Post on 12-Feb-2021
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SCALE INSECTS AND MEALY BUGS (INSECTA: HOMOPTERA: COCCOIDEA)
R.K.VARSHNEY*, M.J. JADHAV AND R.M.SHARMA
Zoological Survey of India, Western Regional Centre, Pune-411044 *Raj Bhawan, Manik Chowk, Aligarh-202001
The Scale insects and mealy bugs fall under the Superfamily Coccoidea, Order
Homoptera. The name refers to their secretion of waxy scales or mealy that serves as a protective
covering. They range in size from 1.5 mm to 25 mm in length. They are economically important
insects, since many species attack different parts of plant and feed by sucking the plant sap. On
the other hand a few species of scale insects are beneficial to mankind, e.g. the cochineal insect
(Dactylopius indicus) yields a carmine dye and the lac insect (Kerria lacca) is the source of
shellac. They are found on nearly all parts of the host plant including the leaves, branches,
trunks, fruits, and roots. They sometimes occur under bark and may cause various kinds of plant
deformities including chlorotic spots, pits, and galls. They cause severe damage to economically
important plants like sugarcane, tea, coffee, cotton, sorghum, vegetables, greenhouse plants and
almost all kinds of fruit trees.
About 7,500 species of Scale insects and Mealy bugs are known at present from the
world which inhabits almost all parts of the globe, in all floral habitats, from the tundra to the
tropics. The Indian diversity accounts to 409 species under 166 genera spread over 14 families
and 14 subfamilies (Varshney; 1992, 2002) and other scattered literature.
SYSTEMATIC ACCOUNT Superfamily: Coccoidea Family: Margarodidae
Subfamily: Margarodinae Tribe: Kuwanini
1. Genus: Kuwania Cockerell in Fernald, 1903
1. Kuwania zeylanica (Green) Fernald, 1903
Distribution: Kerala (Calicut).
Tribe: Margarodini
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2. Genus: Margarodes Guilding, 1829
2. Margarodes formicarum Guilding, 1829
Distribution: South India.
3. Margarodes indicus Green, 1912
Distribution: Mumbai and Bengaluru.
4. Margarodes papillosus Green, 1912
Distribution: Karnataka (Bengaluru, Bellary).
Subfamily: Neomargarodinae
3. Genus: Neomargarodes Green, 1914
5. Neomargarodes niger Morrison, 1928
Distribution: Karnataka (Mysore, Bellary), Tamil Nadu (Coimbatore).
Family: Monophlebidae Subfamily: Monophlebinae
Tribe: Iceryini 4. Genus: Crypticerya Cockerell, 1895
Crypticerya sp.
Distribution: Odisha (Bhubaneshwar, Taptapani, Talcher and Angul).
6. Crypticerya jacobsoni Morrison, 1928
Distribution: Assam (Dibrugarh, Jokai Forest), Karnataka (Bengaluru), Kerala (Cochin).
7. Crypticerya jaihind Rao, 1951
Distribution: Karnataka (Kanara Dist., Mangalore), Kerala (Cochin, Nelliampathy hills, Travancore), Tamil Nadu (Nilambur).
8. Crypticerya kumari Rao, 1951
Distribution: Karnataka (Gopalswami hills).
5. Genus: Icerya Signoret, 1875
Icerya sp.
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Distribution: Odisha (Bhubaneshwar, Taptapani, Dhauli, Konark, Rambha, Gopalpur-on-sea, Kuliana and Baripada).
9. Icerya aegyptiaca (Douglas, 1890)
Distribution: Bihar, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, Tripura, West Bengal.
10. Icerya formicarum Newstead, 1897
Distribution: Andhra Pradesh, Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu, Uttar Pradesh, West Bengal and North-East India.
11. Icerya menoni Rao, 1951
Distribution: Kerala (Maradu).
12. Icerya minor Green, 1908
Distribution: Assam (Haflong), Bihar (Pusa), Uttar Pradesh (Varanasi).
13. Icerya pilosa Green, 1896
Distribution: Andhra Pradesh (Sarola), Bihar (Shahabad, Juneshwargarh farm), Delhi, Haryana (Karnal), Madhya Pradesh (Gwalior), Kerala (Perungaloor, Travancore), Tamil Nadu (Chennai, Hasanoor, Coimbatore, Kanyakumari, Satyamangalam, Coromandel coast), Uttar Pradesh (Aligarh, Meerut, Shahjahanpur) and West Bengal.
14. Icerya pulcher Morrison, 1928
Distribution: Bihar (Pusa).
15. Icerya purchase Maskell, 19879
Distribution: Karnataka, Kerala, Maharashtra, Odisha, and Tamil Nadu.
16. Icerya seychellarum (Westwood, 1855)
Distribution: Andaman Is., Karnataka (Bellary, Kottur), Kerala, Madhya Pradesh, Maharashtra, Odisha, and Tamil Nadu.
17. Icerya travancorensis Rao, 1951
Distribution: Kerala, Tamil Nadu.
18. Icerya zimmermani Green, 1932
Distribution: Bihar (Bhagalpur).
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6. Genus: Steatococcus Ferris, 1921
19. Steatococcus assamensis Rao, 1951
Distribution: Meghalaya (Dawki).
Tribe: Drosichini
7. Genus: Drosicha Walker, 1858
Drosicha sp.
Distribution: Tamil Nadu (Chennai), Bihar (Pusa).
20. Drosicha burmeisteri Cockerell, 1902
Distribution: North India.
21. Drosicha contrahens Walker, 1858
Distribution: North-East India.
22. Drosicha dalbergiae (Green, 1902)
Distribution: Bihar, Punjab (Sutlej Valley) and Uttarakhand (Dehra Dun).
23. Drosicha leachii (Westwood, 1845)
Distribution: Kerala, Uttarakhand (Dehra Dun).
24. Drosicha mangiferae (Green, 1903)
Distribution: Bihar (Bhagalpur, Pusa, Sitamarhi, Dalsingsarai, Santhal Paraganas, Darbhanga, Muzaffarpur), Delhi, Uttar Pradesh (Fatehpur), Uttarakhand (Dehra Dun).
25. Drosicha quadricaudata (Green, 1922)
Distribution: Uttar Pradesh (Varanasi).
26. Drosicha saundersii (Westwood) Foldi, 2001
Distribution: North India.
27. Drosicha stebbingii (Green); Morrison, 1928
Distribution: Bihar (Darbhanga, Saran, Muzaffarpur), Haryana (Karnal), Himachal Pradesh (Shimla), Madhya Pradesh, Rajasthan (Jodhpur), Uttarakhand (Dehra Dun), Uttar Pradesh (Saharanpur, Kanpur, Varanasi), West Bengal (Howrah & Darjeeling Dist.)
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28. Drosicha tectonae (Green); Morrison, 1928
Distribution: Madhya Pradesh (Damoh), Punjab, Uttar Pradesh.
Tribe: Monophlebini
8. Genus: Aspidoproctus Newstead, 1901
29. Aspidoproctus pertinax Newstead, 1901
Distribution: Karnataka (Bengaluru).
9. Genus: Hemaspidoproctus Morrison, 1927
30. Hemaspidoproctus cinereus Morrison, 1927
Distribution: Bihar (Pusa), Karnataka, Kerala, Gujarat, Rajasthan and Tamil Nadu.
10. Genus: Labioproctus Green, 1922
31. Labioproctus poleii Green, 1896
Distribution: Karnataka (Mysore, Coorg), West Bengal (Darjeeling Dist.).
11. Genus: Misracoccus Rao, 1950
32. Misracoccus assamensis Rao, 1950
Distribution: Meghalaya (Shilong).
33. Misracoccus xyliae Rao, 1950
Distribution: Karnataka (Molakalmurn), Kerala (Malabar) and Tamil Nadu (Coimbatore).
12. Genus: Monophlebus Guerin-Meneville, 1827
Monophlebus sp.
Distribution: Tamil Nadu.
34. Monophlebus atripennis Burmeister, 1835
Distribution: North India.
13. Genus: Monophlebidus Morrison, 1927
35. Monophlebidus indicus Morrison, 1927
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Distribution: Karnataka (Bengaluru).
14. Genus: Perissopneumon Newstead, 1900
36. Perissopneumon ferox Newstead, 1900
Distribution: Kerala, Maharashtra, Tamil Nadu, Uttar Pradesh.
37. Perissopneumon phyllanthi (Green, 1923)
Distribution: Andhra Pradesh (Godavari Dist.), Karnataka (Bengaluru), Kerala (Malabar, Walayar Forest) and West Bengal (Kolkata).
38. Perissopneumon tamarinda Rao, 1950
Distribution: Andhra Pradesh (Godavari Dist.) and West Bengal (Kolkata).
15. Genus: Walkeriana Signoret, 1876
39. Walkeriana senex Green, 1896
Distribution: Kerala (Wynaad) and Tamil Nadu.
Family: Ortheziidae
16. Genus: Orthezia Bosc d’ Antic, 1784
40. Orthezia insignis Browne, 1887
Distribution: Karnataka (Bengaluru), Kerala (Wynad), Tamil Nadu (Salem Dist.), Sikkim (Rangpo) and West Bengal.
Family: Tachardiidae Subfamily: Tachardiinae
Tribe: Tachardini
17. Genus: Kerria Targioni-Tozzetti, 1884
Subgenus: (i) Kerria
Kerria sp. Varshney, 2000
Distribution: Odisha (Bhubaneshwar) and Tamil Nadu (Jawalgiri).
41. Kerria (K.) albizziae (Green, 1911)
Distribution: Bihar, Uttarakhand (Dehra Dun), West Bengal.
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42. Kerria (K.) brancheata Varshney, 1966
Distribution: Chhatisgarh, Jharkhand.
43. Kerria (K.) chamberlini Varshney, 1966
Distribution: Rajasthan (Jodhpur).
44. Kerria (K.) chinensis chinensis (Mahdihassan, 1923)
Distribution: North- East India.
45. Kerria (K.) chinensis kydia (Misra, 1930)
Distribution: Assam (Cachar Dist.).
46. Kerria (K.) communis (Mahdihassan, 1923)
Distribution: Andhra Pradesh, Goa, Karnataka, Kerala, Maharashtra, Tamil Nadu.
47. Kerria (K.) ebrachiata (Chamberlin, 1923)
Distribution: Chhatisgarh, Jharkhand, Karnataka (Bengaluru).
48. Kerria (K.) fici fici (Green, 1903)
Distribution: Bihar (Patna), Delhi, Madhya Pradesh, Uttar Pradesh, Jammu & Kashmir, Rajasthan, Tamil Nadu (Coimbatore), West Bengal.
49. Kerria (K.) fici jhasiensis (Misra, 1930)
Distribution: Uttar Pradesh (Jhansi).
50. Kerria (K.) indicola (Kapur, 1958)
Distribution: Bihar.
51. Kerria (K.) lacca lacca (Kerr, 1782)
Distribution: Andhra Pradesh, Bihar, Karnataka, Odisha, Tamil Nadu, Uttar Pradesh and West Bengal.
52. Kerria (K.) lacca ambigua (Misra, 1930)
Distribution: Uttar Pradesh (Guna, Jhansi).
53. Kerria (K.) lacca mysorensis (Mahadihassan, 1923)
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Distribution: Karnataka (Bengaluru, Mysore).
54. Kerria (K.) nagoliensis (Mahadihassan, 1923)
Distribution: Madhya Pradesh (Raipur, Sohagpur).
55. Kerria (K.) nepalensis Varshney, 1977
Distribution: Bihar.
56. Kerria (K.) pusana (Misra, 1930)
Distribution: Bihar.
57. Kerria (K.) laca mysorensis (Mahadihassan, 1923)
Distribution: Karnataka.
58. Kerria (K.) sharda Misra & Sushil, 2000 rangoonensis (Chamberlin, 1925)
Distribution: Odisha.
Subgenus: (ii) Chamberliniella Varshney, 1987
59. Kerria (C.) rangoonensis (Chamberlin, 1925)
Distribution: Assam.
Subfamily: Tachardininae
18. Genus: Paratachardina Balachowsky, 1950
60. Paratachardina mahdihassani (Kondo & Gullan, 2007)
Distribution: Karnataka.
61. Paratachardina mithila Varshney, 1968
Distribution: Meghalaya (Shillong).
62. Paratachardina silvestri (Mahdihassan, 1923)
Distribution: Andhra Pradesh, Karnataka (Bengaluru) and Tamil Nadu.
63. Paratachardina ternate (Chamberlin, 1923)
Distribution: Kerala (Travancore).
64. Paratachardina theae (Green) in Green & Mann, 1907
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Distribution: Assam, Sikkim and West Bengal.
Family: Eriococcidae
19. Genus: Eriococcus Targioni-Tozzetti, 1869
65. Eriococcus araucariae Maskell, 1879
Distribution: Karnataka (Bengaluru) and Tamil Nadu.
66. Eriococcus lagerstroemiae Kuwana, 1907
Distribution: Tamil Nadu.
67. Eriococcus phyllanthi Ferris, 1957
Distribution: Karnataka (Bengaluru).
68. Eriococcus rhodomyrti Green, 1922
Distribution: Maharashtra.
69. Eriococcus transwersus Green, 1922
Distribution: India
70. Eriococcus beesoni Green, 1926
Distribution: Odisha (Ganjam Dist.) and Tamil Nadu
Family: Kermesidae
20. Genus: Kermes Boitard, 1828
71. Kermes himalayensis Greee, 1909
Distribution: North-West India.
21. Genus: Pseudopulvinaria Atkinson, 1889
72. Pseudopulvinaria sikkimensis Atkinson, 1889
Distribution: Assam, Meghalaya, Sikkim, Shillong.
Family: Dactylopiidae
22. Genus: Dactylopius Costa, 1835
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Dactylopius sp.
Distribution: India: Odisha (Bhubaneswar).
73. Dactylopius coccus Costa, 1835
Distribution: India.
74. Dactylopius indicus Green, 1922
Distribution: Andhra Pradesh (Godavari Dist.), Bihar (Patna, Gaya), Himachal Pradesh, Odisha, Tamil Nadu and West Bengal.
75. Dactylopius tomentosus (Lamarck, 1801)
Distribution: Odisha, Karnataka, Maharashtra, Tamil Nadu and West Bengal.
Family: Pseudococidae Sub family: Pseudococcinae
Tribe: Phenacoccinini
23. Genus: Birendracoccus Ali, 1975
76. Birendracoccus saccharifolii (Green, 1908)
Distribution: Andhra Pradesh, Bihar, Delhi, Uttar Pradesh (Aligarh, Meerut) and West Bengal.
24. Genus Brevennia Goux, 1940
77. Brevennia rehi (Lindinger, 1943)
Distribution: Andhra Pradesh, Bihar, Delhi, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu and West Bengal.
25. Genus: Coccidohystrix Lindinger, 1943
Coccidohystrix sp.
Distribution: Punjab (Hoshiarpur).
78. Coccidohystrix insolita (Green, 1908)
Distribution: Andhra Pradesh, Bihar, Gujarat, Goa, Kerala, Haryana, Tamil Nadu and West Bengal.
26. Genus: Helicoccus Sulc, 1912
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79. Helicoccus singularis Avasthi & Shafee, 1982
Distribution: Andhra Pradesh.
27. Genus: Naiacoccus Green, 1919
80. Naiacoccus serpentines Green, 1919
Distribution: Punjab and Uttar Pradesh.
28. Genus: Nipacoccus Sulc, 1945
Nipacoccus sp.
Distribution: Punjab, Karnataka and Odisha.
81. Nipacoccus viridis (Newstead, 1894)
Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Himachal Pradesh, Maharashtra, Kerala, Karnataka, Uttar Pradesh and West Bengal.
29. Genus: Novonilacoccus Ghosh & Ghose, 1987
82. Novonilacoccus oryzae Ghosh & Ghose, 1987
Distribution: West Bengal (Kalyani).
30. Genus: Paraputo Laing, 1929
83. Paraputo taraktogeni Rao, 1950
Distribution: Assam (Dibrugarh).
31. Genus: Phenacoccus Cockerell, 1893
Phenacoccus sp.
Distribution: Kashmir and Uttarkhand (Dehra Dun).
84. Phenacoccus indicus (Avashti & Shafee, 1980)
Distribution: Karnataka (Mysore, Bengaluru).
85. Phenacoccus quaternus (Green, 1912)
Distribution: Tamil Nadu (Coromandel Coast).
32. Genus: Rastrococcus Ferris, 1954
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Rastrococcus sp.
Distribution: Bihar (Patna).
86. Rastrococcus iceryoides (Green, 1908)
Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Himachal Pradesh, Maharashtra, Kerala, Karnataka, Uttar Pradesh and West Bengal.
87. Rastrococcus invadens Williams, 1986
Distribution: India
88. Rastrococcus mangiferae Ferris, 1954
Distribution: Andhra Pradesh, Karnataka, Odisha and Tamil Nadu.
89. Rastrococcus ornatus Ferris, 1954
Distribution: Tamil Nadu.
90. Rastrococcus spinosus (Robinson, 1918)
Distribution: Odisha (Bhubaneswar).
91. Rastrococcus resinophila Green, 1916
Distribution: Himachal Pradesh, Jammu & Kashmir and Uttar Pradesh.
Tribe: Pseudococcinini
33. Genus: Adolesoma Borchsenius, 1948
92. Adolesoma phragmitidis Borchsenius, 1948
Distribution: West Bengal.
34. Genus: Cataenococcus Ferris, 1955
Cataenococcus sp.
Distribution: Andaman Is.
93. Cataenococcus theaecola Das & Ganguli, 1962
Distribution: Assam and West Bengal.
35. Genus: Chorizococcus McKenzie, 1960
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94. Chorizococcus sorghi Williams, 1970
Distribution: Madhya Pradesh.
36. Genus: Crisicococcus Ferris, 1950
95. Crisicococcus dischidae Ezzat & McConnell, 1959
Distribution: Tripura.
37. Genus: Dysmicoccus Ferris, 1950
96. Dysmicoccus breviceps (Cockerell, 1893)
Distribution: Andhra Pradesh, Karnataka, Kerala, Odisha, Tripura and West Bengal.
97. Dysmicoccus carens Williams, 1970
Distribution: Tamil Nadu.
98. Dysmicoccus cocotis (Maskell, 1890)
Distribution: Kerala.
99. Dysmicoccus cucurbitae Avashti & Shafee, 1986
Distribution: Uttar Pradesh.
100. Dysmicoccus triadus Williams, 1987
Distribution: India.
101. Dysmicoccus williamsi Avashti & Shafee, 1981
Distribution: Bihar.
38. Genus: Ferrisia Fullaway, 1923
Ferrisia sp.
Distribution: Odisha (Bhubaneswar, Puri).
102. Ferrisia virgata (Cockerell, 1893)
Distribution: Andhra Pradesh, Bihar, Goa, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu, West Bengal and Uttar Pradesh.
39. Genus: Indococcus Ali, 1968
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Indococcus sp.
Distribution: Odisha.
103. Indococcus pipalae Ali, 1968
Distribution: Bihar and Odisha.
40. Genus: Kermicus Newstead, 1897
104. Kermicus wroughtoni Newstead, 1897
Distribution: India
41. Genus: Kiritshenkella Borchsenius, 1948
105. Kiritshenkella sacchari (Green, 1900)
Distribution: Andhra Pradesh, Bihar, Assam, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu and West Bengal.
42. Genus: Maconellicoccus Ezzat, 1958
106. Maconellicoccus hirsutus Green , 1908
Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu, Uttar Pradesh.
107. Maconellicoccus perforates (De Lotto 1954)
Distribution: Bihar.
43. Genus: Paracoccus Ezzat & McConnell, 1956
108. Paracoccus burnerae Ezzat & McConnell, 1956
Distribution: Uttar Pradesh.
109. Paracoccus nellorensis Avashti & Shafee, 1984
Distribution: Andhra Pradesh.
44. Genus: Planococcoides Ezzat & McConnell, 1956
110. Planococcoides bengalensis Ghosh & Ghose, 1988
Distribution: West Bengal.
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111. Planococcoides robustus Ezzat & McConnell, 1956
Distribution: Andhra Pradesh, Bihar, Gujarat, Karnataka, Kerala, Maharashtra and Uttar Pradesh.
45. Genus: Planococcus Ferris, 1950
Planococcus sp.
Distribution: Tripura.
112. Planococcus citri Ferris, 1950
Distribution: Karnataka, Kerala and Tamil Nadu.
113. Planococcus ficus Fernald, 1903
Distribution: India.
114. Planococcus indicus Avashti & Shafee, 1987
Distribution: Kerala (Ernaculum) and Tamil Nadu (Coimbatore: Botanical Garden)
115. Planococcus lilacinus (Cockerell, 1905)
Distribution: Andhra Pradesh, Karnataka, Kerala, Odisha and Tamil Nadu.
46. Genus: Pseudococcus Westwood, 1840
Pseudococcus sp.
Distribution: Andhra Pradesh, Maharashtra and Odisha.
116. Pseudococcus formiceticola (Newstead, 1900)
Distribution: Maharashtra (Konkan).
117. Pseudococcus longispinus (Targioni-Tozzetti, 1867)
Distribution: Andhra Pradesh, Karnataka, Kerala, Tamil Nadu and West Bengal.
118. Pseudococcus saccharicola Takahashi, 1928
Distribution: Tamil Nadu (Coimbatore).
47. Genus: Saccharococcus Ferris, 1950
119. Saccharicoccus sacchari (Cockerell, 1895)
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Distribution: Andhra Pradesh, Bihar, Delhi, Goa, Haryana, Maharashtra, Karnataka, Kerala, Madhya Pradesh, Odisha, Tamil Nadu and Uttar Pradesh.
48. Genus: Trionymus Berg, 1899
120. Trionymus ceres Williams, 1970
Distribution: Madhya Pradesh.
Tribe: Rhizoecini
49. Genus: Geococcus Green, 1902
121. Geococcus citrinus Kuwana, 1923
Distribution: Tamil Nadu (North Arcot Dist.).
122. Geococcus coffeae Green, 1933
Distribution: Tamil Nadu (Coimbatore).
50. Genus: Rhizoecus Kunckel d’Herculais, 1878
123. Rhizoecus amorphophalli Betram, 1940
Distribution: India.
124. Rhizoecus cocois Williams, 1985
Distribution: Kerala (Thiruvananthapuram Dist.).
125. Rhizoecus cynodontis Green, 1931
Distribution: Andhra Pradesh (Godavari Dist.) and Tamil Nadu (Coimbatore).
51. Genus: Xenococcus Silvestri, 1924
126. Xenococcus annandalei Silvestri, 1924
Distribution: Andhra Pradesh, Karnataka, Odisha (Berkuda Is. In Chilka lake) and Tamil Nadu.
127. Xenococcus scorpioides De Lotto, 1977
Distribution: India.
Tribe: Sphaerococcini
52. Genus: Antonina Signoret, 1875
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128. Antonina anceps Green, 1914
Distribution: Maharashtra (Probably Pune).
129. Antonina graminis (Maskell, 1897)
Distribution: Bihar, Delhi, Karnataka, Odisha, Uttar Pradesh and Tamil Nadu.
130. Antonina maritime Green, 1922
Distribution: Tamil Nadu (Coimbatore) and West Bengal.
131. Antonina zonata Green, 1919
Distribution: Bihar and Tamil Nadu (Coimbatore).
53. Genus: Chaetococcus Maskell, 1898
132. Chaetococcus bambusae (Maskell, 1893)
Distribution: Tamil Nadu (Coimbatore).
Family: Aclerdidae
54. Genus: Aclerda Signoret, 1874
Aclerda sp.
Distribution: Odisha (Bhubaneswar, Taptapani).
133. Aclerda distorta Green, 1909
Distribution: Tamil Nadu (Coimbatore) and West Bengal.
134. Aclerda inermis Ayyar, 1936
Distribution: Andhra Pradesh.
135. Aclerda ischaemum Ayyar, 1930
Distribution: Tamil Nadu (Coimbatore) and Uttar Pradesh.
136. Aclerda tokionis (Cockerell, 1896)
Distribution: Bihar, Odisha and Tamil Nadu (Salem).
Family: Asterolecaniidae Subfamily: Asterolecaniinae
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55. Genus: Asterolecanium Targioni-Tozzetti, 1868
Asterolecanium sp.
Distribution: Tripura (Agartala) and West Bengal.
137. Asterolecanium longum Green, 1909
Distribution: Tamil Nadu (Coimbatore).
138. Asterolecanium minutum Takahashi, 1930
Distribution: West Bengal.
139. Asterolecanium pseudomilliaris Green, 1922
Distribution: Assam and West Bengal.
140. Asterolecanium tenuissimum Green, 1909
Distribution: Kerala (Malabar, Dhoney valley).
56. Genus: Bambusaspis Cockerell, 1902
141. Bambusaspis bambusae (Boisduval, 1869)
Distribution: India.
Family: Cerococcidae
57. Genus: Asterococcus Borchsenius, 1960
142. Asterococcus ramakrishnai Lambdin, 1983
Distribution: Tamil Nadu (Coimbatore), Maharashtra (Mumbai).
58. Genus: Cerococcus Comstock, 1882
Cerococcus sp.
Distribution: Uttar Pradesh (Aligarh).
143. Cerococcus bryoides (Maskell, 1894)
Distribution: Tamil Nadu (Chennai).
144. Cerococcus indicus (Maskell, 1897)
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Distribution: Andhra Pradesh, Bihar, Karnataka, Maharashtra (Mumbai), Odisha and West Bengal.
145. Cerococcus ornatus Green, 1909
Distribution: Tamil Nadu.
Family: Lecanodiaspididae
59. Genus: Anomalococcus Green, 1902
146. Anomalococcus indicus Ayyar, 1930
Distribution: Karnataka, Odisha and Tamil Nadu.
60. Genus: Cosmococcus Borchsenius, 1959
147. Cosmococcus erythrinae Borchsenius, 1959
Distribution: India.
61. Genus: Stictacanthus Lambdin & Kosztarab, 1973
148. Stictacanthus azadirachtae Green, 1909
Distribution: South India and Uttar Pradesh.
Family: Coccidae Subfamily: Ceroplastinae
62. Genus: Ceroplastes Gray, 1928
Ceroplastes sp.
Distribution: Delhi, Odisha and Tripura.
149. Ceroplastes actiniformes Green, 1896
Distribution: Andhra Pradesh, Bihar, Maharashtra, Odisha, Uttar Pradesh, Tamil Nadu and West Bengal
150. Ceroplastes ajmerensis (Avasthi & Shafee, 1979)
Distribution: Rajasthan (Ajmer).
151. Ceroplastes alami Avasthi & Shafee, 1986
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Distribution: Uttar Pradesh (Aligarh) and Tamil Nadu (Coimbatore).
152. Ceroplastes ceriferus (Fabricius, 1798)
Distribution: Bihar, Uttar Pradesh, Himachal Pradesh, Maharashtra (Mumbai), Madhya Pradesh, Tamil Nadu (Coimbatore, Tanjore) and West Bengal.
153. Ceroplastes floridensis Comstock, 1881
Distribution: Assam, Bihar, Uttar Pradesh, Himachal Pradesh, Karnataka, Maharashtra (Mumbai), Madhya Pradesh, Odisha, Tamil Nadu (Coimbatore, Tanjore) and West Bengal.
154. Ceroplastes pseudoceriferus (Fabricius, 1798)
Distribution: Bihar, Delhi, Odisha and Uttar Pradesh.
155. Ceroplastes rubens Maskell, 1893
Distribution: Andhra Pradesh (Chicacole), Assam, Kerala, Tamil Nadu (Coimbatore) and West Bengal.
156. Ceroplastes rusci Signoret, 1972
Distribution: Uttarakhand (Dehra Dun).
63. Genus: Vinsonia Signoret, 1972
157. Vinsonia stellifera Westwood, 1871
Distribution: Andhra Pradesh, Karnataka, Kerala, Uttar Pradesh, Maharashtra (Mumbai), Odisha, Tamil Nadu and West Bengal.
Subfamily: Coccinae Tribe: Coccini
64. Genus: Akermes Cockerell, 1902
158. Akermes montanus (Green, 1908)
Distribution: Himachal Pradesh.
65. Genus: Ceronema Maskell, 1895
159. Ceronema koebeli Green, 1909
Distribution: Tamil Nadu (Coimbatore).
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66. Genus: Coccus Linnaeus, 1758
Coccus spp.
Distribution: Andhra Pradesh, Karnataka, Tripura, Uttar Pradesh and Odisha.
160. Coccus acutissimus Green, 1896
Distribution: Kerala (Malabar) and Tamil Nadu (Coimbatore).
161. Coccus adersi (Newstead, 1917), Ali, 1971
Distribution: Tamil Nadu (Coimbatore).
162. Coccus almoraensis Avasthi & Shafee, 1984
Distribution: Bihar and Uttar Pradesh (Almora).
163. Coccus discrepans Green, 1904
Distribution: Andhra Pradesh, Assam, Bihar and Kerala.
164. Coccus gymnospori Sanders, 1909
Distribution: Andhra Pradesh and Maharashtra (Pune).
165. Coccus hesperidium Linnaeus, 1758
Distribution: Andhra Pradesh, Assam, Bihar, Goa, Gujarat, Karnataka, Kerala, Odisha, Jammu & Kashmir, Tripura and West Bengal.
166. Coccus kosztarabi Avasthi & Shafee, 1984
Distribution: Karnataka.
167. Coccus latioperculatum Green, 1922
Distribution: Tamil Nadu (Coimbatore).
168. Coccus longulus (Douglas, 1887)
Distribution: Andhra Pradesh, Assam, Karnataka and Odisha.
169. Coccus ophiorrhizae Green, 1896
Distribution: Andhra Pradesh (Kurnool).
170. Coccus ramakrishnae (Ayyar, 1919)
22
Distribution: Andhra Pradesh and Himachal Pradesh.
171. Coccus viridis (Green, 1889)
Distribution: Assam, Karnataka, Kerala, Odisha, Tamil Nadu and Tripura.
172. Coccus watti (Green, 1900)
Distribution: Assam, Meghalaya and West Bengal.
173. Coccus radicicola Green, 1921
Distribution: Tamil Nadu (Coimbatore).
174. Coccus olivaceum Green, 1922
Distribution: Tamil Nadu (Coimbatore).
67. Genus: Eriochiton Maskell, 1887
175. Eriochiton tessellates Signoret, 1873
Distribution: Tamil Nadu (Coimbatore).
68. Genus: Eulecanium Cockerell, 1893
Eulecanium sp.
Distribution: Himachal Pradesh.
176. Eulecanium capreae (Linnaeus, 1767)
Distribution: Jammu & Kashmir.
177. Eulecanium coryli (Linnaeus, 1758)
Distribution: Jammu & Kashmir and Uttar Pradesh.
69. Genus: Hemilecanium Newstead, 1906
178. Hemilecanium imbricans (Green, 1903)
Distribution: Tamil Nadu (Coimbatore, Nilgiris) and Karnataka (Mysore, Belur).
70. Genus: Maacoccus Tao et al., 1983
179. Maacoccus bicruciatus Tao et al., 1983
23
Distribution: Tamil Nadu.
71. Genus: Marsipococcus Cockerell & Bueker, 1930
180. Marsipococcus marsupiale (Green, 1904)
Distribution: Kerala and Tamil Nadu.
72. Genus: Paralecanium Cockerell & Parrot, 1899
181. Paralecanium expansum expansum (Green, 1896)
Distribution: Karnataka and Tamil Nadu (Chennai).
182. Paralecanium maritimum Fernald, 1903
Distribution: India.
73. Genus: Parthenolecanium Sulc, 1908
183. Parthenolecanium persicae Fabricius, 1776
Distribution: India.
74. Genus: Saissetia Deplanche, 1859
Saissetia sp.
Distribution: Odisha (Bhubaneswar, Baripada, Puri ).
184. Saissetia coffeae (Walker, 1852)
Distribution: Andhra Pradesh, Assam, Bihar, Goa, Karnataka, Maharashtra, Manipur, Odisha, Tamil Nadu and West Bengal.
185. Saissetia crassum Ali, 1971
Distribution: India.
186. Saissetia depressa King, 1902
Distribution: Karnataka, Kerala and Tamil Nadu.
187. Saissetia filicum Fernald, 1903
Distribution: Karnataka, Odisha and Tamil Nadu.
188. Saissetia miranda De Lotto, 1970
24
Distribution: Tamil Nadu (Palani hills).
189. Saissetia nigra King, 1902
Distribution: Assam, Bihar, Jammu & Kashmir, Tamil Nadu, Uttar Pradesh and West Bengal.
190. Saissetia lieae Cockerell, 1901
Distribution: Assam (Tocklai).
191. Saissetia privigna De Lotto, 1965
Distribution: Andhra Pradesh (Guntur).
75. Genus: Taiwansaissetia Tao et al., 1983
192. Taiwansaissetia formicarii Tao et al., 1983
Distribution: Assam and Karnataka.
Tribe: Pulvinariini
76. Genus: Chloropulvinaria Borchsenius, 1952
193. Chloropulvinaria cellulose Varshney, 1985
Distribution: Bihar, Uttar Pradesh and West Bengal.
194. Chloropulvinaria floccifera (Westwood, 1870)
Distribution: West Bengal.
195. Chloropulvinaria psidii Borchsenius, 1957
Distribution: Andhra Pradesh, Bihar, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, Tripura and West Bengal.
196. Chloropulvinaria venkatramani Varshney, 1992 [nomen novum]
Distribution: Tamil Nadu (Coimbatore).
77. Genus: Micropulvinaria Hodgson, 1968
197. Micropulvinaria maxima thespesiae (Green); Varshney, 1992 [n. comb.]
Distribution: Andhra Pradesh and Tamil Nadu.
78. Genus: Protopulvinaria Cockerell, 1894
25
198. Protopulvinaria mangiferae Steinweden, 1929
Distribution: Bihar, Tamil Nadu and West Bengal.
79. Genus: Pulvinaria Targioni Tozzetti, 1866
Pulvinaria spp.
Distribution: Bihar and Odisha
199. Pulvinaria aligarhensis Avasthi & Shafee, 1980
Distribution: Uttar Pradesh (Aligarh).
200. Pulvinaria burkilli Green, 1908
Distribution: Tamil Nadu (Coimbatore) and West Bengal.
201. Pulvinaria indica Avasthi & Shafee, 1980
Distribution: Andhra Pradesh.
202. Pulvinaria obscura Newstead, 1894
Distribution: Tamil Nadu (Chennai).
80. Genus: Takahashia Cockerell, 1986
203. Takahashia japonica (Cockerell, 1896)
Distribution: Uttar Pradesh.
Subfamily: Filipiinae
81. Genus: Cardiococcus Cockerell, 1903
204. Cardiococcus bivalvata (Green, 1903)
Distribution: Andhra Pradesh and Tamil Nadu.
82. Genus: Ceroplastodes Cockerell, 1893
Ceroplastodes sp.
Distribution: Bihar, Jammu & Kashmir, Odisha and Uttar Pradesh.
205. Ceroplastodes cajani Cockerell, 1900
26
Distribution: Andhra Pradesh, Bihar, Himachal Pradesh, Kerala, Punjab, Tamil Nadu and West Bengal.
206. Ceroplastodes chiton Green, 1909
Distribution: Himachal Pradesh, Uttar Pradesh and West Bengal.
83. Genus: Dicyphoccus Borchsenius, 1959
207. Dicyphococcus castilloae Borchsenius, 1959
Distribution: Tamil Nadu (Coimbatore).
84. Genus: Inglisia Maskell, 1879
208. Inglisia chelonioides Green, 1903
Distribution: Kerala, Maharashtra (Shiroda) and Tamil Nadu (Coimbatore).
85. Genus: Lecanopsis Targioni Tozzetti, 1868
209. Lecanopsis ceylonica Green, 1922
Distribution: Tamil Nadu (Coimbatore).
86. Genus: Metaceronema Takahashi, 1955
210. Metaceronema japonica (Maskell, 1897)
Distribution: West Bengal (Darjeeling).
Superfamily: Diaspidoidea Family: Diaspididae
Subfamily: Parlatoriinae Tribe: Leucaspidini
87. Genus: Leucaspis Signoret, 1869
211. Leucaspis manii (Borchsenius, 1964)
Distribution: Meghalaya.
212. Leucaspis riccae Targioni Tozzetti, 1881
Distribution: Andhra Pradesh (Hyderabad).
88. Genus: Lopholeucaspis Balachowsky, 1953
27
Lopholeucaspis sp.
Distribution: Karnataka (Anekal).
213. Lopholeucaspis cockerelli (Grandpre & De Charmoy, 1899)
Distribution: Tamil Nadu.
214. Lopholeucaspis excoecariae Borchsenius, 1964
Distribution: West Bengal.
215. Lopholeucaspis japonica (Cockerell, 1897)
Distribution: Karnataka (Vasanthapura).
216. Lopholeucaspis menoni Borchsenius, 1964
Distribution: Rajasthan, Uttar Pradesh and West Bengal.
89. Genus: Radionaspis Ferris, 1942
217. Radionaspis indica (Marlatt, 1908)
Distribution: Maharashtra (Pune).
90. Genus: Salicicola Lindinger, 1905
218. Salicicola indiaeorientalis (Lindinger, 1911)
Distribution: Himachal Pradesh (Shimla).
Tribe: Parlatoriini
91. Genus: Apteronidia Berlese, 1895
219. Apteronidia blanchardi (Targioni Tozzetti, 1892)
Distribution: Andhra Pradesh (Hyderabad).
92. Genus: Cryptoparlatoria Lindinger, 1905
220. Cryptoparlatoria pini Takahashi, 1938
Distribution: Himachal Pradesh (Manali).
93. Genus: Genaparlatoria MacGillivray, 1921
28
221. Genaparlatoria pseudaspidiotus (Lindinger, 1905)
Distribution: Andhra Pradesh (Godavari Dist.).
94. Genus: Greeniella Cockerell, 1897
222. Greeniella cornigera (Green, 1896)
Distribution: India.
223. Greeniella dentata (Lindinger, 1911)
Distribution: India.
224. Greeniella ferreae (Rutherford, 1914)
Distribution: India.
225. Greeniella viridis (Lindinger, 1911)
Distribution: Kerala (Travancore).
95. Genus: Gymnaspis Newstead, 1898
Gymnaspis sp.
Distribution: Kerala (Kattuthurai).
226. Gymnaspis affinis Ayyar, 1930
Distribution: Tamil Nadu (Tirunelveli).
227. Gymnaspis cassida Hall & Williams, 1962
Distribution: Tamil Nadu (Coimbatore).
228. Gymnaspis ficus Green, 1919
Distribution: Tamil Nadu (Coimbatore).
229. Gymnaspis mangifera Ayyar, 1924
Distribution: Tamil Nadu (Tirunelveli)
230. Gymnaspis ramakrishnai Green, 1919
Distribution: Tamil Nadu (Tirunelveli).
29
96. Genus: Neoleucaspis Green, 1926
231. Neoleucaspis parallela Green, 1926
Distribution: India.
97. Genus: Parlaspis McKenzie, 1945
Parlaspis spp.
Distribution: Karnataka and Kerala.
232. Parlaspis papillosa (Green, 1919)
Distribution: Kerala (Palghat).
98. Genus: Parlatorearum Lindinger, 1911
Parlatorearum sp.
Distribution: Andhra Pradesh (Cuddapah).
99. Genus: Parlatoreopsis Lindinger, 1911
233. Parlatoreopsis chinensis (Marlatt, 1908)
Distribution: Tamil Nadu (Coonoor, Coimbatore).
100. Genus: Parlatoria Targioni Tozzetti, 1868
Parlatoria sp.
Distribution: Karnataka and Tamil Nadu.
234. Parlatoria artocarpi Green, 1919
Distribution: Kerala (Peria Ghat).
235. Parlatoria atalantiae Green, 1905
Distribution: Tamil Nadu.
236. Parlatoria boycei MacKenzie, 1952
Distribution: West Bengal.
237. Parlatoria camelliae Comstock, 1883
30
Distribution: Tamil Nadu.
238. Parlatoria cinerea Hadden in Doane & Hadden, 1909
Distribution: West Bengal and South India.
239. Parlatoria crotonis Douglas, 1887
Distribution: Bihar
240. Parlatoria crypta McKenzie, 1943
Distribution: Andhra Pradesh, Delhi, Karnataka, Maharashtra and Punjab.
241. Parlatoria marginalis McKenzie, 1945
Distribution: Delhi.
242. Parlatoria morrisoni McKenzie, 1943
Distribution: Delhi.
243. Parlatoria mytilaspiformis Green, 1899
Distribution: India.
244. Parlatoria oleae (Colvee, 1880)
Distribution: Bihar, Kashmir, Maharashtra, Rajasthan and West Bengal.
245. Parlatoria orientalis Rao, 1953
Distribution: Tamil Nadu (Coimbatore).
246. Parlatoria pergandii Comstock, 1881
Distribution: Bihar, Karnataka, Kerala and Tamil Nadu (Coimbatore, Mundathorai, Tirunelvelly)
247. Parlatoria proteus (Curtis, 1843)
Distribution: Karnataka, Tamil Nadu (Coimbatore, Nilgiris), Tripura and West Bengal.
248. Parlatoria vateriae Green, 1919
Distribution: Kerala (Travancore, Kollam).
249. Parlatoria ziziphi (Lucas, 1853)
31
Distribution: Tamil Nadu (Coimbatore, Ramanathapuram, Tanjore) and West Bengal.
101. Genus: Tamilparla Takagi, 1987
250. Tamilparla smilacis Takagi, 1987
Distribution: Tamil Nadu (Coonoor, Nilgiris).
Subfamily: Odonaspidinae Tribe: Rugaspidiotini
102. Genus: Mangaspis Takagi & Kondo, 1997
251. Mangaspis Bengalurunsis Takagi & Kondo, 1997
Distribution: Karnataka (Bengaluru).
103. Genus: Natalaspis MacGillivray, 1921
252. Natalaspis simplex (Green, 1899); Ben-Dov & Takagi, 1974
Distribution: Tamil Nadu (Coimbatore).
104. Genus: Ramachandraspis Rao, 1953
253. Ramachandraspis fenestrata Rao, 1953
Distribution: Tamil Nadu (Coonoor, Nilgiris).
Tribe: Odonaspidini
105. Genus: Berlesaspidiotus MacGillivray, 1921
254. Berlesaspidiotus crenulatus Ben-Dev, 1988
Distribution: Tamil Nadu (Chennai, Palani).
106. Genus: Froggattiella Leonardi, 1900
255. Froggattiella penicillata (Green, 1905); Rutherford, 1915
Distribution: Tamil Nadu (Coimbatore).
107. Genus: Odonaspis Leonardi, 1897
Odonaspis spp.
Distribution: Tripura.
32
256. Odonaspis paucipora Ben-Dov, 1988
Distribution: West Bengal (Kolkata).
257. Odonaspis saccharicaulis (Zehntner, 1897); Fernald, 1903
Distribution: Karnataka, Odisha and Tamil Nadu (Coimbatore).
Subfamily: Aspidiotinae Tribe: Aspidiotini
108. Genus: Abgrallaspis Balachowsky, 1948
258. Abgrallaspis cyanophylli Balachowsky, 1948
Distribution: Karnataka (Bengaluru Kerala and Tamil Nadu (Coimbatore).
109. Genus: Aonidia Targioni - Tozzetti, 1868
259. Aonidia crenulata Green, 1900
Distribution: Assam.
260. Aonidia indica Green, 1919
Distribution: West Bengal.
261. Aonidia loranthi Green, 1896
Distribution: India.
262. Aonidia tentaculata Green, 1919
Distribution: Kerala (Travancore).
110. Genus: Aonidiella Berlese & Leonardi, 1895
263. Aonidiella andersoni (Laing, 1929); McKenzie, 1938
Distribution: Tamil Nadu.
264. Aonidiella aurantii (Maskell, 1879)
Distribution: Andhra Pradesh, Assam, Bihar, Delhi, Kanataka, Kerala, Odisha, Punjab and Tamil Nadu.
265. Aonidiella citrine Nel, 1933
33
Distribution: India.
266. Aonidiella comperei McKenzie, 1937
Distribution: Maharashtra (Mumbai, Colaba).
267. Aonidiella inortata McKenzie, 1938
Distribution: India.
268. Aonidiella misrai Laing, 1929
Distribution: India.
269. Aonidiella orientalis McKenzie, 1937
Distribution: Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Punjab, Tamil Nadu, Tripura and West Bengal.
111. Genus: Aspidiella Leonardi, 1898
270. Aspidiella hartii (Cockerell, 1895)
Distribution: Maharashtra (Pune) and Tamil Nadu (Coimbatore).
271. Aspidiella sacchari MacGillivray, 1921 [comb. new.]
Distribution: Tamil Nadu.
112. Genus: Aspidiotus Bouche, 1833
Aspidiotus spp.
Distribution: Andhhra Pradesh, Odisha and Karnataka.
272. Aspidiotus destructor Signoret, 1869
Distribution: Andhra Pradesh, Kerala, Jammu & Kashmir, Maharashtra (Mumbai), Rajasthan, Tripura, West Bengal and Lakshadweep Is.
273. Aspidiotus hederae (Vallot, 1829)
Distribution: India.
274. Aspidiotus pandani (Boisduval, 1868)
Distribution: India.
34
113. Genus: Borchseniaspis Zahradnik, 1959
275. Borchseniaspis palmae Zahradnik, 1959
Distribution: Karnataka, Kerala and Tamil Nadu.
114. Genus: Brainaspis MacGillivray, 1921
276. Brainaspis kellyi MacGillivray, 1921
Distribution: Delhi.
115. Genus: Clavaspis MacGillivray, 1921
Clavaspis sp.
Distribution: Karnataka.
116. Genus: Chrysomphalus Ashmead, 1880
277. Chrysomphalus aonidum (Linnaeus, 1758)
Distribution: Andhra Pradesh, Assam, Bihar, Kerala, Jammu & Kashmir, Maharashtra (Mumbai), Madhya Pradesh, Gujarat, Tamil Nadu, Tripura and West Bengal.
278. Chrysomphalus dictyospermi (Morgan, 1889)
Distribution: Andhra Pradesh, Assam, Meghalaya, Tamil Nadu and Tripura.
279. Chrysomphalus diversicolor (Green, 1923)
Distribution: Karnataka and Tamil Nadu.
117. Genus: Cryptopalratoreopsis Borchsenius, 1947
280. Cryptopalratoreopsis euphorbiae Hall & Williams, 1962
Distribution: Tamil Nadu (Coimbatore).
281. Cryptopalratoreopsis longispina (Takahashi, 1939)
Distribution: India.
282. Cryptopalratoreopsis spinosissima (Lindinger, 1911)
Distribution: India (Central India).
283. Cryptopalratoreopsis targionipsis (Lindinger, 1911)
35
Distribution: India.
118. Genus: Diaspidiotus Berlese & Leonardi, 1896
284. Diaspidiotus pseudocamelliae Borchsenius, 1966
Distribution: Karnataka.
119. Genus: Duplaspidiotus MacGillivray, 1921
285. Duplaspidiotus tesseratus (Grandpre & de Charmoy, 1899)
Distribution: Karnataka and Maharashtra (Mumbai, Lonavala).
120. Genus: Hemiberlesia Cockerell, 1897
286. Hemiberlesia lataniae (Signoret, 1869)
Distribution: Assam, Odisha, Karnataka, Maharashtra (Mumbai), Tamil Nadu, Tripura and West Bengal.
287. Hemiberlesia rapax (Comstock, 1881)
Distribution: Assam, Odisha, Karnataka, Tamil Nadu, Tripura and West Bengal.
121. Genus: Lindingaspis MacGillivray, 1921
Lindingaspis sp.
Distribution: Tripura.
288. Lindingaspis floriadana Ferris, 1942
Distribution: India.
289. Lindingaspis fusca McKenzie, 1943
Distribution: Maharashtra (Pune) and South India.
290. Lindingaspis rossi (Maskell, 1891)
Distribution: Andhra Pradesh, Maharashtra (Pune) and Madhya Pradesh.
122. Genus: Melanaspis Cockerell, 1897
Melanaspis sp.
Distribution: Odisha.
36
291. Melanaspis glomerata (Green, 1903)
Distribution: Andhra Pradesh, Bihar, Odisha, Gujarat, Tamil Nadu and Uttar Pradesh.
292. Melanaspis smilacis (Comstock, 1883)
Distribution: India.
123. Genus: Metaspidiotus Takagi, 1957
293. Metaspidiotus sp.
Distribution: Karnataka and Goa.
124. Genus: Morganella Cockerell, 1897
294. Morganella cueronsis (Cockerell, 1899)
Distribution: Tamil Nadu and South India.
295. Morganella longispina (Morgan, 1889)
Distribution: Tamil Nadu (Tirunelveli).
125. Genus: Neofurcaspis Green, 1926
296. Neofurcaspis andamanensis Green, 1926
Distribution: Andaman Is.
126. Genus: Octaspidiotus MacGillivray, 1921
297. Octaspidiotus stauntoniae (Takahashi, 1933)
Distribution: Karnataka and Goa.
298. Octaspidiotus australiensis (Kuwana, 1931)
Distribution: India.
299. Octaspidiotus calophylli var. symplocos (Green, 1924)
Distribution: Tamil Nadu (Nilgiris, Coonoor, Kotagiri).
300. Octaspidiotus tamarindi (Green, 1919)
Distribution: Karnataka (Nandi) and Tamil Nadu (Coimbatore).
37
301. Octaspidiotus tripurensis Takagi, 1984
Distribution: Tripura (Agartala).
127. Genus: Parrottia MacGillivray, 1921
Parrottia sp.
Distribution: Karnataka (Devarayanadurga).
302. Parrottia moorei (Green, 1896)
Distribution: Tamil Nadu (Chennai).
128. Genus: Pseudaonidia Cockerell, 1897
Pseudaonidia spp.
Distribution: Tripura.
303. Pseudaonidia duplex (Cockerell, 1896)
Distribution: Assam.
304. Pseudaonidia trilobitiformis (Green, 1896)
Distribution: Andhra Pradesh, Bihar, Kerala, Tamil Nadu and West Bengal.
129. Genus: Quadraspidiotus MacGillivray, 1921
305. Quadraspidiotus perniciosus (Comstock, 1881)
Distribution: Andhra Pradesh, Bihar, Himachal Pradesh, Jammu & Kashmir, Karnataka, Kerala, Maharashtra (Mahabaleshwar), Meghalaya, Odisha, Punjab, Sikkim, Tamil Nadu, Uttar Pradesh and West Bengal.
130. Genus: Semelaspidus MacGillivray, 1921
306. Semelaspidus artocarpi (Green, 1896)
Distribution: Karnataka (Bengaluru, Coorg) and Maharashtra (Mahabaleshwar).
131. Genus: Temnaspidiotus MacGillivray, 1921
307. Temnaspidiotus excisus (Green, 1896)
Distribution: South India and Tripura (Agartala).
38
Subfamily: Diaspidinae Tribe: Protodiaspidini
132. Genus: Anaimalaia Takagi, 1995
308. Anaimalaia scabra Takagi, 1995
Distribution: Tamil Nadu (Anaimalai Hills).
Tribe: Kuwanaspidini
133. Genus: Kuwanaspis MacGillivray, 1921
309. Kuwanaspis annandalei (Green, 1919)
Distribution: Bihar (Parasnath Hills).
134. Genus: Xiphuraspis Borchsenius & Williams, 1963
310. Xiphuraspis spiculata Borchsenius & Williams, 1963
Distribution: Kerala (N. Malabar).
135. Genus: Coronaspis MacGillivray, 1921
311. Coronaspis malabarica Takagi, 1999
Distribution: Kerala (Periyar Tiger Reserve) and Tamil Nadu (Anaimalai Hills).
Tribe: Lepidosaphidini
136. Genus: Acanthomytilus Borchsenius, 1947
312. Acanthomytilus sacchari (Hall, 1923)
Distribution: Bihar, Tamil Nadu (Coimbatore), Uttar Pradesh and West Bengal.
313. Acanthomytilus vermiformis (Takahashi, 1931)
Distribution: Kerala (Dhony Valley).
137. Genus: Andaspis MacGillivray, 1921
Andaspis sp. (nr. leucophloeae Rao)
Distribution: Karnataka (Bengaluru).
314. Andaspis artocarpi Borchsenius, 1967
39
Distribution: Karnataka (Bengaluru).
315. Andaspis betulae (Borchsenius, 1967)
Distribution: Meghalaya (Shilong).
316. Andaspis laingi Rao, 1952
Distribution: Tamil Nadu (Coimbatore, Makkalidurga).
317. Andaspis leucophleae Rao, 1952
Distribution: Tamil Nadu (Coimbatore, Chennai) and Madhya Pradesh.
318. Andaspis meliae (Green, 1919)
Distribution: Andhra Pradesh, Karnataka (Mandya), Tamil Nadu (Coimbatore) and West Bengal (Kolkata).
319. Andaspis numerate Brimblecombe, 1959
Distribution: West Bengal.
320. Andaspis retrusa (Green, 1919)
Distribution: Tamil Nadu.
138. Genus: Aonidomytilus Leonardi, 1903
321. Aonidomytilus albus (Cockerell, 1893)
Distribution: Karnataka (Bengaluru), Tamil Nadu (Coimbatore, Salem, Puttukottai) and Puducherry.
139. Genus: Cornuaspis MacGillivray, 1921
322. Cornuaspis beckii (Newmann, 1869)
Distribution: Assam, Karnataka, Kerala and West Bengal.
323. Cornuaspis piperis (Green, 1908)
Distribution: Western Ghats; Kerala (N. Malabar, Wynaad, Travancore).
140. Genus: Eucornuaspis Borchsenius, 1963
324. Eucornuaspis pinnaeformis Borchsenius, 1966
40
Distribution: West Bengal (Darjeeling).
325. Eucornuaspis pseudomachili Borchsenius, 1964
Distribution: Assam.
141. Genus: Howardia Berlese & Leonardi, 1896
326. Howardia biclavis (Comstock, 1883)
Distribution: Karnataka, Kerala (Wynaad) and Tamil Nadu (Chennai, Anaimalai Hills).
142. Genus: Insulaspis Mamet, 1950
327. Insulaspis citrine Borchsenius, 1964
Distribution: New Delhi.
328. Insulaspis gloverii (Packard, 1869)
Distribution: Bihar, Karnataka and West Bengal.
329. Insulaspis mcgregori (Banks, 1906)
Distribution: Kerala (Thiruvananthapuram Dist.).
330. Insulaspis pallidula Williams, 1969
Distribution: Andhra Pradesh (Godavari Dist.).
331. Insulaspis tapleyi (Williams, 1960); Borchsenius, 1963
Distribution: Karnataka and Uttar Pradesh.
332. Insulaspis tokionis (Kuwana, 1902); Borchsenius, 1963
Distribution: Kerala and Tamil Nadu.
143. Genus: Lepidosaphes Shimer, 1868
333. Lepidosaphes cornutus Green in Ayyar, 1937
Distribution: Kerala and Tamil Nadu (North Arcot Dist., South Arcot Dist., Tanjore, Salem).
334. Lepidosaphes malicola Borchsenius, 1947
Distribution: Jammu & Kashmir (Anantnag).
41
335. Lepidosaphes ulmi (Linnaeus, 1758)
Distribution: India.
144. Genus: Neopinnaspis McKenzie, 1949
336. Neopinnaspis travancorensis (Lindinger, 1911)
Distribution: Kerala (Travancore).
145. Genus: Velataspis Ferris, 1937
337. Velataspis serrulata Ganguli, 1958
Distribution: Assam.
Tribe: Diaspidini
146. Genus: Africaspis MacGillivray, 1921
338. Africaspis chionaspiformis (Newstead, 1910)
Distribution: Tamil Nadu (Coimbatore, Tirunelveli).
147. Genus: Aulacaspis Cockerell, 1893
Aulacaspis spp.
Distribution: Karnataka and Kerala.
339. Aulacaspis gudalura Ferris, 1956
Distribution: Tamil Nadu (Nilgiris).
340. Aulacaspis hedyotidis Takagi, 1985
Distribution: India.
341. Aulacaspis herbae Hall, 1946
Distribution: Tamil Nadu (Coonoor, Nilgiris).
342. Aulacaspis litzeae (Green, 1896)
Distribution: West Bengal (Darjeeling).
343. Aulacaspis loranthi Cockerell, 1902
42
Distribution: Bihar (Parasnath Hill).
344. Aulacaspis madiunensis (Zehtner, 1898)
Distribution: Andhra Pradesh and Tamil Nadu (Chengalpattu, North & South Arcot Dist., Tanjore, Tiruchirapalli).
345. Aulacaspis malayala Varshney, 2002
Distribution: Kerala (Periyar Tiger Reserve).
346. Aulacaspis mangiferae (Newstead, 1908)
Distribution: Andhra Pradesh, Karnataka and Tamil Nadu (Tirunelveli).
347. Aulacaspis orientalis (Green, 1922)
Distribution: Tamil Nadu (Courtallum, Tirunelveli).
348. Aulacaspis rosae (Bouche, 1833)
Distribution: Tamil Nadu (Courtallum, Tirunelveli).
349. Aulacaspis tubercularis (Newstead, 1908)
Distribution: Karnataka, Kerala and Uttar Pradesh.
350. Aulacaspis vitis (Green, 1896)
Distribution: Assam, Bihar, Himachal Pradesh, Odisha, Karnataka and Tamil Nadu.
148. Genus: Chionaspis Signoret, 1869
Chionaspis sp.
Distribution: India.
351. Chionaspis engeddensis Bodenheimer, 1924
Distribution: Punjab.
352. Chionaspis furfura (Fitch, 1857)
Distribution: Jammu & Kashmir.
353. Chionaspis himalaica Takagi, 1985
Distribution: Himachal Pradesh.
43
354. Chionaspis ramakrishnai Rao, 1953
Distribution: Tamil Nadu (Coonoor, Nilgiris).
355. Chionaspis salicis (Linnaeus, 1758)
Distribution: Himalayas and Jammu & Kashmir.
149. Genus: Contigaspis MacGillivray, 1921
356. Contigaspis coimbatorensis Borchsenius & Williams, 1963
Distribution: Karnataka and Tamil Nadu (Coimbatore).
357. Contigaspis indigoferae (Hall, 1928)
Distribution: Tamil Nadu (Hosur).
150. Genus: Dentachionaspis MacGillivray, 1921
358. Dentachionaspis centripetalis Rao, 1953
Distribution: Jammu & Kashmir (Srinagar).
151. Genus: Diaspis Costa, 1828
Diaspis sp.
Distribution: Odisha and Tamil Nadu.
359. Diaspis boisduvali Signoret, 1869
Distribution: Tamil Nadu (Nilgiris).
360. Diaspis echinocacti (Bouche, 1833)
Distribution: Karnataka, Kerala, Maharashtra (Pune), Odisha, Tamil Nadu and Uttar Pradesh.
152. Genus: Duplachionaspis MacGillivray, 1921
361. Duplachionaspis divergens (Green, 1899)
Distribution: Kerala, Punjab and Tamil Nadu.
153. Genus: Fiorinia Targioni- Tozzetti, 1868
Fiorinia spp.
44
Distribution: Andhra Pradesh and Tripura.
362. Fiorinia fioriniae (Targioni Tozzetti, 1867)
Distribution: India.
363. Fiorinia fletcheri Laing, 1929
Distribution: India.
364. Fiorinia frontecontracta Green, 1919
Distribution: Maharashtra (Mumbai).
365. Fiorinia himalaica Takagi, 1975
Distribution: Himachal Pradesh and Uttar Pradesh.
366. Fiorinia japonica Kuwana, 1902
Distribution: West Bengal (Kolkata).
367. Fiorinia multipora Lindinger, 1911
Distribution: Meghalaya.
368. Fiorinia pakistanensis Ahmad, 1970
Distribution: Kashmir.
369. Fiorinia plana Green, 1919
Distribution: Tamil Nadu (Coimbatore, Tirunelveli).
370. Fiorinia proboscidaria Green, 1920
Distribution: Meghalaya.
371. Fiorinia sapindi Green, 1919
Distribution: Maharashtra (Pune).
372. Fiorinia theae Green, 1900
Distribution: Assam, North India and West Bengal (Darjeeling).
154. Genus: Greenaspis MacGillivray, 1921
45
373. Greenaspis decurvata (Green, 1903)
Distribution: Kerala, Odisha, Tamil Nadu and West Bengal.
374. Greenaspis elongata (Green, 1896)
Distribution: Kerala and Tamil Nadu.
155. Genus: Lineaspis MacGillivray, 1921
375. Lineaspis caroli (Green, 1919)
Distribution: West Bengal (Darjeeling).
156. Genus: Myrtaspis Takagi, 1999
376. Myrtaspis syzygii (Takagi, 1985)
Distribution: Uttar Pradesh.
157. Genus: Pinnaspis Cockerell, 1892
Pinnaspis sp.
Distribution: Karnataka (Thalghatpura, Pannayanpalaya, Shivganga).
Pinnaspis sp. [nr. strachani (Cooley, 1899)].
Distribution: West Bengal (Barrackpore) and Karnataka (Gokarna).
377. Pinnaspis aspidistrae (Signoret, 1869)
Distribution: Andhra Pradesh, Karnataka (Shivganga), Kerala (Cochin, Malabar), Madhya Pradesh, Maharashtra (Janjira Estate), Tamil Nadu (Coimbatore, Nilgiris) and Tripura.
378. Pinnaspis buxi (Bouche, 1851)
Distribution: India.
379. Pinnaspis dracaenae (Cooley, 1899)
Distribution: Kerala (South Malabar, Vadanapalle).
380. Pinnaspis exercitata (Green, 1896)
Distribution: Karnataka (Coorg), Odisha, Tamil Nadu (Coimbatore, Chennai) and West Bengal.
381. Pinnaspis fici (Green, 1908)
46
Distribution: Bihar.
382. Pinnaspis minima (Green, 1908)
Distribution: Bihar.
383. Pinnaspis strachani (Cooley, 1899)
Distribution: Andhra Pradesh, Bihar, Karnataka (Bengaluru, Devanahally, Bellari, Vasanthapura, Hoskote, Nandi, Koramangala), Kerala (Kondayur), Jammu & Kashmir, Tamil Nadu (Coimbatore, Nilgiris, Marudamali) and West Bengal.
384. Pinnaspis theae (Maskell, 1891)
Distribution: North India (Assam) and West Bengal (Darjeeling).
385. Pinnaspis uniloba (Kuwana, 1909)
Distribution: Assam and Tamil Nadu (Chennai, Pudukottai).
158. Genus: Poliaspis Maskell, 1880
386. Poliaspis cycadis Comstock, 1883
Distribution: Maharashtra (Kolhapur).
159. Genus: Pseudaulacaspis MacGillivray, 1921
Pseudaulacaspis sp.
Distribution: Karnataka (Kodigehalli, Namadachiluve).
387. Pseudaulacaspis barberi (Green, 1908)
Distribution: Andhra Pradesh (Visakhapatnam, Alamanda), Karnataka (Bengaluru, Bellari) and Tamil Nadu (Tanjore Dist.).
388. Pseudaulacaspis cockerelli (Cooley, 1897)
Distribution: Andhra Pradesh (Godavari Dist.), Karnataka (Bengaluru, Lal Bagh, Koramangala), Kerala (Malabar, Wynaad, Kondayur), Maharashtra (Pune), Tamil Nadu (Hosur, Nilgiris) and West Bengal (Kolkata).
389. Pseudaulacaspis gynandropsidis (Green, 1922)
Distribution: Karnataka (Bengaluru, Jalahalli).
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390. Pseudaulacaspis manni (Green, 1907)
Distribution: Assam, Himachal Pradesh (Kangra), Tamil Nadu (Hosur) and West Bengal (Kolkata, Darjeeling).
391. Pseudaulacaspis megaloba (Green, 1899)
Distribution: Bihar.
392. Pseudaulacaspis pentagona pentagona (Targioni- Tozzetti, 1886)
Distribution: Assam (Maungep, Sonapari), Karnataka (Byadarahalli, Vasanthapura), Meghalaya (Shilong) and Uttar Pradesh (Chaubattia).
393. Pseudaulacaspis pentagona theae (Maskell, 1896)
Distribution: North India.
394. Pseudaulacaspis polygoni (Green, 1899)
Distribution: Karnataka (Mallandur).
395. Pseudaulacaspis pusa (Rao, 1953)
Distribution: Bihar.
396. Pseudaulacaspis subcorticalis (Green, 1905)
Distribution: Karnataka (Jayanthigrama, Kuppehalli, Kobbinahalli).
397. Pseudaulacaspis tenera (Green, 1922)
Distribution: Karnataka (Karkada).
398. Pseudaulacaspis varicosa (Green, 1896)
Distribution: Karnataka (Coorg, Sidapur) and Tamil Nadu (Mundathurai, Nilgiris, Tirunelveli).
399. Pseudaulacaspis venue (Menon & Khan, 1961)
Distribution: Himachal Pradesh (Kangra, Palampur).
160. Genus: Pseudoparlatoria Cockerell, 1892
400. Pseudoparlatoria parlatorioides (Comstock, 1883)
Distribution: India.
48
161. Genus: Rutherfordia MacGillivray, 1921
401. Rutherfordia major (Cockerell, 1894)
Distribution: Kerala (Periyar).
162. Genus: Sinistraspis MacGillivray, 1921
402. Sinistraspis unilateralis (Newstead, 1913)
Distribution: Kerala (Thiruvananthapuram).
163. Genus: Tenuiaspis MacGillivray, 1921
403. Tenuiaspis minuta (Green, 1896)
Distribution: India.
164. Genus: Thysanofiorinia Balachowsky, 1954
404. Thysanofiorinia leei Williams, 1971
Distribution: Tripura (Agartala) and Uttarakhand (Dehra Dun).
405. Thysanofiorinia nephelii (Maskell, 1897)
Distribution: Karnataka (Coorg, Siddapur) and West Bengal (Krishnanagar).
165. Genus: Trullifiorinia Leonardi, 1906
406. Trullifiorinia geloniae (Green, 1900)
Distribution: Tamil Nadu (Ooty).
166. Genus: Unaspis MacGillivray, 1921
Unaspis sp.
Distribution: Karnataka (Nandi).
407. Unaspis acuminate (Green, 1896)
Distribution: Kerala (Chalikudi, Malabar, Peria Ghat) and Tamil Nadu (Madurai).
408. Unaspis atricolor (Green, 1922)
Distribution: Andhra Pradesh (Hyderabad) and Tamil Nadu (Coimbatore, Dharmapuri).
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409. Unaspis permutans (Green, 1899)
Distribution: Kerala (Malabar, Peria Pass).
REFERENCES
Varshney R. K. 1992. A Check list of Scale insects and mealy bugs of South Asia, (Part-1), Rec. zool. Surv. India, Occ. Paper, 139: 152. Varshney R. K. 2002. A Check list of Scale insects and mealy bugs of South Asia, (Part-2), Rec. zool. Surv. India, Occ. Paper, 191: 147. Williams D. J. 1978. The anomalous ant-attended mealy bugs (Homoptera: Pseudococcidae) of South-east Asia, Bull. Br. Mus. nat. (Ent.), 37 (1): 1-72. Y. Ben-Dev, C.J.Hodgson and D.R.Miller. 1997. Changes and Comments on the Taxonomy and Nomenclature of Some Taxa in the Families Coccidae, Eriococcidae and Pseudococcidae (Homoptera: Coccoidae). Phytoparasitica, 25(3): 199-206. Internet Source : http://www.sel.barc.usda.gov/SCALENET/SCALENET.HTM