Cuadernas Geología Ibérica Vol. 8 Págs. 847-865 Madrid 1982

SOME SPANISH CRETACEOUSBIVALVES

PORANNW V. DHoNDT*

ABSTRACT

Ibis paperdiscussesaselectionof Bivalves (Ostreacea,Pectinidae,Limidae, Spondytidae) from SpanishCretaceousdepasits. By usingmodernnanienclature,comparisonof Spanishfaunaswith thosefromdeposits outsideSpain becomespossible.Tentative palaeobiogeogra-phical conclusionsresultingherefram are:

1. Aptian bivalvesfrom EasternSpainare very similar with thasefram EasternFranceand Switzerland.

2. Cenomanianbivalvesfrom Somolinos(Guadalajara)and fromTejada(Burgos) have very many speciesin camnionwith Le Mans(France)and Upper Greensandfaunas as knawn fram Warminsterandotherlacalitiesin England,but nextla thosespeciesa few Tethyanelementsaccur.

3. Santonianbivalve faunas fi-am the Prepyrenees(mainly Léri-da) are very claseta faunasfroin the Provenceandfram the Gasandepositsin Austria: thus they belangto the NorthernTethys.

4. Campanian and Maastrichtiafl deposits of the same reglancontain faunal elementsal the sanietime claselo thaseof the Aqui-taine, ta thoseof Narth Africa, andpassiblyalsoendemiefactors.

RESUMEN

En este trabajo se presentauna selecciónde Bivalvas (Ostreacea,Pectinidae,Limidae, Spondylldae) del Cretácico español.El usa de

* Department of Palaentology,Koninklijk Belgisch Instituut voor Natuur-wetenschappen.Vautierstraat29, B 1040 Brussels,Belgium.

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una nomenclaturamoderna permite establecercomparacionesentrelas faunasespañolasy las de otros países.

A continuaciónse exponenlos resultadosde un intento de con-clusionespaleogeográficas:

1. Los bivalvos aptensesdel Este de Españason muy semejan-tes a los del Este de Franciay Suiza.

2. Los bivalvos cenomanensesde Somolinos (Guadalajara)y deTejada(Burgos) presentanmuchasespeciesen común con Le Mans(Francia) y las faunasde la «Upper Greensand»reconocidaen War-minster y otras localidadesinglesas;sin embargo,junto a ellas hayalgunos elementosdel Tethys.

3. Las faunasde bivalvos santoniensesde los Prepirineos(Léri-da principalmente)se aproximanmucho a las faunasde Provenzayde los depósitosde Gosau,en Austria: así> pues,pertenecenal Tethysseptentrional.

4. Los depósitos del Campaniensey Maastrichtiensede dichasreglonescontienen,además,elementosfaunísticosrelacionadostantocon los de Aquitania como con los del Norte de Africa, y posiblemen-te algunos factoresendémicos.

INTRODUCTION

SpanishCretaceousbivalveshavebeenstudiedfor about150 years.Spanishlocalities and specimensare mentioned in some of the fun-damentalWestern Europeanstratigraphicalworks [f. i. d’Orbigny’sProdrome(1850)].

SpanishCretaceousfaunasare important: the Iberian Penínsulawas placed betweenthe Tethys and WesternEurope (and was befo-re the wideníngof the North Atlantic not evenfar from North Ame.rica), Some Spanisharcas during the Cretaceoushad faunasalmostidentical with Western Europe (such as the Aptian around Morellaor the Cenomanianof Somolinos); other faunaswere purely Tethyan;again other faunal critities were of marginal Tethys affinity (thePrepyreneanSantonian-Campaníanfaunashadcharacteristicsof Aqui-taine, ProvenceandAustria).

Ihe study of the Cretaceousbivalves from Spain has unfortuna-tely often beenundertakenwith a strong provincial bias, by Spanishand by non-Spanishauthors; also some publications on Spanishbí-valves have not been generallyeasily accessiblelo the rest of Iheworld: this results in a nomenclativeusagerather different from theresí of Europe.

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In this paper, no attempt has beenmade at giving a completereview of SpanishCretaceousbivalves: 1 have intentionally limitedmyself to thematerialwhich 1 havebeenableto study(the collectionsand field localities are referred to in appendix). Though they helongto the same palaeogeographicalentities, the PortugueseCretaceousspecieshave beenomitted for practical reasons.The purposeof thepaper is to give a readjustednomenclature;by using those <new’ na-mes, palaeobiogeographicalinterpretation becomescasier.

FAUNAS

For eachspeciesmentioned,the stratotypeandtype areaor loca-lity are indicated, a good figure is referred to, the palaeogeographicdistribution briefly statedandthe occurrencesin Spain usted.

PEcTINIDAF

Camptonectesvirgatus (NILSSON, 1827): originally described fromthe SwedishLate Campanian;fig. in Dhondt (1972a, pl. 2, fig. 1);distribution: ubiquitous from the Mid Cenomanianto terminalMaastrichtian.Occurrence in Spain: Santonian: Lérida: Collades de Bastus(1. Br.).

Chlamysachates(COQUAND, 1865): originally describedfrom the Ap-tian of «Quatre-Dineroset ParrasMartin (Aragón)»: type specimensin the MAFI coIl., Budapest (Hungary); fig. herein pl. 1, fig. 7and 8; distribution: only known from the Aptian of Spain: Ara-gón: Coquand localities (MAFI), Castellón: Morelia (Sem. B.,U. Val.), Olocau del Rey (Sem. E.), Tarragona: Puenta del Aliga,Amettla de Mar (Sem.B.), Teruel: Mas Antonino, Peñarroyade Tas-tavins (Sem. E.).

Chlamyselongata (LAMARCK, 1819): originally described from theCenomanianof Le Mans (France); fig. in Ohondt (1973b, pl. 2);distribution: widely distributed from the Aptian to the terminalCenomanianin temperateseas.Occurrence in Spain: Cenomanian: Navarra: Orobe Alsasna(Sem. E.).

Chlamysespaillaci (D’ORBIGNY, 1847): originally describedfrom the«Senonian» of the Charenteand the Dordogne (France); Hg. ind’Orbigny (1847, pl. 439, figs. 1, 4).

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Occurrence in Spain: Santonian: Lérida: Montsec (Mus. B.),Maastrichtian: Barcelona: Figois-las-Minas(Seni. E.).

Chlamys cf. jaujasi (DEFRANCE, 1825): originally described fromthe Sint Pietersbergnear Maastricht (the Netherlands), LateMaastrichtian;fig. in Dhondt (1973b, pl. 1, fig. 2); distributed intemperateseasfrom latest Cenomanianto terminal Maastrichtian.Probable occurrencein Spain: Upper Coniacian: Navarra: Ola-zagutía (1. E. R).

Chlamys subacuta(LAMARCK, 1819): originally described from theCenomanianof Le Mans (France); fig. in Dhondt (1973b, pl. 4);distributed in temperateand northern Tethys depositsof Albianand Cenomanianage.Occurrencein Spain: Cenomanian:Guadalajara: Somolinos (Sem.B.), Soria: Picofrentes (1. Br.).

Entolium membranaceum(NILSSON, 1827): originally describedfromthe Late Campanianin Kñpinge (Sweden); fig. in Ohondí (1971,pl. 1, fig. 2); widely distributed in temperateseasfrom the latestCenomanianto the terminal Maastrichtian.Occurrencein Spain: TuronianVIII: Navarra: Orogoyen(1. Br.);Coniacian:Navarra: Zudaire(1. Br.); Late Coniacian:Lérida: nearHerbasina(seePons et al., 1981, p. 60) (1. Br.).

Entolium orbiculare (SOWERBY, 1817): originally described fromthe Greensandin Devizes (England) (Cenomanian);fig. in Dhodt(1971, pl. 1, fig. 1) and in B. M. F. (1975, pl. 56, fig. 2); widelydistributed in «boreal»to temperatedepositsfrom the Ryazanianto the Late Cenomanian.Occurrence in Spain: Aptian: Castellón: Alcalá-de-Chivert, NERasfail (Sen-, P~

Merklinia catalaunica(VInAL, 19221): originally describedfrom theCampanianof Gosol (Lérida); fig. Vidal (1921, pl. 2, fig. 4, pl. 4,figs. 4, 5). Ihe paperby Vidal was not noticedoutsideSpain; thespecieswasalso describedasMerklinia perornata(Cottreau,1922),see in Dhondt (1976, pp. 14-18, pl. 2, fig. 2); this speciesis widelydistributed in the Tethys in the Campanian~Maastrichtian:Mada-gascar,North Africa, Spain and Cuba.Occurrencein Spain: Campanian.Maastrichtian:Barcelona: Fal-gars (5cm. E.), Figols-]as-Minas(Sem.R), La Noii, Priorat, Berga(5cm.E.), St.JuliadeCerdanyola,Guardiolade Bergueda(Sem.B3;

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Lérida: Serra-del-Verd, Tuixent (5cm. E.), La Tossa, Salas dePallars(Areny Sandstone)(1. Br.).

Merklinia septemplicata(NILSSON, 1827): eriginally describedfremthe Campanianof Sweden;fig. in Dhondt (1972b, pl. 3, fig. 3 subLyropectenseptemplicatus);widely distributedbut rarely occurringfrom the Santonian to Late Maastrichtian: Sweden, Germany,Belgium-the Netherlands,Austria, Bulgaria.Occurrencein Spain: Campanian: Lérida: Coil d’Ares, MontsecSection (1. Br.), Maastrichtian: Lérida: Serie V Nemes Morts(Univ. Belí. colí. 3. Gallemi).

Mimachlamyscretosa (DEFRANCE, 1822): originally describedfromthe Craie de Meudon (near Paris, France) (Campanian);fig. inDhondt (1973b, pl. 6, fig. 2); widely distributed in temperateseasfrom the Turonian to terminal Maastrichtian.Occurrencein Spain: Santonian: Lérida: Barranede las Cellades(1. Br.), Montsec(1. Br.).

Mimachlamysrobinaldina (D>ORBIGNY, 1847): originally describedfrom the Neocomianof Saint-Dizier (Haute-Mame,France);Hg. inDhondt (1973b, pl. 7, fig. 2); very widely distributed from te Va-langiniante the Late Cenomanian.Occurrencein Spain: Cenomanian: Navarra: Monte Orobe-Alsa-sua (Sem.E.).

NeitheaP) alpina (D’ORBIGNY, 1847): originally describedfrom theUpper Cretaceousof SE France;Hg. in d>ORBIGNY (1847, pl- 446,figs. 4-8); distributed from the Cenomaniante te Campanian(? Maastrichtian)of the Tethys.Occurrencein Spain: Coniacian11-1V: Burgos:Villamartín (1. Br.);Santonian: Lérida: Mentsecd’Ager (Sem.E.).

Neithea(?) dutrugei (COQUAND, 1862): originally describedfrom theCenomanianof Tenoukla (Algeria); fig. in Dhondt (1973a, pl. 4,fig. 3, pl- 5, figs. 4 a, b); occurs from the Cenemanianto teSantonianof the Tethys.Occurrencein Spain: Santonian: Lérida: Tuixent (Mus. 133.

Neitheahispanica(D>ORBTGNY, 1850): originally describedfrom «Lla-ma Oscura prés de Oviedo» (Late Cenomanian); fig. by Roger(1956, fiche 44, PalaeentologiaUniversalis); widely distributed inthe Tethys and in Africa in Late Cenomanianand Turonian de-posits.

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Occurrencein Spain: Asturias: Llama Oscura(Mus. P.), Guadala-jara: Somolinos (Sem. B.).

Neitheaquinquecostata(SOWERBY, 1814): originally describedfromthe Greensandin Chute (England) (Cenomanian);fig. in Dhondt(1973a> pl. 2, figs. 2 a-c), distributed worldwide in Tethys and intemperateseasfrom the Albian to the terminal Maastrichtian.Occurrencein Spain: Cenomanian: Soria: Picofrentes (1. Br.);Upper Cenomanian:Navarra: Ganuza(1. Br.).

Neithearegularis (SCHLOTHEIM, 1813): originally describedfrom theSint PietersbergnearMaastricht(the Netherlands)(Late Maastrich-tian); fig. in Dhondt (1973a, pl. 1, fig. 3, pl. 2, figs. 1 a-d); widelydistributed mainly in temperateseasfrom the latest Cenomanianto the terminal Maastrichtian.Occurrencein Spain: Coniacian: Burgos: Terradillos de Seda-rio (1. Br.), Villamartín (1. Br.>, Lérida: Herbasina (Pons et al.,1981, pp. 60-61, 1. Br.); Santonian: Burgos: Lacazina Limestone:Torme (1. Br.), Lérida: Castello 500 E Sta. Engracia, Tremp (Sem.B.), Clot d’Olsi, Montsec (Univ. Belí.), Collades de Bastus (Univ.Belí.), Montsec(Mus. E.); Campanian~Maastrichtian:Lérida: ArenySandstone:La Tossa(1. Br.).

Neithea sexangularis (D’ORBIGNY, 1847): originally described fromthe Campanian?of Pons(Charente,France);fig. in Dhondt (1973a,pl. 3, fig. 3); occurred iii [he warm temperateseasof the Aquitai-ne (SW France) in Campaniantimes.Occurrencein Spain: Campanian: Lérida: Valí de Lluc.Cornelles(Sem. 13.), Tuixent (Sem. E.); Campanian.Maastrichíian:Lérida:Areny Sandstone:La Tossa (1. Br.).

Neithea sexcostata(WOODWARD, 1983): originally describedfromthe Campanianof Norfolk (England); fig. in Dhondt (1973a, pl. 5,figs.-2a;2 b)andiirWoods(1903,pi. 40, figs. 10-15,pl. 4l,tigs. 1-10);distributed in temperateseasfrom the Late Albian te the terminalMaastrichtian.Occurrencein Spain: Cenomanian:Navarra: Monte Orobe-Alsasua(Sem. 13.).

Neithea ste/anoi (CHOFFAT, 1900): originally described from theHellasianof Bellas (Portugal); fig. in Choffat (1902, pl. 2, figs. 3 a,3 b) (specimenin the Geological Survey of Portugal, Lisbon); spe-cies known from Portugalandpossiblyfrom Texas (USA).Occurrencein Spain: Late Albian: Albacete: La Solanadel Alam-bin, Almanca (Univ. Val.).

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Nejíhea striatocostata(GOLDFUSS, 1833): originally describedfremLate Maastrichtianof Maastricht (the Netherlands);fig. in Dhondt(1973a, pl. 3, figs. 2 a-d); widely distributed in the ?Santonian-Campanian.Maastrichtianof temperateseas.Occurrencein Spain: Santonian: Lérida: Flamicelí-PallaresaSec-tion (Pons el al., 1981, Pp. 23-27, 1. Br.); Campanian: Lérida: Erin-ya (Mus. B.); Maastrichtian: Barcelona:Coil de la Subirana,Falgars(Sem.B.), Figols-las-Minas(Sem.B.).

Nelíhea syriaca (CONRAD, 1852): originally describedfrom the Ap-tian at Abeih in Lebanon; fig. in Woods E 1903, pl. 39, figs. 14-17,sub Pecten(Neithea)nzorriSil;distributedin the Aptian and Albian,mainly from warm temperateand Tethys deposits.Occurrencein Spain: Aptian: Albacete:Almansa(Univ. Val.), Bar-celona: Garraf (Mus. B.), Castellón: Costa de Reinals (Sem. B.),Morelia (Unix’. Val.), Tarragona: Cañellas(Mus. B.), Coya del Vidre,Mas de Barberans(Sem.E.), Marmella (Mus. B., Sem. E.), Puertadel Aliga, Amettla de Mar (Sem. E.).

Nejíhella norabilis (MtYNSTER in GOLDFUSS, 1833): originally des-cribed from the Cenomanianof Essen/Ruhr(GFR); fig. in Dhondt(1973 a, pl. 4, fig. 2); widely distributed from the Mid Albian toterminal Cretaceous.Occurrence in Spain: Cenomanian: Burgos: Valdeporres(1. Br.),Navarra: Monte Orobe-Alsasua(Sem. B.).

Radiopecten cf. quinquenarius(CONRAD, 1853): originally describedfrom the Delawarearid ChesapeakeCanal, Delaware(USA) (LateCampanian);fig. in Wade(1926, pl. 21, figs. 6, 7); this North Ame-rican Campanian-Maastrichtianspecieshas previously not beenrecorded from Europe; it was only known from the Gulf andAtlantic Coastal Plain deposits.Probable occurrencein Spain: Campanian-Maastrichtian:Lérida:Areny Sandstone:La Tossa (1. Br., Unix’. Belí.).

Syncyclonemagreppini (PICTET and RENEVIER, 1858): originallydescribed from the <Mames jaunes de la Perte du Rhóne’ (Am,France,Aptian); fig. in Pictet and Renevier (1858, pl. 19, fig. 4);distributed in Aptian aud Albian Buropean depositsof temperateseas; very close to Syncyclonemainconspicua(Cragin, 1895) fromstrataof similar age in Texas.Occurrencein Spain: Aptian: Barcelona: Garraf (Mus. 13.).

Syncyclonemanilsoni (GOLDFUSS, 1835): originally describedfromthe Maastrichtianof Maastricht (the Netherlands);fig. in Dhondt

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(1971, pl. 4, figs. 1 a, 1 b); widely distributed in temperate,warmtemperateandmarginal Tethysdepositsfrom te ?Cenomaniantethe terminal Maastrichtian.Occurrence.in Spain: Coniacian 1: Burgos: Villamartín (1. Br.).

SFeNDX-LIDAF

Spondylusspinosus (SOWERBY, 1814): originally described fremBrigliten and Northfleet (England) (Lower Senenian); fig. inWoods (1901, pl. 23, figs. 6-11, pl. 24, figs. 1-7); widely distributedin the Turonian-LateCampanianWhite Chalks.Occurrencein Spain: Turonian: Navarra: Ollogeyen (1. Br.); Ce-niaclan: Burgos: Terradillos de Sedane(1. Br.).

LIMIDAB

Lima catalaunica(VIDAL, 1921): originally describedfrem the San-tenian of the Montsech,Lerida; fig. in Vidal (1921, pl. 5, fig. 4);this specieshasse far only beendescribedfrom Catalenia.Other ecclrrencesin Spain: Maastrichtian: Barcelona: La Nou,Priorat, Berga (Sem. 13.), Lérida: Casade las Cots,Gabana(Sem.B.).

Limaria dubisiensis(PICTET and CAMPICHE, 1870): originally des-cribed froni the Valanginian near Sainte Croix, Vaud (Switzer-land); fig. by Pietet and Campiche(1870, pl. 161, figs. 2 and 3);widely distributed in the Neocemianof the Tethys and warmtemperatedepesits.Occurrencein Spain: Cretacicemf.: Valencia: Pont del Garrofer,Oliva (5cm. B.).

Limaria marticensis (MATHERON, 1842): eriginally describedfremMartiguesnearMarseille(France)in depositsof pessiblySantonianage; fig. in Zittel (1866, pl- 16, figs. 1 a-d); distributed in tenerthernTethys from Santoniante Campanian.Occurrencein Spain: Santenian: Lérida: Clot d’Olsi, Mentsec(Unix’. Beil., 1. Br.), Celladesde Bastus(Univ. Belí., 1. Br.), Montsec(Mus. 13.).

Limatula jittoni (sensuD>ORBIGNY, 1847): eriginally describedfromte UpperGreensandof Blackdown(England); fig. in Woods (1904,

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pl. 7, figs. 12-15); distributed in Late Albian and Cenomanian«sandsOccurrencein Spain: Cenemanian:Guadalajara: Somolinos(San-13.), Navarra: Monte Orebe-Alsasua(Sem.B.).

Limatula semisulcata(NILSSON, 1927); originally describedfrom theCampanianof Sweden;fig. in Marquet (1982, pl. 1, fig. 2); distri-buted in Europein temperateand warm temperateseasof Cena-cian te Late Maastrichtianage. Occurrencein Spain: Santonian:Lérida: Colladesde Bastus(Univ. Belí.), Montsec(Mus. 13.), Vila-navade Meja la Llebera(Sem. 13.); Maastrichtian:Lérida: Sensuy(Sem. 134.

Píagiostomacretaceum(WOODS, 1904): eriginally describedfrom theWhite Chalk of SeuthernEngland; Hg. in Woods (1904, pl. 4,figs. 13-15, pl. 5, figs. 1-4); widely distributedmainly in temperatedepositsfrem the Turonian to the Late Maastrichtian.Occurrencein Spain: Santonian: Lérida: Clet d’Olsi, Montsec (1.Br), Maastrichtian: Lérida: Sensuy(Mus. 13.).

Plagiostonzasantonense(D’ORBIGNY, 1847): eriginally describedfromthe <Lower Senonian’at Saintes(France) (Santonian);Hg. in d’Or-bigny (1847, pl- 425, figs. 1, 2); mainly known frem Santeniande-posits in SouthernFrance.Occurrencein Spain: Santonian:Burgos: Villamartín (1. Br.); Lbrida: Colladesde Bastus(Univ. Beil.).

Plagiostomasemiornatum(IYORBIGNY, 1847): originally describedfrom Le Mans (France)(Cenomanian);Hg. in Woods (1904, pl- 3,figs. 14-16, pl. 4, fig. 1); distributed from the Late Albian te theterminal Cenomanianin temperateEuropeandeposits.Occurrencein Spain: Cenomanian:Guadalajara: Somelinos(Sem.13.), Upper Cenomanian:Soria: Picefrentes(1. Br.).

?Pseudolimeacottaldina (D’ORBIGNY, 1847): originally describedfrom the Aptian of Wassy(Haute-Mame,France);fig. in d’Orbigny(1847, pl. 416, figs. 1-5); widely distributed in Late Neocomianand Aptian deposits of temperateand border Tethys EurepeanandWesternAsian seas.Occurrencein Spain: Aptian: Barcelona: Garraf (Mus. 13.), Caste-llón: Anroig-Chert (Sem. 13.), Morelia (Sem. 13<), Tarragona: Coyadel Vidre, Mas de Barberans(Seni. 13.), Marmella (Mus. 13., Sem.13.), Puertadel Aliga (Sem.13.), La Roqueta-Canyelles(Sem.B.).

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OsTRrAca~

Por easier censultation by non-oyster specialists ah the oystergeneraare listed alphabetically,not taking into consideratienwhetherthey belong to the Gryphaeidaeor the Ostreidae.

Aetostreoncouloni (sensu D’ORBIGNY, 1847): according to Pervin.quiére, 1910, Exogyra couloni Defrance, 1821 has beenmisinter-preted by later authors; the Neocomian species,more elongatedand narrow than Aetostreon iatissimum, has generallybeen re-ferred to A. couloni and 1 continue to use the name with thatmeaning, fig. by d>Orbigny (1847, pl. 467, figs. 1-3, non pl. 466);distribution: widely distributed in the warm temperateand Te-thyan Neocomian.Occurrencein Spain: Neocemian: Valencia:Oliva (Sem.134; Hau-terivian: Castellón:Chert (Sem.13.), La Querola,Concentaina(Sem.13.), San Mateo (Sem.13.>.

Aetostreonlatissimum(LAMARCK, 1801): neither in 1801 nor in 1819did Lamarck indicate the locality from where his specimenwascollected; Pervinquiére (1910) suggestedthat it came from theAptian of the Easternside of the Paris Basin (Dept. Haute Mameer Aube); fig. in Pervinquiére(1910, fiche 194, PalaeontologiaUni-versalis, figs. H, Ha, lib, lic) in Stenzel (1971, p. N1117, fig. J 92,1); widely distributed in Aptian warm temperateand Tethyande-posits.Occurrencein Spain: Aptian: Barcelona: C. Almirahí, St. Pére deRibes (Sem. 13.), Castellón: Corachar(Sem.13.), Forcalí (Sem. 13.),Fredes-CostaReinals (Sem. 13.), Olocau del Rey (Sem. 13.), Tarra-gona: Coyadel Vidre, Mas de Barberans(Sem. 13.), Teruel: Cañizardel Olivar (Sem.13.), Castellete(Sem. 13.), Mirambelí (Sem.E.).

Amphidonte obliquatum (PULTENEY, 1813): originally describedfrom the Cenomanianof Melbury (Dorset, England); Hg. in B.M.F.(1975, pl. 55, figs. 4-5); distributed in the Late Aptian to Late Ce-nomaman,generallyknown as Bxogyra conica (Sowerby).Occurrence in Spain: Aptian: Asturias: Nara (Sem. E.), Xixun(Sem. 13.), Cenomanian:Burgos: Sierra de Tejada (Sem. 13.), Gua-dalajara: Somolinos (Sem. 13.), Navarra: Ganuza(I. Br.), Soria:Picofrentes(Sem. 13.).

Ceratostreonflabellatum(GOLDFUSS,1833): originally describedfromSaumur (France)and Boesingfeld (Germany); fig. in Bayle (1878,pl. 133); distributed in Albian-Cenomanian of warm temperateseas.

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Occurrencein Spain: Late Albian: Albacete: Tejar de Bolarin (1.Br.), Cenomanian:Burgos: Tejara (Sem.13.), Valdeporres(1. Br.),Guadalajara: Albendiego (Sem. 13.), Atienza (Sem. 13.), Condemiosde Abaja (Sem. 13.), Somelinos(Sem. 134.

Ceratostreonplicijerum (DUJARDIN, 1837): eriginally describedfremthe <craie tufau’ of Touraine,France (here: Coniacian-Santonian);fig. in Bayle [1878, pl. 134, Hgs. 1, 2, 10, 11, sub Ceratostreonma-theroni (d’Orbigny), in Stenzel(1971, pp. N1118-1119,Hg. 392, 2 a-dsub Ceratostreon spinosum (Mathéren)]; distributed from theTuronian to terminal Maastrichtian in shallow, warm temperateand Tethys deposits.Occurrencein Spáin: Turonian: Soria: Picofrentes(Sem. 13.), Co-niacianII-TV: Burgos: Villamartín (1. Br.), Santonian:Burgos: La-cazinaLimestene: Torme(1. Br.); Villamartín (1. Br.), Lérida: Clotd>Olsi, Mentsec (Unix’. BelL, 1. Br.), Collades de Bastus (Unix’.Belí.), Montsec(Mus. 13.); Maastrichtian: Barcelona: Falgars,Pobladc Lillet (Sem. 13.), Burgos: Terme sectien(1. Br.), Lérida: Cas-tello SantaEngracia,Tremp (Sem.13.), Sensui(Sem.13.), Navarra:Puerto de Olazagutia(1. Br.).

Ceratostreon tuberculiferum (KOCH and DUNKER, 1837): eriginallydescribedfrom Elligser Brink (nearHannover,G. F. R.) (?Aptian);fig. pl. 1, figs. 1-5 arid in Woods (1913, pl. 61, figs. 7-11); distributedfrom the Valanginian to the Aptian (?Albian).Occurrencein Spain: Neocomian: Valencia: Oliva, Pont del Ga-rrefer (Sem. 13.), Aptian: Castellón: near Chert (Sem. 13.), nearForcalí (Sem. 13.), nearFredes(Sem.E.), Mas de Cabra,Todobella(Sem. 13.), nearMorella (Sem. 13.), near Olocaudel Rey (Sem. 13.),Vallibona (Sem. 13.), Zoretadel Maestrazgo(Sem.13.); Cuenca:Ca-sas Nuevas(Sem. B.); Lérida: Enforcaduradel Pedraforca(Sem.13.), Tarragona: Les Clodes, Mas de Barberans(Sem. 13.), nearMarmella (Sem.13.), Tortosa(Sem. 134, Valícañera(Sem. 13.); Te-ruel: Josala Tejeria (1. En, Sem. 13.), Obón, Ermita de SanMiguel(Sem. 13.).

Costagyra olisiponensis(SHARPE, 1850): originally describedfromnear Lisbon (Postugal)(Late Cenomanian-EarlyTuronian); fig. inStenzel(1971, p. N1117, fig. 3 91); distributed in Late Cenomanian-Turonian of the Tethys.Occurrencein Spain: Cenemanian:Burgos: Tejada(Sem.13.), Gua-dalajara: Somolinos (Sem. 13.).

Gryphaeostreacanaliculata (SOWEREY, 1813): originally describedfrom the Upper Greensandof Chute (England) (Cenomanian);

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fig. in Woods (1913, pl. 56, figs. 2-16); very cemmenin certaindeposits frem Aptian te terminal Maastrichtian; is often consí-deredto havelived attachedte mangrovereets.Occurrencein Spain: Aptian: Castellón: in and around Forcall(Sem.13.), Morella (Sem.E.), Tarragona:Puntadel Aliga, Amettladel Mar (Sem. 134.

Ilymatogyrapellicol (VERNEUIL et COLLOMB, 1853): originally des-cribed from «Cuevadel Vidrio prés Tortosa»; fig. pl. 1, figs. 9-12;systematicnote: Coquand(1869, p. 162) consideredOstrea pellicolas a variety of O. boussingaulti d>Orbigny. 1 considerthis mostu.nlikely: the radial ornamentationof O. pellicoi, as statedby deVerneují, is more or less parallel with the longitudinal spiral ofthe valve,whereasin O. boussingaulti(asunderstoodby Coquarid,aCeratostreonspec. close te C. tuberculiferum)a longitudinal foldis intersectedby plicaemore or less at right angleswith the majorfoid. Ilymatogyra pellicoi is se far only known from the Aptianof Spain: Castellón: Mi Roig, Chert (Sem. 13.), in and aroundForcalí (Sem. E.), Morelia (Sem. 13.), Olocau del Rey (Sem. 13.),Todobella(Sem.E.), Zoretadel Maestrazgo(Sem. 13.); Tarragona:Coya del Vidre, Mas de Barberans(Sem. E.), Puentadel Aliga,Amettla de Mar (Sem. 134, Valícañera,La Cenia(Sem.E.); Teruel:Josala Tejeria (Sein.13.).

Ilymatogyra cf. pseudoajricana(CHOFFAT, 1886): originally descri-bed from the Eellasianof Portugal; fig. in Choffat (1886, pl. 4,figs. 1-4); distributedin the Iberian Peninsulaand in North Africa.Occurrencein Spain: Cenomanian:Burgos: Tejada(Sem.13.), Gua-dalajara: Atienza (Sem. 13.), El Atanze (Sem.E.>, Somolinos(Sem.E.); Turonian: Soria: Picofrentes(Sem.134.

IYtGUL,tuJprt~ rttcuzse¿ I~UUVUA1NU, iño¿>: originaIIy clescribecltrom theCampanianof the provinceof Constantine(Algeria); fig. in Stenzel(1971, pp. N1164-1165,fig. J 137); distributed in the Tethys fremthe Turonian to the Maastrichtian.Occurrencein Spain: Campanian:Lérida: Os da Balaguer (Mus.E.).

Pycnodonte biauriculatum (LAMARCK, 1819): originally describedfrom Le Mans, France(Cenomanian);fig. in Bayle (1878, pl. 137,figs. 1-4); distributed in Late Cenomaianstrataof warm temperateand northern Tethys regions.Occurrencein Spain: Burgos: Tejada(Sem.134.

858

Pycnodontevesiculare (LAMARCK, 1806): originally describedfromMeudon nearParis,France(Late Senonian,probably Campanian);fig. in 13. M. F. (1975, pl. 59, figs. 1, 2); widely distributed in tem-perateandTethysseasfrom the Albian tul the terminal Maastrich-tian (Danian?).Occurrencein Spain: Ceniacian: Lérida: Cellada Gassó (Carre-gador de CarreuMarly Limestone,Pons et al., 1981, Pp. 56-59) (1.Br.); Santonian:Barcelona: La Noii (Sem. 13.), Lérida: ColladesdeBastus(1. Br.); Campanian-Maastrichtian:Lérida: Areny Sandsto-ne: La Tossa(1. Br.); Maastrichtian: Barcelona: St. Julia de Cer-danyola (Sem.B.), Lérida: La Rua,Tremp (Sem.13.).

Pycnodontevesiculare.subsp.nildtini (ARKHANGUELSY, 1905): oil-ginally describedfrom the Coniacian-Santonianand from the Ma-astrichtianof the Russianplatform: asmall almost comma-shapedPycnodontevesiculare subspecieswhich at certain timesoccurredin greatnumbers,mainly in the Lower Santonian;Renngartenna-med this subspeciesPycnodontesubvesiculosaRenngarten,1964;his specimensare from the Caucasus.Occurrencein Spain: Santonian:Burgos: Villamartín (1. Br.), Lé-rida: Collado Gasso:Carregaderde Carreu(Pons et al., pp. 56-59,1981) (1. Br.), Maastrichtian: Lérida: Castello 500 m E of SantaEngracia (Sem. 13.).

Pycnodontevesiculosum(SOWERBY, 1822): originally describedfromthe Upper Greensaridof Warminster (England) (Cenomanian);Hg. in Woods (1913, pl- 55, figs. 10-14, pl. 56, fig. 1); known fromthe Late Albian and Cenomanianof temperateseas.Occurrencein Spain: Upper Cenomanian: Navarra: Ganuza (1.Br.).

Rhynchostreonsuborbiculatum(LAMARCK, 1801): originally descri-bedfrom Le Mans (France) (terminalCenomanian);fig. in Stenzel(1971, pp. N 1119-1120, Hg. J 94, 1); widely distributed in LateCenemanian-Turenian(?Coniacian)deposits; generallyknewn asExogyra columba(Lamarck).Occurrencein Spain: Late Cenomanian:Burgos: Tejada(Sem.13.).

DISCUSSION AND CONCLUSIONS

The datadiscussedin this paper give only a very incempleteviewof the SpanishCretaceousbivalves, even of those from the fourgroupsconsidered.Tentatively, a few conchusionscan be indicated:

859

1. Aptian faunasfrem EasternSpain: 9 species,2 of which seemte be endemicte Spain; the other speciesbelong te a typical Tethysassemblage:many speciesare common with EasternFrance, fewerwith Soutliern England: undoubtedsimilarity with the Middle East-WesternAsian Tethys.

2. Cenemanianfaunas from NW Spain:

Species SornoUnos Picofrentes Tejada Navarra

CIII. clongata +CIII. subacata + +Mi. robinaldina +Nei. hispanica +Nei. quinquecostata +Nei. sexcostataNei. notabilis +Lim. fittoni + +Pta. semiornatum + +

Amph. obliquatuni + + + +Cer. /labellatum + +Cost. olisiponensis + +Ilynl. pseudoajricana + + +¡‘ye. biauriculatum +Pvr 1w clrflln vr”i +

Rhy. suborbicutatum +

The faunasfrom Navarra (Monte Orobe andGanuza)only containspecies known from 5. England and the regien around Le Mans(France); the three other regienscontain speciesfrom Tethys faunasas well as from temperateshallow faunas and seemte havebeenatleast partially stratigraphically somewhatyounger.

3. Santonianfaunasfrom Lerida: 14 species,of which 3 are de-finitely Tethyan,9 are common with the faunas of similar age in theCharenteandDordogne,7 alsooccurin the Gosaudepositsof Austria.

4. Campanian-Maastrichtianfaunasof Catalonia: also 14 species,of which ene is endemic, two are definitely Tethyan,ene is NorthAmerican, 9 are common with the faunas from the Charente,7 arealso known from the stratotypical Maastrichtian,4 are known fromthe White Chalk depositsof similar agein Northern Europe.

Hence,the faunasherein considered,mainly from NorthernSpainseem te be closer te these of the berder Tethys region, after Ap-tian times.

860

APPENDIX

List aná abbreviationsof collectionsmentioned

1. Br.: Department of Palaeontology, KBIN, Brussels; MAFI:Magyar Allami Fñldtani Intézet, Budapest;Mus. 13.: Museo geológico,Barcelona(Ciudadela); Mus. P.: Institut de Paléontologie,Muséumnational d’Histoire naturelle, Paris; Sem. R: SeminarioBarcelona,Laboratorio del CSIC.

Unix’. Belí.: Universidad Autónoma Barcelona, Bellaterra, Depar-tamento Paleontología; Unix’. Val.: Universidad Valencia, Paleonto-logía.

Excursionsduring which material was collected

Mid CretaceousEvents, Iberian Field Conference,1977: part IIlead by Y. Wiedmann,part III lead by L. Vilas and his colleagues.

Working Group Coniaciante MaastrichtianStages,Third WorkingSession,Tremp, 1981, lead by Y. M. Ponsandhis colleaguesfor paú 1,and by Y. Wiedmann and M. Lamolda for part II.

ACKNOWLEDGMENTS

1 would like te expressmy sinceregratitudeto themanycolleagueswho gaye guidancein the field and madepossible the collecting ofthematerialon which part of this work is based:especially3. M. Ponsand his colleaguesfrom the University in Bellaterra, L. Vilas and hiscolleaguesfrom the University in Madrid, J. Wiedmann from Tilbin-gen, andM. Lamolda andhis colleaguesfrom Bilbao. Fer permissionte study collections in their care and help in many ways 1 thankL. Vía Boada,5. CalzadaandA. Ros (Barcelona),M. de Renzi (Valen-cia), andAnna Horvath (Budapest).

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863

PLÁTEL

Ños. l-S.——Ceratosrreontuberculi/erurn (Koch and Dunker): Xl, Josala Tejeria,Teruel, Aptian (1. Br).FIG. 6.—Merklinia septemplicata(Nilsson): Xl, Serie y Honxes Morts, Lérida,Maastrichtian (Unix’. BeIl.: colí. J. Callemí).FIG. 7-8.—-Chlamysachates(Coquarid): X 1, Fig. 7: right valve; Fig. 8: leít valve,C< 131063 (Univ. Tiibingen: coil. J. Wiedrnann).Ños. 9-l2.—Ilymatogyrapellicoi (de Verneuil and Collomb): X3, Figs. 9 and 10:Mas de Roc, Forcail,Castellón,Aptian (Sem.B.); Figs. 11 and 12: Mas de Cabra,Todobella, Castellón, Aptian (Senx. B.).

864