:s

60.

61.

62. 63.

64. 65.

66.

67.

68.

69. 70. 71. 72.

73.

' . 74. 71.

(1972) H W ~ I N E N . HANNU: flandnan rcgiond pollcn vvnblagc zmncí in nncm Fdand. 21 p. ............................................................... 5.- (1973) GYIIENIIUG. GOMN: CompvirOn of the Cvtaian divcr teduiique and thc pohgnph ic incrhod. an opcn systcm for mcuuring rhc rcspincory rata in k -c copcpods. 13 p. ....................................................... (1973) Pmcm. P o m s : Ubcr + Biotopvcneilung mldbodcnlebcnder Pscudo- vorpionida (Ararhnoidca) in F i d und bncncich. i i p. ....................... K m h . BJOW: Plcktoccnc Jaguur in No& Amcriu. 23 p.. ........................ (1972) K u ~ h . BJOW: Gcognphicd Variauon in rhc Sizc Li rhc Puma (Fdu concdor

(1973) EIIIKSSON. Amvr W.: Human rwiMing in and m u n d rhc h a n d lslands. 159 p. 21.- (1973) K u m h , B)OW: Transbcringian Rclauonrhips of h t u amo# Linné (Bmwn and Grizzly h). 10 p.. ..................................................... 3.- (1973) Km&. BJOUN: Fouil Glutton (Gvlo guk, (L.)) from Tornnown Cave. South

(1973) SECUSTL~U. S w C.: RNlo of boctom fauna umpling in thc Tvinninnc a m (imcr Bduc). with spcciai rcfcrcnce ot thc soslllcd komdonropomt i thmy. 12 p. (1973) JAUHIAINFN. bu: Agc and dcgm of podzolisauon of wdds on the c d

(1973) Fomm, A": Evolutionvy changa in thc mcrapodiak of f d hom. 18 p. 4,- (1973) HEMYINKI. KAN: Plasma mcmbnna icOlted fmm UnménirC bnin cek. 27 p. 6.- (1973) P m c w . Poms: kitrige zur Kc~tniC dcr spinaenfauna dcr Ostdpcn. 52 p. 15.- (1974) GOLDIN. A N ~ : Thyrouophin in human XIWII. A mcrhodologiul and cliniul

(1974) P m c w . Poms and LONNQWX. BERIT: "he spidcm of somc habitan at he Nato Biologiul Station (u, Finland). 10 p. ...................................

(1974) SAMIKOSKI. JUHANI: CompvirOn of thc Cffcco o€ tcmpcnnuc on conducrion in

3.-

3.- 1.-

h e ) . 8 p . . .......... .............................................. 3.-

Dnon. 8 p .................................................................. 3.-

3.-

pllio of Nonh-Wat Finland. 32 p. ............................................. 7-

rnidy in d i K u a of rhc thymid g h d . 41 p. ...................................... 12.-

1.- 12,- (1974) v. BONSDOUF. CARL-HFNRIK: The suucnuc of snnwd Fomt virus. 13 p.. ........

rhc IympathctK neme ganglion of a hibcrmcing and a non-hibcmacing munmal. 1) p. 5.- 76. (1974) BKUMMU-KORWNKON~O. Mmus: Bunymvcn arbovirus supergroup in Finiand.

12 p.. ........ ........................................ 20.- 77. (1975) SECUSIXAIE. Svm G.: Litcnnrrc on Mnnc biology in thc Baiuc mi publkhcd

¡n thc y r u r 1963-1972. with thc addiuon of some sympos¡um p a p p h t c d in 1973 and 1975. 80 p.. ....................... ................................ 25.-

78. (197>) K ~ W I S . L. G. MAITHIAS: B i d and LMiunohLtodicmKal m d i a of lysozymc in nonnd. gcrmfrcc and lcuicmic no and obscrvationr on thc non-anubactd cfftctroflyrozymconccllrinviuo.41 p. ........... ................... 15.-

W& K u ~ I o w , 195 3. 22 p. ..

dwing thc Fiandrian. 28 p. ...............................

Madcin. 37 p. .......................................... ........... 20.-

79. (1975) BUD. GOw: S d

80. (1971) HICKS. SHULA: Variatio

81. (1976) LOGINOVA. M. M.: Psyllids (ñylloidca, Homoptcn) of

81 1976

Psyiiids (Psylloidea, Homoptera) of the Canary Islands and Madeita M. M. Logioova Zool. INUNU of me Acadcmy of Scicncn. Lningnd. rhc USSR.

I

ISBN 951-653-0>4-0 ISSN 0069-6179

Societas Scientimm Fennica Heisinki - Helsingfon

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Socic~r ScienriuUm Fcnnica ISBN 911-613-054-0

ISSN 0069-6179

i

HUSINGlN WKUSiüüJAPAiNO OY HUSlNKl 1976

AbStnet

LOGINOVA, M.M. (Zool. Institue. Audany of Sckncc. Lcningrd. USSR): PsyUidr CPsyUoida, Homopiem) of the Cpiury isiandr and Mdeha. Commentur. BioL 81.37 pp. (1976). This paper u the fint to contah a suumy and dcsuiption of the psyUid fauna from the nonhern iriandr of MacPronaia. AU the 29 rpecies b t d below ate na>rded from this tenitory for the fmt time.

Authw'r uddmss: DI. M.M. Loginova. 2nd lnrtitute of the Acadany of Sciencu, La@%! 8-16), the USSR.

In this study 1 made use of the collections of Prof. Hakan Lindbcrg, who visited the Canary Islands and Maderia several times (1926-1963). This paper, written in respectifui memory of Prof. Lindberg and dedicated to him, completes my d y s i s of hk mcxt interesting collections of the Psylloidea fauna from north-western Africa, a temtory which up till now has been p d y investigated as rqards this group of insects.

The species dealt with hen inciude psyliids coilected by Dr. Remane from Temrife in 1954. These collections were sent to me for investigation by Prof. W. Wagner (Hamburg), to whom 1 edpress my sincere gratitudc.

Description are given her of two genera and 20 species up tü1 now unknown to the scientific wodd. The holotypes and paratypes are in the keeping of the Zoological Mwum in Helsinki (Finland) and of the Zoological Institute of the Academy of Scienees (Leningrad, the USSR).

The mcasurements are given in millimetres, and ihe abbreviations in the mcasurement list are the same as in the author's paper on the Psyllids of Morocco (LKINOVA, 1972). The geographical names are talten from the original labeh. ' € le collector's name is given when the material was not collected by Rof. Lindberg himself. The Qures are original. The same part of the body and details of its otnictun in members of the tribe Aryfahhi are shown at the same magnifiutionr, so that both the distinguishing morphological characters and the tme &e of the species may be easily grasped and compared.

Aph.IPridac EuphyUura Foerster, 1848

Reprrrntatives of thir genur from thc Oid World show very n m w trophic speculizrtion, b e i resuicted to OIca L and Phü'yreu L. (fam. Olucue), and have a unifom st~cturc The speckr ue dUthyished by the fonn, venition and psttrm of thc fonwíngs and the mile gcnitaiia.

Two rpeciu have becn discoved on the Cuury islands, E.divin0 (Costa), huc oecurring at the wutemmost point of its range, anda ncw rpecier - &.conarie~~& Lo&, describcd in the mision of the tribe Auphyllurini (Loginova, 1973).

Euphyllum divina (Costa) - Gran canaria, Tafiua. 24 d 9, ieg. R. Stort On spccks of O&u. in Europc on O.cumpueu (Loghovi, 1972:4).

Euphylium CIVILV¡UI& Loginova - Tcnerife: Vaik S. 2 111 1950, 3 dd includin a hoiotype. 5 99; A-. Bailadero 700 m. 2 I I I ~ W e s , 28-30 V 1947. 2 69 (on Pmmn iusiraniru~~'Gran Canaria. S v.lar Palmas, 5 111 ¡SSO. I d.

Fom and venation of forcwings like thore in E.o~hina, but even more ¡*e thore in€.oct&iopua Silwstri. 1934; ptcmrtigma without transversal vcins, ükc that of Gacrhiopicu. apcx of dorsal platc in femalc turncd downwuds, genital scgment in male quadrangulu (lateral v~ew). Thus. 8.canarunsU rcprcxntr an intermediate specier bctwetn two morphological g~oups of the palrcuctic specicr of thc gcnus - the grwp Phülyieae, which ir chuicterized by the lack of tnnnersal vcinr on the pterostigma. and the ~ I O U P Olivina, having a longer pterostigna. croswd with transversal vcinr

Agonoscena Enderlein, 19 14 Agonoscena tugionii (Licht.) - Gran Canaria, Tafua. 8 dgcieg. R. Storl). On Pisrucia lenfiscus

(Loginova. 19721).

Srrophingia Enderlein, 19i4 n i e typc specks of thir genus. SericoC (Curtir, 1835). is widerpnad in Europc, extending

nomvards to Central Finland and thc Leningrad redon in the USSR. and living on &#una vulgatis. Tht rccond spccicr of the genus. S.cinemac Hodkinson. 1971, from England. t ivu on Enea cinerea

Two new spccus are dcscribed below. They werc discovcrcd in the mountains of Mideira and Tencrife on arboreous hcathcrs. mostly €rica arbma, which suggests that thc S n u s ir confmd to h u t h c n of the subfamily Elicoidco.

Sfmphin#ia arborca Loginwa, p.nov.; f i 1-1 1. Dcscribed frorn a series of 3 1 spccimenr ftom Mdeira and Tcnerife. nie cmaging inítctr arc green, the predominant coiour of the p e c h e n s from Tenenfe.

Howevsr. Iatei on thc body bccomer yellow. the change beginning with the head and ley and thcn anbracing the thonx and abdomen.

The mijority of the specimens from Madcin u e ycllowish. having no patlun on rhe dorsal pirt of the thorax; thc tarsi, antennal regmentr 8-10, the apicei of the genitrlir in the femak and thc puamuer in thc malc are bmwnish to bmwn Eyes blackish or fulvous-biack. Forrtvingj more or Icír in tcnr ycllow. xidom beuing a faintiy deficd, pak brownish coral-like pattern. mostly in ccll rs, veins light-colourcd.

Body roburt owing o wide. convu, dom*like thorax and short appieued had . The anterior p u t of the scutum of thc meronoNm reprcrnts thc a p u of the u c h fonned by the h u d and thorax (in lateral view). H u d nurower than thorax and preued to it. pmplcuritu g c n c d y invisible. Vertex even, straight poiteriorly. slightly convcx anteriorly. changing gndually into genac. Fweae shdlow. Daucd occlli on lateral mugins of vertcx, at its hind angics, unpakcd ocellus on a very small crimson rhombic frontal sckrite in facial plane of hcad and hardly viribk from above. G e ~ e not swollen. Pronohtm rectangular. its siightiy narrowing sides curled on to pleurd part of thorax. almost hdf as long as pnescutum of mcronotum. Fommgs in both male and female cggshaped, Costa anylarly cuned in cell c; with a tapcring roundcd apu; no costal break; ptcrostigma disiinct, tcminating behind middle of ccll r; cells m i and cu mirlt, more 01 lesr qurl; vcins not strong. Manbrane EO~C.CCOUL with few tranrersai wrinkies, aimost opaque.

Hind wings romewhai shorter than fonwings, mcmbranous, tnnsiuccnt. widest at the roundcd apical m q i n : M and Cu - onc-branched. stem MCu short.

Lcgi short with shon strongly dL ted femon: mxae oí hind legs with a long mcnun t , tibiae basally smmooth. with distinctly düated apiccr having 7 equaliy spaccd saltatorial spinez Fust agmcnt of tusus with two such spincs Fcmak. Geniialia cuncatc. fairly murive. basaily thick (fig. 4); dorsai platc slghtly cancave

rbove. Ventral plate basaliy aimost t w t c as high as dorsal plate. Extcriorvaives wide.

Figx 1-11. Snoph¡@ arbwm Lo&inov& p.n.: 1 - hud. from abovc and below, 2 - antenna, 3 - forcwing of male, 4 - genmltrlir of fanalc. in laterai v h . 5 - gcnitrlir of mak, in l a t d view. 6-7 - paramaes, from inride uid from abovc. 8 - 2nd segment of p e a 9 - htid leg. 10 - apcx of tibia and tusur of hind kg, 11 - e s .

Malr Ami tube with a faintly dkicd C < > I I V ~ posterior mar@; about 1.5 h e r as h@ u parunaes and genital x p e n t Panmercr shple , widut a t bue, ttpering at apu. Penh kiit d o w n w d r at itr wide o p a , ductus short. riluated above pomt w h m a p a bendr dowmvudr

Egg w. narmw, with a shirpcned anterior pole and a thick attaching rtmi at ¡U porwior

Meuurcmcntr of femalc: kngth 1.92-2.50; kngth of fornvings 1.62-2.02. width 0.62-0.80;

0.30-0.37; avcr8gc kngth of dorsal phte 0.50. Mcuunmenta of m a k lcngth 1.67-1.87; length of

length 0.20-0.22; lcngth of antennae 0.30; he¡@ of p m e r e t O. 12

pole crí 11).

width of h u d 0.55-0.65. width of Vertu 0.39-0.42. leiyth 0.22-0.25; l~iyth O f UIkWUC

f o i e w m 1.30-1.50, width 0.55-0.62; width of h a d 0.50-0.55, width of vertu 0.32-0.37.

Madein: SMaddcnn. 4-5 VI1 1957. 1 9 ; Quehadas. 24-26 VI 1957. 2 99. 14-16 v

6 M. M. Locorovr. PIyllids of thc Cvwy LIpD<1I and M d e h

1959. 5 dd including a hobtype, 10 9% Serra de Agua. 20-21 IV 1959, 1 d ton Erica zucinulur); Fontc de Pedra. 23 IV 1959, 2 99. Tenerife: L o s Mercedes, 28-30 V l947,9 99, 1 d ion €rica arborea).

AU the spcimenr from Tenerik are JmaUer thrn t h o r from Madcira Sarborea sp.nov. aispiays a CIOK genetic connection with S.fulbx sp.n., described below; both species strongly rermble E~ricae and E.cinereae.

Smphingmfalkx Loginova. v.nov.; f i s 12-16. Derribed from 13 fcmrles and a male from Madeka Colour at cmergence ir probably grcen. as the abdomen of somc females retainr t h s colour.

Head and tho ru yellow, the colour being more vivid than in S.arboreu; vertcx and dorsum either yellow or orange. with no pattcrn. Membrane of forcwings yeUow.

"he new species differs from Sarborea in the following charactcrs: the vcrtex i s more swollen antcrionly; its tnnsition into the genal cones ir more marked, owing to thc lightercoiour of the gcnae; the frons i s larger and pentagonai. the unpaied ocellus belng invisible from above; ceU m i on the forcwingr ir large, considerabiy larger than cell cu 1.

Thc gedulu of the kmale are longer and more massive, than in Sarborea, the dorsal plate undulate above, thc ventral plate basally as high u the dorsal, with a longcr tapering apcx.

M a k The only male (a holotype) has a damahd genital scgnent, thc T o n of the lower part being obrure. n i c anal tube is wider postcriorly than in E.arborea. and almost straight antcriorly; the paruneres are nurowest a t the b a r . being 1.5 times as wider at the apex, thcy are slightly

.downwards cuned throughout. and widcly roundcd along the apicd margin The sttucture of the apiccs of the puameres ir distinctive in both spccies (vicw from above. f i

16) and thcir posterior outline; the interior sides of the paramereí (posterior view) in Sarborea are pea-shqed with a downwards turned dition. whde in SfalLa thcy are oblonpmii.

Mcuurcmcnts of femalc: length 1.57-1.72; kngth of forewings 1.35-1.40, width 0.50-0.52; width of h e d 0.42-0.47, wMth of veitex 0.27-0.30. length 0.16-0.17; kngth of antennae 0.27; kngth of dorsai piatc 0.4S-O.S2. Measurcments of mak: length 1.50; lcngth of forewings 1.12, width 0.47; width of head 0.42. width of venex 0.26. kngth 0.15; Lcnith of antcnnae 0.27; height of parameres 0.12.

On Erica arbonn Mdeira: Poiso-hieiro, 19 VI 1957. 3 99 (on Caurus); Pico Ruivo, 27-29 Vi 1957, 10 99,

S.fal&x sp.n. ir rclated to Surborea, but it ir mialler, bcing ncarer in rue to S.ericae, from 1 d holotype (on Erica arborea).

which it differs in the rhombic b a r of its forcwings and thc stmcture of the malc gcnitalia

Lisronia Loginova. gen. nov. Body thick; thorax short. bintly archcd above, the highest point of the arch being in the

rutum of the mesonotum (lateral view). h u d m i 4 slightly bcnt downwards nurower than thorax. Uppcr p u t s of h u d and thonx and vcins of forewings covercd with spuse. but very strong. long brown r t i e .

Vertex rtnight porteriorly; Conning a gcntly sloping u c h anteriody. with a süght incirion in the middle. widcly rounded towards the facial pu t . fusing without marked transiüon into faintiy convcx genae. Genre withart düations md cones anteriorly. a mid well-defmed trapuium-lkc frons in facial planc the apex of epicranial suture abow it strongly muked. suture missing in upper planc of vertex. Foveae with the adjoining pprtroCveltex deeply concave. Antennr short. more or ku dutinct, rgments 4, 6, 8 and 9 beuing rhinirii. Antennai foveae small. placcd in facial pkne. Eyer louted in facial pknc, small, slightiy conwx.

Pronotum widc. arch-shapcd. with two pUn of decp dcnts in margins proplcuritcs usually hiddcn undcr thc projccting mcronotum and consisting of 2 sclerites, divided by an oblque - transversal suture; the lower scleritc 2.S times u &rgc as the uppcr (fi. 18). Praesaitum fairly narrow. mcdidly somewhat longer (han pronotum and 213 length of scutum; scutellum luge.

7

F i n 12-16, S e a @ f a k Loginova, sp.n.: 12 - heid. from above and below. 13 - fonwing of kmde, 14 - p n i U of funde, in kteril v w ; 15 - pniWio of mal6 in lrtcril viev. 16 - parameres, from above.

Le@ long and thh, tibmc of hind lees approxhately t w h u long as femora, b d y anooth. abnost undbtcd towrds apat. with 5 d t a t d rpines 1st upncn t of Uur w¡th 2 aich piaU

Fomvbgs oblo-d, 28 times u 10% u their -test widlh, whieh U in centre of whg; venation more 01 ius longiiudinrl Hind whgs only r l i t l y shorter than fornms Cu hu two bnnchu. and hims with M to fom the compound stcm MCu.

Anal tubc of maie sinple. having no proccuu siong the posterior mugin.

Liwonia echklno Lof~inova. $p.nov. i s the type spepaf*s. The foUowing marpholqiul charretera are dlgiostic of the gcnus: a short thkk thonx with a

widc but not ilat prwfutum. the kck of the epicniii.l suture, and long rtrong utae on the upper par( of body and thc rms of the foiewinp.

c h i n c t e n indicaihg that the p u s LirrwrL gcn.n. belonp to the subfmüy A p h d E d b Vondr.. and in which it p t l y rcrmbles the gcncra Str~ph- Enderl. and Aph&Uk& CRWf. are: ihe la& oí gcnd cones. the atruciure of the antennae. the faintly developed menan t s on ihe coxae of the hind l e e thc venation and ~ W c t u r c of ihe forewmgs. and the lrck of procerro on the anal íube of the malc.

LiYonia echidna Loginova, V.nov.: f i 17-30.

ikrr ibcd from 8 males and 3 femalea from CranCanuia. Body light-yellow. fovuc. dents on thc prOnONm and apex of paramena browh Eyes yellow

or brownish diver; antcnnae dirty yellow. rgmcnu 8-10 brown Forcwing mcmbnnous, but fauly dcnr . aimost opaque. faintly yellowish. Hind wings membranous, colourley exccpt for the yellow clawr

Vertcx slightly longer than half its width; paued ocelli on lateral sider of vcrtex. and a seta on cach mar ande. Unpaired ocellus invisible from abovc Svellings behind the eyes widc espccLlly neu ver la . Csnac faintly dhted. clypeur lying in a shdow dcpression behveen than. distinctiy visible in anterior view with 2 long rtae on its anterior m u g a

Proború compressed bctween coxac of fon kgs, itr long terminal put direeted fotwudr Antennac aImost as long as width of head, thc rgmcnu gmdudy thickening from 3 to 8. but rgmcnti 5 and 7 thinner and shorter than thcir neighbourr Tibiae of hind lega with 5 q u d l y spaccd vltalorirl rpincs, brownhh 01 light-colourcd; mcracanu having the shape of a md obtuse Nbercule.

Fonwingr Ion6 rounded. with apex in ceU r s veins rmngly raised. the nodal lme abrnt . M bnnching off at dirtinctly shorter distance from base of wing than R+ Ptcrostigma wide, long; thcn ir a costal bnak. cells m , and cul luga.

Female. Gcnitalia longer than other segmcnu of abdomen, dorsal plate concave above, ventral plate slightly ninower than dorsal plate and half as long (lateral view), iu apex rectüineu, so that the broad cxtenirl vrivcs of the ovipositor are free.

Mde. Anal tube convex anteriody. dightly d b t e d porteriorly. Pumercs somcwhat lowcr, tapering and rounded at apex. termimting in a narrow swelling turned inwudr; the geatcr p u t oí <he interior plate of the parameni d h t e d and httened. above with the outline of an ovai blade. Sccond rgment of penú nlatively short. itr sharpcned apcx turned downwards, ductus long.

Musunments of femak: kngth 1.65-1.72; kngth o í fonwings 1.38-1.39, width 0.45-0.48; width of hcad 0.39. width of wrtex 0.28-0.30. kngth 0.19; length of dorsal platc 0.54-0.57. mclrurcments of mnk: lcngth 1.35-1.51; kngth of fonwings 1.09-1.35, width 0.42-0.45; width of held 0.39-0.43, width of wrtex 0.26-0.28. lcngth 0.16-0.17; kngth of antennu 0.43.

Gmn Canaria, Valle de Fataga. 1 IV 1959.8 dd. a holotypc among thm. 3 99. On Cisrus mmopd~rn . The poor ílying abüity of thcse inrc ts ir widcnt from the mal1 suc of the body. thin tibiae,

faintly dcvcloped mcracants, heavy setac and cqud wingr Thc rnain function of their extremities is walking.

Lirronia echidna sp.n. is morphologically an unusual species; ¡ti genctic connections within the rubfarnily Aphabroidinae are still obscure.

Rhdochlanis Loginova, 1964. Rhodochlanis patvipunctaius LOg-MldCua: So0 Lorcndo. 2 VI 1957,l 9.1 c$ lhku da Cevada.

8 IV 1959. 20 99. 9 dd lhku do Fora. 8 IV 1959.1 d 1 ?;Monte-Terreira da Luta. 9 IV 1959. I 9; P O ~ O Santo, Callieta. 10 IV 1959. I 9.

Hort plant unknown. Besidcs Madeira and South Morocco (Loginova, 1972 : 8-9) collectcd from Alpria (Oran).

Colposcenia Enderiein, i 929 ~oiposcenm viridis ~ g . - Gran Canaria: Gando, 9 111 1963. I 9; ~ o ~ u , 23 II 1949, 2 99;

Maspolomas, 24-26 II 1949. 1 9. Lanzarote : El Golfo. 16 III 1963. 1 9; Haria, 19 111 1949, 3 99. 3 dd. 10-15 111 1963, 2 99. Fuerteventure : Matural. 29 IIl 1%3.2 99. 1 d;Corralejos. 27 111 1963. 1 9; Gran Tarajal. 12-15 111 1949. 2 dd. 30 111 1963.2 ?9,1 d;Chilegua, 4-14 111 1949.3 99.4 dd; Jable, 5-8 111 1949,3 dd.

Prcsumably on TamarUr. The fmds in thc eastcrn p u t of the Canary lslands and in Morocco,

10 M. M. LOCINOVA. Pqiüds of <hc Cuiuy islaodc rnd M+dUn

(Loginwa, 1972: 11)13) have becn supplemented by its discovcry in Portugal: Sintr-a, July 1953. colLHcdopHarrison. 2 dd, 1 9. Thea specimens sent for idcntification by the British Muaum (Londoni. are badly damaged.

Megadicmnia Lodnova, gen.nov.

Hcad as wide as thorax, slightly bcnt downwardr, togcthcr with thorax fonning a wide arch (lateral vicw). Thonx slighily convcx, even; pronotum flat. with broad IamcUae. in middle 2/3 as long Y praerutum, which in its turn i s 2/3 as loni as thc ovil, posteriorly clongatcd scutum. Scutellum of mCIOnONm more 01 les, heut-rhapcd. Propleuritcs of equal sue, dividcd a h o s t diagonally by a suNn. which U directed posteriorly from the anterior ingle of the pronotum; thc small cpkterns triangular. the epimens large. egg.shapcd. with a transvcrsc median constriction. Ver tu Ion& flttencd, with shrllow. dot-lic fwue; togethcr with the anterior lober the ver tu ir h o r t as long Y itr posterior width; it has an ireh-like excision on its posterior margin, andjuts out in front of cach eye in an anguiar procey which i s Ui conIact with a anaU tdedte. situatcd between the base of thc cye and thc mar& of thc antennal fovea Fronr im*i¡le. Unpaircd occilur placcd on v e r t u a t a short distana from its anterior margin. Antcnnae thkk, compuitively Ion& iíorm. the sccond scgment somewhat thinner and shortcr than the fust. Eyer smrll. süghtly COIWCX. bcvclled cuwcd inwuds from posterior mugh to anterior (view from abwe). Thc swellhgs behind the eyes have the fom of vcry nmow lamcllae. Genae faintiy swollen, especidly in front of eyes clypeus hidden in dcprcuion between them and invisible in literal view.

Forewings with anterior and posterior margins paraUcl, dirtinctly bevcUcd towudsCu2 at apex, the rncmbrane dense, unwrinkkd; no costal break; pterostigma devcloped. brancher of M very long. stem of M sliihtly longer than stem of Cu; a U c u ~ smoU, in Iyhg posture. Hind wings somewhat shoner than forewings, membranous. with the costal m@n concave in the m a h part and with a dense welldeveloped clavus; M two-branchcd in fenuk, one-bnnchcd in malc. Cu one-bnnched. fusing with M n e u foot of wing.

Meracants on hind coxae wclidevelopcd, femora 2/3 as long Y tibipc and much thicker. rtrongly düated at apu. Tibue builly smooth. at apex crowncd by 9 cqually spaad saitatorial spines íírst segment of t v s i with 2 su& spinca

Cenilalu of fcmdc massive, thkk. dorsal and ventnl plates of almost the samc Icngth. with apiccr incliied towuds each othcr; exterior valves of wipositor widc.

Anal tubc in mde simple. slightiy dh tcd postcriody; parameres h p k . lamellate. 'Ihc typc spccies of thU genus U Megadicrania tecricepr Loginwa, sp.n.

Megadicrania tectueps Loginova, sp.nov.; figs. 31 -39.

D e r r i k d from a male and femalc from Gran Canaria. Creen at emcrgence, subsequently becoming yellow. Coloration of male green, with pattems on vcrtex, thorax, femon of iegr, upper part of abdomen and lower part of genitai scgment yellow. Antennae dark. dirty ycllow, wiIh rgnen t r 3-8 a1 apex and scgmcnts 9-10 wholly brown. eycs fawn-colourcd, ocelli and two ~ ~ g u h t spots on forc lobes of ver tu red Nmow longitudinal black band posteriorly on each tibu of hind legs sometimes brownish spots on fcmora

Forewingr whitish, opaque, each cell with pat tan of diffusc yeUow spots and mmutc round brown dots the lightcoloured veins striped with short brown dasher, a minute, moIc strongiy coloured d u k brown spot in ceU cu at apex of Cuz.

Cdont ion of funde; general tone light ycllow. pattcrn on body more vivid abwe; genitalia brown at apex; dark band on tibhe of hind lcgs missing. OiherwUe the une as in male.

Vcrtex polygonal. its fon lober furcd bPYlly. h o s t ribshaped along anterior margin, with long thick sctae; transition from vcrtex to genae unmarked. Clypcus faintiy düated, trhnyiar, with depression in middlc of anterior mugin, lying dcep in concave between gcnac. approximately at leve1 of posterior margin of hud . Parapteron sphericd. Forewings flattcned, widc. with angukrly

,/*y-----.- ,+---- .-----.- ~ _....____---.. - I

11

FY. 31-39. Megadicrania tecticepr Loginova, spn.: 31 - h u d and thorañ from above, rhcmatic dnwing, 32 - head and thorax in lateral view, danat ic drawing, 33 - plairiter of prothonx. shcmatic drawing, 34 - fonwing of femole, in lateral vkw, 35 - hind wmg, 36 - gcnitalia of female, in internal view. 37 - genitaih of mde. in lateral vKw. 38 - parameres and genital scgment, in posterior vkw. 39 - parameres from above.

bcnt Corta in CCU c; veins faintly r a i d above m c m b m , rtcm RRr dihtcd. Rs very bng. gently S-rhaped in o p i d hU. pteroitigma terminating bchind middk of -11 r. R dirtinct thmughout itr kngth. Thc brinching off from the root of thc w i y ir disphccd towud, the base i d oaun in thc main third of itr Icngth. stan RRs U approximately quai in icngth to MCu and the root of the wing In all the celli the mcmbrane ir cwered with minutc thkk superticid q k r up to the vcinr Hmd wings m c m b m u s . the cianu alom ir dense; branches of M long here u well u on Ioming.

Position of rhinuia on antcnna imposible to d h r n without miuorcope. The genitah are shown in Figz 36-39. lky are very thick in thc femdc, markedly iniaccd in

the dorsal phte bchind thc Uolitcd and rcgion Anal Nbc of male convcx antcriorly widened portcriody in the rhrpe of a biade. with a

triangular protuberance at the e x ; it u t w h aa high u the genital scpent "he puamenr rcetanguiuly Ismcllatc, thh, with trianpiar projectionr in the lpral third of the poatssior mugin (dorsal visw).

M u u m n c n t of femdc: kngth 3.65; length of forewinga 180. width 1.0; width of h u d 0.72. width of vertcx poatcdody 0.40, anteriorly 0.52, length dong epicnnirl auture 0.35. with lobe; 0.52; kngth of ante- 1.45; length of genitoln 0.87; height a t base 0.57. Mururrmenta of male: kngth 3.0; kngth of forewings 2.32. width 0.85; width of hed 0.66. widths of vertex 0.35 Ud 0.45 rerpcctively: kngthr 0.32 and 0.50; le@ of antcnnae 1.40, height of uul bubc 0.37, of paramuu 0.30. of genital sepnent 0.20, length of genital rcgnent 0.30.

Gran Canaria: Buiduna, 5 IV 1%3.1 d a holotype; Tafua, 1 9 (kg. R. Stod). "he spcekr and genur derribcd above undoubtcdly bcbng to the family Apliobrtdoc nK

unque rt"cture and lpck of any biolo&d in fma t ion make it diffmlt to idcntify itaplcr: in thc family. Howeve~, aich spe& luaimr as the strongiy developcd fon lober pad tlatmrr of the vcrtcx. dense rhombic fonwinga with long M baiichcr and genital stiucture rugcrt a connection bchvccn Megudkmnia pn.n. and Ihe genur E u p h y / h Frst. It ir probabk that thc new gen= ir endcmic to the CÍnary IrLnds.

PSyW8C

Diapbrina LOw, 1879.

Fuerteventure, Martural. 23 111 1%3.1 d. 1 9 . Islas Cpnvis. L o b q 28 111 1963.1 d. Dbphwh ContuUrO LO&- L a m o t e : Oir~h. 19 111 1963. 1 d; El G ~ O , 16 111 1963,25 d9.

Hort plant unknown ücrribed from Morocm(Loginaa. 1972; 15-16).

Atytaina Frst. 1848. Representativer of thu genus are disiributcd di over the world in regions with I subtropical

ciimrte mortly occunuig on l N b s of the Leguminosac. More than 30 pecics havc becn reported (HeilopHaf~iron, 1960), but in my opinion thc genur is hctcrogencous and in urgent nced of m i r i o n

The main chuactcrr diagnostic of the p n u s Arytaina: i/ exterior wltatorial vine on furt ugment of tusus of hind lcgs. 21 vey wide hcd (wider than thorax). with sphcriul eycs. 3/ gmduil tranaition fmm vcrtex to gcnd cones, which are not contipous u thcii baacs n c u

4/ Iack of pterortigma on the forewinga (in somc cases it is Ndimentuy) and of costal break, 51 rtrong thorn on base of hind tibiac. %o ncw rpecics and onc subspcaes fmm thc Cuiory Ishndr are dercribed bebw.

the unpaircd occllus,

Arytaina mtbwqa Loginwa. v.nov.; fuk 40-50.

ücrribcd from 136 spccimcnr from Tencrife. Body yellow, coloui varying in intensity betwcen spccimens. somc onnge or rcddirh, the h u d

COMMENTATlONE3 BKuw;icrre 81 13

uaiaily bcing pdc. D o d pattcrn Wre that in A.gen&fue Latr.. dark brown; thonx klow, feanon of I q , and abdominal rletiter h n h g ths m e qota, whwi Liaoue with qo. F b t Uuw .II- ugmcnta yellow browr the colair dulcning progedvely bi the fdowin# aepwntr Eyes crimrobfukous, occlli redduh. Facial and cervical p u t of hud , ki m e cuca incLding the clypeus blck-brown

Figi 40-50. Aryiaina nubffqp7 Loginwa, va.: 40 - hcad, from above and bclow, 41 - antenna. 42 - clypeus, 43 - forewing of mdc, 44 - geniWu of female, in iatcnl vbw, 45 - genitalia of mal+ in lateral VKW. 46-47 - puamercs, from inride d in antctio~.intcrior vkw. 48 - apcx of penis. 49 - hind leg. 50 - egg.

Forewing colwrleu. scldom faintly whitish, having yellow patcher in centre of u c h ceU exccpt ccil c íFiS 43); thcrc prtchu often diminishcd to smrli "tonycr", from which r k a series 01 minute biown marginal rpincr Veinr dark brown

The cdont ion of thc body, $Le and rmcturc u e very Iike thoít ofA.gcnistue, but the h u d is much widei than thc t h o r u and slopa downwuds, thc facial piane u widcr and thc Iiom U almost mund. The trirnplu genal coner are a p W y h t t cncd Ud tum downwuds. whereu in. A.gcniutue thcy are parakl to the wrtcx. l l ~ ~ wrtex has dccpcr foverc. its posterior angles are faintly d h t e d and incline downwards, projccthg in thc form of rounded loks, u c h of which has an occllur in ita centre. The unpWed occüuí U invisible from above, since the fore loks of the vertcx are contigous; the anterior m a r a of the vcrtcx ir gcntly archcd, but not trianpiar like that of A.genistuc.

Forewings narrow, with anterior and posterior margins more 01 les paralle4 widcly rounded apkaily. 3 times u Ion8 as their width; pterortigmr Iacking. but the region whcre R fusines with the costal m a r a dightly dh ted . SupcrfKkl spher in cell NZ around the anal íuture, marginal q~inu foming n w o w üner in c c h m 1. m2 and cu 1.

Ccnicrlir of femile as in A.gcnistue Ceniiilii of mok differcnt in ccttain detlür: anal tube eithcr higher 01 as high as genital

rgmcnt a t ita bar ; its protruding apcx having the fom of a wide Mg. The gen id tegmcnt ir k g e r than in A.genirtue. and extended longitudinrliy. The parameres have a different outlinc, with a l~tudinrl weliiig interiorly, ao thit they are contipous at two points on the a p a , one below the other (iir 45-47). The penu rernnbler that ofA.genistue.

M u u m n e n t r of femile: length 3.5-4.0; length of foming 2.9-3.25. width 1.0-1.2; width of head 1.05-1.12. width of vutcx 0.6-0.67, length 0.25-0.3, len@h of g c d wncs 0.15-0.2; kngth of antcnnac 1.95-2.22; kngth of geniwk 0.75-0.85. biurunmcntr of nuk: length

vutcx 0.55-0.62, length of pn.1 cones 0.15-0.2; length of antennac 1.97-2.22; average heyht of anal tubc 0.37, genítrl t e v e n t 0.35-0.36. h e w t of panmms 0.20.

T e n d e : Pico di Teyde. 36 d? (kg. R. Sto&, 53 d?, Muding 8 holotype d (kg. R. Fny); Teyde, Canada 20 V 1947.9 d ? (oa Spurtffyliarr nubbnur and C y h s ) ; Las Cañadas. 14-15 11 1940, 18 d? (on Spmocyfinrr nubbenur); Las CaWas 2300m. 16-17 IV 1950, 8 dv ; Canadas 2300-2500 m. 28-30 III 19S4, 5 dd 0%. Dr. R Remane); Lu C l i u n y 3 IV 1959,7 d9.

Both the charactus similar to tharc of Agenistur and tharc diffcrentlting Anubhugu are cmphisizcd in thc deiuiption. The new pccicr w u collected on SpartffytinrZ nubigcnus and Cyfiorrrp., which are probrbly its host plan&

3.3-3.62; length O f fünwiryr 2.72-3.0, ~ id th 0.95-1.12; width O f h u d 0.97-1.1, width of

A l y a h drviu Loginovp. sp.nov.; fiis 51-63.

D c r r i k d from 87 pecimcns from Tcnerile and Fuerteventura. Young spccimcnr geen 01 iight yellow. pattern on herd and thonx yellow to o n n p . Md

cpicrankl íuhlre, and facid and ccn iu i puta of h u d brown. Cdour darkeniq with igc; pattern on upper p u t of body becoming brown. and Iowu part of thonx. tibue of kgr and abdominal sckriter becoming covercd with bmwn spots, so that imaUy only n m w s t n p bctwecn pattern on h u d and thorax, gend coner and manbrane betwccn abdominal sckriter remrin unpotted.

Figs 51-63. Alytuinu d d a Loginova. rp.n.: 51 - hcod, from abovc m d below. 52 - antenna, 53 - clypeur and proboscis, in Ia tud vicw. 54 - proboscis. from below. 55 - fmwing of femrle, 56 - genitalia of female, in lateral view. 57 - geniWia of male. in laten1 vicw. 58-59 - puuncres, from inside ind behind. 60-61 - paruncres, from behind and above, 62 - apcx of penis, 63 - hind leg.

Figs 64-66. Arytuhu devio insuhrir Loginon. subsp.n.: 64 - genitrlir of mak, in lateral VKw, 65 - gcnitalir. in posterior vuw, 66 - paruneres, írom ibove.

1s

16 M. M. Locarovrr. Psyllids ofrhc Cauq kLiidt d Midelln

Forewings coiourieu or palc yeiiow, fairiy oftcn with yeliow patches in centre of ccils, Umilu to thor in A.nubh@ sp.n.. but s n l e r and paler. Vcins brownuh with dUtinct dark brown spotr at apicer, and on marginal vcin under a series of mmute marghd spkr Antcnnac. cyes and ocelli coioured iikc thora of A.nubhaga

A.&& combina the morphologiul charactsls of A.grn&&e and A.nubivugu The hcad U Wre that of A.genisiue, the vcrtcx triangular antenorly, the fiQd p h e betwocn the vcrtex and short, almost contiguous. genal canes U narrow; thc unpaind oeellur U visible from abovc However, thc heid ir not wider than the thorax, which resanbks that of A.nublvqpr. but U mon convcx above; the rutum of the meronotum ir iongcr in rclation to the rizc of the pronotum and pracrrutum.

Forcwings diitincily widened towardr the apkal third (of the whoie Icngth) with avwy wideiy rwndcd apial mugin, which u not typicai of the rcprcrantativer of the gcnus; Ndimenlaxy ptcrortigma, fonncd by the diiaüon of C and R, which is not alwayr strongly mukcd, cell cu1 high; no minuto aiperíiiiai sphes.

Gcnitaiu of femalc as in A.nubhagu, but narrower at apex. Mak. A d tube Wre that in prccediag specks, genital rgment shorter, llmost sphericll. iowcr

than anal tube. A p W blnde of pcnis, tumed downwardr, shorter and widcr than inl.gen&tue and A.nubivuga. Thc parameres compliuted, hteraily ruembiii those of A.nubivau and p o r t e r i y thor of Ay//un&rita (Zctt.), Uncc thc posterior margins of the pacamere lunellre uedüated and fonn widc bbes, denrely cowrcd with small r t a c (fig. 58-59).

Murumnentr of female: kngIh 3.1-3.5; lcngth of forewingr 2.6-295, width 1.12-1.17; width of hcrd 0.77-0.85. width of vertex 0.45-0.47, kngth 0.2-0.22, kngth of pnal conw 0.07-0.1; icngth of antcnnae 1.42-1.75. Muinments of male: kqth 2.8-3.1; kngth of foccwings 2.35-2.55. width 1.0-1.05; width of h u d 0.75-0.76, width of vertex 0.45, kngth 0.2, Icngth of genal cones 0.07; lcngth of antenm 1.55-1.52.

Tenerife: Valle de Santiago. 12-13 V 1947.13 do; Aqua Mansa, 15 V 1947.2 99, l d and 8 II 1949. 1 d; Teydc. Canadas. 20 V 1947.41 d9, a hoiotype d among them; Bermejo 1400 m, 21 V 1947. 1 ¿ and I I II 1949.22 d9; Las Cyíadas 14-15 II 1949.2dd; V M o r 1500 m, 27 11 1950, 3 99, 1 ¿ (on Cytisuspmiiferus. AdenowpusvimsusJ. Fuerteventure, LaOüvo. 16 111 1949,19.

A.devm v.nov. differs from ail thc known representatives of thc gcnus in the IIUTOW hcpd, the forewings, widcning towards the apex and hiving a N d h e n W ptcrostigma, and the d o d y more convex thorax. Ther features are ahared with the rcprcrantatives of the pnur ArytlinillP and tcstify to a genetic rciation ktwecn thue two genera. A.dcvfo sp.n. re~scmbiesA.nubivugu and has ken collected from the m e stations in a numkr of placer

AryiaiM devia insularir Loginova. subsp.nov.: ti@. 64-66.

i h e rue. sttuctum and coloration of the body are iike thosc of A.ddu sp.n. i h e structure of the panmeres u dutinciive; the posterior marginr of the paramcre plates are only dighliy widcned at thc b w , whüe in A.devm thcy have iugc rectangular almost cintiguous bhdes (cf. figr 60 and 65).

The rpecimens on whkh thU aibrpecies was separatcd were found on the W s of Paima and Gran CuurL, where the nominal species was not discovered.

La Palma, El Paso, 6 VIII. 2 dd (kg. R. Firy). 26 V 1947, 1 d. Gran Canaria: Valle de Tejcda, 7 VI 1947.1 d; Santa Brigida, 21 11 i949,l 9.

Avtainil& Ramirex-Comez, 1972.

For the nomencbtbc explanation regarding the denomhation. compodtion and diagnosis of the genus, LCC my paper: Loginwa 1972: 17-18. Thc materia¡ collccted by Prof. Lindberg in thc Canary Idandr has inuoduccd 10 ncw mcmbers, so that the genus now comprks 18 specic$.

lk rpcckr from tbe Cuiuies iad the two rcpnantativer fmm bíomcco ampose a d o m i group of pnstially nhtd rpccia, c m r i z e d by a seria of common m o r p h o ~ features: short, b l u t genal cona mt widely ap8rt; wide. pcatago~i, b u s fmnt, unifonn atNctum of nuk gcnitdia Howcvei. them 12 lpecks us clurly divided into two morphdq¡d w p i . mosW by i diffennce in the structPn of the funde pnitrlb

m f i t group amtuna four exclurivcly insular rpccWr: A.pboreidu. A.n&whm& LdivLlcnr and A.rnudim spp.a The fanale has iong, murive the body b i U s and the proborb considerably b q e r üua usual

nic sccond p u p mntillu six rpecier f r a the CanaricrA.&luio. A.quilonc, A.p¡k&&, Aprognatu, A.iruirbu and A . u m h t u spp.n.. and two morown rpccks, A.hakun1 b& uid A.colF.r. L w The chancten &tinyiJiLig thus spcckr M: rel.tivdy short pdWL in the f sm, le. with more 01 iur rtmigly deveioped bldbiikc projcciionr i t the interior margbr of the venW platc. theu ai i l l cr size iad a proboscia of more uaud lengih.

i h e remlinmg six representativa d the genus have no speciai peailiuitier terti@& so súoogly to thcir morphdo&ai unifomiity; their origin aiso differs f r a that of the abova<nantioacd BrWPS

Sincc I am dcscribicy a largc serica of ncw rpefks ofArytuhüb, I should point out th.t beforc I mived Prof. Lindbag's coktions in 196s. thcy wcre for many ymrs in thc k p h g of Dr. G. HedopHurkoa in E~gland. Howevu, the coktions nmrined unaortcd uid WCIC not provided with any nota bcyond a fcw hand-wdtten IaWs with gencric names.

Among thc spccicr described herc. úbeh inrn'bcd "Lindber@" occuned twia. Thb genetic dcnomination was been acapted in science as LMbetgia nomrudum (HerlopHarrison, 1949). Howevcr, the author bimrcll chsnged it to Lindbergicfb in 1961, sina at thb time t b forma denomination happcncd to be pnoccupied (Lindbetgia Rbdel. 1958).

Dr.HetbpHurUon gavc the d-nosis of the p u s Lindbergidb in hU work on the South Afriua Prylüdr (1961). having c h e d that thc type rpecitr of the pnur would be dcrciibed from the coktions of P r d L d b q from the Canuiu. No doubt he ¡atended to describe one of thc *¡es ofAry&iní//8 dncribcd in thb paper.

At the u m e time, whcn crccting the p u s Lhdbe@e& ihe aminent sckntlt dercribcd tbs subgcnus h k o b d b q í e & He&-Hui. for the South Afrian spe& He pointcd out t h t itr type

comspondmg p n u k charictcrirtics, apart from having two ( i t d of one) dtatorkl sphieron thc tarsus of thc hind lcgs (l%l: 510).

Owhg to the untimely duth of Dr. HerlOpHPnion. the tvpe rpceies of L W b e W b w u not dcsaikd. and its only representative rcmains L (POuraUndbe~db) pnmWr0 Hcrl.-H*n. whkh has become the typc species of the gcnur, as rlnrdy mcntioncd in scicnMi p a p a (CrpeM. 1970).

Judging by its dcsdption (the rpeciu ir dcrcribed f r a i femile from Cipe Penhmh), L (P.) primiriro has charactcrs of the genen AcItsb HerL-Hm. andpSya0 Ceoffr. and its propu pLes u most probably in one of them. However, thh probiem cannot be solved withcut i thonwgh study and rcinvwtiption of the type specimen; mcanwhüe the exirtuice of the monotypk (snur Lindbe@ib must be acknowkdgc. aithoug it U not ckarly distinguishabk from the two othcr genera.

i shouid Iike to commcncc the ducription of thU youp of pccks from the Cuvry Irluid8 by pouiting out the churters they havc in common (rathu than those dtgiostic of thc separate rpecrí Ud by emphrgzing the distinctive StNCtUE of these hects.

Thc body ir grcen. but becomes yeiiow with age; the pattcrn on the back, though remlinmg iight-coioured. Lc not always dktinctly markcd. and doer not atúact attention by its wtüanely vivid yellow or orange coiour.

i h e head is as widc as the thorax, or somcwhat wider. and short; thc vertex consklidurbiy shorter than hall its width (fonc third of its width), thc vuw ir divides from thc genal ron- by a hqh step. owing to which the facial piane of the hud U evcn.

Thc genai cones are short, with blunt apicer; se( wide aput. so that the wide pen ,yonil frontal sclerite i s alwiyr w d defined. nie gend cones and vertex are almost completely, or at krst

SpCCieS, L (RJ @Iitba, iS unlü<e thC homonymous p0up Of lpecks fi- the cliur*i. bUt hl l

antcriorly mered with rtae. The proboscU is often vcry long. projecting fu beyond the anterior mar& of thc head (latual view), however in the mjority of the rpecks it U of more unid length.

The elongaiion of the proboscu should appuently be considered an adaptive phenomenon, depending on thc depth of the vcucls from whkh the insccts are sucking and on the ande at which the hud of the insect U inclincd when it U stabbing the tissue of the plant.

Representativcs of othcr genera of the tribe Arytohini show a similar kngthening of the proboreir Amblyrhina cognoto LOw has a proboris as long as that of Aryfohilb proborcideo sp.n.. whüc in a rclatcd speckr Ambiyrhino mculnfo Low, the probosck U ffen m e w h a t shorter than usual. Thb suwstr thit the lenglh of the probosck has not thc tuonomic vrhic attribtcd to it by HerlopHumoh

Thonx mioothiy convcx. pronotum approximately twke as wide as length at sides, postcriody aipcrimpord over the pracmtum of thc mcronotum. whkh is relatively longer in thc species of thU g a i p than in the other representatives of the genus, praexuNm * twke as iong as pronotum. whkh U bent forvuds and downwards

Forewhp manbranais, widest in thc apkal third of the whole Iength, wiih a widcly rounded apicpl mar&. pteroítigma WCU dcvdoped. denser than wing membrane. bordered intcriorly by vividly muked R.

in the furt four rpecics, thc females have long massive genital¡; in the remiining six new spccecu>. thcy havc short. more or lus conaükc genitalia.

Arytainillo proborcideo Loginova. sp.nov.: fyrr 67-77.

Dcrribcd from 59 spccimcns from the utpnds Tenerife. Comen, Gran Canaria and Fuerteventura.

In thc oldest spccimens, besides the general yebwing, thc amcd Ud facial puts of the head and the anterior put of the pronotum have become bmwn-blck; thc abdominal steinites ue oftcn bmwnirh. Anlcnnac duty ycibw, scgmcnts 4-6 bmwn at apcx, rcemsnu 6-10 whoUY bmwn.

Forewinga hy.l& with nmow yeUm prtches in SUS of apical half of wing and around anal aiturc; the pitcha may diminirh to mmute lime& from which u*e a scr¡cs oímarginal sphiu; veinr d u k brown, bordered with shart lightsolaircd dashcs Superficial spines dark, occupymg only central part of ceilr. k k i n g in c e U c.

Femalc. Ccnitaiia mrírne, long, strktly ainute. M i k . Anal tube ahost undüatrd postcriody. slightly h u c r than paramms, and genital

-ent. whkh h widely roundcd Wow. Paramenr with a dktuict bi.de along anterior margin. whkh U coverrd with long dense rctu. N m w triangular apices of paramerca touching only at theu posterior andes (view from above. fig. 75).

Meuurcments of lemale: kngth 2.87-3.60; kngth of forcwings 2.37-3.07, width 1.0-1.25; width of hed 0.75-0.9. width of vcrtex 0.42457, kngth 0.15-0.18, length of genal ames 0.04-0.06; kngth of antennac 1.5-1.7; kngth of dorsal p l te 0.92-1.37. Mcasunmcntr of mik: kngth 2.8-3.15; bngth of fowings 2.5-2.62. width 0.95-1.07; width of head 0.77-0.85, width of wrtex 0.45-052. length 0.15-0.17. kngth of g e d cones 0.05; icngth of antennac

Tenuife: Pico di Teyde, 2 99 O-. R. Ficy); Teyde. Canadas, 20 V 1947. 20 d9 (on Adenocorpur virconrs); Las Cañadas, 14-15 II 1949. 19 d?; Las Cuudas 2300 m. 16-17 iV 1950, 1 d. 2300-2500 m. 28-30 III 1954, 12 d9. inchiding a holotypc d Ocg. R. Remane); Viflor 1500 m. 27 II 1950, 1 9. Comen. Hormiya, 1 d O-. R. Frey). Cnn Canaria, Moya, 1 ? (kg. R. Stod). Fuerteventura, 10 111 1949. 1 d.

A series of 5 fanaics from Tcmrife (Agua M a n a Porao, 2 V 1947; lns Canarir. 3 V 1947) from the colkction of the R i j h u r u m van Nituurlgkc Historie in Leyden (Hohd) , which w u scnt to the author for identirication. is also designated as paratypiul

Arytohillo proborcideo sp.n. rescmbla A.n&oIinmla sp.n., dcscnbcd below. but diffcrs in its forcwings whkh are more widcly roundcd at the apex, lack supcrficll spincs in ccll c and have

1.52-1.62; hejht of u l ~ l tube 0.32-0.37, hejht O f P M ~ C E S 0.30-0.37.

F&s 67-77. Arylohüla proboscideo Loginova, q.n: 67 - hud, from above and baow;68 - i lyi lum of antenna. 69 - clypeus and probos& 70 - probos&, from below, 71 - fomwing of female. 72 - gcnitaüa of female, m lateral vinv. 73 - geniWY of malee, in htcralvicw; 74-75 - parameres, from inside and above; 76 - apcx of p e a 77 - hind kg.

strong duk veins with bght-colaired dasher; the form of the puuneres U also diffeicnt in thae

Host plant unknown; it may be Adenocorprr v k c u w from a tcrier of insccts ms collected two rpecrr

Arytoinilia nipalineata Loginova, sp.nov.; TÍr 78-00.

Dcscribed fmm 46 spccimens from the idandr Comna, Gran CnauL and Tenerife. Brown pigncntation not uually apperuiiy when the body becomes yellow. Fomwing

Difiers from A.proboscidea in thc very smiU downwards t u d pnal conas and in having 2-3 tnnducent 01 whitish. ycllow patchu in thc aUs dimmer and shorter than in Aproboocidcn.

pairs of long dark setae on thc facial planc of thc hud.

,

20

! r , E -

W Fig% 78-88. Aryrainilla n&aiineota Loginova. sp.n.: 78 - head. from above and below, 79 -

antenna. 80 - clypeur and probork 81 - forewing of femalc. 82 - genitalia of fanalc. in lateral view. 83 - genital¡¡ of malc, in lateral view, 84-85 - parameres, from inside rnd abovc. 86 - apex of penis, 87 - hind Ieg, 88 - e a .

The forewing has rtrong ribshaped dilatcd light-coloured veins, dcnsely bordered with dark brown dashes, except for thc costal mugin; the minute superfill spincs are Iike thore of A.proborcidro. but are also prcsent in ccll c. forming a mal1 spot at thc base of the pterostigma.

í h c gcnitalu of the femalc ue alike in the hvo spccies. but A.n&alinmra has a small blade-iike düation on thc interior margin of the ventral platc.

Male. Anal tube Iikc that of A.pmboscidea, approximatcly o í same height as parameres and

21

1

genital rcgncnt Paramuer. withait biadoüke projsctions anteriorly, nurowing gradual& towardr apcx. whue they fonn wide triui<uiU projcctions, himed inwardr and contipious at thni interior marginr ( v h from above. fw 85L

Mwurcmentr of fande: length 3.0-3.17; leylth of forewingr 2.45-3.05. width 1.27-1.5;

0.02-0.05; 1 ~ 8 t h of rntcnnac 1.52-1.78; hgth of d o d p t t c 0.9-1.1. Maaurcmentr of mak: Iength 2.8-3.35; kngth of fomings 2.37-2.8, width 0.95-1.12; width of hcad 0.77-0.85, width

heiat of anal eibe 0.35-0.42. heigth of parameres 0.37-0.42 Tenerife: Los Mercedei 1 9 (kg. R Frey), 28-30 V 1947, 1 9.2 IV 1959.4 dd, 3 99; LÍ

Esperanza, 31 IV 1947,l d. 2 99 (onAdcnocarprvircocur) ; m p n lcod, 9 V 1947.1 d, &den de Cuhar. 3 IV 1959. 1 9. Gran Canaria: Cnu de Tejeda 1450 m, 11-13 111 1950,Z dd. 6 99 (on Adcnwprpul; Los Lagunetu. 1 IV 1949. 10 dd. 10 99. includiq a holotype d (on C y h a ) . La Gomua: Vok de Ir Rosa, 19 111 1950.1 d, 2 99; Vdtc Humigua, p u s super, 18 111

charactcdtics rlnady mentioned It rpparcntly livcr in Adcnoempta rnd Cytisur

Aryiainillo dwidcnr Loginova, v.nov.; fgs. 89-97.

width Of h u d 0.82-0.90, width of VU~CX 0.5-0.55, kqth 0.15-0.17, kqth Of *MI COIICS

Of VCRCX 0.47-0.51, lwyth 0.15, kiyh of CIUI CO~CI 0.02-0.05; length O f a t a n = 1.45-1.9;

1950,ld AryrmnjUa n ~ i n c a t a sp.r diffen from A.probosci&a sp.n. in itr lnger body, beaidcs ths

Derribcd f r m 73 ~cc imens frm the Ulondr Palma, Tenmfe and Gran Cuurt No brown colontion developr during thc yellowhg of the inscctg prttun on back u u d y

indistinct; no yellow patchu in ab of forewingr Antennae of oidu insccts b r m from ugmmt 3 onwardr

AdivYlrnr sp.n morphologiully runnbks the two pr&i specicr, but its pioboscb u not elonyted. F o m i n g distinctly widening lo the ap"pI third of the wholc wing kngth WMely rounded at the apicrl mar& and cuning inwards 10 Cu2; v e h thh, liit-colaircd or bmwa, unbordered; cell cut highcr and largcr than in prcecding ípccier Supcrficll s p h s Wrs &ore b A.nigdinc#ra sp.n. but thiu: spot 0eaipir.s more than half of c d c

Genitalh of fanalc l ik thw of A.proborcidco sp.r MIlc AMI lube straightu and wider. much lower than thc iUgc parameres. Genital rtmcnt

considerably longu than itr height but not spherroL Parammr tapering gr8dualJy apkad, bcat inside. anteriorly with d@Uy piotnidmg but dirtinct tri8nguiU biade. their posterior m e bavlly düatcd (posterior view). Apex of penirvery dirtinctivc (fw 95-%).

Mcasurementr offemak:kngth 3.15-4.12;kngthoffonwings255-3.5,width 1.1-1.37;width of h u d 0.85-0.97, width of vertai 0.5-0.57. length 0.17-0.22. length of gcnai cones 0.07-0.12; length of UltCMOe 1.5-2.15; length of dorsal phte 1.0-1.25 Measuranentr of mrle: lengm 3.12-3.5; length of forewingr 2.5-29. width 1.0-1.13; width ofhcad 0.83-0.9, width ofvatcx 0.5-0.54, kngth 0.2-0.22. length of gcnrl coner 0.07-0.1; length of antennae 1.7-1.95; aeiitit

La Palma, Caldera, 25 V 1947, 1 d. Tenenfe: Teyde, Canadas. 20 V 1947,ll dd, inchdmg a holotype. 11 9 9 ( o n CyriW p/ifccnu); ViUe de Santiago. 12-13 V 1947, 1 d.3 99.20-21 II 1950. 1 Y ; Aqur Mansa, 15 V 1947, 2 99; Benmejo 1400 m, 21 V 1947.4 dd, 2 99 (m Gmtninc). 11 II 1949, 2 99; Portillo 2000 m. 17 IV 1950.1 d; ViUe de Manso, pan super 1000 m. 21 II 1950. 1 9. ion Cytiarrpmfifms);Cvudrr 2300-2500 m. 23-30 III 1954,lg &. h. Remane). Gnn Canaria: Atalaya. 25 VI. 3 dd, 5 99 (kg. R. Frey); Cruz de Tejcda 1450 m,

2 dd. 9 99; Las Lagunetas. 1 IV 1949. 5 dd. 4 93; Santa Brigida, 21 11 1949, 1 9; Maspalom~<. 1 IV 1959.1 9; Bandomr 500 m. 5 111 1950.1 d. 1 9.

A series of 6 femaies and 4 males cokcted on Tenerife (Aqua Mansa, Portülo, 2 V 1947. on @tisus and Eriu arborca. I q r Dr. C.O. van Rqtucn Alten). from thc collcction of thc Rijlumuseum van Nahiurlijke Historie in Leyden, HoUand. ir designated as paratypiul

The hort plants of thú specws Lc most probably C y h r pm/ifCN& from whtch it has romethu been recorded.

of uul nibe 0.4-0.45. hei&t O f p ~ n m e n r 0.55-0.57.

11-13 III 1950.1 d, 1 9.1 IV 1947.1 9.1 IV 1950,l ¿, 1 9; Vdle de Tejed& 28-29 III 1949.

22

Figx 89-97. Arytainilh dividenr Loginovq sp.n.: 89 - hcad, from above uid below. 90 - forewing of mde. 91 - hind wing of male, 92 - genitalia of femaic. in lateral VKW. 93 - genitalia of male, in lateral VKW; 94 - paruneres from inride. 95-96 - apex of penk 97 - parmeres, from abovc.

I

c

.b

CQWIENTAllONES Bwxociw 81 23

A ~ l o i n U b modka LOgnovr, rpsov.; í&r 90-103.

Dcscribcd f m n 9 m d a Ud femiltr from the irLndr PJma lad Hiena In iu coloration, &e and stNciun gratly n a m b h A.dlrldrnr q.a. from which it chbfly

diffur in the rtruc~~rc of the gcniwh. erpefiüly thois of the mala Femalc. Dorvl pkte nrnowed, erpefLUy ilmg intefior m8rgin. in the p h a whcm it puwr

into the ipicil thiid of the length. Intcrku muginr of ventnl phtc widcned in rhipe of long and m o w zigh t-anglcd bladek rtretching on to Mes of d d phte.

M e Puunuu pntly tapcrhg from base towuda blrde in anterior m q h , bu1 strongly mowing ia upper part, u) that they f o m bnnchcs, contipour at uitulor m q h r of trbnphr rpeg drawn d e . quite diffuent from in A.dwi&ns (fw 101-102;vkw from above). PortuiOr @r of the piromem undilated. thcir interior mUpns thur having the outhe of an obbng oval in portenor vKW; in A.diuAdenr thc lowu port of the ovil u conridembly MXIOWC~ owag to the diiation of the posterior marginr

Muainmentr of female: le@ 3.3-3.75; length of forewings 2.82-2.97. width 1.1-1.15;

0.07-0.1; icngth of antclu~~c 1.77-1.9; kngth of dorsal phtc 1.0-1.05. Maaircments of mrb: iength 3.0-3.25; iength of fonwingr 255-27. width 1.05-1.07; width of h u d 0.8-0.82. width

wylth of hud 0.85-0.87. width of vertex 0.5-0.51. h g t h 0.21-0.22. kqth of @ CO~W

Figx 98-103. Alylainüh m o d h Loginova, sp.n: 98 - herd. from above and bclow, 99 - fomwing of male, iüü - genitaiia of malc. in hterdview. 101-102 - puunercs, from inridemd above. 103 - apex of pcnir

of vertex 0.45-0.5. kngth 0.16-0.2. length of genal cones 0.07; length of antennae 1.62-1.77; heigth of anal iube 0.4-0.42. heigth of parameres 0.5-0.55.

La Palma: Caldera, 25 V 1947. 1 d; supra El Paso 600 m, 4 IV 1950.1 d, 1 9; supn El Paso. El Pindi 1300 m, 1 9. Hierro, Guuuou, 26 III 1950.3 dd. including a hoiotype, 2 99.

Found on Palma with A.dividens, but evidently more common than it on the island. judgimg by the collectionr Hori plant uncertíin.

Aryiuinüb diluia Loginwa, sp.nov.; Ti& 104-112.

Desdbed from IS males and fmaler from Gran Canaria Ud Tcncrüe. UliuUy uniformly strawyellow. the dorsal pnern Lclting, forcwings colourlcu or whitish with

brown veinr; faint yellow patches sometimes found in centre of cells, crpecLlly beride a Iuier of marginal spinu Firrt 4 antennal segmentr dirty ycllow; 3-4 apicdly, 5-10 wholly duk-brown.

F b 104-112. ArYfUinüb diüm Loginova. sp.n.: 104 - hcad, from above below, 10s - antenna. 106 - forewing of femalc. 107 - genitaih of femalc. in hteral vkw. 108 - g e n i w of malc. in lateral view. 109- 110 - parameres, from insidc and above. 1 1 1 - apuc of pcnir, 112 - e@.

o !

I Y

FomvLyr widert ki.ond fust thud of their lcngth, minute supcrfkhl rpiner miss@ in ccll e; in the res1 of the c e b i k y approuh the veins. but lave wide rpuwlcrr bordcrr

Femilc Donrl p l ia wids; slightiy clevated in uul puf itr top süghtly bcnt u p w d r Ventral piate basaüy twyC as he u doraal, considenbly Bata, with bldbüke dilations i l q intoikr marginr

M i l c Anal tube WIOIY, porterioriy faintly düated up l o hilf itr heighf lower than puunt%~~. Panmeres simply lamdiate, widenky at buc and apuc. rpially f o m b a hook-iikc foiwud cuned branch (vicw f r o i above. fi 110).

Merruremcntr of f d : kngth 242-275; km of forsaring 2.02-2.1. width 0.9-0.97; width of heid 0.62-0.76. width of V ~ C X 0.37-0.47, imfi 0.15-0.17, lS@ O f @?IIAl COMS 0.05-al; kngth of antcnna 1.25-1.45; kqth of dorsal plite 0.65-0.75. M ~ m c n t a ofmik: icngth 225-267; lcngtb of forwings 1.87-2.2; width 0.75-0.9; width ol head 0.61-0.69. width of vertex 0.37-0.42. Ln(th 0.12-0.16, length of genal cones 0.03-0.05; length of rntennie

Tenaife. Lu Merccdc~, 28-30 V 1947, 1 9. G m Cpnuk: TiniuiS. S. BPrthdwis, 14 111 ae iqúa, 28-29 111 1 9 9 . i d; CNZ dc Tejeda 1450 m, 11-13 111 1950.1 9 , 1

A.di&& sp.n cI& ruembkr the spceier desaihd bclow cQ.quifans sp.n.1. Hort p h t

1.25-1.4; hcifii of uul tube 0.27-0.32. height of ~UMICICS 0.35-0.37.

1950,19; IV 1959.4 dd. inchidhg a holotype. 4 99; Suita B W , 31 111 1959,2 dd.

unknown

AryiamüiaequimsLoQnon, spnov.; fm 113-121.

Dcscribcd from 23 malu and femiler from Gran Canaria, GreerryeiioV to mwtud yellow; b d dong antaior mugn of vutex. its foV- sclaites of

mctanotum and abdomen brown Patkm on muoaoúun in dder rpechnenr conrYtny of o m p bonds and rpotr Uppa putr of hud and thonx having dbtinct ügbtcdoumd uty whkh are deiuer and loiyr t h r in A&& q.n, rnd 2 pWr of the longcst setae w g e d cona uurlly l i g h t - c o l d inned d b r o w n

Foiavinl, Wrc thac of A.di&&, parhes oftcn ofcuniiy in ccll, of apial put, csll c da; rninute upuficll rpim in uch eeU. in cell c fomiiry ovil rpot in middle.

Fe&. Donrl p h k FnnUed ntar anur. fainüy concave abow (lateral vim). Triangular wntnl plate b d y twke u h¡@ u dorsal, with triangular bLder in middlc of interior mu&r embmbg sidcr of d o d phtc

Mak. Anal tubc *puLig ap"d. almort as high as psnmuu lod genital scgment. Param~rrr iameüate. (heir nuroriy apicca turned fonvud and inwud their s t ~ ~ t u i e ir vmy pcniiur. the posterior andca of thai a p i w be@ contiguwr (rim from above. Fig. 119). The structum of the

width of h u d 0.76471, width of vatex 0.45-0.49, ie~@th 0.15-0.17. length of wai cones 0.07; luigth of antemnc 1.42-1.6; lcngth of dorsai platc 0.7-0.75. Munuementa of mile: failth

lpcxof thepn**hpecullr. M u a i m C a t r Of tnnrle: la@ 267-3.12: kryh of fütCWing 225-2.55, ~ i d t h 0.97-1.05;

255-3.0; k@ Of f O W i i U , 2.1-2.45. width 0.85-1.05; width of hed 0.78-0.8, ndth of Vertu< 0.42-0.43. kyth 0.16-0.18; kryth O f p d COlKI 0.05-0.07; ICQth O f Mtainie 1.4-1.52;h&htüf ~aibuO.27-0.28.hd~toípuun~rtt0 .3 .

Gran Camia. Valb de TejmJi, 28-29 111 1949.1 9,2 dd(on C)Msus); Tirajana S Butholomc, 14 III 1950. 3 ¿¿, 1 9; CNZ de Tejeda, 1 IV 1959.1 d, 1 9; Santa B~igida, 31 111 1959.5 99; Atairya, 26 VI, 4 dd. mClud@ a hdotype, 5 99 (kg. R. Fw) .

Host plant u n h o m ; colltcted once from Qtirur A.equifonr *.a. wu tonittimu rrcded from the y m c gcogqhiul dbtrietr as A.di&lo sp.n; the two spceics are most probrbly rympatrlc. llley are clorcly relatrd.

Arytainilb pileobta Loginovp, sp.nov.; f i 122-130.

ücrribcd from 22 males and females from Tencrife. Coiourcd iikc A.cquiianr sp.h, but in oider specimcns the veas with brown pigment are

snler . when yellow pitchcs occur on the forcwing. they are found oniy beside series of marginal spiner Setic on thorax short and sparrr on uppcr p r r t Forcwings oval, wkkst apierliy to middle, erch ecU with minute superfxll spiner

Femilc. Do& piate concave ibove. no1 dh tcd anur Ventral platc triangulu, with a vcry süght düaüon in middle of interna¡ m q i n , ike that in A.dilufa sp.n.

Mak. A ~ r l tube nurow. like i h o r of A.diluta and A.cquitanr spp.n., but bwcr thm PUU~CKS uid genital rgmcnt. Parameres similar in form to those of A.hakoni Log., A.cognaia

’

T

i

21

F b 122-130. Arytainübpücohio Loghwa, sp.n.: 122 - hud. from r h e uid bdow, 123 - anterI4 124 - forrwmg of femalc. 125 - gcnitrüi of f a n d q in hted vkw. 126 - p i t r l l r o f mak. in Ltenl view. 127-128 - pinmeier. fmm inde Ud .bovs. 129 - .pcx of p& 130 - CS.

Log. and A.innrba sp.n.; their anterior mugin hwing I m i l trLnykr p r o a s bililly to the Iapering apcx and c o v d with hrge pecic; i p k d margin rmight, h e b e of puunercr contiguair at posterior angier of rpicei (vicw from ibove; Tic 128).

Muniiemaitr of femde: iength 2.5-2.87; length of fomwinga 2.05-237, width 0.87-1.02; width of h u d 0.7-0.76. width of vcrtex 0.41-0.49, kngth 0.15-0.18, iength of genal conea 0.07-0.1; iength of antennie 1.35-1.5; iength of dorui ph te 0.65-0.67. Meuinmentc ofmak: kngth 2.5-257; kngth of fonwmgs 2.02-2.25, width 0.85-0.95; width of herd 0.66-0.7, width

Tencrife: La Mcncdu, 1 9 (kg. R. Fny. N 1325). 28-30 V 1947,l ¿, 3 IV 1949,19,12 iV 1954. 7 ¿¿, inciuding i hoiotypc 5 99 (iq. R Remane); Orotiva, 3 d¿. 1 Q (*r. R Fny); Bailadcro 700 m, 23 IV 1950,l ¿. 2 99 (on Lwrici).

O f mteñ 0.4-0.43, h f i h 0.15-0.17, k W h Of ~ n r l W(KI 0.07; k W h Of ~ l t e ~ u u ~ I.45-1J; height üf mal Nbe 0.25-0.27, he@t of P.IMIWW 0.22-0.25.

In the SlNChire of udi p u t of the body, Apikobta ?.n. bcars a grcat rcsembluice lo two representatives of the p n u s from Morocco. A.cognara Lo& and A.hakoni Log, and 10 A.ineuba sp.n from Mideir Thcr rpocies u e p n c t i d i y cioriy related. Unfortunatcly their host piant rcmhs unknown, but thcir food source may be some kpminous ShNbL

ArifainUlo incuba Loginova, *.nov.; f i 131-140.

Dcsaibcd from 19 males and femalcs from Mdeira. Only onc of the males ir grccn. the othcr Ipecimens are unifomly yeiiow without any pattern

on the thorax Ud abdanen. Antennae yellow or dirty yeiiow, up lo 3rd segmcnt, thc coiour then grduiuy ch- apicad to dark brown. F o n w m p hyaiine. scldom faintiy whitish, no yeilow patches in ecllr, vcins brown

Hcad U wide as tlrorax. dorsal sctac abscnt. Fonwings with anterior and posterior margins more or ku po~Uel . m w i y roundcd at ap ia l m8rgin with apcx in middlc of ceU n Minute upef l c id Ip inrs in iu dlr Gcnai cones smiu ind WOW compucd with thorc of the othcr qwcks and havhg the l r p e o€ obluraangicd tukmiiu.

Fe-. Dorsai piale rweUcd in anal rcgion, bu1 tapcriiy y d u i u y . d o d y aimost straight. Ventral piatc M e . b d y more than twiec u @e u dorsal. the wide biader on ik interior margins not w e k d u in A q u i t a ~ but apprerrcd LO ddes of dorsal piatc (htenl VKW).

Maie. Anal tube wide. resembling that of A.cognata and A.liobni, more than twice as wide as puuneres at ihcir base. lowu than pupmerer. Puuneres tapcring ipicad, with only f h r l y defined projstioní. not distendiry the interior marginK thek apicai mapins with acutc-andcd projections birned fonvardr; thc hdLe in contact owing 10 contiguity of uppu parts of apiul m y i n s (view from abwe; Fb. 138).

Meas~mncnts of femaie: kngth 3.25-3.75; iength of forewings 2.75-3.1. width 1.07-1.2; width of h u d 0.77-0.87. width of vertcx 0.47-0.52. length 0.17-0.22, iength of gcnai cones 0.06-0.07; length of antennac 1.72-1.9; iength oí dorsai piate 0.75-0.85. Mcuunments of male: iength 3.0-3.35; iength of forewiny 2.55-282. width 1.0-1.12; width of hcad 0.75-0.85; width d vertcx 0.45-0.5. length 0.2-0.22, lcngth of p d cones 0.07; iength of antennae 1.77-1.85;

Madeira: Queimdas, 24-26 VI 1957,4 bd, including a hobtypc, 7 99;Ro&, 24 IV 1959, 3 óó, 2 99; Boaventura, 17 v 1959. i d. 2 99.

As alrcady mcntioncd. A.incubo sp.n. is v u y cioseiy reiatcd to A.cognuta Lq., A.hahni Log and Apileobia sp.n.

he@t Of uul tube 0.27-0.3, hciJit O f D u U n e n S 0.35-0.37.

Aryrainüb pmgnata Loginova ?.nov.; fis 141-142.

ücrribed frum a malc and 3 females from Gran Canaria. üody iight yellow. vertex and oftcn pattern on meronotum orange; abdominal acierites

sometimes browtwpottcd. ventral piate and dorvl platc in puticulor builly brown Hed and thoru wilh üght-coioured rctae abovc. h u d U wide as thorpx. C e d cones comprntivdy @e. Ud more than haif iength of vertu<; u i t c ~ a e shortu than in rciatcd sppecks of the genus, ieu than twice as ion( as width of hcad. Forcwinp evenly rounded at apiul mugin. widcst in apiul third.

FigS 131-140. Aryrainilb incuba Loginova, 9.n: 131 - hcad. from above and below, 132 - hcad. in anterior vicw. 133 - antcnna, 134 - íorewhg of mais 135 - geniWi of fcmale. in laterai view, 136 - genitaiu of maie, in lateral view. 137-138 - parameres from inside and above, 139 - apex of penir. 140 - hind 1%.

Figs 141-142. Atyioinüb pr0dMt.s Loginovs. sp.n.: 141 - genitllu of maie. in iatenl view, 142 - paruncreí. from abovc.

l I F

.

Female. Dorsal pktc concave above. widcst in anal region. Ventral plate without dht ions on convex interior marginr

Mile. A n J tube m o w . with apex somewhat bent backwards about 1.5 times u wide as parameres a t thcir bua. Parameres ükc hose of A.incuba sp.n., but more shatply tapering from third of thcir height towardr apex, Iacking projections along anterior margin and denrely covercd with long retae both anteriorly and porteriorly; lamellse of parameres contiguous almoot throughout more or ICU stmight apical margin (vicw from abwe; fig. 142).

Mcasurements of femalc: Icngth 2.67-2.75; Icngth of forewings 2.25-2.3, width 0.92-0.95; width of h a d 0.67-0.7, width of vertex 0.4-0.42, length 0.17-0.18. kngth of g n J cones 0.1-0.12; length of antennae 1.15; lcngth of dorsal plate 0.7-0.75. Mcasurements of male: length 262; length of foming 2.15, width 0.82; width of head 0.67, width of vertex 0.4. length 0.17, length of genal cones 0.1; length of antennae 1.25; height of anal tube 0.35. height ofparameres 0.27.

Gran Canuia: Valle de Tejedo. 28-29 III 1949, 1 d a holotypc, 2 99; Tinjana. S Bartholorne, 14 III 1950, 1 9.

A.pugMta sp.n. h u thc grcatest reaunblancc toA.pi1eolota sp.n. Host plan1 unknown.

Arytainüh umbonata Loginon. sp.nov.; fig. 143-152.

ücscribed from 71 insrets from Madeh Body grcm to pak yellow with oranp dorsal pattcrn and orangc abdomcnal scleritcs, often

bcaring blown spotr Forewing hyalinc. in ohicr insccts usually yellowish at apex; coloration rtnngthcning towards apical mugin; veinr brown.

Hcad u wide u thorax, p n d cones very short, 1/3 or 1/4 length of vcrtcx. d h c t e d h o s t downwardr No dorsal setac. Fominga with anterior and posterior m y i n s more or leu parallel. widcst in a p i d Ihird, dightly cuning inwarda to Cu, along the apical m y i n . Minute aiperfictl spincs not approaching veins, foming andl wrl spot in cell c.

Fcmaie. Dorsal plate düated in anai region. dightly concave above. Vmtn l pkte dutinguishcd by long and narrow blades along interior m y i n s and rtrongly diiated pprt in middle of lower mugin (lateral view).

Malc. Anal lube strongly düated and bladc-üke postaiorly, widcst in fust third of hcight, higher than thc parameres, and as high as genital rgment. Parameres rectuigulu (lateral view), thkk posterior view. with dent interiorly in middk of anterior margin. Apex of parameres with long widc projection. its widc bhdc being extended vutically. Parameres apicilly conIiguousat vertical rnargins of iherc projections (dorsai and anterior view).

Measurements of female: lcngth 2.9-3.37; length of forewings 2.5-2.7. width 1.0-1.1; width of hcad 0.71 0.77.width of vcrtex 0.43-0.47. Icngth O. 17-0.2. Icngth of pna l cones 0.05-0.07; lcngth oí aniennae 1.5-1.75; Icngth of dorsal phte 0.6-0.75. Musurementr of mrle: lcngth 2.75-3.05; length of forewings 2.2-2.5. width 0.85-0.95; width of head 0.75-0.77, width of vertcx 0.42-0.47, lengtb 0.17-0.2.len&th of genal cones 0.05-0.06; length of antennae 1.52-1.8; height of anal Nbe 0.3, height of parameres 0.17.

Madcua: Sena de Aqua, 2 VI1 1957. 7 dd, 6 99, 16-17 Vil 1957, 4 dd, including a holotype. 3 99, 20-21 IV 1959. 14 dd, 16 99(kg. Lindberg and Panelius. on Lyrminowe); Curral das FreirasVvco Gil. 8 V 1959.7 dd, 13 99 0%. Lindberg and Paneliud; Ponta de Pargo, 23 IV 1959.19.

A.umbonaru sp.n. rcsernblcsA.incuba sp.n. in the sbucture of the fonwings, h a d and genilalia, but ir clearly distinguishcd from it by the pcculk struchire of the ventral platc in thc female and the parameres in thc male.

Aylia Ceoffry, 1762.

Thrcc spccies of ibis numerous genuí. two of them new, wcre discovmd in the Canary Idandr

31

Jp 161 u F i n 143-152. Arytainilh umbonota Loginova, ip.n.: 143 - hud. from above and below, 144

- antauu; 145 - fmwing of fmirie. 146 - genitilir. of femdc, in lateral v h . 147 - ganitllk of mde, in iaterai view, 148-149 - puunwes, from inside and above, 150 - apcx oí peak 151 - hind 1%. 152 - egg.

Psy& arhntica Loginovr. ap.nw.; fw 153-160.

Duuibad fran a series of 184 inrtcts from Madeira and Tenerifc. lñe insecu am p n at cmagencc, bu t m n change 10 yellow, then in yellowsrgangc-bnwq

abdomen latcnlly and genitlür sometimes crlnaon. Vertcx and dorsal pat tun vivid o-. uid mow stripes bemeen thc budi of pattern ivory-coloured; thonx ventrally m d IatuiUy with brown-bkk spots, legs yellow to brown. Forewingr tnnaiuccnt. colourleu to dirty ycllowkh. with yellow band Nning along a p i d third of cüvus, ptcrostigrna brownish, minute superficial sphier dark.

.

32 M. M. ~OWNOVA. F'quidrof rheCanq kluidraod

Hcad u wide as thorax. bent down through mgle of 40' or wcn more; gcnd cones u long as, or more often lonpr thra vertex, rct not v u y vide aput. narrowly conical, rtrongly conave extermlly. directed forwudr and downwardr Forrwingr widcrt in apKI1 thud, widcly rounded aiong apiul margin, pterortigma loq, with dutinct R; minute superfiial piner in u e h c e 4 leavins spaee Jong veinr

Femaic GeNtllh cuneate. dorsal pbte di&hUy concave above.

F& 1531-160. Psyih atlontica Lo&ova, íp.n.: 153 - hud. Crom rbwe and below, 154 - forewiy of mile. 155 - genitllk of femde. in bieril view. 156 - gcnitslll of mak in hterd view, 157-158 - puamcres, from insidc and above. 159 - apex of penis. 160 - hind leg.

GMUEMATMWES BKwa;icna 81 33

M J C A d tuk Sshapcd. aincd. not wide. p l n m e m Wrs thw o f h a t b p p í n q u Log., but with apvrl part morc disUncUy tapering and extended backwards rnd inwrrdr The p u u n a u in

Mwurrmenta of fem.b: kqth 3.0-3.75; le@ of fomings 2.55-3.0, width 1.02-1.27;

0.2-0.25; kngth of antennac 1.07-1.25; length of dorsal pbte 0.75-0.9. Meuwmcab of mrh: Iength 285-3.25; length of fomings 237-275, width 0.92-1.07; width of h u d 0.66-0.74, width of vertur 0.4-0.45, I e n N 0.17-0.2,kn~th of g c d cones 0.2-0.22; height of uul (uk

M~deira: Pie0 Ruivo, 27-29 VI 1951,22 dd, 22 99 (onEricaarbm);Poim-A&ito. 19 VI 1957, 12 dd. 8 99 (on Lartus); QuQmdu, 24-26 VI 1957.1 9,14-16 V 1959,l d, 1 9 (m solir); Ribein Frb. 3 V, 1 d.5 %?(kg. StorA); Boanntun. 17 V 1959,4 dd, 8 99; Sem de Aqua, 20-21 IV 1959.1 9; Ribclra Brava, 20 IV 1959.20 dbincludiq a hobiype. 29 %!ton Prunur); S o Vicente, 24 IV 1959, 1 9. T e n d e : Laa M e d e 30 1 1949, 1 9.12 IV 1954.2 99 (ki. R. Rcmw); V i r 1500 m. 27 11 1 9 5 O d f r u 1300 m. 19 V 1950. 1 9:

them rpc~Wr ~ V C dLtinct ~tlhier b W y rnd po~ariorfy (cC. Lodnov.. 1966: 22-23).

width of h e d 0.72-0.8, width of v e r t u 0.45-0.47, 1- 0.17-0.2, h g t h üf ~CMI concg

0.25-0.27; hei@t oí p ~ r u n f ~ ~ ~ 0.17-0.22

Valle de S Andri4 2 III 1950. 1 d; G A d i l k , 26 lI~i95& 1 d; Bm.dcL Inf&, 24 11 1950,ZO 66,21 Y Y (m M&).

In tbe complex of wUow-fe#tiiy spcckr of the p n u s Psyk thb wier ir the ninth rcpnlcntatlve of thc group ftwphih (Loginovr 1967). It mort d o s l y rsrmbka R p n ~ p i n q ~ ~ SchuL and Rrubpropinqua Log. DUtinctive fatures in the stNcturc of the hud, forcwinp iiid gcniwL rnd its irobted ara indicate tlutP.ut&ntica sp.n. ir an indepcndent rpccicr Its hwt phnt u clurly thc wiuow.

PIylh exima Loginova, ?.nov.; fig~. 161-169.

' Ducnbcd from 29 males and fanaiu from Tencrife, 26 of thm cobcted by Dr. R. Remane in 1954.

Somc specimcnr pm-onng with a brown abdomen, thc mapdty much darkrcolw~. h c d Ud thm omgc with a brown p i t t m above and rpotr on rider Ud bclow. Vutex iloq cpiuullil suture. pmnotum. prvrcutal uyicr of mesonotum üght yeüow. p n i l m n u yeüow to omge Antennae to 5-6th regnuit duty ydlow, regmentr 3 - 6 ipicdly brovn. m a e rpieri segments brown b k k . Le@ dirty yeUow with brovn l a n a Forewinga transpucnt. ydowish. veinr, minute superficial pincs and a p i d hrlf of drvur on both pain of wings brown Ventnl rleriter of abdomen browmblrck, the hte rqmentd mcmbruie reddish 01 onnge. uul tuk rnd parameres of mile dirty yellow.

vertiully d o w m d s , almort u loq as vertex. sct wide 8pUt. nurowly c o n M a r t U i d y roncavc Antuuue rlmost 1.5 t ima u loiy u h a d Forewing widert in ipkd third thm waüy munded. ptemstignr wide. b@y almort u wide u adjoining rcgion of ceU r, tetqimlosing at level of bifurution of M. minute superficial rpiaer dcnscly oct. but not clore lo veinr (4 162).

Hud U Wide UiOnX. StrOngIy bent f0Wudr ( t h r ~ g h Of 30'). g C d cones dincted

Fermlc. Both platcr tapcring rpiud. donrl pira EOIL~.YC rbovc Milc. A d tube convex ~ t u i o r l y , m u v c postaiorly, u high as parmeres, but lower thm

the hemkphedd genitll regment Paramera u narrow Lunellre with a nun>wed ipur. dnwn fonvrrds (lateral view and view fmm above), thkk posterior view. ilong posterior mugin curyinl a comb of 1-2 liner of very rtrons brown pbteller Apiul p u t of penis having d&ht oval dilrtion. ductur long.

Mcuuranents of femak: kwth 265-3.0; kqth of forewbgr 2.27-2.65, width 0.95-1.0; width of he& 0.65-0.67, width of Vatu( 0.4-0.42. l~ngth 0.17-0.2, kngth of genal CWI 0.17-0.2; length of antcnnae 0.97-1.07; length of donil pbtc 0.62-0.75. Measuranentr ofmile: kngth 2.45-2.67. kngth of forewings 2.07-2.17. width 0.85-0.9; width Of h e d 0.65; widfhof vemx 0.37. length 0.15-0.17. kngth of g c d cones 0.15-0.17; length of antennre 0.92-1.05; height of a d tube and paramcru 0.25.

.

34

Figs 161-169. Prylh uima Loginova, sp.n.: 161 - hud. from abovc and bclow. 162 .- forcwirq of femalc. 163 - genitJir of female. in lateral vkw. 164 - donrlphte. from ab<rvc, 165 - gcnitaüa of mde, in lateral vicw. 166-168 - parameres, from inride, bchmd uid abwe, 169 - apex of penit

Tende:- 12 1V 1954, 10 dd, a holotype unong thcm. 6 99 (ieg. Remane);

Host plan1 unknown. i t ir difnfult lo disover a rciationship lo any other hown rpccics; the blon&&&c. 17 11 1949.1 9; Anya-00 m, 23 IV 1954.1 d. 1 9.

stnicturr of thc genitrlu Lc unqua.

Rylh ahtema F6rrt.- Tenerüe. Aren- 15 V 1947. 2 dd. 1 9.

Puerto de la CNZ. 25-29 111 1959. 1 ¿(kg. Remane); Las

35

T ~ z a F h t e r , 1848.

7iioza unlcoc 4) - Yadeira, PnL Fonnou pr. FuncW 28 IV 1959,19 d9. On Urda dioicn, U urew < I c m M b k In me& fomt Widely dirtributcd hi the Pilierrctk,

no1 recordcd.from A f n u

~wzaolwirFkr-PortoSmto.6-12VII 1957.1 d.Madein:Flcdo Buro, 4 Vil 1957.15 d9;

1957. 4 d9, 14-16 V 1959, 3 99 (on Urtica rp.); Roruio. 24 IV 1959, 1 d. L. P a h , Lor L h q 23 V 1947,l d. La Gomcra, El Cedro u lo00 m. 23 111 1950,l 9. Te~eiife: Tacomate, 6 69 (kg. R Sto&; Orotrva, 8 dQ (is. R. S t d ) ; Las Msrccdy 8 d9 (kg. R S t d ) , 28-30V 1 9 4 7 , 2 9 9 , 3 I V I 9 4 9 , 1 0 ¿ ~ ; & ~ ~ 1 9 &g.R.Stod).31V1947,1 d;PkodiTeydq 1 9 ilcg. R. Fny); Buru>u S Antonio, 21 V 1947, 5 d 9 . 4 II 1949. 5 d9 (onLmuur);- Aguim, 17 11 1949.17 ¿%Puerto de la CNZ, 2-4 11 1949,l 9; Fuente fria 1300 m, 19 IV 1950,

m V u e i t a de T y u i w 700 m, 16-17 11 1950. 3 99. Gran Canaria: Moya, 14 d9 (iet R

On Laumr nobilk. Lauur canaden* the nymphr cause malformationr in the fouYq putfflkrly in young shootr; the marginr of the luvu curl downwudq forming gaüs, in whkh the nymphs develop. Therr are hvo pnuationr a ycpi íin the Crima, the Caucuur wíthin the borda of the USSR), and the adult inaectr hibematc. Widely dirtlibutcd throughout the M e d t u r i o u n corsts of Europe up to tbe Cauaniq reportcd fmm Algcria. Known in mon northun reghnr up to En- and Swcden, but only in hot-houset Importcd to North and South Americo, A put.

Sena de Aqw. 16-19 VII 1957,4 d9; Poiro-Arieiro. 16 VI 1957. 8 d9; QUC~IU~Y 24-26 VI

1949,2 dd. Std);CmzIle ieJw& 1 rP

h9u oblqua Wmr) - Porto Santo. Bokka, 10-16 IV 1959,ll d%P.Cutello - P.F.di0. IV 1959. 1 9. Madeira: Canical. 29 IV 1959. 14 d9 (on Cñenopdum ai6um); RibeiraFdo, 14 V 1959. 1 9; San V h t e , 1-2 VI1 1957, 1 d (onA@&x). La +p Vaüe Hennigua, para upu. 18 111 1950. 8 d9; Supn San Sebutian. 21 111 1950.1 9. encn e: Cuimu. 12 1 1949.31 d9; Cnndi , 23-24 1 1949. 25 d0 ion Chenopodium d e ) ; Puerto de h out, 1-8 V

1950.6 d9; Bur. del Infierno, 24 U 1950,12 dQ; Tunaimo 600 m, 22 11 1950.1 9 ; Coja-Tejina 500 m. 23 11 1950. 1 9; aipn Adcje Querta de la Puor 1200 m. 25 11 1950, 1 ¿; ciwlu 2300-2500 m. 28-30 111 1954. 1 9 0% Rmm). Cna Cuwi.. Lu P h w 5-6 VI 1947,4 d9 (on Cñenopudium); Condo. 9 111 1963. 5 d9; Gddu. 23 U 1949. 10 d9; Mrrpdomy

10-15 111 1963, 14 69; F m s 18 111 1963, 8 d9; P d u ~ del Qiacha, 13,22 111 1963,l 9. Fuuteventurr, Chikgua, 4-14 111 1949,l d.

On onnuai rpc ier of Aorpu and Climopodium Centnl and Southem Europe, Klukhstui. Middk Asia of thc USSR. Morocw.

1947.4 d9. 25-29 111 1954. 1 9 (1% Rmiu>s); Si- 12 11 1949.1 ¿;PII&O de S JUM. 23 11

24-26 11 1949,7 d9; Suih B&i& 21 U 1949.1 9; ANCU, 2 IU 1949.1 d. L M Z U O ~ C H ~ ~ ~ ,

On his numerous travels, Profesor Hlkon Lindberg, m outstanding IutUdkt and man of science, cnthuriasticaiiy coiiseted reprerntatives of various inrct goupr His collections of psyliiis from Morocco, thc Canuy . Isiands and Madein wbn undoubtedly not intended to give a complete picturc of the fauna of thh lpoup in thor areas, being evidentiy more or l e s incidentil. Nevythcles, they are of a p a t significance as the first matcriai offeriq infonnation on this intensting BIOUP in the extreme south-mst of the Palaearctic. The collections from the Canaries and Mpdeira

.

are especirlly valuable. since they give some conccption of ihe distinctive character and weaith of the fauna of thoa isiands.

Rylloideu are phytophagou wcking ihcects of s d &.e. They show very narrow trophic spccidization, being restricted to ccrtain p h t ~ , SO and their divertity and genetic differentiation are thus primarüy detcrmined by the -ter of the p n m t vegetation and its genesir in a @ven territory.

Macaronesia ir an independent floral Mon closely COMCCted with the Meditema- man district and the Palaeotropics and having some boreal and neotropical elements in its flora (Iijinsky, 1937). The Canaries. at the eastem liMt of the *on, lie in the zone with an arid climate and semidesert psunmophytous wgetation, almost wholly devoid of arboreous species. The Westem Canariea are mountahous and have a luxuriant vegetation. Altogether, 1300 spccies of v w ñ l l u @ts grow on the Canaries, 806 of which are aboriginal, haif endemic md 490 imported.

In the lower zone (up to 700-800 m above sea-levei) the vegetation has a subtropical-African character. Four vertical zones can be tnced in the mountainous region - the laurel forests at a height of 700-800 m are gradually nplpced by the maquis; at a height of 1600-2100 m are forests ofpimcs cana&nsis, and at a height of 2000-32000 m the dopes are covered with scnibby u p h d derrts, with bushes of Spartocytisus supmbiur

Among the most common laurel spccies are Persea úuika, Lautus crmorirnuí and Ocoreu faiens; the maquis eonrut mortiy of arboreous heathen, su& u Ericu uhomu, of üex. cümbing Convolvulus cmuviensir, and ieguminous shnibs, e.& Cyricus pmli/cnrs and Adenocurpus viscosur Iñc climate in Madeira ir more equabie than that of the C w i e s , though it is

ninier and the average annual temperature ir somewhat lower. There are no pine foreats and mountainous deserts. A zonc with ihra distiishable phytocoenoses foUowr the zone of laurel forests: there grow ccrellr, upiand shrubs. including Vpcemium mudemse, and hcaiths with Erica arboreu and other phts .

Thirty four pcr a n t of the flon of Madeira is endemic; thesc endemic specier are cither relict spacies from Madeúa or nlated to South European species. nie p h t s growing on open land in both Madein and-the Canaries are dortly endemic.

Considention of the zoopopphical composition of the North Macaronesian isiands is rertiicted by the meagreneu of the information avrilable on the food, biology and distribution of these.

Ea& of the 29 species treated here are reported from these isiands for the first time, threc species have recently been described by the author from Morocco and seven are more or leu widely distributed in Europe and Asia.

The most typid feature of the fauna is its Palaearctic - Mediterranean chancter: the gnat majority of the spccies (24-25) are representatives of the humid subtropical Mediterranean region. Thiu U not surprishg, since the hbtoriul connections and preant procimity of the idands to the coasts of Europe and Africa have resuited in great similarlities betwan thcir climate, vegetation and fauna.

Howcver, of the 20 new speciea, Psyl& atiantica sp.n. aionc differs iittie from its cloaest Europan-Asian relations, i.e. the species of the group Itheophila, and Aryfainil& pileohta and A.innrba spp.n. are geneticaily closely connectcd with two

I

species from Morocco. A . h i and A.cogmta me rcmahng 17 speeies are strüungiy difiercnt. and the rmjority teem to be endgnic to Uie Cuurics and Mide¡¡. Thur, the psylüd fauna of the irluidr ir mnrrkible for its diatinctive diuicter and l y e numbcr of endcmic specks and genen.

A m ~ n g the 10 spccier known from the Paiaearctic, Tdom urricoc ir a tnnrp.lre- arctic forest speciu. and Triozu obliqu U llro widely diatributed in that zoogeognphi- cal region. but OCEUR mortly in the uid mnc. Thra rpocks osighate from the uid zone, Rhodochlanis puw@uncturur. DiaphonnO conlinuri and Co@osceniu viridis and until now were biown oniy from Southem M o r o c a .

Two of thepe thm specier are reported from the deserts of the eartem Uiuids LPNIrote and Fuerteventure, while Cvirkiis U alro reported from Cmn Cinuir md from Portugi. Thc npnsentativ« of the gcnus RhodochhnU are xeported from annual dtwortp of the Owno- which are haiophytous and often occu beyond the zone of deserta and stcpper. nie host p h t for R puwipuncrurus ir not yet known, the speeier is recorded from Morocco and Algeria.

The remaining fiw rpccics are of Meditemnean origjn, their faod sourcu are typicai subtropicai arborsJ piants (Oleu. purcicis Lowus etc); like most of the new species, thcy are widely distributed in South Europe and North Africa, and m m of thmi penetrate far into the arid subtropics.

The genetic composition of the Ayl&ideu fauna from the Canary Isluids and Madeira ir thu fairiy homogtneous, the grut majority of the species being of subtropiul origin. There are no boreai speciea; the fact that 10 !% of the species originate from the arid zone indicates a conncction with the Sachara depert fauna. niiS connection ir l e s marked on the i h d s Ytuated fpither from the c o a of South Morocco.

Litmhim

Copcner A.L 1970. Southern Afhcm Pguidic (Homoptcra). 1 - J.entSoc.rth.Af~, vol 33. No. 2:

Hesbpffmhon C. 1951: Thc Alytiinini of thc SubhUy PIYlliay H a n i p t c r r H a n o p ~

HedopHoniron C. 1961: Hempfui (Homoptcn): Psywdra - South Afr.Anim.Lüe, voL8.

IlíjimkyA.P. 1937: Vegetition of the world. Aad.ScL USSR. M o r c c n w - h W 252-270. Loginovu M.M. 1966: New rnd littiekoown psyUidr (Homoptsn) frm the USSR - Tniuctbiu

~ @ O W M.M. 1967: Renw on pd8cuctic prYuidr of íhe enur h ~ u i Cwífr. (Homopt~,

Lw'novu MM. 1972: On the fauna of Pglloidu (Homoptera) fmn Moroeco. - CommenLBioL.

195-200.

FmJV P ~ u i d . e - Al lh M& N8t H h . SCK 12. ~014:417-454.

uppSal8: 487-532

of Zooiogid Inbiutc of Acadany of Scknce USSR. 37: 3-35.

Plylloidu), uiaciitcd wiIh willowr - AnnZoolqici, 34, No. 7, Wunw: 427-461.

47. H ~ l P n k i - H e l l l n ~ ~ ~ ~ : 3-31.