Aquatic Ecosystems and Invertebrates of the

Grand Staircase-Escalante National MonumentCooperative Agreement Number JSA990024

Annual Report of Activities for 2000

Mark Vinson

National Aquatic Monitoring Center

Department of Fisheries and Wildlife

Utah State University

Logan, Utah 84322-5210

www.usu.edu/buglab

1 April 2001

i

Table of contentsPage

Foreword . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . iIntroduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1Study area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1Long-term repeat sampling sites . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2Methods

Locations and physical habitat . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3Aquatic invertebrates

Qualitative samples . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3Quantitative samples . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

Laboratory methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4Results

Sampling locations . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5Habitat types . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6Water temperatures . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8Aquatic invertebrates . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8

Literature cited . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13Appendices

1. Aquatic invertebrates collected in the major habitatsA. Alcove pools . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14B. Hanging gardens . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15C. Posy Lake . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16D. Ponds and reservoirs . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17E. Springs and seeps . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18F. Rivers and streams . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19G. Tinajas . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21

2. Aquatic invertebrates sampling locations . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22

cover photo: Brooke Bushman sampling within drainage tinajas near the Highway 12 overlook.

i

ForewordThe work described in this report was conducted by the National Aquatic Monitoring Center, Utah State

University, Logan, Utah. The following people participated in the field collection of the samples, the

identification of the invertebrates, and the writing of this report. I thank them for all their help and good

spirit.

Dave Axford

Dan Barnes

Brooke Bushman

Leslie Ogden

Matt Tagg

Erin Thompson

Dan Zamecnik

1

MonthOCT DEC FEB APR JUN AUG

Pre

cipi

tatio

n (in

ches

)

0.000.250.500.751.001.251.501.752.00

MonthOCT DEC FEB APR JUN AUG

Pre

cipi

tatio

n (in

ches

)

0.000.250.500.751.001.251.501.752.00

Figure 1. Long-term means + SE in monthly precipitation at Escalante, Utah.

IntroductionThe Grand Staircase-Escalante National Monument was established on 18 September 1996 and

comprises more than 760,000 hectares of geological and ecological diversity in southern Utah. In 1998

we began to survey and monitor the Monument’s aquatic habitats with the goals of collecting basic

faunistic and ecological data and long-term biomonitoring data. Sampling should continue through at

least 2004. Our sampling strategy has been twofold:

1) qualitatively sample as many locations as possible; and

2) repeatedly sample across seasons and years a subset of perennial habitats collecting both

quantitative and qualitative samples.

The purpose of this report is to describe where, when, and how we have sampled through

December 2000. Similar reports were prepared for samples collected in 1999 and 2000. Thee reports

can be obtained from our web site: http://www.usu.edu/buglab/gsenm.htm or from the Grand

Staircase-Escalante National Monument, 180 West 300 North, Kanab, Utah 84741, 435-644-4300.

Study areaElevations within and adjacent to the National Monument range from 1100 to 3000 m. At

Escalante, Utah, elevation 1616 m, annual air temperatures vary widely from <10 to 100 F (–30 to 37 C).

The mean annual precipitation is about 12 inches (30 mm). Winters are cold and windy and summers

are characterized by hot days and cool nights. Moisture falls predominantly as snow between January

and May (Figure 1). Early summer is usually dry, whereas intense localized thunderstorms are common

in late summer.

2

MonthJan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec JanW

ater T

empe

ratur

e (C)

05

101520253035

Disc

harg

e (CF

S)

01002003004005002000

300040005000

MonthJan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec JanW

ater T

empe

ratur

e (C)

05

101520253035

Disc

harg

e (CF

S)

01002003004005002000

300040005000

Figure 2. Discharge and water temperature for the Escalante River near its mouth in 1951.

Streamflow patterns are atypical for the Rocky Mountains with peaks occurring in response to

both spring snowmelt and late summer monsoonal rain events. Monsoonal stormflows can be 50 times

greater than mean annual flows and 2-10 times greater than spring peak flows (Figure 2). Water

temperatures range from near zero to >33 C (Figure 2). Stream substrates vary widely and are often

either bedrock or sand.

Long-term repeat sampling sitesThe following sites were identified as locations where long-term seasonal collections will be

made. Samples collected on rivers near tributary streams would be taken upstream of the tributary

stream. This list will likely be amended as we become more familiar with the Monument.

1. Escalante River near Escalante

2. Escalante River near Mamie Creek

3. Escalante River near Sand Creek

4. Escalante River near Calf Creek

5. Escalante River near Boulder Creek

6. Escalante River near the Gulch

7. Escalante River near Harris Wash

8. Escalante River near Coyote Gulch

9. Mamie Creek at mouth

10. Sand Creek at mouth

11. Calf Creek at mouth

12. Boulder Creek at Highway 12 bridge

3

13. Boulder Creek at mouth

14. Deer Creek at Burr Trail bridge

15. The Gulch at Burr Trail bridge

16. The Gulch at mouth

17. Pine Creek at Lower Box Canyon trailhead

18. Paria River at Highway 12 bridge

19. Harris Wash near trailhead

20. Harris Wash at mouth

21. Coyote Gulch near Hamblin Arch

22. Coyote Gulch at mouth

MethodsLocations and physical habitat

The latitude and longitude of each site was determined using hand held global positioning units.

Elevation was determined using an altimeter or a 7.5 min topographic map. Water and air temperatures

were determined at the time of sampling using thermometers. The conductivity of the water was

determined using an Oakton TDS Testr 40 hand held meter and the pH was determined using an Oakton

pH Testr 2. Substrate particle size distributions were estimated by pebble counts (Wolman 1954). Over-

stream shade was estimated using a spherical densiometer. Substrate particle size distributions were

estimated by pebble counts (Wolman 1954). Over-stream shade was estimated using a spherical

densiometer. In 2000 we placed continuous recording thermographs (Hobo temps, Onset Inc.) in the

following locations:

Escalante River near the town of Escalante Escalante River at the Highway 12 bridge

Calf Creek at the mouth near Highway 12 Boulder Creek at the Highway 12 bridge

Deer Creek at the Burr Trail bridge The Gulch at the Burr Trail bridge

Pine Creek near the Upper and Lower Box Canyon trail heads

Tinajas off of Highway 12 near the panoramic overlook

Aquatic invertebrates

Qualitative samples—The objective of qualitative invertebrate collections was to collect as many

different kinds of invertebrates living at a site as possible. Qualitative collections of invertebrates were

done at all sites. Samples were collected with a kicknet (457 x 229 mm) with a 500 micron mesh net and

by hand picking invertebrates from woody debris and large boulders. All major habitat types (e.g., riffles,

pools, back waters, macrophyte beds) were sampled and all samples were composited to form a single

sample from each site. In addition, tinajas were sampled with a zooplankton net (130 mm diameter, 80

micron mesh). Generally, 10 zooplankton tows were made across each tinaja at a variety of sampling

depths and composited into a single sample.

Quantitative samples—The objective of quantitative invertebrate sampling was to collect the more

4

common invertebrates at a site and estimate their relative abundances. Quantitative samples were

collected using a rectangular kicknet (457 x 229 mm) with a 500 micron mesh net. Samples were

collected by disturbing the area immediately upstream of the front of the net with our hands and

scrubbing individual substrate particles within the sampling area and allowing the invertebrates and

detritus to wash downstream into the net. The area of each sample was approximately 0.18 m2 (0.45 m

wide x 0.4 m deep). Four samples were collected in 4 different riffles and composited to make a sample

of approximately 0.7 m2 (4 kicknets x 0.18 m2 ) for each location.

Laboratory methods—The general procedures followed for processing invertebrate samples were

similar to those recommended by the United States Geological Survey (Cuffney et al. 1993) and are

described in greater detail and rationalized in Vinson and Hawkins (1996). Qualitative samples were

processed in the entirety, i.e., all the organisms were removed and identified. Quantitative samples were

sub-sampled if the sample appeared to contain more than 500 organisms. Sub-samples were obtained

by pouring the sample into an appropriate diameter 250 micron sieve, floating this material by placing

the sieve within an enamel pan partially filled with water and leveling the material within the sieve. The

sieve was then removed from the water pan and the material within the sieve was divided into equal

parts. One side of the sieve was then randomly chosen to be processed and the other side was set

aside. The sieve was then placed back in the enamel pan and the material in the sieve again leveled

and split in half. This process was repeated until approximately 500 organisms remained in one-half of

the sieve. This material was then placed into a petri dish and all organisms were removed under a

dissecting microscope at 10-30 power. Additional sub-samples were taken until at least 500 organisms

were removed. All organisms within a sub-sample were removed. During the sorting process the

organisms were separated into Orders. When the sorting of the sub-samples was completed, the entire

sample was spread throughout a large white enamel pan and searched for 10 minutes to remove any

taxa that might not have been picked up during the initial sample sorting process. The objective of this

"big/rare" search was to provide a more complete taxa list by finding rarer taxa that may have been

excluded during the sub-sampling process. These rarer bugs were placed into a separate vial and

tracked separately from the bugs removed during the sub-sampling process. All the organisms removed

during the sorting process were then identified. Once the data had been entered into a computer and

checked, the unsorted portion of the sample was discarded. The identified portion of the sample was

placed in 70% ethanol, given a catalog number, and was retained.

5

#

##

#

###

####

##

#

#

#

###

####

#

##

#

##

#

##

#

#

#######

#

###

##

#

#

#####

##

##

#

##

###

# ##

##

#

#

#

###

##

#

#

###

##

#

#

#

#

#

#

#

#

#

#####

##

##

#

##

#

##

#

#

###

#### #######

#

#

#

#

#

#

#

#

#

#

#

#

###

#

####

#

#

###

##

#

#

#

#

#

#

###

#####

##

###

#

###

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

###

## #

#

#

#

#

#

#####

##

#

#

##

#

##

##

#

###

#

#

###### ## ## #

#

#

#

#

#

#

#

##

#

#

#

#

#

#

#

#

#

## #

#

##

#

#

##

##

# #

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

#

######

##

##

##

##########

###

###

##

#

###

#

###

##

####

##

##

#####

5000000 0 5000000 10000000 Miles

N

EW

S

# ALCOVE POOL# POND# SPRING# STREAM# TINAJA

Figure 3. Aquatic invertebrate sampling locations.

ResultsAquatic invertebrate sampling locations

To date we have sampled 136 locations within and adjacent to the Grand Staircase - Escalante

National Monument (Figure 3). The majority of these sites have been perennial streams (74), but we

also sampled alcove pools (7), hanging gardens (1), Posy Lake, natural and man-made ponds, wetlands

(2), and reservoirs (2), limnocrene (3) and rheocrene springs (3), seeps (6), and tinajas (17). The

general location and specific geographic coordinates for each sampling location are presented in

Appendix 2.

6

Calf Creek Paria River

Wetland pond, naturalLivestock pond, man-made

Aquatic habitatsThis desert landscape contains many unique ephemeral and perennial aquatic habitats, including

streams, alcove pools, natural and man-made ponds, springs, tinajas (rock pools), and hanging gardens

(Table 1). Alcove pools were defined as perennial and ephemeral pools located beneath bedrock pour-

offs. Hanging gardens are assemblages of aquatic and semi-aquatic plants and animals occurring at

small seeps on canyon walls. Springs were separated into rheocrenes (flowing water springs),

limnocrenes (non-flowing water springs), and seeps (small water sources occurring on the canyon floor).

Tinajas are perennial and ephemeral water filled depressions in and outside of defined bedrock

drainages.

Perennial streams are

flowing water habitats

that have water year

around. Perennial

streams can vary

considerably with

respect to their flow

and water temperature

regimes, substrates,

and riparian vegetation

communities.

Ephemeral streams are

streams that flow only

part of the year.

Streamflow may be

caused by snowmelt or

rain storms. These

streams vary

considerably with

respect to how long

they maintain flow and

their size.

Natural ponds and

wetlands. These

habitats vary

considerably with

respect to their size

and the length of time

they maintain surface

water.

Man-made ponds vary

with the respect to their

size, how long they

maintain water, and the

degree of livestock

use.

7

Alcove pool

Hanging garden

Tinaja, within a clearlydefined drainage

Tinajas, outside of aclearly defineddrainage

Spring system

Spring, near AlveyWash

Alcove pools have

been defined as

perennial and

ephemeral pools

located beneath

bedrock pour-offs.

These have contained

high species richness

of invertebrates and

amphibians.

Hanging gardens are

located up above the

high water level within

slot canyons and along

rock walls near

waterfalls and alcove

pools.

Tinajas, within

drainages. These

habitats vary with

respect to their size,

the length of time they

maintain surface water,

their shading and the

quantity of emergent

wetland vegetation.

Tinajas, outside

defined drainages.

These habitats vary

with respect to their

size, the length of time

they maintain surface

water, and the quantity

of wetland vegetation

associated with them.

Rheocrenes - flowing

water springs and

seeps that may flow

into perennial or

ephemeral streams or

may disappear into the

ground a few meters

from their source.

Limnocrenes - non-

flowing water, lentic

spring ecosystems that

commonly resemble

small wetlands.

8

-5

0

5

10

15

20

25

0

5

10

15

20

25

30

Wat

er te

mpe

ratu

re (C

)

0

5

10

15

20

25

30

Month 2000

Jan Feb Mar Apr May Jun Jul Aug Sep Oct 0

5

10

15

20

25

0

5

10

15

20

25

Boulder Creek at HWY 12

Escalante River near USGS gage

Deer Creek near campground

Calf Creek near mouth

Pine Creek at Lower Box trailhead

Figure 4. Water temperatures recorded during year 2000.

Water temperatures

Continuous water temperatures were

recorded at several locations during year

2000 (Figure 4). Additional data was

collected in 2000 than is shown in Figure

4. This data is not presented because the

thermographs were switched in September

2000, so the data collected between

September and December2000 is still in

the field. This data will be retrieved when

the thermographs are switched in April

2001. Minimum water temperatures

occurred in the winter and were near 0 C

for all locations. Maximum water

temperatures occurred in the summer and

varied from about 22 C in lower Pine

Creek to more than 25 C in Calf Creek and

Deer Creek. All sites exhibited

considerable diel variation in water

temperature. In 2001 we will collecting

data at these same sties as well as several

new sites including the head of Calf Creek,

so that we can determine the degree of

warming that occurs along the entire length

of Calf Creek.

Aquatic invertebrates

An impressive number of aquatic invertebrates have been collected from the many unique habitats

present in the Monument. To date we have collected and processed 345 samples from 136 locations.

We have identified 43,436 individual invertebrates from 27 orders, 94 families, and 177 different genera

(Table 2). Diversity was highest among insects. We have found 35 Coleoptera, 31 Diptera (excluding

Chironomidae), 14 Ephemeroptera, 10 Hemiptera, 18 Odonata, 14 Plecoptera, and 31 genera of

Trichoptera (Table 2). Non-insects were less diverse, but we have also not yet identified many of these

taxa to genus, so the results are more preliminary. Nonetheless, we found 11 genera of Crustacea and 7

9

Number of Sampling Locations0 25 50 75 100 125 150

Cum

ulat

ive

Taxa

Ric

hnes

s (O

TUs)

0255075

100125150175200225250275300

All Habitat TypesStreams

Other habitats1:1 line

Number of Sampling Locations0 25 50 75 100 125 150

Cum

ulat

ive

Taxa

Ric

hnes

s (O

TUs)

0255075

100125150175200225250275300

All Habitat TypesStreams

Other habitats1:1 line

Figure 5. Taxa accumulation curves to show the rate whichnew taxa are being encountered.

genera of Mollusca. Species accumulation curves suggest that we are a long ways from finding all of the

different aquatic invertebrate taxa in the Monument (Figure 5). With each new location that we sample

we find new taxa, and there is little indication that this relationship is flattening out.

Table 2. Summary of aquatic macroinvertebrate collections made through December 2000 by major

habitat type. OTU = operational taxonomic unit and represents a mix of taxonomic levels, because not

all individuals can be identified to a consistent taxonomic level.

MeasureAll

locationsAlcovepools

Hanginggardens

PosyLake

Wetlands,Ponds,

ReservoirsSprings,Seeps

Rivers,Streams Tinajas

Number of samples 345 10 1 2 10 13 257 52

Total OTU taxa richness 294 59 13 19 54 63 252 85

Total invertebrate genera 177 35 8 11 31 37 154 47

Total insect genera 156 31 8 5 23 34 142 38

Coleoptera genera 35 13 4 0 8 10 29 12

Diptera (non-midge) genera 31 5 2 1 4 10 29 8

Ephemeroptera genera 14 2 1 0 2 2 14 2

Hemiptera genera 10 4 1 1 3 4 8 6

Megaloptera genera 1 0 0 0 0 0 1a 0

Odonata genera 18 6 0 2 3 2 16 5

Plecoptera genera 14 0 0 0 0 0 14 0

Trichoptera genera 31 1 0 1 3 5 29 5

Crustacea genera 11 1 0 5 2 1 4 6

Mollusca genera 7 2 0 1 5 1 6 1aAdult collected on Lake Powell, larvae collected near San Juan arm of Lake Powell.

10

Figure 6. The blindsubterranean amphipod, Stygobromus.

Some notable taxa that have been collected include a blind

subterranean amphipod (Figure 6) from a spring near Alvey Wash and

several species of Notonecta from throughout the Monument. The

amphipod belongs to the Stygobromus hubbsi group, and is likely a

new undescribed species. Several interesting fairy shrimp were

collected and are currently being identified by Dr. Tim Graham. A list

of all taxa collected and identified to date are shown in Table 3.

Plans for 2001Our plans for 2001 include sampling in the spring, summer, and fall at our long-term sampling sites.

We plan to locate and sample more tinajas and we want to do more exploration in the southwestern part

of the monument than we have in previous years. We will be deploying additional thermographs so that

we can begin to investigate how temperature may be influencing invertebrate assemblages and how

geomorphic features and past vegetation management activities influence water temperature.

11

Table 3. Aquatic organisms identified from all samples collected through January 2000. The numbers

refer to the number identified and in our collection. All identifications should be considered provisional

until verified.

Annelida Hirudinea 14 Oligochaeta 541 Haplotaxida Tubificidae 16 Lumbriculida Lumbriculidae 8Arthropoda Arachnida Trombidiformes 220 Crustacea 181 Amphipoda 2 Crangonyctidae Stygobromus hubbsi group 1 Gammaridae Gammarus lacustris 4 Hyalellidae Hyallela azteca 380 Anostraca Branchinectidae Branchinecta packardi 17 Calanoida 4 Diaptomidae Diaptomus 49 Temoridae Epischura nevadensis 2 Cladocera 139 Bosminidae Bosmina longirostris 9 Daphnidae 90 Daphnia 107 Daphnia pulex 71 Daphnia rosea 258 Daphniidae Ceriodaphnia 1 Ceriodaphnia laticaudata 8 Ceriodaphnia quadrangula 285 Copepoda 119 Cyclopoida 1297 Decapoda 5 Cambaridae Orconectes virillis 3 Isopoda 1 Notostraca Triopsidae Triops longicaudatus 3 Entognatha Collembola 8 Insecta Coleoptera 4 Amphizoidae Amphizoa 1 Curculionidae 1

Dryopidae Helichus 127 Dytiscidae 49 Agabus 166 Coptotomus 3 Deronectes 70 Eretes sticticus 1 Hydroporous 1 Hygrotus 6 Laccophilus 8 Neoclypeodytes 10 Oreodytes 31 Rhantus 92 Thermonectus mormoratus 11 Elmidae 569 Cleptelmis 1 Heterelmis 6 Heterlimnius 253 Microcylloepus 116 Microcylloepus pusillus 16 Narpus 23 Optioservus 701 Ordobrevia 8 Stenelmis 1 Zaitzevia 88 Gyrinidae Gyrinus 57 Haliplidae Haliplus 1 Peltodytes 18 Helophoridae Helophorus 10 Hydraenidae Hydraena 2 Ochthebius 5 Hydrophilidae 26 Ametor 1 Berosus 22 Hydrobius 3 Hydrochara 1 Lacobius 9 Paracymus 11 Tropisternus 9 Lampyridae 1 Scirtidae Elodes 15 Staphylinidae 1 Diptera 56 Athericidae Atherix 31 Ceratopogonidae 28 Atrichopogon 4 Bezzia 26

Culicoides 17 Dasyhelea 269 Probezzia 257 Chironomidae 680 Chironominae 3755 Orthocladiinae 6617 Tanypodinae 742 Culicidae 16 Culiseta 164 Dixidae Dixa 19 Dixella 9 Dolichopodidae 7 Empididae 21 Chelifera 63 Clinocera 2 Hemerodromia 16 Ephydridae 7 Muscidae 77 Limnophora 8 Psychodidae Maruina 3 Pericoma 41 Simuliidae 47 Prosimulium 24 Simulium 3529 Stratiomyidae 21 Caloparyphus 50 Euparyphus 5 Myxosargus 4 Nemotelus 1 Stratiomys 4 Tabanidae 8 Chrysops 1 Tabanus 17 Tipulidae 6 Antocha 69 Dicranota 85 Erioptera 1 Hesperoconopa 8 Hexatoma 66 Limonia 25 Tipula 189 Ephemeroptera 18 Ameletidae Ameletus 23 Baetidae 99 Baetis 3852 Baetis bicaudatus 30 Baetis tricaudatus 6161 Callibaetis 188 Paracloeodes 1 Caenidae

12

Caenis 1 Ephemerellidae 147 Drunella coloradensis 14 Drunella doddsi 63 Drunella grandis 255 Ephemerella 116 Ephemerella inermis 422 Heptageniidae 229 Cinygmula 280 Epeorus 349 Heptagenia 25 Nixe 10 Rhithrogena 162 Leptophlebiidae Paraleptophlebia 8 Tricorythidae Tricorythodes 414 Hemiptera 17 Belostomatidae Lethocerus 4 Corixidae 48 Corisella 4 Neocorixa 21 Gelastocoridae Gelastocoris oculatus 1 Gerridae 52 Aquarius 33 Gerris 14 Naucoridae 7 Ambrysus 60 Ambrysus mormona 19 Notonectidae 39 Notonecta 93 Notonecta kirbyi 37 Notonecta lobata 2 Notonecta undulata 3 Veliidae 6 Microvelia 1 Rhagovelia 6 Lepidoptera 1 Pyralidae Petrophila 29 Megaloptera Corydalidae Corydalus cognatus 1 Odonata 26 Aeshnidae 1 Aeshna 1 Aeshna umbrosa 67 Anax walsinghami 1 Calopterygidae Hetaerina 4 Hetaerina americana 3 Coenagrionidae 22 Argia 123 Argia plana 1 Argia vivida 36 Coenagrion 81 Enallagma 30

Enallagma cyathigerum 9 Ischnura 10 Telebasis salva 3 Cordulegastridae Cordulegaster diadema 4 Cordulegaster dorsalis 5 Gomphidae 7 Erpetogomphus 2 Erpetogomphus compositas 2 Gomphus 1 Ophiogomphus 1 Ophiogomphus severus 11 Lestidae Archilestes 5 Archilestes grandis 19 Lestes 1 Libellulidae 6 Libellula 18 Libellula saturata 17 Paltothemis lineatipes 1 Pantala flavescens 1 Tramea 3 Plecoptera 112 Capniidae 25 Utacapnia 69 Chloroperlidae 126 Suwallia 8 Sweltsa 124 Leuctridae 5 Nemouridae 214 Malenka 80 Prostoia 43 Zapada 33 Perlidae Hesperoperla pacifica 21 Perlodidae 48 Isogenoides 56 Isogenoides zionensis 8 Isoperla 96 Isoperla longiseta 113 Megarcys 3 Megarcys signata 2 Skwala 47 Pteronarcyidae 17 Pteronarcella badia 112 Taeniopterygidae 9 Taenionema 108 Trichoptera 78 Brachycentridae 45 Brachycentrus 311 Brachycentrus americanus 58 Brachycentrus occidentalis 8 Micrasema 587 Glossosomatidae 1 Glossosoma 1 Helicopsychidae 8 Helicopsyche 165 Helicopsyche borealis 9 Hydropsychidae 90

Arctopsyche 2 Hydropsyche 1735 Parapsyche 23 Smicridea 1 Hydroptilidae 115 Hydroptila 20 Leucotrichia 7 Neotrichia 1 Ochrotrichia 1 Lepidostomatidae Lepidostoma 72 Lepidostoma neodinarthrum 3 Leptoceridae 6 Oecetis 4 Triaenodes 2 Ylodes 2 Limnephilidae 18 Amphicosmoecus canax 1 Dicosmoecus 1 Hesperophylax 115 Limnephilus 77 Onocosmoecus 1 Philopotamidae 15 Chimarra 62 Dolophilodes 2 Wormaldia 14 Phryganeidae Agrypnia 1 Psychomyiidae 1 Tinodes 82 Rhyacophilidae 2 Rhyacophila 129 Sericostomatidae Gumaga 9 Uenoidae Neophylax 13 Neothremma 6 Oligophlebodes 97 Ostracoda Podocopida 237 Candoniidae Candona 1Mollusca Gastropoda 2 Basommatophora Ancylidae Ferrissia 1 Lymnaeidae 22 Fossaria 5 Radix 6 Stagnicola 2 Physidae Physella 130 Planorbidae 12 Gyraulus 5 Pelecypoda Veneroidea Sphaeriidae Pisidium 49

13

Nematoda 38Nematomorpha 17Platyhelminthes Turbellaria 28

Vertebrata Amphibia Anura 11 Bufonidae Bufo 3 Bufo woodhousei 8

Pelobatidae Spea intermontana 70 Osteichthyes 2 Cypriniformes Cyprinidae 9

Literature Cited

Cuffney, T.F., Gurtz, M.E. and Meador, M.R.. 1993. Methods for collecting benthic invertebrate samples

as part of the National Water-Quality Assessment Program: United States Geological Survey

Open-File Report 93-406.

Vinson, M.R. and C.P. Hawkins. 1996. Effects of sampling area and subsampling procedures on

comparisons of taxa richness among streams. Journal of the North American Benthological Society

15:393-400.

Wolman, M.G. 1954. A method of sampling coarse river-bed material. Transactions of the American

Geophysical Union 35:951-956.

14

Appendix 1A. List of taxa collected in 10 samples taken in alcove pools within and

adjacent to the Grand Staircase - Escalante National Monument.

Annelida Oligochaeta Haplotaxida TubificidaeArthropoda Arachnida Trombidiformes Crustacea Amphipoda Hyalellidae Hyallela azteca Decapoda Insecta Coleoptera Dytiscidae Agabus Deronectes Laccophilus Rhantus Thermonectus mormoratus Gyrinidae Gyrinus Haliplidae Haliplus Peltodytes Hydraenidae Hydraena Hydrophilidae Berosus Lacobius Paracymus Scirtidae Elodes

Diptera Ceratopogonidae Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Culicidae Dixidae Dixella Simuliidae Simulium Tabanidae Chrysops Tipulidae Limonia Ephemeroptera Baetidae Baetis Baetis tricaudatus Callibaetis Hemiptera Belostomatidae Lethocerus Corixidae Neocorixa Gerridae Aquarius Notonectidae Notonecta Notonecta kirbyi Notonecta lobata

Odonata Aeshnidae Aeshna umbrosa Coenagrionidae Argia Coenagrion Enallagma Gomphidae Lestidae Archilestes grandis Libellulidae Libellula Libellula saturata Trichoptera Hydroptilidae Ochrotrichia Ostracoda Podocopida Candoniidae Candona Mollusca Gastropoda Basommatophora Physidae Physella Pelecypoda Veneroidea Sphaeriidae Pisidium Platyhelminthes Turbellaria

15

Appendix 1B. List of taxa collected in a single sample taken in a hanging garden

adjacent to the Grand Staircase - Escalante National Monument near Lake Powell.

Arthropoda Insecta Coleoptera Dytiscidae Agabus Elmidae Zaitzevia Hydraenidae Hydraena Hydrophilidae Scirtidae Elodes Staphylinidae Diptera Chironomidae Chironominae Dixidae Dixella Stratiomyidae Caloparyphus Ephemeroptera Baetidae Baetis Hemiptera Gerridae Aquarius

16

Appendix 1C. List of benthic taxa and zooplankton collected 13 June 1999 in Posy

Lake within the Dixie National Forest updrainage from the Grand Staircase - Escalante

National Monument.

Annelida OligochaetaArthropoda Crustacea Amphipoda Gammaridae Gammarus lacustris Hyalellidae Hyallela azteca Calanoida Temoridae Epischura nevadensis Cladocera Daphnidae Daphnia rosea Daphniidae Ceriodaphnia laticaudata Ceriodaphnia quadrangula Cyclopoida Insecta Diptera Ceratopogonidae Bezzia

Chironomidae Chironominae Tanypodinae Hemiptera Corixidae Corisella Odonata Aeshnidae Aeshna umbrosa Coenagrionidae Telebasis salva Trichoptera Limnephilidae Limnephilus Ostracoda PodocopidaMollusca Pelecypoda Veneroidea Sphaeriidae Pisidium

17

Appendix 1D. List of taxa collected in 10 samples taken in ponds, wetlands, and

reservoirs within and adjacent to the Grand Staircase - Escalante National Monument.

Annelida OligochaetaArthropoda Crustacea Amphipoda Hyalellidae Hyallela azteca Cladocera Daphnidae Daphnia Daphnia rosea Cyclopoida Insecta Coleoptera Amphizoidae Amphizoa Dytiscidae Agabus Deronectes Hygrotus Oreodytes Haliplidae Peltodytes Helophoridae Helophorus Hydrophilidae Paracymus Diptera Ceratopogonidae Bezzia

Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Muscidae Simuliidae Simulium Stratiomyidae Myxosargus Ephemeroptera Baetidae Baetis Baetis bicaudatus Baetis tricaudatus Callibaetis Hemiptera Corixidae Corisella Neocorixa Gerridae Notonectidae Notonecta Odonata Coenagrionidae Enallagma cyathigerum Lestidae Archilestes grandis Libellulidae Libellula

Trichoptera Hydropsychidae Hydropsyche Hydroptilidae Hydroptila Phryganeidae Agrypnia Ostracoda PodocopidaMollusca Gastropoda Basommatophora Lymnaeidae Radix Stagnicola Physidae Physella Planorbidae Gyraulus Pelecypoda Veneroidea Sphaeriidae Pisidium NematodaVertebrata Amphibia Anura Pelobatidae Spea intermontana

18

Appendix 1E. List of taxa collected in 13 samples taken in springs and seeps within and

adjacent to the Grand Staircase - Escalante National Monument.

Annelida OligochaetaArthropoda Arachnida Trombidiformes Crustacea Amphipoda Hyalellidae Hyallela azteca Decapoda Insecta Coleoptera Dytiscidae Agabus Deronectes Laccophilus Oreodytes Rhantus Elmidae Microcylloepus Hydrophilidae Berosus Lacobius Paracymus Tropisternus Diptera Ceratopogonidae Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Culicidae Culiseta

Dixidae Dixella Empididae Clinocera Simuliidae Simulium Stratiomyidae Stratiomys Tabanidae Tabanus Tipulidae Hexatoma Limonia Tipula Ephemeroptera Baetidae Baetis Baetis tricaudatus Callibaetis Hemiptera Corixidae Gelastocoridae Gelastocoris oculatus Gerridae Aquarius Gerris Notonectidae Notonecta kirbyi Veliidae Lepidoptera Pyralidae Petrophila Odonata Coenagrionidae

Argia Argia vivida Lestidae Archilestes grandis Trichoptera Helicopsychidae Helicopsyche Limnephilidae Limnephilus Philopotamidae Chimarra Psychomyiidae Tinodes Sericostomatidae Gumaga Ostracoda PodocopidaMollusca Gastropoda Basommatophora Lymnaeidae Physidae Physella NematodaTerrestrial - bugVertebrata Amphibia Anura Pelobatidae Spea intermontana Osteichthyes Cypriniformes Cyprinidae

19

Appendix 1F. List of taxa collected in 257 samples taken in rivers and streams within

and adjacent to the Grand Staircase - Escalante National Monument.

Annelida Hirudinea Oligochaeta Haplotaxida Tubificidae Lumbriculida LumbriculidaeArthropoda Arachnida Trombidiformes Crustacea Amphipoda Crangonyctidae Stygobromus hubbsi group Gammaridae Gammarus lacustris Hyalellidae Hyallela azteca Decapoda Cambaridae Orconectes virillis Isopoda Entognatha Collembola Insecta Coleoptera Curculionidae Dryopidae Helichus Dytiscidae Agabus Deronectes Eretes sticticus Hydroporous Laccophilus Oreodytes Rhantus Thermonectus mormoratus Elmidae Cleptelmis Heterelmis Heterlimnius Microcylloepus Microcylloepus pusillus Narpus Optioservus Ordobrevia Stenelmis Zaitzevia Gyrinidae

Gyrinus Haliplidae Peltodytes Helophoridae Helophorus Hydraenidae Ochthebius Hydrophilidae Ametor Berosus Hydrobius Hydrochara Lacobius Paracymus Tropisternus Lampyridae Scirtidae Elodes Diptera Athericidae Atherix Ceratopogonidae Atrichopogon Culicoides Dasyhelea Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Culicidae Culiseta Dixidae Dixa Dixella Dolichopodidae Empididae Chelifera Clinocera Hemerodromia Ephydridae Muscidae Limnophora Psychodidae Maruina Pericoma Simuliidae Prosimulium Simulium Stratiomyidae

Caloparyphus Euparyphus Myxosargus Nemotelus Stratiomys Tabanidae Tabanus Tipulidae Antocha Dicranota Erioptera Hesperoconopa Hexatoma Limonia Tipula Ephemeroptera Ameletidae Ameletus Baetidae Baetis Baetis bicaudatus Baetis tricaudatus Callibaetis Paracloeodes Caenidae Caenis Ephemerellidae Drunella coloradensis Drunella doddsi Drunella grandis Ephemerella Ephemerella inermis Heptageniidae Cinygmula Epeorus Heptagenia Nixe Rhithrogena Leptophlebiidae Paraleptophlebia Tricorythidae Tricorythodes Hemiptera Belostomatidae Lethocerus Corixidae Neocorixa Gerridae Aquarius Gerris

20

Naucoridae Ambrysus Ambrysus mormona Notonectidae Notonecta Notonecta kirbyi Notonecta undulata Veliidae Microvelia Rhagovelia Lepidoptera Pyralidae Petrophila Megaloptera Corydalidae Corydalus cognatus Odonata Aeshnidae Aeshna Aeshna umbrosa Anax walsinghami Calopterygidae Hetaerina Hetaerina americana Coenagrionidae Argia Argia plana Argia vivida Coenagrion Enallagma Ischnura Cordulegastridae Cordulegaster diadema Cordulegaster dorsalis Gomphidae Erpetogomphus Erpetogomphus compositas Gomphus Ophiogomphus Ophiogomphus severus Lestidae Archilestes Archilestes grandis Libellulidae Libellula Paltothemis lineatipes Pantala flavescens Tramea Plecoptera Capniidae

Utacapnia Chloroperlidae Suwallia Sweltsa Leuctridae Nemouridae Malenka Prostoia Zapada Perlidae Hesperoperla pacifica Perlodidae Isogenoides Isogenoides zionensis Isoperla Isoperla longiseta Megarcys Megarcys signata Skwala Pteronarcyidae Pteronarcella badia Taeniopterygidae Taenionema Trichoptera Brachycentridae Brachycentrus Brachycentrus americanus Brachycentrus occidentalis Micrasema Glossosomatidae Glossosoma Helicopsychidae Helicopsyche Helicopsyche borealis Hydropsychidae Arctopsyche Hydropsyche Parapsyche Smicridea Hydroptilidae Hydroptila Leucotrichia Neotrichia Lepidostomatidae Lepidostoma Lepidostomaneodinarthrum Leptoceridae Oecetis Triaenodes Ylodes

Limnephilidae Amphicosmoecus canax Dicosmoecus Hesperophylax Limnephilus Onocosmoecus Philopotamidae Chimarra Dolophilodes Wormaldia Psychomyiidae Tinodes Rhyacophilidae Rhyacophila Sericostomatidae Gumaga Uenoidae Neophylax Neothremma Oligophlebodes Ostracoda PodocopidaMollusca Gastropoda Basommatophora Ancylidae Ferrissia Lymnaeidae Fossaria Radix Physidae Physella Planorbidae Gyraulus Pelecypoda Veneroidea Sphaeriidae Pisidium NematodaNematomorphaPlatyhelminthes TurbellariaTerrestrial - bugVertebrata Amphibia Anura Bufonidae Bufo Pelobatidae Spea intermontana

21

Appendix 1G. List of taxa collected in 52 samples taken in tinajas (rock pools) within

and adjacent to the Grand Staircase - Escalante National Monument. Annelida Hirudinea OligochaetaArthropoda Arachnida Trombidiformes Crustacea Anostraca Branchinectidae Branchinecta packardi Calanoida Diaptomidae Diaptomus Cladocera Bosminidae Bosmina longirostris Daphnidae Daphnia Daphnia pulex Daphniidae Ceriodaphnia Ceriodaphnia quadrangula Copepoda Cyclopoida Notostraca Triopsidae Triops longicaudatus Insecta Coleoptera Dytiscidae Agabus Coptotomus Deronectes Laccophilus Neoclypeodytes Oreodytes Rhantus Elmidae Microcylloepus pusillus Gyrinidae Gyrinus

Helophoridae Helophorus Hydrophilidae Berosus Tropisternus Diptera Ceratopogonidae Bezzia Culicoides Dasyhelea Probezzia Chironomidae Chironominae Orthocladiinae Tanypodinae Culicidae Culiseta Ephydridae Simuliidae Simulium Tabanidae Tipulidae Dicranota Tipula Ephemeroptera Baetidae Baetis tricaudatus Callibaetis Heptageniidae Hemiptera Corixidae Corisella Neocorixa Gerridae Aquarius Gerris Naucoridae Ambrysus mormona Notonectidae Notonecta Notonecta kirbyi

Veliidae Lepidoptera Odonata Aeshnidae Aeshna umbrosa Coenagrionidae Enallagma Lestidae Archilestes Archilestes grandis Lestes Libellulidae Libellula saturata Trichoptera Brachycentridae Brachycentrus Micrasema Helicopsychidae Helicopsyche borealis Hydropsychidae Hydropsyche Limnephilidae Limnephilus Ostracoda PodocopidaMollusca Gastropoda Basommatophora Physidae Physella NematodaNematomorphaTerrestrial - bugVertebrata Amphibia Anura Bufonidae Bufo woodhousei Pelobatidae Spea intermontana

22

Appendix 2. Aquatic invertebrate sampling locations. Station notations with BLMand

NPS designations were sites sampled by BLM personnel in Escalante and National

Park Service personnel from the Glen Canyon National Recreation Area. These

samples were processed by the BugLab, Logan, Utah.

Station Location Latitude Longitude Elevation (m)

ES-001 Escalante River near Escalante Cemetery trailhead 37.776 111.579 1737ES-002 Tinaja, down drainage from HWY 12 below overlook 37.754 111.443 1737ES-003 Vinson Creek, trib to Deer Creek, off Burr Trail 37.848 111.374 1707ES-004 Deer Creek, downstream from campground 37.855 111.355 1646ES-005 Steep Creek at mouth 37.864 111.312 1567ES-006 Gulch Creek upstream from Burr Trail bridge 37.865 111.313 1579ES-007 Calf Creek, just downstream from upper falls 37.855 111.451 1676ES-008 Tinaja, near Phipp's pictographs 37.759 111.419 1524ES-009 Calf Creek, at mouth 37.776 111.419 1439ES-010 Deer Creek, below tributary ES-003 37.838 111.367 1701ES-011 Tinaja near Deer Creek 37.838 111.366 1777ES-012 Tinaja between Hwy 12 and Boulder Creek 37.810 111.400 1679ES-013 Tinaja between Hwy 12 and Boulder Creek 37.809 111.397 1747ES-014 Tinaja between Hwy 12 and Boulder Creek 37.809 111.396 1630ES-015 Boulder Creek, off Hwy 12 near Calf Creek CG 37.812 111.394ES-016 Little Death Hollow Creek midway down 37.759 111.215 1765ES-017 Tinaja near Highway 12, down drainage from ES-002 37.749 111.444 1860ES-018 Tinaja near Highway 12, down drainage from ES-018 37.750 111.444 1878ES-019 Alcove pool downstream from upper Calf Creek Falls 37.854 111.451 1420ES-020 Tenmile Creek 37.680 111.485ES-021 Alvey's Wash Creek 37.719 111.629 1675ES-022 Upper Alvey Wash Creek 37.695 111.631 1933ES-023 North Creek 37.778 111.708 1910ES-024 Birch Creek 37.764 111.696 1910ES-025 Escalante River at Birch Creek and North Creek 37.765 111.684 1882ES-026 No-name Reservoir off Pine Creek Road 37.805 111.601 1868ES-027 Pine Creek below lower box trailhead 37.816 111.608 1720ES-028 Upper Valley Creek, west of Escalante 37.747 111.703 1720ES-029 Harris Wash Creek near trailhead 37.634 111.341 1720ES-030 Un-Named Creek between Sand and Mamie Creeks 37.777 111.473 1720ES-031 Mamie Creek at mouth 37.781 111.505 1720ES-032 Escalante River upstream from Mamie Creek 37.781 111.523 1720ES-033 Tinaja near Escalante River, between Mamie & Sand 37.778 111.495 1720ES-034 Sand Creek at mouth 37.775 111.458 1720ES-035 Spring near Sand Creek 37.776 111.459ES-036 Lake Creek 37.923 111.524 2097ES-037 Tinaja upstream from upper Calf Creek Falls 37.867 111.462 1853ES-038 Calf Creek at source 37.866 111.462 1701ES-039 Calf Creek above upper falls 37.854 111.453 1798ES-040 Tinaja south of Deer Creek, west of The Gulch 37.821 111.343 1695ES-041 Tinaja south of Deer Creek, west of The Gulch 37.819 111.342 1707ES-044 Coyote Gulch downstream from Hamblin Arch 37.420 111.040 1274ES-045 Seep in Coyote Gulch 37.422 111.003 1158ES-046 Coyote Gulch at mouth 37.428 111.981 1122

23

ES-047 Harris Wash, 0.5 near NRA boundary 37.642 111.290 1494ES-048 Harris Wash - Upper 37.637 111.319 1506ES-049 The Gulch at mouth 37.725 111.296 1512ES-050 Escalante River upstream from Harris Wash 37.663 111.215 1390ES-051 Harris Wash at mouth 37.663 111.215 1390ES-052 Escalante River at bridge north of Escalante town 37.776 111.593 1768ES-053 Pleasant Creek at Pleasant Creek campground 38.102 111.336 2670ES-054 Wetland off Burr Trail, trib to Vinson Creek 37.860 111.381 1807ES-055 Tinaja West of MeGath Point 37.835 111.465 1945ES-056 Tinaja below Early Weed Bench 37.541 111.186 1433ES-057 Tinaja East of Haymaker Bench 37.790 111.392 1719ES-060 Cat Well 37.486 111.253 1524ES-061 Orins Well 37.456 111.236 1520ES-062 Tinaja at Dance Hall Rock 37.360 111.097 1537ES-063 Ampitheater Tinaja at Dance Hall Rock 37.357 111.101 1512ES-064 Seep to Forty Mile Creek 37.342 111.018 1150ES-065 Hidden Falls Pool in Forty Mile Creek 37.343 111.014 1124ES-066 Spring off Forty Mile Creek 37.344 111.014 1176ES-067 Forty Mile Creek 37.342 111.006 1176ES-068 Seep at Fifty Mile Creek, 4 mi. from Cave Pt. TH 37.317 110.976 1184ES-069 Fifty Mile Creek 37.320 110.962 1184ES-070 North Creek, upstream from reservior 37.847 111.758 2099ES-071 Lost Creek, east of Twin Lakes 37.914 111.699 2766ES-072 Posy Lake 37.936 111.694 2499ES-073 Hungry Creek, 1 mile SE of Posy Lake 37.930 111.684 2687ES-074 Deep Creek, 2 miles E of Posy Lake 37.936 111.666 2616ES-075 Blue Spring Creek at Hell's Backbone Road 37.963 111.656 2422ES-076 Grimes Creek near Hell's Backbone Road 37.970 111.574 2496ES-077 Death Hollow Seep, upstream from Mail Trail 37.844 111.521 1977ES-078 Death Hollow Creek at Boulder Mail Trail 37.844 111.521 1977ES-079 Tinaja between Sand Creek and Death Hollow 37.860 111.492 1851ES-080 Sand Creek upstream from Boulder Mail Trail 37.867 111.496 2008ES-081 Lake Creek at confluence with Sand Creek 37.868 111.495ES-082 Sand Creek at Lake Creek confluence 37.867 111.495 1782ES-083 Tinaja near Sand Creek mouth 37.863 111.490 1875ES-084 Reservoir off Burr Trail 37.920 111.140 2005ES-085 Reservoir 2 off Burr Trail Rd. 37.874 111.106 2046ES-086 Frisky Creek near source 38.013 111.373 2754ES-087 Upper Steep Creek 37.873 111.323 1743ES-088 Lower Steep Creek 37.866 111.317 1701ES-089 Water Canyon Creek 37.915 111.292 1798ES-090 Water Canyon Creek upstream from The Gulch 37.904 111.285 1777ES-091 Large pond off Burr trail 37.887 111.398 1859ES-092 Small pond off Burr trail 37.894 111.406 1875ES-093 Sheep Creek upstream from spillway 37.496 112.066 1829ES-094 Yellow Creek upstream from old homestead 37.535 112.063 1719ES-095 Paria River upstream from Cannonville Bridge 37.569 112.051 1768ES-096 Pine Creek near upper trailhead 37.963 111.652 2341ES-097 Pine Creek between upper and lower trailheads 37.934 111.643 2195ES-098 Hungry Creek upstream from Pine Creek 37.917 111.645 2134ES-099 Pine Creek at Lower Box Trailhead 37.864 111.635 1731ES-100 Ribbon Canyon Alcove 1 37.255 110.842 1247ES-101 Ribbon Canyon Hanging Garden 37.249 110.832 1247

24

ES-102 Ribbon Canyon Alcove 2 37.259 110.839 1247ES-103 Bown's Canyon Alcove Pool 37.359 110.875 1170ES-104 Bown's Canyon lower 37.374 110.878 1209ES-105 Bown's Canyon upper 37.380 110.886 1228ES-106 Iceberg Canyon Alcove 37.293 110.734 1140ES-107 Iceberg Canyon Creek 37.286 110.732 1136ES-108 Iceberg Canyon Creek, pool at head of left fork 37.295 110.704 1189ES-109 Cottonwood Canyon Alcove 37.242 110.854 1134ES-110 Cottonwood Canyon Seep 37.242 110.855 1131ES-111 Death Hollow Creek, 2 mi upstream Escalante River 37.805 111.511 1707ES-112 Death Hollow Creek, at mouth 37.782 111.505 1646ES-113 Calf Creek, downstream from bridge, near HWY 12 37.793 111.413 1631ES-114 Calf Creek, downstream from Lower Falls 37.829 111.419 1722ES-115 Alcove pool near Upper Calf Creek Falls 37.815 111.418 1692ES-116 Tinaja upstream from Upper Calf Creek Falls 37.867 111.462 1853ES-117 Escalante River at confluence with Coyote Gulch 37.428 111.981 1122ES-118 Lake Powell, Red Canyon 37.716 110.458ES-119 Escalante River above Sand Crk. 37.776 111.459 1619ES-120 Escalante River at Hwy 12 Bridge 37.776 111.419 1521ES-121 Boulder Creek at Hwy 12 37.901 111.437 2003ES-122 Tinaja off trail to Upper Calf Creek Falls 37.855 111.448 1825ES-123 Calf Creek, big pool upstream Up Calf Creek Falls 37.856 111.453 1638ES-124 Willis Creek above Skutumpah Road 37.483 112.097 1757ES-126 Sheep Creek below spillway 37.493 112.064 1798ES-127 Tinaja A, between HWY 12 and Boulder Creek 37.810 111.399 1793ES-128 Tinaja B, between HWY 12 and Boulder Creek 37.809 111.396 1762ES-129 Tinaja C, between HWY 12 and Boulder Creek 37.810 111.396 1719ES-129B Tinaja D, between HWY 12 and Boulder Creek 37.810 111.396 1719ES-130 Boulder Creek, near Calf Creek Campground 37.811 111.395 1756ES-131 Tinaja E, between HWY 12 and Boulder Creek 37.811 111.394 1734ES-132 Tinaja off Hwy 12 between miles 71 & 72, west side 37.752 111.445 1625ES-133 Tinaja, updrainage of HWY 12, near mile 71-72 37.754 111.444 1710ES-135 Escalante River at Boulder Creek confluence 37.758 111.350 1573ES-136 Boulder Creek at confluence with Escalante River 37.758 111.350 1573ES-BLM-001 Drip Tank Creek 1512ES-BLM-002 Chimney Rock CreekES-BLM-003 Brown Spring 1890ES-BLM-004 Cottonwood Creek 1585ES-BLM-005 Three Mile Creek 2164ES-NPS-001 Escalante River 37.500 111.042 1262ES-NPS-002 Escalante River 37.333 110.917 1085ES-NPS-SJ1 San Juan River 37.197 109.975 1195ES-NPS-SJ2 San Juan River 37.195 109.982 1195ES-NPS-SJ3 San Juan River 37.208 109.979 1195ES-NPS-SJ4 San Juan River 37.212 109.981 1195