophonus) 110

simplex (Molops) 140, 141

simplex meridionalis (Molops) 140, 141

simplex simplex (Molops) 140

simulator (Dyschirius) 48

Sinechostictus 10, 73

Sinechostictus s.str. 73

skrimshiranus (Stenolophus) 120

skutariensis (Harpalus) 104

smaragdinus (Harpalus) 104

solitaris (Harpalus) 103

sordidum (Agonum) 135

sordidum sordidum (Agonum) 135

sordidus (Platynus) 135

spartanus (Molops) 140

spartanus parnassicola (Molops) 140

spartanus valonensis (Molops) 140

Speluncarius 12, 160

Speluncarius s.str. 160

Sphaerotachys 75

SPHODRINA 171

SPHODRINI 165

spinibarbis (Leistus) 17, 18

spinibarbis punctatus (Leistus) 18

spinibarbis rufipes (Leistus) 18

spinipes (Zabrus) 185

spinipes insignis (Zabrus) 185

splendidum (Bembidion) 58

splendidum pincum (Bembidion) 58

spoliatus (Chlaenius) 90

spoliatus spoliatus (Chlaenius) 90

spreta (Amara) 177

spurcaticornis (Anisodactylus) 94

steindachneri (Molops) 147

stenocephalus (Duvalius) 82

stenocephalus (Trechus) 81

Stenochoromus 147

STENOLOPHINA 12, 115

Stenolophus 119

Stenolophus s.str. 119

INDEX 241

sterbai (Carabus) 34

stictus (Ophonus) 113

stierlini (Asaphidion) 53

stolfai (Bembidion) 65, 66

Stomis 160

Stomis s.str. 160

strenuus (Argutor) 154

strenuus (Pterostichus) 154

strepens (Brachinus) 42

striatopunctatus (Poecilus) 150

striatul sic! (Bembidion) 66

striatum (Bembidion) 66

strumosus (Dyschirius) 46

strupii (Duvalius) 82

sturanyi (Duvalius) 81, 82

sturanyi (Molops) 147

sturanyi (Nebria) 19

sturanyi matejkai (Duvalius) 82

sturanyi munelensis (Molops) 146

sturanyi stenocephalus (Duvalius) 81, 82

sturanyi sturanyi (Duvalius) 82

subaeneus (Apristus) 127

subchalybaeus (Harpalus) 101

subcostatum (Bembidion) 68

subcostatum javurkovae (Bembidion) 68, 69

subcostatum spartanum (Bembidion) 69

subcostatum vau (Bembidion) 68, 69

subcylindricus (Harpalus) 102

subfasciatum (Bembidion) 71

submarginatus (Licinus) 129, 130

subnotatus (Trechus) 85

subnotatus ljubetensis (Trechus) 86

subnotatus subnotatus (Trechus) 85

subquadratus (Acinopus) 97, 98

subquadratus (Ophonus) 110

subrugosus (Carabus) 35

subsinuatus (Harpalus) 100

substriatum (Bembidion) 62

substriatus (Dyschiriodes) 50

substriatus (Dyschirius) 50

substriatus (Notiophilus) 25

substriatus priscus (Dyschirius) 50

sulphuripes (Harpalus) 100

sulphuripes sulphuripes (Harpalus) 100

sutomorensis (Carabus) 30

suturalis (Acupalpus) 117

suturalis (Neotrechus) 84

suturalis (Trechus) 84

suturalis pentheri (Neotrechus) 84

suturalis pretneri (Neotrechus) 83

suturalis suturalis (Acupalpus) 117

sycophanta (Calosoma) 29

sylvatica (Cicindela) 27

sylvatica sylvatica (Cicindela) 27

Syntomus 12, 127

Synuchidius 171

SYNUCHINA 173

Synuchus 173

Synuchus s.str. 173

TACHYINA 73

TACHYINI 10

Tachypus 36

Tachys 75

Tachyta 75

Tachyta s.str. 75

Tachyura 76

Tachyura s.str. 76

Taenidia 26

Talanes 71

Tapinopterus 12, 161

Tapinopterus s.str. 161

tardus (Harpalus) 103

tarsicus (Sinechostictus) 73

Tarulus 123

taurica (Amara) 179

tenebrioides (Zabrus) 185

tenebrioides tenebrioides (Zabrus) 185

tenebrosus (Harpalus) 98

tenebrosus var. centralis (Harpalus) 98

tenellum (Bembidion) 59

tenellum tenellum (Bembidion) 59

terminale (Bembidion) 71

terminatus (Chlaenius) 90

Terminophanes 71

terricola (Laemostenus) 173

terricola (Pristonychus) 173

terricola (Scarites) 51

terricola punctatus (Laemostenus) 173

terricola terricola (Scarites) 51

testaceum (Bembidion) 60

testaceum testaceum (Bembidion) 60

Testedium 71


tethys (Bembidion) 70

tetracolum (Bembidion) 69

tetracolum tetracolum (Bembidion) 69

teutonus (Acupalpus) 120

teutonus (Stenolophus) 120

Thalassophilus 88

thessalicus tomoricensis (Tapinopterus) 163

thoreyi (Agonum) 136

thoreyi (Platynus) 136

thoreyi thoreyi (Agonum) 136

tibiale (Bembidion) 54

tibialis (Amara) 177

Tomocarabus 36

tomoricensis (Amara) 179, 180

tomoricensis (Molops) 145

tomoricensis (Nebria) 21

Trachycarabus 37

transfuga (Molops) 143

transversalis (Cicindela) 27

transversalis transversalis (Cicindela) 27

TRECHINA 79

TRECHINAE 7, 52, 187

TRECHINI 79

TRECHINITAE 10

TRECHODINA 88

Trechus 85

Trechus s.str. 85

Trepanes 72

Trichochlaenius 92

Trichotichnus 12, 115

Trichotichnus s.str. 115

tricolor (Bembidion) 56

tricuspidata (Amara) 181

tricuspidatus (Carterus) 95

tricuspidatus (Ditomus) 95

trimaculata (Lebia) 126

triseriatus (Harpalus) 105

trisignata (Cylindera) 28

trisignata trisignata (Cylindera) 28

tristis (Chlaenius) 90

tristis (Harpalus) 104

tristis (Trechus) 87

tristis tristis (Chlaenius) 90

truncatellus (Metabletus) 128

truncatellus (Syntomus) 128

turcicus (Platynus) 138

uliginosus (Elaphrus) 38

unipustulatus (Badister) 128

ustulatum (Bembidion) 69

valbonensis (Trechus) 85

valonensis (Nebria) 23

vaporariorum (Cymindis) 123

varicolor (Bembidion) 56

varicolor varicolor (Bembidion) 56

variolosus (Carabus) 31

variolosus nodulosus (Carabus) 31

varium (Bembidion) 63

varium dyrrhachii (Bembidion) 63

velutinus (Chlaenius) 91

venustus (Laemostenus) 172

verbasci (Bradycellus) 118

vernalis (Harpalus) 102

vernalis cursor (Pterostichus) 151

versicolor (Poecilus) 151

vestitus (Chlaenius) 90

victoris (Molops) 143, 144

viduus moestus (Platynus) 133

viduus var. moestus (Platynus) 133

vignai (Duvalius) 82

violaceus (Carabus) 32

violaceus azurescens (Carabus) 32

violaceus azurescens m. merditana (Carabus) 33

violaceus azurescens m. merditanus (Carabus) 33

violaceus azurescens m. zabljakensis (Carabus) 32

violaceus azurescens n. zabljakensis (Carabus) 32

violaceus azurescens var. shljebensis (Carabus) 32

violaceus dryas (Carabus) 33

violaceus dryas var. korabensis (Carabus) 33

violaceus dryas var. merditanus (Carabus) 33

violaceus exasperatus (Carabus) 33

violaceus merditanus (Carabus) 33

violaceus shardagensis (Carabus) 33

violaceus var. korabensis (Carabus) 33

violaceus var. shardagensis (Carabus) 33

violaceus vlasuljensis (Carabus) 33

violaceus zabljakensis v. merditanus (Carabus) 33

viridicupreum (Agonum) 135

viridicupreum viridicupreum (Agonum) 135

viridicupreus (Platynus) 135

viridulus (Harpalus) 107

vivalis (Synuchus) 173

vivalis vivalis (Synuchus) 173

INDEX 243

vlasuljensis (Carabus) 33

vseteckai (Bembidion) 62

vseteckai dissimile (Bembidion) 62

walteri (Pterostichus) 156

weiratheri (Carabus) 30

weiratheri (Molops) 140

weiratherianus (Carabus) 35

wetterhallii (Masoreus) 92

wetterhallii wetterhallii (Masoreus) 92

winkleri (Carabus) 35

winkleri (Zabrus) 182

winklerianus (Molops) 144

Xenocelia 180

zabljakensis (Carabus) 32

ZABRINA 181

ZABRINI 174

Zabrus 181

Zabrus s.str. 185

Zezea 181

zhalovi (Duvalius) 82

ZUPHIINA 185

ZUPHIINI 185

Zuphium 186

Zuphium s.str. 186

B.v. gueorguiev annotated catalogue of the carabid beetles of albania coleoptera carabidae v. 64...

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Transcript of B.v. gueorguiev annotated catalogue of the carabid beetles of albania coleoptera carabidae v. 64...