Activity 3. Cnidarians

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    Cnidaria (Grknide, nettle + L. aria; like orconnected with

    With more than 9,000 species Cnidocytes = which contain stinging

    organelles (cnidae)

    Cnidae come in several types includingthe common nematocysts

    Nematocysts are used only bycnidarians

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    Phylum Coelenterata

    Gr. koilos, hollow, + enteron, gut + L. ata,

    characterized by

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    BODY PLANS: Have two body forms MEDUSA & POLYPS

    Have Radial Symmetry

    Known for Cnidocytes contain stinging cells such as nematocyst

    REPRODUCTION: Reproduce Sexually & Asexually

    Free-swimming larva - planula

    CNIDARIAN CLASSES:

    o Anthozoa sea anemones and corals (polyps; no medusa stage)

    o Hydrozoa fire corals, Portuguese man-of-war, hydras

    o Scyphozoa jellyfish medusa form is dominant but may begin as polyp

    o Cubozoa

    box jellyfish & sea wasp

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    Entirely aquatic (mostly marine)

    With oral and aboral ends

    Radial symmetry or biradial symmetryaround a longitudinal axis Radial symmetry = body parts are arranged

    concentrically around an oral-aboral axis

    Biradial symmetry = is basically a type of radialsymmetry in which only two plane through theoral-aboral axis divide the animal into mirrorimages

    No definite head

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    Exoskeleton or endoskeletonchitinous,calcareous, or protein components in

    some Body with 3 layers: epidermis and

    gastrodermis , mesoglea (acellular)

    Gastrovascular cavity orcoelenteron(often branched or dividedwith septa) = with a single opening thatserves as mouth and anus

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    Extensive tentacles usually encircling themouth or oral region

    Special stinging-cell organelles calledcnidae, in particular nematocyst, ineither epidermis or gastrodermis or inboth

    Monoecious and dioecious

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    Two basic forms:

    Polyps

    Or hydroid form

    Adapted to asedentary or sessile

    life

    Medusae

    Or jellyfish form

    Adapted for afloating or free-swimming existence

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    Polyps

    Have tubular bodies

    Mouth surrounded by tentacles (defines theoral end)

    Mouth leads into a blind gut orgastrovascular cavity

    Aboral end is usually attached toasubstratum by a pedal disc or other device

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    Reproduceasexually by:

    Budding

    Fission Pedal laceration

    Zooids

    buds that remains attached to the polyp:gastrozooids (for feeding)dactylozooids (for defense)gonozooids (for making sexually reproducing stages)

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    Usually free-swimming

    Have bell-shaped or

    umbrella-shaped bodies Tetramerous symmetry (body parts

    arranged in fours)

    Mouth is usually centered on theconcave side

    Tentacles extend from the rim of theumbrella

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    Jellylike layer of mesoglia is thickerconsisting the bulk of the animal and

    making it more buoyant

    medusae arecommonly called jellyfishes

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    Algae frequently live as mutuals intissues of cnidarians, notably in some

    freshwater hydras and reef-buildingcorals.

    The presence of algae in reef-buildingcorals limits the occurrence of coral reefsto relatively shallow, clear water wheresunlight is sufficient for photosyntheticrequirements of the algae

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    Hydras as food for mollusc and flatworms

    Live as commensals on shells and other

    surface of host (sea anemones andcertain hydroids)

    Economic value of reef

    Reef provide substantial amounts of food forhuman (fish and other animals associatedwith reefs

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    Includes:branching, plantlike hydroids

    Flowerlike sea anemones Architects of the ocean floor- gorgonian

    corals (sea whips, sea fans, etc)

    Stony coralswhose thousand of years ofcalcareous housebuilding great reefs

    and coral islands.

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    Class Hydrozoa Hydra

    Obelia

    Goneonemus

    Class Scyphozoa Jellyfishes

    Class Anthozoa Sea anemone

    Stony corals

    Class Cubozoa

    The basis of classification is the adult forms whether polyp or medusa.

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    Fresh water members of this class livesingly, but the marine forms live singly or

    form polymorphic colony. Though only polyp or medusa is found

    but sometimes both polyp or medusamay be found in the same animal.

    Gastrovascular cavity is directlyconnected with mouth.

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    Alternation of generation is found in thelife cycle.

    Ciliated planula larva is found. No stomodeum, velum present in

    medusa.

    Radial canals in medusa are notbranched.

    Gametes develop from ectoderm.

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    Kingdom Animalia

    Phylum Cnidaria

    Class Hydrozoa OrderHydroida

    SuborderHydrida

    Family Hydridae Genus Hydra

    Species: Hydra vulgaris

    http://animaldiversity.ummz.umich.edu

    http://animaldiversity.ummz.umich.edu/site/accounts/pictures/Animalia.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Cnidaria.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydrozoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydroida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydrida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydridae.htmlhttp://animaldiversity.ummz.umich.edu/http://animaldiversity.ummz.umich.edu/http://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydridae.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydrida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydroida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Hydrozoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Cnidaria.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/pictures/Animalia.html
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    Kingdom Animalia (animals) Eumetazoa (metazoans)

    Phylum Cnidaria (cnidarians)

    Class Hydrozoa (hydralike animals,hydroids, and hydrozoans)

    OrderHydroida (medusae)

    SuborderHydrida

    Family Hydridae Genus Hydra

    Species Hydra utahensis

    http://animaldiversity.ummz.umich.edu/site/accounts/classification/Animalia.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Eumetazoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Cnidaria.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydrozoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydroida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydrida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydridae.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydra.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydra.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydridae.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydrida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydroida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydroida.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydrozoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Hydrozoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Cnidaria.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Cnidaria.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Eumetazoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Eumetazoa.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Animalia.htmlhttp://animaldiversity.ummz.umich.edu/site/accounts/classification/Animalia.html
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    Class Hydrozoa Order Hydroida

    Suborder Anthomedusae

    Freshwater species

    Hydranths lack achitinous covering

    With no medusa stage

    Length up to 25-30 mm

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    ZOOIDSMostly HYDRANTHS(Gastrozooids

    feeding polyp

    STALK

    BASE (basal disc

    or pedal)

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    http://www.geochembio.com/biology/organisms/hydra/

    http://www.geochembio.com/biology/organisms/hydra/http://www.geochembio.com/biology/organisms/hydra/
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    Class Hydrozoa

    Order Hydroida

    Suborder Leptomedusae

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    Dioecious

    Gametes are fertilized in the water and thezygote develops into a planula larva whichsettles into substrate and metamorphose intocolony

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    Kingdom: Animalia

    Subkingdom: Eumetazoa

    Phylum: Cnidaria Subphylum: Medusozoa

    Class: Hydrozoa

    Subclass: Leptolinae Order: Leptomedusae

    Suborder: Proboscidoidea Family: Campanulariidae

    Genus: ObeliaPeron and Lesueur, 1810

    Species: Obelia dichotoma

    Obelia geniculata Obelia longissima

    Obelia bidentata

    http://en.wikipedia.org/wiki/Obelia

    http://en.wikipedia.org/wiki/Animalhttp://en.wikipedia.org/wiki/Eumetazoahttp://en.wikipedia.org/wiki/Cnidariahttp://en.wikipedia.org/wiki/Medusozoahttp://en.wikipedia.org/wiki/Hydrozoahttp://en.wikipedia.org/wiki/Leptolinaehttp://en.wikipedia.org/wiki/Leptomedusaehttp://en.wikipedia.org/w/index.php?title=Proboscidoidea&action=edit&redlink=1http://en.wikipedia.org/wiki/Campanulariidaehttp://en.wikipedia.org/wiki/Obeliahttp://en.wikipedia.org/wiki/Obeliahttp://en.wikipedia.org/wiki/Campanulariidaehttp://en.wikipedia.org/w/index.php?title=Proboscidoidea&action=edit&redlink=1http://en.wikipedia.org/wiki/Leptomedusaehttp://en.wikipedia.org/wiki/Leptolinaehttp://en.wikipedia.org/wiki/Hydrozoahttp://en.wikipedia.org/wiki/Medusozoahttp://en.wikipedia.org/wiki/Cnidariahttp://en.wikipedia.org/wiki/Eumetazoahttp://en.wikipedia.org/wiki/Animal
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    The medusa of Gonionemus. hydrozoa

    http://webs.lander.edu/rsfox/invertebrates/gonionemus.html

    http://webs.lander.edu/rsfox/invertebrates/gonionemus.htmlhttp://webs.lander.edu/rsfox/invertebrates/gonionemus.html
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    Anthozoans include

    corals, sea anemones,sea pens, and seapansies.

    These animals are eithersolitary or colonialpolyps that liveattached to a substrate(surface).

    Of the 6,000 knownanthozoan species,corals comprise about

    2,500 species.

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    Sea anemones and corals have polypsand no medusae

    Corals are colonial and secrete calciumcarbonate skeletons. Coral reefs are theaccumulation of these skeletons.

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    The Class Anthozoa is further divided into three subclasses:Octocorallia, Zoantharia, and Tabulata (extinct colonialcorals).

    Subclass Octocorallia. Polyps are characterized byhaving eightpinnate (side- branching) tentacles.Octocorallians include gorgonian corals, sea pens, seapansies, organ- pipe corals, and soft corals (orderAlcyonacea). Most are colonial.

    Subclass Zoantharia. Polyps are characterized by havingtentacles in multiples of six. Zoantharian tentacles arerarely pinnate. Black corals and reef-building corals (orderScleractinia) are members of this subclass. Reef-buildingcorals are also known as "hard corals" or "stony corals".Zoantharians may be either solitary or colonial.

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    Classification:

    Kingdom Animalia (animals)Phylum Coelenterata (corals, jellyfish, sea anemoneshydroids)

    Class Anthozoa meaning "flower-like animals"(corals and sea anemones)

    Order Actiniaria.

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    Diadumene dia

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    The sea anemone is considered to be the flower

    of the sea. Sea anemones look like plants, but

    they are really meat-eating animals.

    Sea anemones come in different sizes and many

    different colors.The sea anemone can attach

    itself to rocks or coral. In the center of the seaanemone is their mouth. In order for the sea

    anemone to eat, they must wait for their food to

    swim by. Then they sting it with their tentacles,

    and then push it into their mouth.

    Sea anemones spend most of their lives in one

    place. They can attach themselves to rocks.

    Some even bury themselves in the mud.

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    Class AnthozoaSubclass Zoantharia (Hexacorallia)

    OrderScleractiniaGenus Meandrina (Brain coral)

    Genus Astrangia (Eyed coral)Genus Fungia (Mushroom coral)Genus Acropora (Staghorn, elkhorn, Antler coral)Genus Porites (reef coral)Genus Oculina

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    This Class contains the largerjellyfishes,with some reaching two metres acrossthe bell and with tentacles 30 metres

    long.

    They are mainly marine and free floating,

    though they can "swim" by pulsations ofthe bell. In this Class the polyp stage iseither reduced or absent. There arearound 215 species.

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    Class Scyphozoa

    Order Semaeostomeae

    http://www.discoverlife.org/nh/tx/Cnidaria/Jellyfish/

    Aurelia(Desor, 1848) Sea nettle

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm

    http://www.discoverlife.org/nh/tx/Cnidaria/Jellyfish/http://www.discoverlife.org/20/q?search=Aureliahttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://www.discoverlife.org/20/q?search=Aureliahttp://www.discoverlife.org/nh/tx/Cnidaria/Jellyfish/
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    Aurelia sp. is a typicalexample

    The lappets come in pairs,and between them is a senseorgan called a rhopaliumcontaining a statocysts togive the animal information

    on its equilibrium andorientation, sensory pits, and,in some species, ocelli (simpleeyes).

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm

    Aureliarhopallium,a multifunction sensory organ

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm
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    The oral arms captureprey.

    The tentacles are armedwith nematocysts (seeillustration on the right), asis the entire body surface.

    Aurelia sp. has relativelyshort tentacles, and feedson plankton. The planktonis passed to the mouth bycilia.

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm
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    Aurelia, early strobila stageAurelia, late strobila stage

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm
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    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm

    Aurelia, ephyra stage (young medusa

    http://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htmhttp://cas.bellarmine.edu/tietjen/AnimalDiversity/Lab4%20Porifera%20and%20Cnidaria.htm
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    Sea Nettle scientific classification:Kingdom: AnimaliaPhylum: CnidariaClass: Scyphozoa

    Order: SemaeostomeaeFamily: PelagiidaeGenus: ChrysaoraSpecies: Chrysaora fuscescens

    Pacific Sea Nettle (Chrysaora fuscescens), Montery BayAquarium

    Dec 2009, Photo by Mark Leavitt

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    The cubozoan body is shaped like a square bell, with themouth suspended inside it on a tube (the manubrium) thatleads upward to the stomach, which is inside the top part

    of the bell. Extending around the inside bottom of the bellis a ring of tissue called the velarium, and at the bottomcorners of the bell are muscular fleshy pads (pedalia), withone or more tentacles connected to each pedalium. Foursensory structures called rhopalia are located near thecenter of each of the four sides of the bell. Box jellyfishhave eyes that are surprisingly complex, including regionswith lenses, corneas, and retinas; however, box jellyfish donot have a brain, so how the images are interpretedremains unknown. Like all cnidarians, box jellyfish possessstinging cells that can fire a barb and transfer venom(Waggoner and Collins 2000).

    http://www.newworldencyclopedia.org/entry/Eyehttp://www.newworldencyclopedia.org/entry/Eyehttp://www.newworldencyclopedia.org/entry/Eye
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    Cubozoans are agile and active swimmers, unlike themore planktonic jellyfish. They have been commonlyobserved to swim a meter in just five to ten seconds,and there are unconfirmed reports of largespecimens of Chironex fleckeri swimming as fast astwo meters in one second (Waggoner and Collins2000). The high speeds are achieved through a kindof squirting action in which the medusa contractswhile the velarium at the bottom contracts evenmore, producing a constricted opening through

    which the water is forcefully expelled.

    Box jellies can be found in many tropical areas,including nearAustralia, the Philippines, Hawaii,and Vietnam.

    http://www.newworldencyclopedia.org/entry/Australiahttp://www.newworldencyclopedia.org/entry/Philippineshttp://www.newworldencyclopedia.org/entry/Hawaiihttp://www.newworldencyclopedia.org/entry/Vietnamhttp://www.newworldencyclopedia.org/entry/Australiahttp://www.newworldencyclopedia.org/entry/Philippineshttp://www.newworldencyclopedia.org/entry/Hawaiihttp://www.newworldencyclopedia.org/entry/Vietnamhttp://www.newworldencyclopedia.org/entry/Vietnamhttp://www.newworldencyclopedia.org/entry/Hawaiihttp://www.newworldencyclopedia.org/entry/Philippineshttp://www.newworldencyclopedia.org/entry/Australia
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    Kingdom:AnimaliaPhylum:CnidariaClass:CubozoaWerner,1975Order:CubomedusaeHaeckel 1877

    http://www.newworldencyclopedia.org/entry/Animalhttp://www.newworldencyclopedia.org/entry/Cnidariahttp://www.newworldencyclopedia.org/entry/Ernst_Haeckelhttp://www.newworldencyclopedia.org/entry/Ernst_Haeckelhttp://www.newworldencyclopedia.org/entry/Cnidariahttp://www.newworldencyclopedia.org/entry/Animal